treatments-xml/data/03/C0/3B/03C03B6BFF9EFFFDFF7B2BC0FD3EF82A.xml

<document id="2CE0FBCA59FDB4F2663F466DB17321DB" ID-CLB-Dataset="2431" ID-DOI="10.11646/zootaxa.5093.1.1" ID-GBIF-Dataset="0faf68be-3777-4466-8cd5-9eb8ca72a239" ID-ISSN="1175-5326" ID-Zenodo-Dep="5901006" ID-ZooBank="C2F5E368-CB15-4207-9944-E52C190FBF20" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1643101690477" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2022" docId="03C03B6BFF9EFFFDFF7B2BC0FD3EF82A" docLanguage="en" docName="zootaxa.5093.1.1.pdf" docOrigin="Zootaxa 5093 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus (Euconnus) fustiger" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="FFF94313FF9BFFF6FFEC2B57FFE6FF93" masterDocTitle="Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="6" updateTime="1699292035325" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="E0F85FB12B0902D4338AD587BD63D540" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9F1C10A2DFD62669A42D1BAF08BCB7BA">
<mods:title id="C6D326C2D26BCB2054E4EA0CE966A9F3">Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="4D24A560CF2CE47F38DDB3577DA0A086" type="personal">
<mods:role id="17B71866D07FEDE1C1DD0CE2D042508A">
<mods:roleTerm id="1D79BE441517020093E5BD2A44C050FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F4E2BC57C33A83A7D0380F059684389">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="D98EE85A6B6148B41C1E150C6A859E82">text</mods:typeOfResource>
<mods:relatedItem id="2553FAB7A1A173CF5BEDE07B055E6118" type="host">
<mods:titleInfo id="9ED3BEDD43E98C8FA980CF5E863F520D">
<mods:title id="0C54A7F11F72F0DD97E68CE631036875">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF69C168964A6DDD74EA4F69AFCAA556">
<mods:date id="6095DC7DEB0CA9B02D03D70EE7D2F419">2022</mods:date>
<mods:detail id="937757F4A8CC2F55BEC714E260AD9F38" type="pubDate">
<mods:number id="A81B598BB482F49968A02D310E29858F">2022-01-25</mods:number>
</mods:detail>
<mods:detail id="BC2421B92D93852AAD942D7E29C9843D" type="volume">
<mods:number id="04D4DB8FA3407958483ADB72EC927800">5093</mods:number>
</mods:detail>
<mods:detail id="1935EBD54D11820A2558779D6685D06B" type="issue">
<mods:number id="79E74BBFE4891EF389AF2BE86518EE76">1</mods:number>
</mods:detail>
<mods:extent id="D250ECD176C1071498F02307567D6F37" unit="page">
<mods:start id="652E7CD704AC67098DE7F34D5B9412CC">1</mods:start>
<mods:end id="94E121F311245BC3164B1CBE46BD26BA">37</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7DC8F03C4C077B946ED7D7321A0B3A5A">journal article</mods:classification>
<mods:identifier id="27A5EB6761FFD2C80E7F52E5B3756B0E" type="CLB-Dataset">2431</mods:identifier>
<mods:identifier id="E8D7606E98C4741FE9F5DAD4BA06C550" type="DOI">10.11646/zootaxa.5093.1.1</mods:identifier>
<mods:identifier id="34D7443E47E2A0E09ACFFDB88B390EE8" type="GBIF-Dataset">0faf68be-3777-4466-8cd5-9eb8ca72a239</mods:identifier>
<mods:identifier id="355F0DCE6E6697710004FF7FDBF1CDC8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3C04333A9DF04A943F8A5A9AFCC48662" type="Zenodo-Dep">5901006</mods:identifier>
<mods:identifier id="504E0AB8340F9EB26B4C05823581FB84" type="ZooBank">C2F5E368-CB15-4207-9944-E52C190FBF20</mods:identifier>
</mods:mods>
<treatment id="03C03B6BFF9EFFFDFF7B2BC0FD3EF82A" ID-DOI="http://doi.org/10.5281/zenodo.5905974" ID-GBIF-Taxon="192597242" ID-Zenodo-Dep="5905974" LSID="urn:lsid:plazi:treatment:03C03B6BFF9EFFFDFF7B2BC0FD3EF82A" httpUri="http://treatment.plazi.org/id/03C03B6BFF9EFFFDFF7B2BC0FD3EF82A" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6">
<subSubSection id="C373D9F6FF9EFFF3FF7B2BC0FD93FF22" box="[151,629,151,177]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BD68A7DFF9EFFF3FF7B2BC0FD93FF22" blockId="5.[151,629,151,177]" box="[151,629,151,177]" pageId="5" pageNumber="6">
<heading id="D09E3D11FF9EFFF3FF7B2BC0FD93FF22" bold="true" box="[151,629,151,177]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="4C69F1FEFF9EFFF3FF7B2BC0FD93FF22" ID-CoL="9G922" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[151,629,151,177]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fustiger" subGenus="Euconnus">
<emphasis id="B91D566FFF9EFFF3FF7B2BC0FD93FF22" bold="true" box="[151,629,151,177]" pageId="5" pageNumber="6">
<emphasis id="B91D566FFF9EFFF3FF7B2BC0FEF5FF22" bold="true" box="[151,275,151,177]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FEC82BC0FE46FF22" bold="true" box="[292,416,151,177]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE5D2BC0FDE8FF22" bold="true" box="[433,526,151,177]" italics="true" pageId="5" pageNumber="6">fustiger</emphasis>
(Sharp)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF9EFFF3FF2B2B81FEFFFDD6" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BD68A7DFF9EFFF3FF2B2B81FC2CFF7D" blockId="5.[151,1436,214,581]" box="[199,970,214,238]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2B81FC2CFF7D" box="[199,970,214,238]" pageId="5" pageNumber="6">
<treatmentCitation id="0AC8AC6CFF9EFFF3FF2B2B81FDA9FF7D" author="Sharp, D." box="[199,591,214,238]" page="128" pageId="5" pageNumber="6" year="1874">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2B81FDA9FF7D" ID-CoL="8VXQY" authority="Sharp, 1874: 128" authorityName="Sharp" authorityPageNumber="128" authorityYear="1874" box="[199,591,214,238]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF9EFFF3FF2B2B81FE7EFF7D" box="[199,408,214,238]" italics="true" pageId="5" pageNumber="6">
Scydmaenus 
<emphasis id="B91D566FFF9EFFF3FEA72B81FE7EFF7D" box="[331,408,214,238]" pageId="5" pageNumber="6">fustiger</emphasis>
</emphasis>
<bibRefCitation id="EFF8F78CFF9EFFF3FE722B81FDA9FF7D" author="Sharp, D." box="[414,591,214,238]" pageId="5" pageNumber="6" pagination="105 - 130" refId="ref26336" refString="Sharp, D. (1874) The Pselaphidae and Scydmaenidae of Japan. Transactions of the Entomological Society of London, 1, 105 - 130. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00160. x" type="journal article" year="1874">Sharp, 1874: 128</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Redescribed by 
<treatmentCitation id="0AC8AC6CFF9EFFF3FD102B81FC20FF7D" author="Hoshina, H." box="[764,966,214,238]" page="97" pageId="5" pageNumber="6" year="2019">
<bibRefCitation id="EFF8F78CFF9EFFF3FD102B81FC20FF7D" author="Hoshina, H." box="[764,966,214,238]" pageId="5" pageNumber="6" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina, 2019b: 97</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2BA1FCC3FE9E" blockId="5.[151,1436,214,581]" box="[199,805,245,269]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2BA1FCC3FE9E" box="[199,805,245,269]" pageId="5" pageNumber="6">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2BA1FD92FE9E" ID-CoL="9G922" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[199,628,245,269]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fustiger" subGenus="Euconophron">
<emphasis id="B91D566FFF9EFFF3FF2B2BA1FECDFE9E" box="[199,299,246,269]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FED62BA2FE25FE9E" box="[314,451,245,269]" italics="true" pageId="5" pageNumber="6">Euconophron</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE3E2BA2FDF9FE9E" box="[466,543,245,269]" italics="true" pageId="5" pageNumber="6">fustiger</emphasis>
(Sharp)
</taxonomicName>
; 
<treatmentCitation id="0AC8AC6CFF9EFFF3FD932BA2FCC7FE9E" author="Franz, H." box="[639,801,245,269]" page="55" pageId="5" pageNumber="6" year="1975">
<bibRefCitation id="EFF8F78CFF9EFFF3FD932BA2FCC7FE9E" author="Franz, H." box="[639,801,245,269]" pageId="5" pageNumber="6" pagination="57 - 107" refId="ref24521" refString="Franz, H. (1975) Zweiter Beitrag zur Kenntnis der Scydmaenidenfauna Sudostasiens. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abteilung I, 183, 57 - 107." type="journal article" year="1975">Franz, 1975: 55</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2A43FEAFFED9" blockId="5.[151,1436,214,581]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2A43FEAFFED9" pageId="5" pageNumber="6">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2A43FD80FEBF" ID-CoL="9G922" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[199,614,276,300]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fustiger" subGenus="Eupentarius">
<emphasis id="B91D566FFF9EFFF3FF2B2A43FECDFEB8" box="[199,299,276,299]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FED52A43FE53FEB8" box="[313,437,276,299]" italics="true" pageId="5" pageNumber="6">Eupentarius</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE282A43FDF7FEBF" box="[452,529,276,300]" italics="true" pageId="5" pageNumber="6">fustiger</emphasis>
(Sharp)
</taxonomicName>
; 
<treatmentCitation id="0AC8AC6CFF9EFFF3FD9C2A43FC82FEB8" author="Jaloszynski, P." box="[624,868,275,300]" page="156" pageId="5" pageNumber="6" year="2021">
<bibRefCitation id="EFF8F78CFF9EFFF3FD9C2A43FC82FEB8" author="Jaloszynski, P." box="[624,868,275,300]" pageId="5" pageNumber="6" pagination="145 - 160" refId="ref25844" refString="Jaloszynski, P. (2021 a) Taxonomy of &quot; Euconnus complex &quot;. Part XXII. Status of subgenera Allonapochus Franz, Dimorphoconnus Franz, Eupentarius Reitter, and Myrmecodytes Croissandeau (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4966 (2), 145 - 160. https: // doi. org / 10.11646 / zootaxa. 4966.2.3" type="journal article" year="2021">Jałoszyński, 2021a: 156</bibRefCitation>
</treatmentCitation>
, implied, by placing 
<taxonomicName id="4C69F1FEFF9EFFF3FBD42A43FB27FEBF" ID-CoL="9G88P" box="[1080,1217,276,300]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Euconophron">
<emphasis id="B91D566FFF9EFFF3FBD42A43FB27FEBF" box="[1080,1217,276,300]" italics="true" pageId="5" pageNumber="6">Euconophron</emphasis>
</taxonomicName>
as junior synonym of 
<emphasis id="B91D566FFF9EFFF3FF2B2A64FEA5FED9" box="[199,323,307,330]" italics="true" pageId="5" pageNumber="6">Eupentarius</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2A05FB55FEFA" blockId="5.[151,1436,214,581]" box="[199,1203,337,361]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2A05FB55FEFA" box="[199,1203,337,361]" pageId="5" pageNumber="6">
<treatmentCitation id="0AC8AC6CFF9EFFF3FF2B2A05FDC3FEFA" author="Sharp, D." box="[199,549,337,361]" page="47" pageId="5" pageNumber="6" year="1886">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2A05FDC3FEFA" ID-CoL="3BWPD" authority="Sharp, 1886: 47" authorityName="Sharp" authorityPageNumber="47" authorityYear="1886" box="[199,549,337,361]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF9EFFF3FF2B2A05FE9CFEFA" box="[199,378,338,361]" italics="true" pageId="5" pageNumber="6">Euconnus Lewisii</emphasis>
<bibRefCitation id="EFF8F78CFF9EFFF3FE6D2A06FDC3FEFA" author="Sharp, D." box="[385,549,337,361]" pageId="5" pageNumber="6" pagination="46 - 51" refId="ref26383" refString="Sharp, D. (1886) The Scydmaenidae of Japan. Entomologist's Monthly Magazine, 23, 46 - 51." type="journal article" year="1886">Sharp, 1886: 47</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Placed as junior synonym of 
<taxonomicName id="4C69F1FEFF9EFFF3FCBA2A05FC59FEFA" ID-CoL="9G922" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[854,959,337,361]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF9EFFF3FCBA2A05FC59FEFA" box="[854,959,337,361]" italics="true" pageId="5" pageNumber="6">
E. 
<emphasis id="B91D566FFF9EFFF3FC9E2A06FC59FEFA" box="[882,959,337,361]" pageId="5" pageNumber="6">fustiger</emphasis>
</emphasis>
</taxonomicName>
by 
<treatmentCitation id="0AC8AC6CFF9EFFF3FC092A06FB49FEFA" author="Hoshina, H." box="[997,1199,337,361]" page="97" pageId="5" pageNumber="6" year="2019">
<bibRefCitation id="EFF8F78CFF9EFFF3FC092A06FB49FEFA" author="Hoshina, H." box="[997,1199,337,361]" pageId="5" pageNumber="6" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina, 2019b: 97</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2A27FD01FE14" blockId="5.[151,1436,214,581]" box="[199,743,367,392]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2A27FD01FE14" box="[199,743,367,392]" pageId="5" pageNumber="6">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2A27FDDCFE1B" ID-CoL="8VRNW" authority="(Euconnus) Lewisii Sharp" authorityName="Lewisii Sharp" box="[199,570,368,392]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Euconnus">
<emphasis id="B91D566FFF9EFFF3FF2B2A27FECDFE14" box="[199,299,368,391]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FED62A27FE78FE14" box="[314,414,368,391]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE412A27FE10FE14" box="[429,502,368,391]" italics="true" pageId="5" pageNumber="6">Lewisii</emphasis>
Sharp
</taxonomicName>
; 
<treatmentCitation id="0AC8AC6CFF9EFFF3FDA92A27FD05FE14" author="Csiki, E." box="[581,739,367,392]" page="50" pageId="5" pageNumber="6" year="1919">
<bibRefCitation id="EFF8F78CFF9EFFF3FDA92A27FD05FE14" author="Csiki, E." box="[581,739,367,392]" pageId="5" pageNumber="6" pagination="106" refId="ref24424" refString="Csiki, E. (1919) Pars 70: Scydmaenidae. In: Schenkling, S. &amp; Junk, W. (Eds.), Coleopterorum Catalogus. Vol. 7. W. Junk, Berlin, pp. 106." type="book chapter" year="1919">Csiki, 1919: 50</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2AD8FCE9FE56" blockId="5.[151,1436,214,581]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2AD8FCE9FE56" pageId="5" pageNumber="6">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2AD8FDA5FE35" ID-CoL="3BWPD" authority="(Sharp)" baseAuthorityName="Sharp" box="[199,579,398,422]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lewisii" subGenus="Napochus">
<emphasis id="B91D566FFF9EFFF3FF2B2AD8FECDFE35" box="[199,299,399,422]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FED52AD9FE78FE35" box="[313,414,398,422]" italics="true" pageId="5" pageNumber="6">Napochus</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE402AD9FE08FE35" box="[428,494,398,422]" italics="true" pageId="5" pageNumber="6">lewisii</emphasis>
(Sharp)
</taxonomicName>
; 
<treatmentCitation id="0AC8AC6CFF9EFFF3FDA12AD9FCF2FE35" author="Hoshina, H." box="[589,788,398,422]" page="21" pageId="5" pageNumber="6" year="2004">
<bibRefCitation id="EFF8F78CFF9EFFF3FDA12AD9FCF2FE35" author="Hoshina, H." box="[589,788,398,422]" pageId="5" pageNumber="6" pagination="17 - 22" refId="ref24606" refString="Hoshina, H. (2004 a) A new species and taxonomical note on the genus Euconnus (Coleoptera: Scydmaenidae) from Honshu, Japan. Memoirs of the Research and Education Center for Regional Environment, University of Fukui, 11, 17 - 22." type="journal article" year="2004">Hoshina, 2004a: 21</bibRefCitation>
</treatmentCitation>
; back to 
<taxonomicName id="4C69F1FEFF9EFFF3FC802AD8FC36FE35" ID-CoL="8VRNT" box="[876,976,399,422]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF9EFFF3FC802AD8FC36FE35" box="[876,976,399,422]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EEB14FF9EFFF3FC3A2AD8FBEDFE35" box="[982,1035,399,422]" pageId="5" pageNumber="6" sensu="stricto">s. str.</taxonomicNameLabel>
in 
<treatmentCitation id="0AC8AC6CFF9EFFF3FBC52AD9FAFAFE35" author="Jaloszynski, P." box="[1065,1308,398,422]" page="268" pageId="5" pageNumber="6" year="2021">
<bibRefCitation id="EFF8F78CFF9EFFF3FBC52AD9FAFAFE35" author="Jaloszynski, P." box="[1065,1308,398,422]" pageId="5" pageNumber="6" pagination="255 - 270" refId="ref25908" refString="Jaloszynski, P. (2021 b) Taxonomy of &quot; Euconnus complex &quot;. Part XXIII. Status of Napochus Thomson, Pycnophus Casey, and Filonapochus Franz revisited (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5026 (2), 255 - 270. https: // doi. org / 10.11646 / zootaxa. 5026.2.6" type="journal article" year="2021">Jałoszyński, 2021b: 268</bibRefCitation>
</treatmentCitation>
(implied, by placing 
<emphasis id="B91D566FFF9EFFF3FEF42AFAFE9BFE56" box="[280,381,429,453]" italics="true" pageId="5" pageNumber="6">Napochus</emphasis>
as junior synonym of 
<taxonomicName id="4C69F1FEFF9EFFF3FD8D2AF9FD23FE56" ID-CoL="8VRNT" box="[609,709,430,453]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF9EFFF3FD8D2AF9FD23FE56" box="[609,709,430,453]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EEB14FF9EFFF3FD272AFAFCE5FE56" box="[715,771,429,453]" pageId="5" pageNumber="6" sensu="stricto">s. str.</taxonomicNameLabel>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2A9BFACDFE77" blockId="5.[151,1436,214,581]" box="[199,1323,459,484]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2A9BFACDFE77" box="[199,1323,459,484]" pageId="5" pageNumber="6">
<treatmentCitation id="0AC8AC6CFF9EFFF3FF2B2A9BFCEDFE70" author="Franz, H." box="[199,779,459,484]" page="56" pageId="5" pageNumber="6" year="1976">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2A9BFCEDFE70" authority="Franz, 1976: 56" authorityName="Franz" authorityPageNumber="56" authorityYear="1976" box="[199,779,459,484]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="miyawakianus" subGenus="Euconophron">
<emphasis id="B91D566FFF9EFFF3FF2B2A9BFECDFE70" box="[199,299,460,483]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FED62A9BFE25FE77" box="[314,451,460,484]" italics="true" pageId="5" pageNumber="6">Euconophron</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE3E2A9BFD85FE77" box="[466,611,460,484]" italics="true" pageId="5" pageNumber="6">miyawakianus</emphasis>
<bibRefCitation id="EFF8F78CFF9EFFF3FD852A9BFCEDFE70" author="Franz, H." box="[617,779,459,483]" pageId="5" pageNumber="6" pagination="51 - 60" refId="ref24555" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Ergebnisse einer auf Einladung der Internationalen Vereinigung fur Vegetationskunde nach Japan durchgefuhrten Forschungsreise. Entomologische Blatter fur Biologie und Systematik der Kafer, 72 (1), 51 - 60." type="journal article" year="1976">Franz, 1976: 56</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized with 
<taxonomicName id="4C69F1FEFF9EFFF3FC362A9BFBDEFE77" ID-CoL="3BWPD" box="[986,1080,460,484]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF9EFFF3FC362A9BFBDEFE77" box="[986,1080,460,484]" italics="true" pageId="5" pageNumber="6">E. lewisii</emphasis>
</taxonomicName>
by 
<treatmentCitation id="0AC8AC6CFF9EFFF3FBB32A9BFAC1FE77" author="Hoshina, H." box="[1119,1319,460,484]" page="21" pageId="5" pageNumber="6" year="2004">
<bibRefCitation id="EFF8F78CFF9EFFF3FBB32A9BFAC1FE77" author="Hoshina, H." box="[1119,1319,460,484]" pageId="5" pageNumber="6" pagination="17 - 22" refId="ref24606" refString="Hoshina, H. (2004 a) A new species and taxonomical note on the genus Euconnus (Coleoptera: Scydmaenidae) from Honshu, Japan. Memoirs of the Research and Education Center for Regional Environment, University of Fukui, 11, 17 - 22." type="journal article" year="2004">Hoshina, 2004a: 21</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B2ABCFC34FD91" blockId="5.[151,1436,214,581]" box="[199,978,490,514]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B2ABCFC34FD91" box="[199,978,490,514]" pageId="5" pageNumber="6">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B2ABCFD85FD91" box="[199,611,490,514]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="miyawakainus" subGenus="Euconophron">
<emphasis id="B91D566FFF9EFFF3FF2B2ABCFECDFD91" box="[199,299,491,514]" italics="true" pageId="5" pageNumber="6">Euconnus</emphasis>
(
<emphasis id="B91D566FFF9EFFF3FED62ABDFE25FD91" box="[314,451,490,514]" italics="true" pageId="5" pageNumber="6">Euconophron</emphasis>
) 
<emphasis id="B91D566FFF9EFFF3FE3E2ABDFD85FD91" box="[466,611,490,514]" italics="true" pageId="5" pageNumber="6">miyawakainus</emphasis>
</taxonomicName>
, misspelling in 
<treatmentCitation id="0AC8AC6CFF9EFFF3FCE82ABDFC28FD91" author="Hoshina, H." box="[772,974,490,514]" page="97" pageId="5" pageNumber="6" year="2019">
<bibRefCitation id="EFF8F78CFF9EFFF3FCE82ABDFC28FD91" author="Hoshina, H." box="[772,974,490,514]" pageId="5" pageNumber="6" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina, 2019b: 97</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF2B295DFD41FDB2" blockId="5.[151,1436,214,581]" box="[199,679,521,545]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB79AD53FF9EFFF3FF2B295DFD41FDB2" box="[199,679,521,545]" pageId="5" pageNumber="6">
<treatmentCitation id="0AC8AC6CFF9EFFF3FF2B295DFDB0FDB2" author="Reitter, E." box="[199,598,521,545]" page="19" pageId="5" pageNumber="6" year="1891">
<taxonomicName id="4C69F1FEFF9EFFF3FF2B295DFDB0FDB2" ID-CoL="3BX6X" authority="Reitter, 1891: 19" authorityName="Reitter" authorityPageNumber="19" authorityYear="1891" box="[199,598,521,545]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schonfeldti">
<emphasis id="B91D566FFF9EFFF3FF2B295DFE45FDB2" box="[199,419,521,545]" italics="true" pageId="5" pageNumber="6">Euconnus Schönfeldti</emphasis>
<bibRefCitation id="EFF8F78CFF9EFFF3FE46295EFDB0FDB2" author="Reitter, E." box="[426,598,521,545]" pageId="5" pageNumber="6" pagination="297 - 305" refId="ref26141" refString="Reitter, E. (1891) Neue Coleopteren aus Europa, der angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannten Arten. Zwolfter Theil. Deutsche Entomologische Zeitschrift, (1), 17 - 36. Reitter, E. (1907) Verzeichnis der von Dr. F. Eichelbaum im Jahre 1903 in Deutsch-Ostafrika gesammelten Scydmaeniden (Col.). Wiener Entomologische Zeitung, 26 (7 - 9), 297 - 305. https: // doi. org / 10.5962 / bhl. part. 8894" type="journal article" year="1891">Reitter, 1891: 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; 
<emphasis id="B91D566FFF9EFFF3FD8D295EFD41FDB2" bold="true" box="[609,679,521,545]" pageId="5" pageNumber="6">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF9EFFF3FF7B297CFEFFFDD6" blockId="5.[151,1436,214,581]" box="[151,281,555,581]" pageId="5" pageNumber="6">
(
<figureCitation id="135296F8FF9EFFF3FF73297CFEF7FDD6" box="[159,273,555,581]" captionStart-0="FIGURES 9–12" captionStart-1="FIGURES 13–18" captionStart-2="FIGURES 19–24" captionStart-3="FIGURES 25–30" captionStart-4="FIGURES 31–49" captionStartId-0="7.[151,264,1752,1776]" captionStartId-1="8.[151,264,1829,1853]" captionStartId-2="9.[151,264,1573,1597]" captionStartId-3="10.[151,264,1931,1955]" captionStartId-4="12.[151,264,1318,1342]" captionTargetBox-0="[163,1428,389,1726]" captionTargetBox-1="[160,1428,680,1800]" captionTargetBox-2="[155,1426,193,1538]" captionTargetBox-3="[160,1420,680,1899]" captionTargetBox-4="[162,1427,190,1288]" captionTargetId-0="figure-80@7.[151,1436,381,1729]" captionTargetId-1="figure-238@8.[151,1436,669,1806]" captionTargetId-2="figure-15@9.[151,1436,181,1549]" captionTargetId-3="figure-269@10.[151,1436,669,1908]" captionTargetId-4="figure-21@12.[151,1436,181,1294]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="12" captionText-0="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." captionText-1="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." captionText-2="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." captionText-3="FIGURES 25–30. Euconnus fustiger, male. Dorsal habitus (25) and historical labels (26) of lectotype male of Euconnus schoenfeldti; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views." captionText-4="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi-0="http://doi.org/10.5281/zenodo.5901012" figureDoi-1="http://doi.org/10.5281/zenodo.5901014" figureDoi-2="http://doi.org/10.5281/zenodo.5901018" figureDoi-3="http://doi.org/10.5281/zenodo.5901022" figureDoi-4="http://doi.org/10.5281/zenodo.5901024" httpUri-0="https://zenodo.org/record/5901012/files/figure.png" httpUri-1="https://zenodo.org/record/5901014/files/figure.png" httpUri-2="https://zenodo.org/record/5901018/files/figure.png" httpUri-3="https://zenodo.org/record/5901022/files/figure.png" httpUri-4="https://zenodo.org/record/5901024/files/figure.png" pageId="5" pageNumber="6">Figs 9–49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF9EFFF0FF7B2939FAC6FB70" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BD68A7DFF9EFFF3FF7B2939FC5BFA33" blockId="5.[151,1437,621,2016]" pageId="5" pageNumber="6">
<emphasis id="B91D566FFF9EFFF3FF7B2939FE44FD14" bold="true" box="[151,418,621,648]" pageId="5" pageNumber="6">Type material studied.</emphasis>
<materialsCitation id="3B018020FF9EFFF3FE442939FF09FD67" ID-GBIF-Occurrence="3454958315" collectionCode="NHM" collectorName="G. Lewis." country="Japan" location="Japan" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B91D566FFF9EFFF3FE442939FC64FD14" bold="true" box="[424,898,621,648]" pageId="5" pageNumber="6">
<typeStatus id="54D234DFFF9EFFF3FE442939FDF3FD1B" box="[424,533,622,648]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF9EFFF3FDD6293AFC9BFD14" authority="Sharp" authorityName="Sharp" authorityYear="1874" box="[570,893,621,648]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF9EFFF3FDD6293AFCCCFD14" bold="true" box="[570,810,621,648]" italics="true" pageId="5" pageNumber="6">
Scydmaenus 
<emphasis id="B91D566FFF9EFFF3FD3D293AFCCCFD14" box="[721,810,621,647]" pageId="5" pageNumber="6">fustiger</emphasis>
</emphasis>
Sharp
</taxonomicName>
:
</emphasis>
sex unknown (probably 
<specimenCount id="9D6F41F4FF9EFFF3FB7B2939FB4CFD1B" box="[1175,1194,622,648]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
) (
<figureCitation id="135296F8FF9EFFF3FB282939FAEEFD1B" box="[1220,1288,622,648]" captionStart="FIGURES 9–12" captionStartId="7.[151,264,1752,1776]" captionTargetBox="[163,1428,389,1726]" captionTargetId="figure-80@7.[151,1436,381,1729]" captionTargetPageId="7" captionText="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." figureDoi="http://doi.org/10.5281/zenodo.5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
), three labels (
<figureCitation id="135296F8FF9EFFF3FF4C29C5FF10FD3F" box="[160,246,658,684]" captionStart="FIGURES 9–12" captionStartId="7.[151,264,1752,1776]" captionTargetBox="[163,1428,389,1726]" captionTargetId="figure-80@7.[151,1436,381,1729]" captionTargetPageId="7" captionText="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." figureDoi="http://doi.org/10.5281/zenodo.5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
): “ 
<collectingCountry id="F37ECAEDFF9EFFF3FEF929C5FEBDFD3F" box="[277,347,658,684]" name="Japan" pageId="5" pageNumber="6">Japan</collectingCountry>
. / 
<collectorName id="269CEFABFF9EFFF3FE9429C5FE0EFD3F" box="[376,488,658,684]" pageId="5" pageNumber="6">G. Lewis.</collectorName>
/ 1910-320.” [white with horizontal orange line, printed], “ 
<taxonomicName id="4C69F1FEFF9EFFF3FB6129C5FAFBFD3F" authorityName="Latreille" authorityYear="1802" box="[1165,1309,658,684]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ 
<emphasis id="B91D566FFF9EFFF3FADA29C5FA6AFD3F" box="[1334,1420,658,684]" pageId="5" pageNumber="6">fustiger</emphasis>
/ Type. D.S.” [white with black frame, handwritten and printed], “Type / H.T.” [circular label with red margin, printed] (
<collectionCode id="ED7812B8FF9EFFF3FF73298DFF01FD67" box="[159,231,730,756]" pageId="5" pageNumber="6">NHM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B018020FF9EFFF3FF16298DFE15FCF3" ID-GBIF-Occurrence="3454958303" collectionCode="NHM" country="Japan" location="Nagasaki" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Nagasaki" typeStatus="lectotype">
<emphasis id="B91D566FFF9EFFF3FF16298DFC77FD60" bold="true" box="[250,913,729,756]" pageId="5" pageNumber="6">
<typeStatus id="54D234DFFF9EFFF3FF16298DFE96FD67" box="[250,368,730,756]" pageId="5" pageNumber="6" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF9EFFF3FE74298DFD57FD60" authority="Sharp" authorityName="Sharp" authorityYear="1886" box="[408,689,729,756]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF9EFFF3FE74298DFDB9FD67" bold="true" box="[408,607,729,756]" italics="true" pageId="5" pageNumber="6">Euconnus lewisii</emphasis>
Sharp
</taxonomicName>
(here designated):
</emphasis>
<specimenCount id="9D6F41F4FF9EFFF3FC75298DFC48FD67" box="[921,942,730,756]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
(
<figureCitation id="135296F8FF9EFFF3FC52298DFBF4FD67" box="[958,1042,730,756]" captionStart="FIGURES 13–18" captionStartId="8.[151,264,1829,1853]" captionTargetBox="[160,1428,680,1800]" captionTargetId="figure-238@8.[151,1436,669,1806]" captionTargetPageId="8" captionText="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." figureDoi="http://doi.org/10.5281/zenodo.5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
), mounted on thick cardboard (Fg. 14) with annotation: “ 
<taxonomicName id="4C69F1FEFF9EFFF3FE6429A9FE1FFC8B" box="[392,505,766,792]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ lewisius / D.S. / 
<collectingRegion id="49AD449FFF9EFFF3FD2829A9FCCAFC8B" box="[708,812,766,792]" country="Japan" name="Nagasaki" pageId="5" pageNumber="6">Nagasaki</collectingRegion>
/ 8.4.91 {below crossed out 9.... illegible} / Lewis”, and two labels (
<figureCitation id="135296F8FF9EFFF3FEF02875FE89FCAF" box="[284,367,802,828]" captionStart="FIGURES 13–18" captionStartId="8.[151,264,1829,1853]" captionTargetBox="[160,1428,680,1800]" captionTargetId="figure-238@8.[151,1436,669,1806]" captionTargetPageId="8" captionText="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." figureDoi="http://doi.org/10.5281/zenodo.5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
) “Sharp Coll. / 1905-313.” [darkened white, printed], circular label “Syn- / type” [white with blue margins, printed] (
<collectionCode id="ED7812B8FF9EFFF3FE4F2811FE0DFCF3" box="[419,491,838,864]" pageId="5" pageNumber="6">NHM</collectionCode>
)
</materialsCitation>
. 
<emphasis id="B91D566FFF9EFFF3FE132811FBB0FCCC" bold="true" box="[511,1110,837,864]" pageId="5" pageNumber="6">
<typeStatus id="54D234DFFF9EFFF3FE132811FD49FCF3" box="[511,687,838,864]" pageId="5" pageNumber="6" type="paralectotype">Paralectotypes</typeStatus>
of 
<taxonomicName id="4C69F1FEFF9EFFF3FD342811FC14FCCC" authority="Sharp" authorityName="Sharp" authorityYear="1886" box="[728,1010,837,864]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF9EFFF3FD342811FC46FCF3" bold="true" box="[728,928,837,864]" italics="true" pageId="5" pageNumber="6">Euconnus lewisii</emphasis>
Sharp
</taxonomicName>
(3 exx):
</emphasis>
<materialsCitation id="3B018020FF9EFFF3FB8C2811FA7EFC5F" ID-GBIF-Occurrence="3454958325" collectionCode="NHM" country="Japan" location="Lewis" municipality="Type" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Nagasaki" typeStatus="lectotype">
<specimenCount id="9D6F41F4FF9EFFF3FB8C2811FB93FCF3" box="[1120,1141,838,864]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
(
<figureCitation id="135296F8FF9EFFF3FB6A2811FB3FFCF3" box="[1158,1241,838,864]" captionStart="FIGURES 9–12" captionStartId="7.[151,264,1752,1776]" captionTargetBox="[163,1428,389,1726]" captionTargetId="figure-80@7.[151,1436,381,1729]" captionTargetPageId="7" captionText="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." figureDoi="http://doi.org/10.5281/zenodo.5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
), mounted in the same way as 
<typeStatus id="54D234DFFF9EFFF3FEC7283DFE73FC17" box="[299,405,874,900]" pageId="5" pageNumber="6" type="lectotype">lectotype</typeStatus>
, on thick cardboard (
<figureCitation id="135296F8FF9EFFF3FD6D283DFD34FC17" box="[641,722,874,900]" captionStart="FIGURES 9–12" captionStartId="7.[151,264,1752,1776]" captionTargetBox="[163,1428,389,1726]" captionTargetId="figure-80@7.[151,1436,381,1729]" captionTargetPageId="7" captionText="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." figureDoi="http://doi.org/10.5281/zenodo.5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="5" pageNumber="6">Fig. 12</figureCitation>
) with annotation: “ 
<taxonomicName id="4C69F1FEFF9EFFF3FC44283DFBFFFC17" box="[936,1049,874,900]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ lewisius 
<collectingMunicipality id="6BB21007FF9EFFF3FB7D283DFB2FFC17" box="[1169,1225,874,900]" pageId="5" pageNumber="6">Type</collectingMunicipality>
/ D.S. / 
<collectingRegion id="49AD449FFF9EFFF3FAC9283DFA6BFC17" box="[1317,1421,874,900]" country="Japan" name="Nagasaki" pageId="5" pageNumber="6">Nagasaki</collectingRegion>
/ 8.4.91. 
<location id="8EB6DCA6FF9EFFF3FF0728D9FED2FC3B" LSID="urn:lsid:plazi:treatment:03C03B6BFF9EFFFDFF7B2BC0FD3EF82A:8EB6DCA6FF9EFFF3FF0728D9FED2FC3B" box="[235,308,910,936]" country="Japan" municipality="Type" name="Lewis" pageId="5" pageNumber="6" stateProvince="Nagasaki">Lewis</location>
”, and three labels (
<figureCitation id="135296F8FF9EFFF3FDE628D9FDBAFC3B" box="[522,604,910,936]" captionStart="FIGURES 9–12" captionStartId="7.[151,264,1752,1776]" captionTargetBox="[163,1428,389,1726]" captionTargetId="figure-80@7.[151,1436,381,1729]" captionTargetPageId="7" captionText="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." figureDoi="http://doi.org/10.5281/zenodo.5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="5" pageNumber="6">Fig. 12</figureCitation>
) “Sharp Coll. / 1905-313.” [darkened white, printed], circular label “Type / H.T.” [white with red margin, printed], and circular label “Syn- / type” [white with blue margins, printed] (
<collectionCode id="ED7812B8FF9EFFF3FAA428E5FA69FC5F" box="[1352,1423,946,972]" pageId="5" pageNumber="6">NHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B018020FF9EFFF3FF7B2881FB6AFBAB" ID-GBIF-Occurrence="3454958310" collectionCode="NHM" country="Japan" location="Nagasaki" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Nagasaki" typeStatus="lectotype">
<specimenCount id="9D6F41F4FF9EFFF3FF7B2881FF4AFC63" box="[151,172,982,1008]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
, mounted on a thick cardboard narrower than that for 
<typeStatus id="54D234DFFF9EFFF3FCCE2881FC6FFC63" box="[802,905,982,1008]" pageId="5" pageNumber="6" type="lectotype">lectotype</typeStatus>
male, annotated: “ 
<taxonomicName id="4C69F1FEFF9EFFF3FB8C2881FB37FC63" box="[1120,1233,982,1008]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ lewisius / var. / 
<collectingRegion id="49AD449FFF9EFFF3FF7B28ADFF19FB87" box="[151,255,1018,1044]" country="Japan" name="Nagasaki" pageId="5" pageNumber="6">Nagasaki</collectingRegion>
/ 9.4.91 {“9” in “91” upside down, appearing as “6”}, and two labels “Sharp Coll. / 1905-313.” [darkened white, printed], circular label “Syn- / type” [white with blue margins, printed] (
<collectionCode id="ED7812B8FF9EFFF3FBD12F49FB62FBAB" box="[1085,1156,1054,1080]" pageId="5" pageNumber="6">NHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B018020FF9EFFF3FB702F49FA7FFB37" ID-GBIF-Occurrence="3454958322" collectionCode="NHM" collectorName="G. Lewis." country="Japan" location="Nagasaki" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Nagasaki" typeStatus="paralectotype">
<specimenCount id="9D6F41F4FF9EFFF3FB702F49FB31FBAB" box="[1180,1239,1054,1080]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
of unknown sex (abdomen destroyed, but no genital preparation attached), mounted on modern card, with four labels: “ 
<taxonomicName id="4C69F1FEFF9EFFF3FAF02F15FA6BFBCF" box="[1308,1421,1090,1116]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ lewisius” [white, handwritten], circular label “Syn- / type” [white with blue margins, printed], “ 
<collectingRegion id="49AD449FFF9EFFF3FB3F2F31FAD8FB13" box="[1235,1342,1126,1152]" country="Japan" name="Nagasaki" pageId="5" pageNumber="6">Nagasaki</collectingRegion>
/ 13.II.- 21.IV.81” [white, printed], and “ 
<collectingCountry id="F37ECAEDFF9EFFF3FE102FDDFDA4FB37" box="[508,578,1162,1188]" name="Japan" pageId="5" pageNumber="6">Japan</collectingCountry>
. / 
<collectorName id="269CEFABFF9EFFF3FDB42FDDFD20FB37" box="[600,710,1162,1188]" pageId="5" pageNumber="6">G. Lewis.</collectorName>
/ 1910-320.” [white with horizontal orange line, printed] (
<collectionCode id="ED7812B8FF9EFFF3FAA52FDDFA77FB37" box="[1353,1425,1162,1188]" pageId="5" pageNumber="6">NHM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B018020FF9EFFF3FF7B2FF9FB62FA83" ID-GBIF-Occurrence="3454958337" collectionCode="NHMW" country="Japan" county="Mt. Kasuga" determinerName="H. Franz" location="Typus" municipality="Honshu" pageId="5" pageNumber="6" specimenCount="2" specimenCount-male="2" stateProvince="Nara" typeStatus="holotype">
<emphasis id="B91D566FFF9EFFF3FF7B2FF9FD55FB5B" bold="true" box="[151,691,1197,1224]" pageId="5" pageNumber="6">
<typeStatus id="54D234DFFF9EFFF3FF7B2FF9FEE2FB5B" box="[151,260,1198,1224]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF9EFFF3FEDC2FF9FD48FB5B" authority="Franz" authorityName="Franz" box="[304,686,1197,1224]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF9EFFF3FEDC2FF9FDB1FB5B" bold="true" box="[304,599,1197,1224]" italics="true" pageId="5" pageNumber="6">Euconnus miyawakianus</emphasis>
Franz
</taxonomicName>
:
</emphasis>
<specimenCount id="9D6F41F4FF9EFFF3FD522FF9FD35FB5B" box="[702,723,1198,1224]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
(
<figureCitation id="135296F8FF9EFFF3FD0A2FF9FCDDFB5B" box="[742,827,1198,1224]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
), four labels (
<figureCitation id="135296F8FF9EFFF3FC0E2FF9FBDFFB5B" box="[994,1081,1198,1224]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="5" pageNumber="6">Fig. 20</figureCitation>
), “ 
<collectingCountry id="F37ECAEDFF9EFFF3FBB62FF9FB7BFB5B" box="[1114,1181,1198,1224]" name="Japan" pageId="5" pageNumber="6">Japan</collectingCountry>
1974 / lg.H.FRANZ” [whte, printed], “ 
<collectingCounty id="62B7F2F1FF9EFFF3FEBB2F85FE33FB7F" box="[343,469,1234,1260]" pageId="5" pageNumber="6">Mt.Kasuga</collectingCounty>
b. / 
<collectingRegion id="49AD449FFF9EFFF3FDE52F85FDA4FB7F" box="[521,578,1234,1260]" country="Japan" name="Nara" pageId="5" pageNumber="6">Nara</collectingRegion>
, 
<collectingMunicipality id="6BB21007FF9EFFF3FDAA2F85FD46FB7F" box="[582,672,1234,1260]" pageId="5" pageNumber="6">Honshu</collectingMunicipality>
” [white, printed], “ 
<taxonomicName id="4C69F1FEFF9EFFF3FC952F85FC0CFB7F" box="[889,1002,1234,1260]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ (
<taxonomicName id="4C69F1FEFF9EFFF3FBE52F85FB42FB7E" box="[1033,1188,1234,1261]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Euconophron">Euconophron</taxonomicName>
) / miyawakianus” / 
<specimenCount id="9D6F41F4FF9EFFF3FA6B2F85FA7AFB7F" box="[1415,1436,1234,1260]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
det. 
<determinerName id="7E0A66B9FF9EFFF3FF532FA1FEFAFA83" box="[191,284,1270,1296]" pageId="5" pageNumber="6">H.Franz</determinerName>
m.” [white, handwritten and printed], “ 
<location id="8EB6DCA6FF9EFFF3FD0C2FA1FCCFFA83" LSID="urn:lsid:plazi:treatment:03C03B6BFF9EFFFDFF7B2BC0FD3EF82A:8EB6DCA6FF9EFFF3FD0C2FA1FCCFFA83" box="[736,809,1270,1296]" country="Japan" county="Mt. Kasuga" municipality="Honshu" name="Typus" pageId="5" pageNumber="6" stateProvince="Nara">Typus</location>
” [red, handwritten] (
<collectionCode id="ED7812B8FF9EFFF3FBF52FA1FB9DFA83" box="[1049,1147,1270,1296]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="6" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B018020FF9EFFF3FB7C2FA1FB11FAA7" ID-GBIF-Occurrence="3454958318" collectionCode="NHMW" country="Japan" county="Mt. Kasuga" determinerName="H. Franz" location="Typus" municipality="Honshu" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Nara" typeStatus="paratype">
<emphasis id="B91D566FFF9EFFF3FB7C2FA1FE5EFAA5" bold="true" pageId="5" pageNumber="6">
<typeStatus id="54D234DFFF9EFFF3FB7C2FA1FB1BFA83" box="[1168,1277,1270,1296]" pageId="5" pageNumber="6" type="paratype">Paratype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF9EFFF3FACA2FA1FE72FAA7" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF9EFFF3FACA2FA1FEDBFAA7" bold="true" italics="true" pageId="5" pageNumber="6">Euconnus miyawakianus</emphasis>
Franz
</taxonomicName>
: 
<specimenCount id="9D6F41F4FF9EFFF3FE4F2E4CFE5EFAA5" box="[419,440,1307,1334]" pageId="5" pageNumber="6" type="female">♀</specimenCount>
</emphasis>
, same data as for holotype, except for red label “ 
<typeStatus id="54D234DFFF9EFFF3FC112E4DFB97FAA7" box="[1021,1137,1306,1332]" pageId="5" pageNumber="6" type="allotype">Allotypus</typeStatus>
” (
<collectionCode id="ED7812B8FF9EFFF3FB602E4DFB08FAA7" box="[1164,1262,1306,1332]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="6" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B018020FF9EFFF3FAE82E4DFC5FFA33" ID-GBIF-Occurrence="3454958336" collectionCode="HNHM" collectorName="Coll. Reitter" country="Japan" location="Japan" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B91D566FFF9EFFF3FAE82E4DFD30FAC4" bold="true" pageId="5" pageNumber="6">
<typeStatus id="54D234DFFF9EFFF3FAE82E4DFA9CFAA7" box="[1284,1402,1306,1332]" pageId="5" pageNumber="6" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF9EFFF3FF7B2E69FE1FFAC4" authority="Reitter" authorityName="Reitter" authorityYear="1891" box="[151,505,1341,1368]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schoenfeldti">
<emphasis id="B91D566FFF9EFFF3FF7B2E69FE46FACB" bold="true" box="[151,416,1341,1368]" italics="true" pageId="5" pageNumber="6">Euconnus schoenfeldti</emphasis>
Reitter
</taxonomicName>
(here designated):
</emphasis>
<specimenCount id="9D6F41F4FF9EFFF3FD302E69FD17FACB" box="[732,753,1342,1368]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
(
<figureCitation id="135296F8FF9EFFF3FCEC2E69FCB5FACA" box="[768,851,1342,1369]" captionStart="FIGURES 25–30" captionStartId="10.[151,264,1931,1955]" captionTargetBox="[160,1420,680,1899]" captionTargetId="figure-269@10.[151,1436,669,1908]" captionTargetPageId="10" captionText="FIGURES 25–30. Euconnus fustiger, male. Dorsal habitus (25) and historical labels (26) of lectotype male of Euconnus schoenfeldti; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views." figureDoi="http://doi.org/10.5281/zenodo.5901022" httpUri="https://zenodo.org/record/5901022/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
), three labels (
<figureCitation id="135296F8FF9EFFF3FC182E69FBA1FACA" box="[1012,1095,1342,1369]" captionStart="FIGURES 25–30" captionStartId="10.[151,264,1931,1955]" captionTargetBox="[160,1420,680,1899]" captionTargetId="figure-269@10.[151,1436,669,1908]" captionTargetPageId="10" captionText="FIGURES 25–30. Euconnus fustiger, male. Dorsal habitus (25) and historical labels (26) of lectotype male of Euconnus schoenfeldti; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views." figureDoi="http://doi.org/10.5281/zenodo.5901022" httpUri="https://zenodo.org/record/5901022/files/figure.png" pageId="5" pageNumber="6">Fig. 26</figureCitation>
), “ 
<taxonomicName id="4C69F1FEFF9EFFF3FB882E69FB33FACB" box="[1124,1237,1342,1368]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ Schönfeldti m. / Japonia 1890” [darkened white, handwritten], “Typus 1891. / 
<taxonomicName id="4C69F1FEFF9EFFF3FCA72E35FC0DFAEF" authorityName="Reitter" authorityYear="1891" box="[843,1003,1378,1404]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schonfeldti">E. Schönfeldti</taxonomicName>
/ Reitt. / 
<collectorName id="269CEFABFF9EFFF3FBB82E35FB07FAEF" box="[1108,1249,1378,1404]" pageId="5" pageNumber="6">Coll. Reitter</collectorName>
” [white with red frame, printed and handwritten], and “ 
<collectingCountry id="F37ECAEDFF9EFFF3FDAF2ED1FD6EFA33" box="[579,648,1414,1440]" name="Japan" pageId="5" pageNumber="6">Japán</collectingCountry>
” [white, printed] (
<collectionCode id="ED7812B8FF9EFFF3FCB92ED1FC57FA33" box="[853,945,1414,1440]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="6">HNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD68A7DFF9EFFF0FF2B2EFEFDFCFC33" blockId="5.[151,1437,621,2016]" lastBlockId="6.[151,1437,151,2037]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B91D566FFF9EFFF3FF2B2EFEFDF6FA50" bold="true" box="[199,528,1449,1475]" pageId="5" pageNumber="6">Additional material studied</emphasis>
(123 exx): 
<emphasis id="B91D566FFF9EFFF3FD7C2EFEFC71FA50" bold="true" box="[656,919,1449,1475]" pageId="5" pageNumber="6">
HONSHU: 
<collectingRegion id="49AD449FFF9EFFF3FCFB2EFEFC71FA50" box="[791,919,1449,1475]" country="Japan" name="Aichi" pageId="5" pageNumber="6">Aichi Pref.</collectingRegion>
</emphasis>
: 4 exx, Misawa-chô, Seto City, 
<date id="FFD7ACBDFF9EFFF3FB172EFDFA9AFA57" box="[1275,1404,1450,1476]" pageId="5" pageNumber="6" value="1999-10-07">10.07.1999</date>
, T. Mizusawa leg. (cPJ); 
<emphasis id="B91D566FFF9EFFF3FE602E9AFDF1FA74" bold="true" box="[396,535,1485,1511]" pageId="5" pageNumber="6">
<collectingRegion id="49AD449FFF9EFFF3FE602E9AFDF1FA74" box="[396,535,1485,1511]" country="Japan" name="Chiba" pageId="5" pageNumber="6">Chiba Pref.</collectingRegion>
</emphasis>
: 2 exx., Okuno, Ichihara City, 16– 
<date id="FFD7ACBDFF9EFFF3FC732E99FBC7FA7B" box="[927,1057,1486,1512]" pageId="5" pageNumber="6" value="2001-04-18">18.04.2001</date>
, T. Shimada &amp; S. Hatsushiba leg. (cPJ); 
<emphasis id="B91D566FFF9EFFF3FF0C2EA6FE4FF998" bold="true" box="[224,425,1521,1547]" pageId="5" pageNumber="6">
<collectingRegion id="49AD449FFF9EFFF3FF0C2EA6FE4FF998" box="[224,425,1521,1547]" country="Japan" name="Fukushima" pageId="5" pageNumber="6">Fukushima Pref.</collectingRegion>
</emphasis>
: 
<specimenCount id="9D6F41F4FF9EFFF3FE542EA5FE14F99F" box="[440,498,1522,1548]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Hibara, Kitashiobara V., 
<date id="FFD7ACBDFF9EFFF3FCF92EA5FC70F99F" box="[789,918,1522,1548]" pageId="5" pageNumber="6" value="2003-10-26">26.10.2003</date>
, T. Watanabe leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FB422EA5FB01F99F" box="[1198,1255,1522,1548]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Koishigahama, Aizuwakamatsu City, 30.05– 
<date id="FFD7ACBDFF9EFFF3FE352D41FDBCF9A2" box="[473,602,1558,1585]" pageId="5" pageNumber="6" value="1999-01-06">01.06.1999</date>
, T. Shimada leg. (cPJ); 
<emphasis id="B91D566FFF9EFFF3FCB12D42FC11F9BC" bold="true" box="[861,1015,1557,1583]" pageId="5" pageNumber="6">
<collectingRegion id="49AD449FFF9EFFF3FCB12D42FC11F9BC" box="[861,1015,1557,1583]" country="Japan" name="Gunma" pageId="5" pageNumber="6">Gunma Pref.</collectingRegion>
</emphasis>
: 3 exx, Hôshi Spa, Niiharu-mura, 
<quantity id="4C912798FF9EFFF3FA9E2D41FF49F9C7" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="m" value="600.0">600 m</quantity>
, 
<date id="FFD7ACBDFF9EFFF3FF562D6DFEDDF9C7" box="[186,315,1594,1620]" pageId="5" pageNumber="6" value="2001-10-20">20.10.2001</date>
, P. Jałoszyński leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FDB12D6DFD73F9C7" box="[605,661,1594,1620]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Akaya-goe, Minakami-machi, 
<quantity id="4C912798FF9EFFF3FC182D6DFBDAF9C7" box="[1012,1084,1594,1621]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="m" value="550.0">550 m</quantity>
, 
<date id="FFD7ACBDFF9EFFF3FBAB2D6DFB2EF9C7" box="[1095,1224,1594,1620]" pageId="5" pageNumber="6" value="2001-10-20">20.10.2001</date>
, P. Jałoszyński leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FF0C2D09FEFCF9EB" box="[224,282,1630,1656]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Mt. Asamakakure-yama, 
<quantity id="4C912798FF9EFFF3FDAF2D09FD7DF9EB" box="[579,667,1630,1657]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="6" unit="m" value="1450.0">1450 m</quantity>
, Kurabuchi vill., 
<date id="FFD7ACBDFF9EFFF3FC8D2D09FC07F9EB" box="[865,993,1630,1656]" pageId="5" pageNumber="6" value="2002-11-10">11.10.2002</date>
, S. Arai leg. (cPJ); 
<emphasis id="B91D566FFF9EFFF3FB522D0AFA99F9E4" bold="true" box="[1214,1407,1629,1655]" pageId="5" pageNumber="6">
<collectingRegion id="49AD449FFF9EFFF3FB522D0AFA99F9E4" box="[1214,1407,1629,1655]" country="Japan" name="Hiroshima" pageId="5" pageNumber="6">Hiroshima Pref.</collectingRegion>
</emphasis>
: 9 exx., near Mitaka Dam, Nishinômi-jima, 
<date id="FFD7ACBDFF9EFFF3FD8D2DD5FD07F90E" box="[609,737,1666,1693]" pageId="5" pageNumber="6" value="2002-08-25">25.08.2002</date>
, S. Arai leg. (cPJ); 2 exx., Mt. Hôdai-yama, 
<quantity id="4C912798FF9EFFF3FB262DD5FAF4F90F" box="[1226,1298,1666,1692]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="m" value="400.0">400 m</quantity>
, Nishinômijima, 
<date id="FFD7ACBDFF9EFFF3FF342DF1FEBFF952" box="[216,345,1702,1729]" pageId="5" pageNumber="6" value="2002-08-25">25.08.2002</date>
, S. Arai leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FDD42DF1FD94F953" box="[568,626,1702,1728]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Miyama-no-taki, 
<quantity id="4C912798FF9EFFF3FCA92DF1FC69F953" box="[837,911,1702,1728]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="6" unit="m" value="240.0">240 m</quantity>
, Kure City, 22– 
<date id="FFD7ACBDFF9EFFF3FBA82DF1FB23F953" box="[1092,1221,1702,1728]" pageId="5" pageNumber="6" value="2002-08-23">23.08.2002</date>
, S. Arai leg. (cPJ); 
<emphasis id="B91D566FFF9EFFF3FF7B2D9EFECEF970" bold="true" box="[151,296,1737,1764]" pageId="5" pageNumber="6">
<collectingRegion id="49AD449FFF9EFFF3FF7B2D9EFECEF970" box="[151,296,1737,1764]" country="Japan" name="Hyogo" pageId="5" pageNumber="6">Hyôgo Pref.</collectingRegion>
</emphasis>
: 5 exx, Miki City, Shizimi-machi, 
<date id="FFD7ACBDFF9EFFF3FD512D9DFCD8F977" box="[701,830,1738,1764]" pageId="5" pageNumber="6" value="2002-03-19">19.03.2002</date>
, S. Nagashima leg. (cPJ); 
<emphasis id="B91D566FFF9EFFF3FB832D9EFAEBF970" bold="true" box="[1135,1293,1737,1763]" pageId="5" pageNumber="6">
<collectingRegion id="49AD449FFF9EFFF3FB832D9EFAEBF970" box="[1135,1293,1737,1763]" country="Japan" name="Ibaraki" pageId="5" pageNumber="6">Ibaraki Pref.</collectingRegion>
</emphasis>
: 4 exx., Mt. Tsukuba, 
<quantity id="4C912798FF9EFFF3FEE82DB9FE62F89B" box="[260,388,1774,1801]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="6" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500–700 m</quantity>
, 
<date id="FFD7ACBDFF9EFFF3FE7C2DB9FDF6F89B" box="[400,528,1774,1800]" pageId="5" pageNumber="6" value="2001-02-12">02.12.2001</date>
, P. Jałoszyński leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FCDB2DB9FC89F89B" box="[823,879,1774,1800]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9EFFF3FBF02DB9FB7CF89B" box="[1052,1178,1774,1800]" pageId="5" pageNumber="6" value="2002-08-06">08.06.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FB052DB9FAC7F89B" box="[1257,1313,1774,1800]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9EFFF3FF282C45FEA4F8BF" box="[196,322,1810,1836]" pageId="5" pageNumber="6" value="2002-10-14">14.10.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FE792C45FE29F8BF" box="[405,463,1810,1836]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9EFFF3FD6F2C45FCE7F8BF" box="[643,769,1810,1836]" pageId="5" pageNumber="6" value="2003-07-19">19.07.2003</date>
(cPJ); 2 exx., same data but 
<date id="FFD7ACBDFF9EFFF3FBBD2C45FB28F8BF" box="[1105,1230,1810,1836]" pageId="5" pageNumber="6" value="2003-07-20">20.07.2003</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FACE2C45FABCF8BF" box="[1314,1370,1810,1836]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
same data but 
<date id="FFD7ACBDFF9EFFF3FF162C61FE9FF8C3" box="[250,377,1846,1872]" pageId="5" pageNumber="6" value="2006-03-06">03.06.2006</date>
(cPJ); 2 exx, Mt. Tsukuba, foot of the mountain, 
<date id="FFD7ACBDFF9EFFF3FC432C61FBD6F8C3" box="[943,1072,1846,1872]" pageId="5" pageNumber="6" value="2004-03-04">03.04.2004</date>
, P. Jałoszyński leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FAB12C61FA71F8C3" box="[1373,1431,1846,1872]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Mt. Tsukuba, 
<quantity id="4C912798FF9EFFF3FEDC2C0DFE45F8E7" box="[304,419,1882,1909]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="6" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</quantity>
, SE slope, along stream in bamboo- 
<taxonomicName id="4C69F1FEFF9EFFF3FCDE2C0DFC27F8E7" box="[818,961,1882,1908]" class="Pinopsida" family="Cupressaceae" genus="Cryptomeria" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D566FFF9EFFF3FCDE2C0DFC27F8E7" box="[818,961,1882,1908]" italics="true" pageId="5" pageNumber="6">Cryptomeria</emphasis>
</taxonomicName>
forest with 
<taxonomicName id="4C69F1FEFF9EFFF3FBAA2C0DFB60F8E7" box="[1094,1158,1882,1908]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D566FFF9EFFF3FBAA2C0DFB60F8E7" box="[1094,1158,1882,1908]" italics="true" pageId="5" pageNumber="6">Alnus</emphasis>
</taxonomicName>
trees, 
<date id="FFD7ACBDFF9EFFF3FB222C0DFAA8F8E7" box="[1230,1358,1882,1908]" pageId="5" pageNumber="6" value="2001-06-04">06.04.2001</date>
, leg. P. Jałoszyński (cPJ); 1 ex, same data but 
<date id="FFD7ACBDFF9EFFF3FDAB2C29FD20F80B" box="[583,710,1918,1944]" pageId="5" pageNumber="6" value="2002-04-23">23.04.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FCF92C29FCA8F80B" box="[789,846,1918,1944]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, Mt. Tsukuba, 
<quantity id="4C912798FF9EFFF3FC192C29FB8EF80B" box="[1013,1128,1918,1945]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="6" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</quantity>
, 
<date id="FFD7ACBDFF9EFFF3FB982C29FB12F80B" box="[1140,1268,1918,1944]" pageId="5" pageNumber="6" value="2001-07-04">07.04.2001</date>
, P. Jałoszyński leg. (cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FEE22CF5FEA2F82F" box="[270,324,1954,1980]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
same data but 
<date id="FFD7ACBDFF9EFFF3FE042CF5FD80F82E" box="[488,614,1954,1981]" pageId="5" pageNumber="6" value="2002-05-04">05.04.2002</date>
(cPJ); 5 exx, same data but 
<date id="FFD7ACBDFF9EFFF3FC772CF5FBFEF82F" box="[923,1048,1954,1980]" pageId="5" pageNumber="6" value="2002-02-11">02.11.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FB892CF5FB7BF82F" box="[1125,1181,1954,1980]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, hill near Mt. Tsukuba, bamboo forest at low alt., 
<date id="FFD7ACBDFF9EFFF3FE512C91FDD8F873" box="[445,574,1990,2016]" pageId="5" pageNumber="6" value="2001-09-29">29.09.2001</date>
, leg. P. Jałoszyński (cPJ); 2 exx, same data but 
<date id="FFD7ACBDFF9EFFF3FBBD2C91FB28F873" box="[1105,1230,1990,2016]" pageId="5" pageNumber="6" value="2001-06-10">06.10.2001</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9EFFF3FAF22C91FAB1F873" box="[1310,1367,1990,2016]" pageId="5" pageNumber="6" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9DFFF0FF162BC0FE91FF22" box="[250,375,151,177]" pageId="6" pageNumber="7" value="2001-11-10">11.10.2001</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FE242BC0FDE7FF22" box="[456,513,151,177]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same data but 11.2001 (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FCB12BC0FC71FF22" box="[861,919,151,177]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9DFFF0FBAB2BC0FB22FF22" box="[1095,1220,151,177]" pageId="6" pageNumber="7" value="2001-11-13">13.11.2001</date>
(cPJ); 3 exx, same data but 
<date id="FFD7ACBDFF9DFFF0FF102BECFE9CFF46" box="[252,378,187,213]" pageId="6" pageNumber="7" value="2002-01-13">13.01.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FE222BECFDEFFF46" box="[462,521,187,213]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9DFFF0FD2C2BECFCD8FF46" box="[704,830,187,213]" pageId="6" pageNumber="7" value="2002-01-19">19.01.2002</date>
(cPJ); 3 exx, same data but 
<date id="FFD7ACBDFF9DFFF0FB662BECFAEEFF46" box="[1162,1288,187,213]" pageId="6" pageNumber="7" value="2002-06-07">06.07.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FAB02BECFA71FF46" box="[1372,1431,187,213]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Tsukuba City env., 04.2007, leg. P. Jałoszyński (cPJ); 3 exx., suburbs of Tsukuba City, bamboo- 
<taxonomicName id="4C69F1FEFF9DFFF0FB2C2B88FAA9FF6A" box="[1216,1359,223,249]" class="Pinopsida" family="Cupressaceae" genus="Cryptomeria" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D566FFF9DFFF0FB2C2B88FAA9FF6A" box="[1216,1359,223,249]" italics="true" pageId="6" pageNumber="7">Cryptomeria</emphasis>
</taxonomicName>
forest, 
<date id="FFD7ACBDFF9DFFF0FF7B2A54FEF0FE8E" box="[151,278,259,285]" pageId="6" pageNumber="7" value="2001-10-11">10.11.2001</date>
, leg. P. Jałoszyński (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FDD22A54FD91FE8E" box="[574,631,259,285]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Shishitsuka ad Tsuchiura, lowland degenerated forest, 
<date id="FFD7ACBDFF9DFFF0FB0B2A54FA81FE8E" box="[1255,1383,259,285]" pageId="6" pageNumber="7" value="2002-09-14">14.09.2002</date>
, leg. P. Jałoszyński (cPJ); 
<emphasis id="B91D566FFF9DFFF0FE6B2A70FDAFFED3" bold="true" box="[391,585,294,321]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FE6B2A70FDAFFED3" box="[391,585,294,321]" country="Japan" name="Kanagawa" pageId="6" pageNumber="7">Kanagawa Pref.</collectingRegion>
</emphasis>
: 3 exx., Mt. Takamatsu-yama, Atsugi City, 
<date id="FFD7ACBDFF9DFFF0FBD02A70FB5AFED2" box="[1084,1212,295,321]" pageId="6" pageNumber="7" value="2001-11-04">11.04.2001</date>
, H. Mizushima leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FF0E2A1CFEFCFEF6" box="[226,282,331,357]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Sagamiko env., 
<date id="FFD7ACBDFF9DFFF0FE0F2A1DFD82FEF6" box="[483,612,330,357]" pageId="6" pageNumber="7" value="2002-05-25">25.05.2002</date>
, leg. P. Jałoszyński (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FC7B2A1CFC35FEF6" box="[919,979,331,357]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9DFFF0FB662A1CFAEEFEF6" box="[1162,1288,331,357]" pageId="6" pageNumber="7" value="2002-07-14">14.07.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FAB02A1CFA71FEF6" box="[1372,1431,331,357]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9DFFF0FED12A39FE5DFE1A" box="[317,443,366,393]" pageId="6" pageNumber="7" value="2002-05-10">05.10.2002</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FDE12A39FDA1FE1A" box="[525,583,366,393]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same data but 
<date id="FFD7ACBDFF9DFFF0FD162A39FC9EFE1B" box="[762,888,366,392]" pageId="6" pageNumber="7" value="2003-01-11">01.11.2003</date>
(cPJ); 4 exx. same data but 
<date id="FFD7ACBDFF9DFFF0FB522A39FADAFE1B" box="[1214,1340,366,392]" pageId="6" pageNumber="7" value="2003-04-19">19.04.2003</date>
(cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FA622A39FF5DFE3E" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Manazuru Penins., 
<date id="FFD7ACBDFF9DFFF0FE442AC5FDC1FE3F" box="[424,551,402,428]" pageId="6" pageNumber="7" value="2006-04-11">04.11.2006</date>
, leg. P. Jałoszyński (cPJ); 6 exx. Yokosuka-shi, Oppama-chô, 
<date id="FFD7ACBDFF9DFFF0FB152AC5FA9EFE3F" box="[1273,1400,402,428]" pageId="6" pageNumber="7" value="1998-11-08">11.08.1998</date>
, T. Mizusawa leg. (cPJ); 9 exx., same data but 
<date id="FFD7ACBDFF9DFFF0FD6F2AE1FCE7FE43" box="[643,769,438,464]" pageId="6" pageNumber="7" value="1999-12-01">12.01.1999</date>
(cPJ); 2 exx, Kamakura, hills around city, 
<date id="FFD7ACBDFF9DFFF0FB0A2AE1FA81FE43" box="[1254,1383,438,464]" pageId="6" pageNumber="7" value="2002-09-03">09.03.2002</date>
, leg. P. Jałoszyński (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FE6F2A8DFE5AFE67" box="[387,444,474,500]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Zigoku-sawa, Mt. Koma-yama, Ohiso town, 06.-5.2002, S. Arai leg. (cPJ); 
<emphasis id="B91D566FFF9DFFF0FAFB2A8DFA73FE60" bold="true" box="[1303,1429,473,500]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FAFB2A8DFA73FE60" box="[1303,1429,473,500]" country="Japan" name="Nara" pageId="6" pageNumber="7">Nara Pref.</collectingRegion>
</emphasis>
: 
<specimenCount id="9D6F41F4FF9DFFF0FF7B2AAAFF36FD8B" box="[151,208,509,536]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Mt. Obako-dake, 
<quantity id="4C912798FF9DFFF0FE4D2AAAFE1EFD8B" box="[417,504,509,536]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="m" value="1300.0">1300 m</quantity>
, near Nosegawa-mura, 
<date id="FFD7ACBDFF9DFFF0FCEC2AAAFC66FD84" box="[768,896,509,535]" pageId="6" pageNumber="7" value="2006-03-26">26.03.2006</date>
, P. Jałoszyński leg. (cPJ); 
<emphasis id="B91D566FFF9DFFF0FB4B2AAAFAAFFD84" bold="true" box="[1191,1353,509,535]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FB4B2AAAFAAFFD84" box="[1191,1353,509,535]" country="Japan" name="Saitama" pageId="6" pageNumber="7">Saitama Pref.</collectingRegion>
</emphasis>
: 3 exx, Mt. Ryogami-san, Kiyotaki-goya, 
<quantity id="4C912798FF9DFFF0FDF42976FD89FDAF" box="[536,623,545,572]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="m" value="1300.0">1300 m</quantity>
, date unknown, T. Shimada leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FBC82976FBBBFDAF" box="[1060,1117,545,572]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, 
<collectingRegion id="49AD449FFF9DFFF0FB852976FB4AFDA8" box="[1129,1196,545,571]" country="Japan" name="Shiga" pageId="6" pageNumber="7">Shiga</collectingRegion>
, Ranzan, 
<date id="FFD7ACBDFF9DFFF0FAF42976FA7EFDA8" box="[1304,1432,545,571]" pageId="6" pageNumber="7" value="2002-11-18">18.11.2002</date>
, K. Toyoda leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FE7B2912FE2EFDCC" box="[407,456,581,607]" pageId="6" pageNumber="7" type="generic">1ex.</specimenCount>
, Mt. Shioyama, Ranzan, 
<date id="FFD7ACBDFF9DFFF0FD032912FC89FDCC" box="[751,879,581,607]" pageId="6" pageNumber="7" value="2000-06-17">17.06.2000</date>
, K. Toyoda leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FB922912FB5FFDCC" box="[1150,1209,581,607]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Marugami-no-taki, Ryougami-mura, 
<date id="FFD7ACBDFF9DFFF0FEB2293EFE39FD10" box="[350,479,617,643]" pageId="6" pageNumber="7" value="1999-01-08">01.08.1999</date>
, S. Arai leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FD28293EFD19FD10" box="[708,767,617,643]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Kasuga-jinja, Tamagawa vil., 
<date id="FFD7ACBDFF9DFFF0FB8F293EFB02FD10" box="[1123,1252,617,643]" pageId="6" pageNumber="7" value="1999-02-10">02.10.1999</date>
, K. Toyoda leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FF3329DBFEF1FD34" box="[223,279,652,679]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Mt. Arima-yama, Naguri vill., 
<quantity id="4C912798FF9DFFF0FD9B29DBFD28FD34" box="[631,718,652,679]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="1200.0">1200 m</quantity>
, 
<date id="FFD7ACBDFF9DFFF0FD3529DBFCBFFD34" box="[729,857,652,679]" pageId="6" pageNumber="7" value="2004-08-05">08.05.2004</date>
, S. Arai leg. (cPJ); 
<emphasis id="B91D566FFF9DFFF0FBC329DBFB21FD35" bold="true" box="[1071,1223,652,678]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FBC329DBFB21FD35" box="[1071,1223,652,678]" country="Japan" name="Tochigi" pageId="6" pageNumber="7">Tochigi Pref.</collectingRegion>
</emphasis>
: 2 exx. Nikko City env., 
<date id="FFD7ACBDFF9DFFF0FF3829E7FEB2FD59" box="[212,340,688,714]" pageId="6" pageNumber="7" value="2006-09-23">23.09.2006</date>
, leg. P. Jałoszyński (cPJ); 
<emphasis id="B91D566FFF9DFFF0FD9229E7FCECFD59" bold="true" box="[638,778,688,714]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FD9229E7FCECFD59" box="[638,778,688,714]" country="Japan" name="Tokyo" pageId="6" pageNumber="7">Tokyo Pref.</collectingRegion>
</emphasis>
: 2 exx., Nippara Valley near Okutama, 
<date id="FFD7ACBDFF9DFFF0FB2729E7FAAAFD59" box="[1227,1356,688,714]" pageId="6" pageNumber="7" value="2003-03-08">03.08.2003</date>
, leg. P. Jałoszyński (cPJ); 
<emphasis id="B91D566FFF9DFFF0FE802983FDC1FD7E" bold="true" box="[364,551,723,750]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FE802983FDC1FD7E" box="[364,551,723,750]" country="Japan" name="Yamagata" pageId="6" pageNumber="7">Yamagata Pref.</collectingRegion>
</emphasis>
: 
<specimenCount id="9D6F41F4FF9DFFF0FDDB2983FD94FD7D" box="[567,626,724,750]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Mt. Haguro, broad-leaved primary forest, 
<date id="FFD7ACBDFF9DFFF0FB8E2983FB04FD7D" box="[1122,1250,724,750]" pageId="6" pageNumber="7" value="2010-09-19">19.09.2010</date>
, T. Lackner leg. (cPH); 
<emphasis id="B91D566FFF9DFFF0FF0729A0FDA7FC82" bold="true" box="[235,577,759,785]" pageId="6" pageNumber="7">
KYUSHU: 
<collectingRegion id="49AD449FFF9DFFF0FE9A29A0FDA7FC82" box="[374,577,759,785]" country="Japan" name="Kagoshima" pageId="6" pageNumber="7">Kagoshima Pref.</collectingRegion>
</emphasis>
: 
<specimenCount id="9D6F41F4FF9DFFF0FDBE29AFFD6BFC81" box="[594,653,760,786]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Ohsumi Lake, Kanoya City, Ohsumi Pen., 
<date id="FFD7ACBDFF9DFFF0FB6629AFFAEDFC81" box="[1162,1291,760,786]" pageId="6" pageNumber="7" value="2001-08-24">24.08.2001</date>
, S. Arai leg. (cPJ); 
<specimenCount id="9D6F41F4FF9DFFF0FF0C284CFEFFFCA5" box="[224,281,795,822]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Kirishima-jingu, 
<quantity id="4C912798FF9DFFF0FE0B284CFDD7FCA5" box="[487,561,795,822]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="7" unit="m" value="450.0">450 m</quantity>
, Makizono town, 
<date id="FFD7ACBDFF9DFFF0FD11284CFC98FCA6" box="[765,894,795,821]" pageId="6" pageNumber="7" value="2001-08-27">27.08.2001</date>
, S. Arai leg. (cPJ); 
<emphasis id="B91D566FFF9DFFF0FBB5284CFA99FCA6" bold="true" box="[1113,1407,795,821]" pageId="6" pageNumber="7">
SHIKOKU: 
<collectingRegion id="49AD449FFF9DFFF0FB1C284CFA99FCA6" box="[1264,1407,795,821]" country="Japan" name="Ehime" pageId="6" pageNumber="7">Ehime Pref.</collectingRegion>
</emphasis>
: 
<specimenCount id="9D6F41F4FF9DFFF0FA62284CFF5DFCCA" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Saragamine, Toon-shi, 100-years old dec. mountain forest, 
<date id="FFD7ACBDFF9DFFF0FCBE2868FC35FCC9" box="[850,979,831,858]" pageId="6" pageNumber="7" value="2018-05-16">16.05.2018</date>
, leg. P. Jałoszyński (cPJ); 
<emphasis id="B91D566FFF9DFFF0FB1F2868FA73FCCA" bold="true" box="[1267,1429,831,857]" pageId="6" pageNumber="7">
<collectingRegion id="49AD449FFF9DFFF0FB1F2868FA73FCCA" box="[1267,1429,831,857]" country="Japan" name="Kagawa" pageId="6" pageNumber="7">Kagawa Pref.</collectingRegion>
</emphasis>
: 2 exx., Nagara-dam, Ayakami-chô, 
<date id="FFD7ACBDFF9DFFF0FDC42835FD4FFCEE" box="[552,681,866,893]" pageId="6" pageNumber="7" value="2003-05-05">05.05.2003</date>
, S. Nagashima leg. (cPJ). Additionally, specimens from 
<collectingRegion id="49AD449FFF9DFFF0FAC42834FA7BFCEE" box="[1320,1437,867,893]" country="Japan" name="Saga" pageId="6" pageNumber="7">Saga Pref.</collectingRegion>
(Kyushu) were also seen (
<collectionCode id="ED7812B8FF9DFFF0FE5628D1FDE8FC33" box="[442,526,902,928]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="7">NSMT</collectionCode>
).
</paragraph>
<paragraph id="8BD68A7DFF9DFFF0FF2B28FDFAC6FB70" blockId="6.[151,1437,151,2037]" pageId="6" pageNumber="7">
<emphasis id="B91D566FFF9DFFF0FF2B28FDFE5EFC57" bold="true" box="[199,440,938,964]" pageId="6" pageNumber="7">Emended diagnosis.</emphasis>
Body middle-sized (BL ~ 
<quantity id="4C912798FF9DFFF0FD0A28FDFC92FC56" box="[742,884,938,965]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="6" pageNumber="7" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
), thick bristles distributed on tempora and sides of pronotum; antennal club tetramerous, sharply delimited from compactly assembled proximal portion of funicle; vertex bulging posterodorsad, with rounded posterior margin; tempora in dorsal view much longer than eyes; pronotum subconical, broadest shortly in front of base, with weakly rounded sides strongly converging anterad; anterior pronotal margin much shorter than posterior margin; pronotum with two pairs of variously distinct small antebasal pits, lacking transverse groove, with sublateral carinae barely discernible or lacking; elytra densely setose; male legs and abdomen unmodified; aedeagus in lateral view with apical projections directed dorsad or distodorsad, thin-walled and stout median lobe, and endophallic structures situated in distal half of median lobe, darkly sclerotized, symmetrical, with lateral pair of strongly elongate structures forming a broad letter “V” in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF9DFFFEFF2B2FBCFB98FDF2" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BD68A7DFF9DFFF0FF2B2FBCFADFFABA" blockId="6.[151,1437,151,2037]" pageId="6" pageNumber="7">
<emphasis id="B91D566FFF9DFFF0FF2B2FBCFE93FA96" bold="true" box="[199,373,1259,1285]" pageId="6" pageNumber="7">Redescription.</emphasis>
Body of male (
<figureCitation id="135296F8FF9DFFF0FDC72FBBFD62FA95" box="[555,644,1260,1286]" captionStart="FIGURES 13–18" captionStartId="8.[151,264,1829,1853]" captionTargetBox="[160,1428,680,1800]" captionTargetId="figure-238@8.[151,1436,669,1806]" captionTargetPageId="8" captionText="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." figureDoi="http://doi.org/10.5281/zenodo.5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="6" pageNumber="7">Figs 13</figureCitation>
, 
<figureCitation id="135296F8FF9DFFF0FD7E2FBBFD56FA95" box="[658,688,1260,1286]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="6" pageNumber="7">19</figureCitation>
, 
<figureCitation id="135296F8FF9DFFF0FD522FBCFD3AFA95" box="[702,732,1259,1286]" captionStart="FIGURES 25–30" captionStartId="10.[151,264,1931,1955]" captionTargetBox="[160,1420,680,1899]" captionTargetId="figure-269@10.[151,1436,669,1908]" captionTargetPageId="10" captionText="FIGURES 25–30. Euconnus fustiger, male. Dorsal habitus (25) and historical labels (26) of lectotype male of Euconnus schoenfeldti; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views." figureDoi="http://doi.org/10.5281/zenodo.5901022" httpUri="https://zenodo.org/record/5901022/files/figure.png" pageId="6" pageNumber="7">25</figureCitation>
, 
<figureCitation id="135296F8FF9DFFF0FD062FBBFCE1FA95" box="[746,775,1260,1286]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="6" pageNumber="7">31</figureCitation>
) moderately slender, strongly convex, BL 
<quantity id="4C912798FF9DFFF0FB1C2FBCFA7EFA95" box="[1264,1432,1259,1286]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" metricValueMax="1.53" metricValueMin="1.33" pageId="6" pageNumber="7" unit="mm" value="1.43" valueMax="1.53" valueMin="1.33">1.33–1.53 mm</quantity>
; cuticle glossy, pigmentation variable, from light to very dark brown, vestiture of setae lighter than cuticle.
</paragraph>
<paragraph id="8BD68A7DFF9DFFF0FF2B2E64FF39F945" blockId="6.[151,1437,151,2037]" pageId="6" pageNumber="7">
Head rhomboidal, distinctly elongate and broadest at eyes, HL 
<quantity id="4C912798FF9DFFF0FC672E64FBD6FADD" box="[907,1072,1331,1358]" metricMagnitude="-4" metricUnit="m" metricValue="3.2499999999999996" metricValueMax="3.5" metricValueMin="3.0" pageId="6" pageNumber="7" unit="mm" value="0.32499999999999996" valueMax="0.35" valueMin="0.3">0.30–0.35 mm</quantity>
, HW 
<quantity id="4C912798FF9DFFF0FB9E2E64FAFEFADD" box="[1138,1304,1331,1358]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" pageNumber="7" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.30 mm</quantity>
; tempora in dorsal view about three times as long as eyes and strongly converging posterad; vertex and frons confluent, weakly and evenly convex, posterior margin of vertex rounded, slightly bulging posterodorsad; supraantennal tubercles barely marked; frons between antennae steeply declining; clypeus unmodified. Eyes moderately large, finely faceted, distinctly but not strongly projecting laterad from the head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, fine and sparse; setae short and sparse, suberect, tempora and posterior margin of vertex densely covered with thick bristles directed laterocaudad. Antennae moderately slender, with compact proximal portion of funicle and moderately loosely assembled, strongly broadened and sharply delimited tetramerous club, AnL 
<quantity id="4C912798FF9DFFF0FF3D2D06FE91F9FF" box="[209,375,1617,1644]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.8" metricValueMin="5.0" pageId="6" pageNumber="7" unit="mm" value="0.54" valueMax="0.58" valueMin="0.5">0.50–0.58 mm</quantity>
; antennomeres 1 and 2 each elongate, 3–6 each about as long as broad or indistinctly elongate, 7 slightly narrower than 6, indistinctly elongate, 8 much broader than 7, slightly transverse, 9 and 10 each strongly transverse and indistinctly broader, but distinctly shorter than 8, 11 indistinctly narrower than 10, about as long as broad.
</paragraph>
<paragraph id="8BD68A7DFF9DFFF0FF2B2DB6FCA6F866" blockId="6.[151,1437,151,2037]" pageId="6" pageNumber="7">
Pronotum subconical with weakly rounded sides, broadest just in front of base, PL 
<quantity id="4C912798FF9DFFF0FBBD2DB7FB12F968" box="[1105,1268,1760,1787]" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="6" pageNumber="7" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.33–0.38 mm</quantity>
, PW 
<quantity id="4C912798FF9DFFF0FAC02DB7FF20F88D" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="6" pageNumber="7" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.33–0.38 mm</quantity>
; anterior margin nearly straight and much shorter than posterior margin, anterior corners weakly marked, sides of pronotum weakly, evenly rounded and strongly converging anterad; posterior corners distinctly obtuse-angled, well-marked; posterior margin nearly straight. Pronotal base with two lateral pairs of tiny and shallow pits showing some variability among studied specimens, with a tendency toward reduction (all pits can be barely discernible or distinct, and inner or outer pair may be barely discernible), transverse groove lacking, sublateral carinae indistinct or lacking. Disc dorsally covered with fine, inconspicuous punctures and moderately dense thin suberect setae, sides with dense thick bristles that obscure the shape of pronotum.
</paragraph>
<paragraph id="8BD68A7DFF9CFFF1FF2B2BC0FAE7FE8E" blockId="7.[151,1437,150,357]" pageId="7" pageNumber="8">
Elytra oval, broadest slightly in front of middle, EL 
<quantity id="4C912798FF9CFFF1FCCD2BC0FC2EFF22" box="[801,968,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="7" pageNumber="8" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.70–0.80 mm</quantity>
, EW 
<quantity id="4C912798FF9CFFF1FBE62BC1FB57FF22" box="[1034,1201,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.3" metricValueMin="5.3" pageId="7" pageNumber="8" unit="mm" value="0.58" valueMax="0.63" valueMin="0.53">0.53–0.63 mm</quantity>
, EI 1.21–1.37; basal impressions short and shallow, humeral calli moderately strongly elevated, elongate, each delimited from adscutellar region by shallow elongate impression running posterolaterad, elytral apices separately rounded. Punctures on elytral disc fine, inconspicuous; setae long, moderately dense, suberect. Hind wings long, functional.
</paragraph>
<paragraph id="8BD68A7DFF9CFFF1FF2B2A70FD3EFED2" blockId="7.[151,1437,150,357]" box="[199,728,295,321]" pageId="7" pageNumber="8">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="8BD68A7DFF9CFFF1FF2B2A1CFE20FEF6" blockId="7.[151,1437,150,357]" box="[199,454,331,357]" pageId="7" pageNumber="8">Abdomen unmodified.</paragraph>
<caption id="DF16DAF5FF9CFFF1FF7B2D8FFE31F8AB" ID-DOI="http://doi.org/10.5281/zenodo.5901012" ID-Zenodo-Dep="5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,264,1752,1776]" targetBox="[163,1428,389,1726]" targetPageId="7">
<paragraph id="8BD68A7DFF9CFFF1FF7B2D8FFE31F8AB" blockId="7.[151,1437,1752,1848]" pageId="7" pageNumber="8">
<emphasis id="B91D566FFF9CFFF1FF7B2D8FFEAFF963" bold="true" box="[151,329,1752,1776]" pageId="7" pageNumber="8">FIGURES 9–12.</emphasis>
<taxonomicName id="4C69F1FEFF9CFFF1FEBD2D8EFDECF963" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[337,522,1752,1776]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF9CFFF1FEBD2D8EFDECF963" box="[337,522,1752,1776]" italics="true" pageId="7" pageNumber="8">
Euconnus 
<emphasis id="B91D566FFF9CFFF1FE512D8FFDECF963" box="[445,522,1752,1776]" pageId="7" pageNumber="8">fustiger</emphasis>
</emphasis>
</taxonomicName>
, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of 
<taxonomicName id="4C69F1FEFF9CFFF1FEFE2DABFE05F887" authorityName="Sharp" authorityYear="1874" box="[274,483,1788,1812]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF9CFFF1FEFE2DABFE05F887" box="[274,483,1788,1812]" italics="true" pageId="7" pageNumber="8">
Scydmaenus 
<emphasis id="B91D566FFF9CFFF1FE7A2DABFE05F887" box="[406,483,1788,1812]" pageId="7" pageNumber="8">fustiger</emphasis>
</emphasis>
</taxonomicName>
, compared to paralectotype female of 
<taxonomicName id="4C69F1FEFF9CFFF1FC872DAAFBF1F887" authorityName="Sharp" authorityYear="1886" box="[875,1047,1788,1812]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF9CFFF1FC872DAAFBF1F887" box="[875,1047,1788,1812]" italics="true" pageId="7" pageNumber="8">Euconnus lewisii</emphasis>
</taxonomicName>
(11) with its annotated mounting card and set of historical labels (12).
</paragraph>
</caption>
<paragraph id="8BD68A7DFF9CFFFEFF2B2C31FE34FE66" blockId="7.[151,1437,1894,2029]" lastBlockId="8.[151,1437,151,645]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Aedeagus (
<figureCitation id="135296F8FF9CFFF1FEAF2C31FE24F812" box="[323,450,1894,1921]" captionStart="FIGURES 13–18" captionStartId="8.[151,264,1829,1853]" captionTargetBox="[160,1428,680,1800]" captionTargetId="figure-238@8.[151,1436,669,1806]" captionTargetPageId="8" captionText="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." figureDoi="http://doi.org/10.5281/zenodo.5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="7" pageNumber="8">Figs 15–18</figureCitation>
, 
<figureCitation id="135296F8FF9CFFF1FE202C31FDF3F813" box="[460,533,1894,1920]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="7" pageNumber="8">21–24</figureCitation>
, 
<figureCitation id="135296F8FF9CFFF1FDF32C31FD8EF813" box="[543,616,1894,1920]" captionStart="FIGURES 25–30" captionStartId="10.[151,264,1931,1955]" captionTargetBox="[160,1420,680,1899]" captionTargetId="figure-269@10.[151,1436,669,1908]" captionTargetPageId="10" captionText="FIGURES 25–30. Euconnus fustiger, male. Dorsal habitus (25) and historical labels (26) of lectotype male of Euconnus schoenfeldti; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views." figureDoi="http://doi.org/10.5281/zenodo.5901022" httpUri="https://zenodo.org/record/5901022/files/figure.png" pageId="7" pageNumber="8">27–30</figureCitation>
, 
<figureCitation id="135296F8FF9CFFF1FD9D2C31FD5FF813" box="[625,697,1894,1920]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="7" pageNumber="8">32–48</figureCitation>
) stout, AeL 
<quantity id="4C912798FF9CFFF1FCD22C31FC04F812" box="[830,994,1894,1921]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="7" pageNumber="8" unit="mm" value="0.2" valueMax="0.25" valueMin="0.15">0.15–0.25 mm</quantity>
; median lobe thin-walled and extremely prone to distortions during preparation, in ventral view broadest in sub-basal region, ventral apical plate lacking distal projection; distolateral regions of median lobe in ventral view variable, from rounded and inconspicuous to forming a pair of variously shaped, elongate lobes projecting distad and bearing tiny setae (in many studied specimens setae were broken off during preparation and only setal sockets could be recognized), in lateral view distal region of median lobe forming variously shaped, often fin-like distodorsal projection; endophallic structures symmetrical in intact aedeagi, but preparation may cause various displacements and endophallus may appear slightly asymmetrical; constant structures are a pair of long lateral sclerites forming a letter “V”, these are in fact composed of two elongate structures each, and one pair may be displaced toward middle of median lobe (as in 
<figureCitation id="135296F8FF93FFFEFBD32A70FB77FED1" box="[1087,1169,295,322]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="8" pageNumber="9">Fig. 36</figureCitation>
), also median subapical subconical structure is constant, but it can rotate and in ventral view appears as a highly variable elongate or circular object, and in some cases is indiscernible; entire endophallus can be everted (also artificially, during preparation) so that median lobe becomes easily distorted and its distal region flips ventrad (as in 
<figureCitation id="135296F8FF93FFFEFBDB2AC4FB6DFE3E" box="[1079,1163,403,429]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="8" pageNumber="9">Fig. 22</figureCitation>
). Parameres slender and short, not reaching apex of median lobe, in lateral view strongly bent and each with one apical seta. See also remarks on genital variability below.
</paragraph>
<paragraph id="8BD68A7DFF93FFFEFF2B2AA8FB98FDF2" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
Female. Externally slightly stouter than male, but otherwise indistinguishable. All females dissected during the present study are wingless. BL 
<quantity id="4C912798FF93FFFEFDF32975FD20FDAE" box="[543,710,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.445" metricValueMax="1.58" metricValueMin="1.31" pageId="8" pageNumber="9" unit="mm" value="1.445" valueMax="1.58" valueMin="1.31">1.31–1.58 mm</quantity>
; HL 
<quantity id="4C912798FF93FFFEFD102975FC44FDAE" box="[764,930,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="8" pageNumber="9" unit="mm" value="0.32999999999999996" valueMax="0.35" valueMin="0.31">0.31–0.35 mm</quantity>
, HW 
<quantity id="4C912798FF93FFFEFC0E2975FB6EFDAE" box="[994,1160,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="8" pageNumber="9" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.30 mm</quantity>
, AnL 
<quantity id="4C912798FF93FFFEFB272974FA97FDAE" box="[1227,1393,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="6.0" metricValueMin="4.8" pageId="8" pageNumber="9" unit="mm" value="0.54" valueMax="0.6" valueMin="0.48">0.48–0.60 mm</quantity>
; PL 
<quantity id="4C912798FF93FFFEFF7B2910FEDAFDF2" box="[151,316,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="8" pageNumber="9" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.33–0.38 mm</quantity>
, PW 
<quantity id="4C912798FF93FFFEFE942910FDF8FDF2" box="[376,542,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="8" pageNumber="9" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.33–0.38 mm</quantity>
; EL 
<quantity id="4C912798FF93FFFEFDBE2910FD11FDF2" box="[594,759,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="7.55" metricValueMax="8.299999999999999" metricValueMin="6.8" pageId="8" pageNumber="9" unit="mm" value="0.755" valueMax="0.83" valueMin="0.68">0.68–0.83 mm</quantity>
, EW 
<quantity id="4C912798FF93FFFEFCD82911FC3FFDF2" box="[820,985,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="6.05" metricValueMax="6.8" metricValueMin="5.3" pageId="8" pageNumber="9" unit="mm" value="0.605" valueMax="0.68" valueMin="0.53">0.53–0.68 mm</quantity>
, EI 1.16–1.33.
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF93FFFEFF2B293DFD03FD16" box="[199,741,618,645]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BD68A7DFF93FFFEFF2B293DFD03FD16" blockId="8.[151,1437,151,645]" box="[199,741,618,645]" pageId="8" pageNumber="9">
<emphasis id="B91D566FFF93FFFEFF2B293DFEBFFD17" bold="true" box="[199,345,618,644]" pageId="8" pageNumber="9">Distribution</emphasis>
. 
<collectingCountry id="F37ECAEDFF93FFFEFE8B293CFE4DFD16" box="[359,427,619,645]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
: Honshu, Kyushu, Shikoku.
</paragraph>
</subSubSection>
<caption id="DF16DAF5FF93FFFEFF7B2C72FB4FF8F1" ID-DOI="http://doi.org/10.5281/zenodo.5901014" ID-Zenodo-Dep="5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,264,1829,1853]" targetBox="[160,1428,680,1800]" targetPageId="8">
<paragraph id="8BD68A7DFF93FFFEFF7B2C72FB4FF8F1" blockId="8.[151,1436,1829,1890]" pageId="8" pageNumber="9">
<emphasis id="B91D566FFF93FFFEFF7B2C72FEB1F8AD" bold="true" box="[151,343,1829,1854]" pageId="8" pageNumber="9">FIGURES 13–18.</emphasis>
<taxonomicName id="4C69F1FEFF93FFFEFEB32C71FDFFF8AD" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[351,537,1830,1854]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF93FFFEFEB32C71FDFFF8AD" box="[351,537,1830,1854]" italics="true" pageId="8" pageNumber="9">
Euconnus 
<emphasis id="B91D566FFF93FFFEFE202C71FDFFF8AD" box="[460,537,1830,1854]" pageId="8" pageNumber="9">fustiger</emphasis>
</emphasis>
</taxonomicName>
, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of 
<taxonomicName id="4C69F1FEFF93FFFEFEA12C1DFE1CF8F1" authorityName="Sharp" authorityYear="1886" box="[333,506,1866,1890]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF93FFFEFEA12C1DFE1CF8F1" box="[333,506,1866,1890]" italics="true" pageId="8" pageNumber="9">Euconnus lewisii</emphasis>
</taxonomicName>
; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views.
</paragraph>
</caption>
<caption id="DF16DAF5FF92FFFFFF7B2D72FC3EF9F2" ID-DOI="http://doi.org/10.5281/zenodo.5901018" ID-Zenodo-Dep="5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1573,1597]" targetBox="[155,1426,193,1538]" targetPageId="9">
<paragraph id="8BD68A7DFF92FFFFFF7B2D72FC3EF9F2" blockId="9.[151,1436,1573,1633]" pageId="9" pageNumber="10">
<emphasis id="B91D566FFF92FFFFFF7B2D72FEBEF9AE" bold="true" box="[151,344,1573,1597]" pageId="9" pageNumber="10">FIGURES 19–24.</emphasis>
<taxonomicName id="4C69F1FEFF92FFFFFE8F2D71FDF8F9AE" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[355,542,1573,1597]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF92FFFFFE8F2D71FDF8F9AE" box="[355,542,1573,1597]" italics="true" pageId="9" pageNumber="10">
Euconnus 
<emphasis id="B91D566FFF92FFFFFE3D2D72FDF8F9AE" box="[465,542,1573,1597]" pageId="9" pageNumber="10">fustiger</emphasis>
</emphasis>
</taxonomicName>
, male. Dorsal habitus (19) and historical labels (20) of holotype male of 
<taxonomicName id="4C69F1FEFF92FFFFFAD42D71FECEF9F2" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF92FFFFFAD42D71FECEF9F2" italics="true" pageId="9" pageNumber="10">Euconnus miyawakianus</emphasis>
</taxonomicName>
; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views.
</paragraph>
</caption>
<subSubSection id="C373D9F6FF92FFFDFF2B2DC7FD3EF82A" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BD68A7DFF92FFFCFF2B2DC7FCD8FD8A" blockId="9.[151,1436,1679,2030]" lastBlockId="10.[151,1437,150,645]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B91D566FFF92FFFFFF2B2DC7FEDDF939" bold="true" box="[199,315,1680,1706]" pageId="9" pageNumber="10">Remarks.</emphasis>
<bibRefCitation id="EFF8F78CFF92FFFFFEAD2DD8FE1AF939" author="Hoshina, H." box="[321,508,1679,1706]" pageId="9" pageNumber="10" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina (2019b)</bibRefCitation>
redescribed this species and in the “Specimens examined” section listed: “ 
<typeStatus id="54D234DFFF92FFFFFADD2DD8FA7AF93A" box="[1329,1436,1679,1705]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF92FFFFFF5B2DE4FE46F95D" authorityName="Sharp" authorityYear="1874" box="[183,416,1715,1742]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF92FFFFFF5B2DE4FE46F95D" box="[183,416,1715,1742]" italics="true" pageId="9" pageNumber="10">
Scydmaenus 
<emphasis id="B91D566FFF92FFFFFEA62DE3FE46F95D" box="[330,416,1716,1742]" pageId="9" pageNumber="10">fustiger</emphasis>
</emphasis>
</taxonomicName>
, 
<specimenCount id="9D6F41F4FF92FFFFFE5C2DE3FE21F95D" box="[432,455,1716,1742]" pageId="9" pageNumber="10" type="male">♂</specimenCount>
, Nagasaki (preserved in Natural History Museum, London (...) 
<typeStatus id="54D234DFFF92FFFFFB722DE3FAE3F95D" box="[1182,1285,1716,1742]" pageId="9" pageNumber="10" type="syntype">Syntypes</typeStatus>
of 
<taxonomicName id="4C69F1FEFF92FFFFFAC22DE3FF06F962" authorityName="Sharp" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF92FFFFFAC22DE3FF06F962" italics="true" pageId="9" pageNumber="10">Euconnus lewisii</emphasis>
</taxonomicName>
, 
<specimenCount id="9D6F41F4FF92FFFFFF032D8FFECCF961" box="[239,298,1752,1778]" pageId="9" pageNumber="10" type="male">2♂♂</specimenCount>
, Nagasaki”. The 
<typeStatus id="54D234DFFF92FFFFFE012D80FDA9F962" box="[493,591,1751,1777]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF92FFFFFD9A2D80FCB8F961" authorityName="Sharp" authorityYear="1874" box="[630,862,1751,1778]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF92FFFFFD9A2D80FCB8F961" box="[630,862,1751,1778]" italics="true" pageId="9" pageNumber="10">
Scydmaenus 
<emphasis id="B91D566FFF92FFFFFCE42D8FFCB8F961" box="[776,862,1752,1778]" pageId="9" pageNumber="10">fustiger</emphasis>
</emphasis>
</taxonomicName>
preserved at NHM is a specimen of unknown sex, which has been dissected in the past (by Hoshina?), and the abdomen has been damaged. The specimen is not accompanied by any genital preparation. Its stout elytra suggest that it may in fact be a female, as it is similar to females included in the type series of 
<taxonomicName id="4C69F1FEFF92FFFFFDA62C13FD52F8CE" authorityName="Sharp" authorityYear="1886" box="[586,692,1859,1885]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF92FFFFFDA62C13FD52F8CE" box="[586,692,1859,1885]" italics="true" pageId="9" pageNumber="10">E. lewisii</emphasis>
</taxonomicName>
(compare 
<figureCitation id="135296F8FF92FFFFFCC32C14FC2EF8CD" box="[815,968,1859,1886]" captionStart="FIGURES 9–12" captionStartId="7.[151,264,1752,1776]" captionTargetBox="[163,1428,389,1726]" captionTargetId="figure-80@7.[151,1436,381,1729]" captionTargetPageId="7" captionText="FIGURES 9–12. Euconnus fustiger, presumable female (9) and female (11). Dorsal habitus (9) and historical labels (10) of holotype of Scydmaenus fustiger, compared to paralectotype female of Euconnus lewisii (11) with its annotated mounting card and set of historical labels (12)." figureDoi="http://doi.org/10.5281/zenodo.5901012" httpUri="https://zenodo.org/record/5901012/files/figure.png" pageId="9" pageNumber="10">Fig. 9 and 11</figureCitation>
), but this is not possible to prove by any existing evidence. As for the 
<typeStatus id="54D234DFFF92FFFFFE372C3FFDDBF811" box="[475,573,1896,1922]" pageId="9" pageNumber="10" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C69F1FEFF92FFFFFD8E2C3FFD2FF812" authorityName="Sharp" authorityYear="1886" box="[610,713,1895,1921]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF92FFFFFD8E2C3FFD2FF812" box="[610,713,1895,1921]" italics="true" pageId="9" pageNumber="10">E. lewisii</emphasis>
</taxonomicName>
, at NHM there are 
<specimenCount id="9D6F41F4FF92FFFFFC4D2C30FBA9F812" box="[929,1103,1895,1921]" pageId="9" pageNumber="10" type="generic">four specimens</specimenCount>
: 
<specimenCount id="9D6F41F4FF92FFFFFBB52C3FFB2FF812" box="[1113,1225,1895,1922]" pageId="9" pageNumber="10" type="female">2 females</specimenCount>
; 
<specimenCount id="9D6F41F4FF92FFFFFB3F2C3FFA8AF812" box="[1235,1388,1895,1922]" pageId="9" pageNumber="10" type="generic">one specimen</specimenCount>
that has been remounted and at least an attempt at dissection has been made, judging from a condition of the abdomen, but this specimen is not accompanied by a genital preparation; and 
<specimenCount id="9D6F41F4FF92FFFFFC6C2CE7FC03F85A" box="[896,997,1967,1994]" pageId="9" pageNumber="10" type="male">one male</specimenCount>
(designated here as 
<typeStatus id="54D234DFFF92FFFFFB2B2CF8FAD7F85A" box="[1223,1329,1967,1993]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
; 
<figureCitation id="135296F8FF92FFFFFAD02CF8FF52F87D" captionStart="FIGURES 13–18" captionStartId="8.[151,264,1829,1853]" captionTargetBox="[160,1428,680,1800]" captionTargetId="figure-238@8.[151,1436,669,1806]" captionTargetPageId="8" captionText="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." figureDoi="http://doi.org/10.5281/zenodo.5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="9" pageNumber="10">Figs 13‒ 18</figureCitation>
) with a genital preparation attached. The aedeagus of the latter is illustrated in the present paper (
<figureCitation id="135296F8FF92FFFFFB0D2C84FA87F87D" box="[1249,1377,2003,2030]" captionStart="FIGURES 13–18" captionStartId="8.[151,264,1829,1853]" captionTargetBox="[160,1428,680,1800]" captionTargetId="figure-238@8.[151,1436,669,1806]" captionTargetPageId="8" captionText="FIGURES 13–18. Euconnus fustiger, male. Dorsal habitus (13) and annotated mounting card with historical labels (14) of lectotype male of Euconnus lewisii; aedeagus (distorted!) in ventral (15, 17) and lateral (16, 18) views." figureDoi="http://doi.org/10.5281/zenodo.5901014" httpUri="https://zenodo.org/record/5901014/files/figure.png" pageId="9" pageNumber="10">Figs 15‒18</figureCitation>
), and it differs in some details from the one illustrated by 
<bibRefCitation id="EFF8F78CFF91FFFCFD332BC0FC71FF22" author="Hoshina, H." box="[735,919,151,177]" pageId="10" pageNumber="11" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina (2019b</bibRefCitation>
: figs 3‒5) for 
<taxonomicName id="4C69F1FEFF91FFFCFBD92BC0FB4CFF22" authorityName="Sharp" authorityYear="1874" box="[1077,1194,151,177]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFBD92BC0FB4CFF22" box="[1077,1194,151,177]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFBB82BC0FB4CFF22" box="[1108,1194,151,177]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
. However, I do agree with 
<bibRefCitation id="EFF8F78CFF91FFFCFF232BECFE6CFF45" author="Hoshina, H." box="[207,394,187,214]" pageId="10" pageNumber="11" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina (2019b)</bibRefCitation>
that 
<taxonomicName id="4C69F1FEFF91FFFCFE532BECFDD5FF46" authorityName="Sharp" authorityYear="1874" box="[447,563,187,213]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFE532BECFDD5FF46" box="[447,563,187,213]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFE312BECFDD5FF46" box="[477,563,187,213]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
is a senior synonym of 
<taxonomicName id="4C69F1FEFF91FFFCFCDA2BECFC7BFF46" authorityName="Sharp" authorityYear="1886" box="[822,925,187,213]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF91FFFCFCDA2BECFC7BFF46" box="[822,925,187,213]" italics="true" pageId="10" pageNumber="11">E. lewisii</emphasis>
</taxonomicName>
, as there are no external differences that could be used to distinguish the 
<typeStatus id="54D234DFFF91FFFCFE522B88FDC6FF6A" box="[446,544,223,249]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C69F1FEFF91FFFCFDAA2B88FD5AFF6A" authorityName="Sharp" authorityYear="1874" box="[582,700,223,249]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFDAA2B88FD5AFF6A" box="[582,700,223,249]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFD8A2B88FD5AFF6A" box="[614,700,223,249]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
and the 
<typeStatus id="54D234DFFF91FFFCFCF12B88FC99FF6A" box="[797,895,223,249]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="4C69F1FEFF91FFFCFC4A2B88FBE8FF6A" authorityName="Sharp" authorityYear="1886" box="[934,1038,223,249]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF91FFFCFC4A2B88FBE8FF6A" box="[934,1038,223,249]" italics="true" pageId="10" pageNumber="11">E. lewisii</emphasis>
</taxonomicName>
. I also agree with 
<bibRefCitation id="EFF8F78CFF91FFFCFB0C2B88FA7AFF6A" author="Hoshina, H." box="[1248,1436,223,249]" pageId="10" pageNumber="11" pagination="17 - 22" refId="ref24606" refString="Hoshina, H. (2004 a) A new species and taxonomical note on the genus Euconnus (Coleoptera: Scydmaenidae) from Honshu, Japan. Memoirs of the Research and Education Center for Regional Environment, University of Fukui, 11, 17 - 22." type="journal article" year="2004">Hoshina (2004a)</bibRefCitation>
that 
<taxonomicName id="4C69F1FEFF91FFFCFF252A54FE36FE8E" authority="Franz" authorityName="Franz" box="[201,464,259,285]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF91FFFCFF252A54FE6FFE8E" box="[201,393,259,285]" italics="true" pageId="10" pageNumber="11">E. miyawakianus</emphasis>
Franz
</taxonomicName>
is identical with 
<taxonomicName id="4C69F1FEFF91FFFCFD782A54FD1AFE8E" authorityName="Sharp" authorityYear="1886" box="[660,764,259,285]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF91FFFCFD782A54FD1AFE8E" box="[660,764,259,285]" italics="true" pageId="10" pageNumber="11">E. lewisii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C69F1FEFF91FFFCFCD82A54FC4CFE8E" authorityName="Sharp" authorityYear="1874" box="[820,938,259,285]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFCD82A54FC4CFE8E" box="[820,938,259,285]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFCB82A54FC4CFE8E" box="[852,938,259,285]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
, as I have not found any external characters to justify a separate position of 
<taxonomicName id="4C69F1FEFF91FFFCFDEC2A70FD27FED2" box="[512,705,295,321]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF91FFFCFDEC2A70FD27FED2" box="[512,705,295,321]" italics="true" pageId="10" pageNumber="11">E. miyawakianus</emphasis>
</taxonomicName>
. Interestingly, 
<bibRefCitation id="EFF8F78CFF91FFFCFC872A70FC1FFED2" author="Franz, H." box="[875,1017,295,321]" pageId="10" pageNumber="11" pagination="51 - 60" refId="ref24555" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Ergebnisse einer auf Einladung der Internationalen Vereinigung fur Vegetationskunde nach Japan durchgefuhrten Forschungsreise. Entomologische Blatter fur Biologie und Systematik der Kafer, 72 (1), 51 - 60." type="journal article" year="1976">Franz (1976</bibRefCitation>
: fig. 5) illustrated the aedeagus of 
<taxonomicName id="4C69F1FEFF91FFFCFA682A70FED1FEF6" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF91FFFCFA682A70FED1FEF6" italics="true" pageId="10" pageNumber="11">E. miyawakianus</emphasis>
</taxonomicName>
as strikingly different from what is illustrated here (
<figureCitation id="135296F8FF91FFFCFC672A1CFBE9FEF6" box="[907,1039,331,357]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="10" pageNumber="11">Figs 21‒24</figureCitation>
), based on the same male 
<typeStatus id="54D234DFFF91FFFCFAD62A1CFA7AFEF6" box="[1338,1436,331,357]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(the only 
<typeStatus id="54D234DFFF91FFFCFEE42A38FE8EFE1A" box="[264,360,367,393]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
is a female). It seems that Franz’s illustration was based on the aedeagus oriented in euparal not ventrally or dorsally up, but rotated in such a way that the apical region was directed more towards observer. Remounting from euparal into 
<collectingCountry id="F37ECAEDFF91FFFCFE182AE0FDAEFE42" box="[500,584,439,465]" name="Canada" pageId="10" pageNumber="11">Canada</collectingCountry>
balsam might have also distorted the aedeagus. However, shapes of ventral and dorsal apical regions and endophallic structures in 
<taxonomicName id="4C69F1FEFF91FFFCFD1F2A8CFC81FE66" authorityName="Sharp" authorityYear="1874" box="[755,871,475,501]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFD1F2A8CFC81FE66" box="[755,871,475,501]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFCFD2A8CFC81FE66" box="[785,871,475,501]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
are highly variable and those in the type specimens of 
<taxonomicName id="4C69F1FEFF91FFFCFF5A2AA8FEF8FD8A" authorityName="Sharp" authorityYear="1886" box="[182,286,511,537]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF91FFFCFF5A2AA8FEF8FD8A" box="[182,286,511,537]" italics="true" pageId="10" pageNumber="11">E. lewisii</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C69F1FEFF91FFFCFEB82AA8FDF5FD8A" box="[340,531,511,537]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF91FFFCFEB82AA8FDF5FD8A" box="[340,531,511,537]" italics="true" pageId="10" pageNumber="11">E. miyawakianus</emphasis>
</taxonomicName>
fall within this variability.
</paragraph>
<paragraph id="8BD68A7DFF91FFFCFF2B2974FED7FD16" blockId="10.[151,1437,150,645]" pageId="10" pageNumber="11">
<taxonomicName id="4C69F1FEFF91FFFCFF2B2974FDFFFDAE" authority="Reitter" authorityName="Reitter" authorityYear="1891" box="[199,537,547,573]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="schoenfeldti">
<emphasis id="B91D566FFF91FFFCFF2B2974FE25FDAE" box="[199,451,547,573]" italics="true" pageId="10" pageNumber="11">Euconnus schoenfeldti</emphasis>
Reitter
</taxonomicName>
, whose 
<typeStatus id="54D234DFFF91FFFCFD9E2974FD3FFDAE" box="[626,729,547,573]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
male is here illustrated (
<figureCitation id="135296F8FF91FFFCFC002974FB8AFDAE" box="[1004,1132,546,573]" captionStart="FIGURES 25–30" captionStartId="10.[151,264,1931,1955]" captionTargetBox="[160,1420,680,1899]" captionTargetId="figure-269@10.[151,1436,669,1908]" captionTargetPageId="10" captionText="FIGURES 25–30. Euconnus fustiger, male. Dorsal habitus (25) and historical labels (26) of lectotype male of Euconnus schoenfeldti; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views." figureDoi="http://doi.org/10.5281/zenodo.5901022" httpUri="https://zenodo.org/record/5901022/files/figure.png" pageId="10" pageNumber="11">Figs 25‒30</figureCitation>
) for the first time, does not differ from 
<taxonomicName id="4C69F1FEFF91FFFCFEF52910FE68FDF2" authorityName="Sharp" authorityYear="1874" box="[281,398,583,609]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFEF52910FE68FDF2" box="[281,398,583,609]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFED42910FE68FDF2" box="[312,398,583,609]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
in any features, including the aedeagus, and this name is placed as one more junior synonym of 
<taxonomicName id="4C69F1FEFF91FFFCFF5A293CFECDFD16" authorityName="Sharp" authorityYear="1874" box="[182,299,619,645]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFF5A293CFECDFD16" box="[182,299,619,645]" italics="true" pageId="10" pageNumber="11">
E. 
<emphasis id="B91D566FFF91FFFCFF39293CFECDFD16" box="[213,299,619,645]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF16DAF5FF91FFFCFF7B2CDCFC27F854" ID-DOI="http://doi.org/10.5281/zenodo.5901022" ID-Zenodo-Dep="5901022" httpUri="https://zenodo.org/record/5901022/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1931,1955]" targetBox="[160,1420,680,1899]" targetPageId="10">
<paragraph id="8BD68A7DFF91FFFCFF7B2CDCFC27F854" blockId="10.[151,1436,1931,1992]" pageId="10" pageNumber="11">
<emphasis id="B91D566FFF91FFFCFF7B2CDCFEBEF830" bold="true" box="[151,344,1931,1955]" pageId="10" pageNumber="11">FIGURES 25–30.</emphasis>
<taxonomicName id="4C69F1FEFF91FFFCFE8E2CDBFDFBF837" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[354,541,1932,1956]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF91FFFCFE8E2CDBFDFBF837" box="[354,541,1932,1956]" italics="true" pageId="10" pageNumber="11">
Euconnus 
<emphasis id="B91D566FFF91FFFCFE3C2CDBFDFBF837" box="[464,541,1932,1956]" pageId="10" pageNumber="11">fustiger</emphasis>
</emphasis>
</taxonomicName>
, male. Dorsal habitus (25) and historical labels (26) of lectotype male of 
<taxonomicName id="4C69F1FEFF91FFFCFAD42CDBFEECF854" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="schoenfeldt">
<emphasis id="B91D566FFF91FFFCFAD42CDBFEECF854" italics="true" pageId="10" pageNumber="11">Euconnus schoenfeldt</emphasis>
</taxonomicName>
i; aedeagus (distorted!) in ventral (27, 29) and lateral (28, 30) views.
</paragraph>
</caption>
<paragraph id="8BD68A7DFF90FFFDFF2B2BC0FBFEFA52" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">
All studied 
<typeStatus id="54D234DFFF90FFFDFEA12BC0FE9BFF22" box="[333,381,151,177]" pageId="11" pageNumber="12">type</typeStatus>
specimens of 
<taxonomicName id="4C69F1FEFF90FFFDFDCE2BC0FD7EFF22" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[546,664,151,177]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF90FFFDFDCE2BC0FD7EFF22" box="[546,664,151,177]" italics="true" pageId="11" pageNumber="12">
E. 
<emphasis id="B91D566FFF90FFFDFDAE2BC0FD7EFF22" box="[578,664,151,177]" pageId="11" pageNumber="12">fustiger</emphasis>
</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C69F1FEFF90FFFDFD4B2BC0FCF7FF22" authorityName="Sharp" authorityYear="1886" box="[679,785,151,177]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lewisii">
<emphasis id="B91D566FFF90FFFDFD4B2BC0FCF7FF22" box="[679,785,151,177]" italics="true" pageId="11" pageNumber="12">E. lewisii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C69F1FEFF90FFFDFCCC2BC0FC06FF22" box="[800,992,151,177]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="miyawakianus">
<emphasis id="B91D566FFF90FFFDFCCC2BC0FC06FF22" box="[800,992,151,177]" italics="true" pageId="11" pageNumber="12">E. miyawakianus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C69F1FEFF90FFFDFBCD2BC0FB2EFF22" authorityName="Reitter" authorityYear="1891" box="[1057,1224,151,177]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="schoenfeldti">
<emphasis id="B91D566FFF90FFFDFBCD2BC0FB2EFF22" box="[1057,1224,151,177]" italics="true" pageId="11" pageNumber="12">E. schoenfeldti</emphasis>
</taxonomicName>
have similar body lengths, ranging from 
<quantity id="4C912798FF90FFFDFE7C2BECFDABFF46" box="[400,589,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="11" pageNumber="12" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.40 to 1.50 mm</quantity>
. Over 120 additional specimens from all over Japanese mainland (collected in 17 prefectures) were examined during the present study and results confirm that 
<taxonomicName id="4C69F1FEFF90FFFDFBCB2B88FB7CFF6A" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1063,1178,223,249]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF90FFFDFBCB2B88FB7CFF6A" box="[1063,1178,223,249]" italics="true" pageId="11" pageNumber="12">
E. 
<emphasis id="B91D566FFF90FFFDFBA82B88FB7CFF6A" box="[1092,1178,223,249]" pageId="11" pageNumber="12">fustiger</emphasis>
</emphasis>
</taxonomicName>
is a broadly distributed species, with a relatively narrow range of body length (extremely small specimens measure 
<quantity id="4C912798FF90FFFDFB612A54FB17FE8E" box="[1165,1265,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="11" pageNumber="12" unit="mm" value="1.31">1.31 mm</quantity>
, and the largest ones 
<quantity id="4C912798FF90FFFDFF3F2A71FEDBFED2" box="[211,317,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="11" pageNumber="12" unit="mm" value="1.58">1.58 mm</quantity>
). The studied specimens show some variability in pigmentation (from light to dark brown), and in proportions of body parts. For instance, the elytral index among males can be as low as 1.21 or as high as 1.37, and among females 1.16‒1.33. Moreover, the two pairs of antebasal pronotal pits are highly variable. In most specimens, all four pits are discernible, although they can be equally distinct, equally indistinct, or the outer or inner pair nearly obliterated. In a few specimens all antebasal pits are so minute that barely discernible under 100× magnification of stereomicroscope and in such cases the pronotum is virtually devoid of any antebasal sculpture. The most unusual, however, is variability found in the structures of the aedeagus (examples are shown in 
<figureCitation id="135296F8FF90FFFDFB802AA8FB09FD8A" box="[1132,1263,511,537]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">Figs 32‒49</figureCitation>
). The aedeagus in this species group is extremely thin-walled, and therefore prone to distortion during preparation. The distal sclerotized region in erected or semi-erected aedeagus is flipped distoventrad (see 
<figureCitation id="135296F8FF90FFFDFBDE2910FB63FDF2" box="[1074,1157,583,609]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1573,1597]" captionTargetBox="[155,1426,193,1538]" captionTargetId="figure-15@9.[151,1436,181,1549]" captionTargetPageId="9" captionText="FIGURES 19–24. Euconnus fustiger, male. Dorsal habitus (19) and historical labels (20) of holotype male of Euconnus miyawakianus; aedeagus (distorted!) in ventral (21, 23) and lateral (22, 24) views." figureDoi="http://doi.org/10.5281/zenodo.5901018" httpUri="https://zenodo.org/record/5901018/files/figure.png" pageId="11" pageNumber="12">Fig. 22</figureCitation>
), and when the aedeagus is fully erected the endophallus is everted through the ostium, and then usually also the median lobe is distorted (narrowed or swollen). This condition causes the endophallic sclerites to displace in such a way that it is not possible to make sure that an erected aedeagus is identical with one preserved in a resting position. Maceration in KOH or NaOH strongly promotes artificial erection and may lead to obtaining preparations of variously erected aedeagi, which look so different that several distinct species can be erroneously identified in a sample collected in the same spot. Among specimens examined during the present study, I found variability (true variability, and not caused by inappropriate preparation method) in: (1) the shape of the lateral distal region of the median lobe, which can form distinct, projecting lobes, differing in shape and size in various specimens; (2) shape of the dorsal apical region in lateral view; (3) shape and arrangement of endophallic structures; and (4) shape of dorsal apical plate. The lateral elongate, darkly sclerotized endophallic structures (visible e.g., in 
<figureCitation id="135296F8FF90FFFDFC7E28F8FC03FC5A" box="[914,997,943,969]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">Fig. 32</figureCitation>
) are in fact composed of two pairs of elongate sclerites, which in resting position overlap and appear as one pair. However, the narrower and usually more basally and mesally situated pair of sclerites can move mesad and become clearly discernible (as in 
<figureCitation id="135296F8FF90FFFDFAD028A0FA76FB81" box="[1340,1424,1015,1042]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">Fig. 36</figureCitation>
). This may appear as a species-specific difference, which it is not. Moreover, the thick median subconical structure with truncate apex (well-visible in 
<figureCitation id="135296F8FF90FFFDFDF72F68FD8CFBCA" box="[539,618,1087,1113]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">
Fig. 
<quantity id="4C912798FF90FFFDFDA22F68FD8CFBCA" box="[590,618,1087,1113]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="11" pageNumber="12" unit="in" value="36.0">36</quantity>
</figureCitation>
in subapical region, presumably functioning as a distal flagellar armature with the true gonopore) is integrated with the movable endophallus and any rotation inside the median lobe changes remarkably the shape of this element in ventral view, from elongate and distinct (
<figureCitation id="135296F8FF90FFFDFBC22FD0FB66FB31" box="[1070,1152,1159,1186]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">Fig. 36</figureCitation>
) to entirely indiscernible (e.g., 
<figureCitation id="135296F8FF90FFFDFF342FFCFEC2FB56" box="[216,292,1195,1221]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">Fig 38</figureCitation>
), and intermediary states can also be found (e.g., in 
<figureCitation id="135296F8FF90FFFDFC992FFCFC23FB56" box="[885,965,1195,1221]" captionStart="FIGURES 31–49" captionStartId="12.[151,264,1318,1342]" captionTargetBox="[162,1427,190,1288]" captionTargetId="figure-21@12.[151,1436,181,1294]" captionTargetPageId="12" captionText="FIGURES 31–49. Euconnus fustiger, male. Dorsal habitus of specimen from Kanagawa Pref. (31); aedeagus in ventral (32, 34, 36, 38, 40, 42, 44, 46, 48) and lateral (33, 35, 37, 39, 41, 43, 45, 47, 49) views of specimens from prefectures: Hiroshima (32–35), Saga (36–37, 42–43), Hyôgo (38–41), Kanagawa (44–45), and Ibaraki (46–49), showing examples of variability." figureDoi="http://doi.org/10.5281/zenodo.5901024" httpUri="https://zenodo.org/record/5901024/files/figure.png" pageId="11" pageNumber="12">Fig. 32</figureCitation>
this median element is displaced close to middle of aedeagus and appears as a short, round sclerotization). Shapes of all structures strongly change with even slight rotations of the aedeagus in solid medium, which additionally complicates comparisons. I was not able to find any correlation of the observed variability with geographic factors, and often specimens collected from the same spot and externally indistinguishable show differences in genital structures. I conclude that 
<taxonomicName id="4C69F1FEFF90FFFDFB402E6CFAC5FAC6" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1196,1315,1339,1365]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF90FFFDFB402E6CFAC5FAC6" box="[1196,1315,1339,1365]" italics="true" pageId="11" pageNumber="12">
E. 
<emphasis id="B91D566FFF90FFFDFB212E6CFAC5FAC6" box="[1229,1315,1339,1365]" pageId="11" pageNumber="12">fustiger</emphasis>
</emphasis>
</taxonomicName>
is either a single species showing a great variability in male genital structures, or a complex of many local populations that still maintain a limited gene flow, and such populations have a potential to evolve into separate species if they become physically separated. This phenomenon requires comprehensive genetic studies.
</paragraph>
<paragraph id="8BD68A7DFF90FFFDFF2B2E9CFA69F9C2" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">
Previously, this species was placed in the subgenera 
<taxonomicName id="4C69F1FEFF90FFFDFCCE2E9CFBB3FA76" authority="Reitter, 1909" authorityName="Reitter" authorityYear="1909" box="[802,1109,1483,1509]" class="Insecta" family="Staphylinidae" genus="Euconophron" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B91D566FFF90FFFDFCCE2E9CFC5FFA76" box="[802,953,1483,1509]" italics="true" pageId="11" pageNumber="12">Euconophron</emphasis>
<bibRefCitation id="EFF8F78CFF90FFFDFC2E2E9CFBB3FA76" author="Reitter, E." box="[962,1109,1483,1509]" pageId="11" pageNumber="12" refId="ref26236" refString="Reitter, E. (1909) Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. II. Band. K. G. Lutz, Stuttgart, 392 pp." type="book" year="1909">Reitter, 1909</bibRefCitation>
</taxonomicName>
(currently a junior synonym of 
<emphasis id="B91D566FFF90FFFDFF592EB8FED8F99B" box="[181,318,1519,1544]" italics="true" pageId="11" pageNumber="12">Eupentarius</emphasis>
Reitter, 1907) (by 
<bibRefCitation id="EFF8F78CFF90FFFDFDFF2EB8FD7BF99A" author="Franz, H." box="[531,669,1518,1545]" pageId="11" pageNumber="12" pagination="57 - 107" refId="ref24521" refString="Franz, H. (1975) Zweiter Beitrag zur Kenntnis der Scydmaenidenfauna Sudostasiens. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abteilung I, 183, 57 - 107." type="journal article" year="1975">Franz (1975</bibRefCitation>
, 
<bibRefCitation id="EFF8F78CFF90FFFDFD442EB8FD07F999" author="Franz, H." box="[680,737,1519,1546]" pageId="11" pageNumber="12" pagination="51 - 60" refId="ref24555" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Ergebnisse einer auf Einladung der Internationalen Vereinigung fur Vegetationskunde nach Japan durchgefuhrten Forschungsreise. Entomologische Blatter fur Biologie und Systematik der Kafer, 72 (1), 51 - 60." type="journal article" year="1976">1976</bibRefCitation>
)) or 
<emphasis id="B91D566FFF90FFFDFCF42EB8FC6EF99A" box="[792,904,1519,1545]" italics="true" pageId="11" pageNumber="12">Napochus</emphasis>
<bibRefCitation id="EFF8F78CFF90FFFDFC622EB8FBD8F99A" author="Thomson, C. G." box="[910,1086,1518,1545]" pageId="11" pageNumber="12" refId="ref26483" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. 1. Berlingska Boktryckeriet, Lund, 290 pp. https: // doi. org / 10.5962 / bhl. title. 138677" type="book" year="1859">Thomson, 1859</bibRefCitation>
(currently a junior synonym of 
<taxonomicName id="4C69F1FEFF90FFFDFF7B2D44FEE3F9BF" box="[151,261,1555,1580]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF90FFFDFF7B2D44FEE3F9BF" box="[151,261,1555,1580]" italics="true" pageId="11" pageNumber="12">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EEB14FF90FFFDFEE22D44FEADF9BE" box="[270,331,1555,1581]" pageId="11" pageNumber="12" sensu="stricto">s. str.</taxonomicNameLabel>
) (by 
<bibRefCitation id="EFF8F78CFF90FFFDFE652D44FDC2F9BE" author="Csiki, E." box="[393,548,1555,1581]" pageId="11" pageNumber="12" pagination="106" refId="ref24424" refString="Csiki, E. (1919) Pars 70: Scydmaenidae. In: Schenkling, S. &amp; Junk, W. (Eds.), Coleopterorum Catalogus. Vol. 7. W. Junk, Berlin, pp. 106." type="book chapter" year="1919">Csiki (1919))</bibRefCitation>
. After redefining 
<emphasis id="B91D566FFF90FFFDFD012D44FBF3F9BE" box="[749,1045,1555,1581]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C69F1FEFF90FFFDFD012D44FC65F9BE" box="[749,899,1555,1581]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Euconophron">Euconophron</taxonomicName>
/Eupentarius
</emphasis>
(
<bibRefCitation id="EFF8F78CFF90FFFDFBC92D44FB1CF9BE" author="Jaloszynski, P." box="[1061,1274,1555,1581]" pageId="11" pageNumber="12" pagination="450 - 460" refId="ref25610" refString="Jaloszynski, P. (2017 b) Taxonomy of &quot; Euconnus complex &quot;. Part XI. Status of Euconophron and Scopophus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4236 (3), 450 - 460. https: // doi. org / 10.11646 / zootaxa. 4236.3.2" type="journal article" year="2017">Jałoszyński 2017b</bibRefCitation>
; d, 2021a; 
<bibRefCitation id="EFF8F78CFF90FFFDFA942D44FF37F9C2" author="Vit, S." pageId="11" pageNumber="12" pagination="185 - 192" refId="ref26524" refString="Vit, S. (2005) Addenda to the scydmaenid fauna of the Canary Islands (La Gomera, Gran Canaria) with emphasis on Euconophron (Coleoptera: Scydmaenidae, Cyrtoscydmini). Vieraea, 33, 185 - 192." type="journal article" year="2005">Vít 2005</bibRefCitation>
) and 
<taxonomicName id="4C69F1FEFF90FFFDFEE32D60FE9BF9C3" box="[271,381,1591,1616]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF90FFFDFEE32D60FE9BF9C3" box="[271,381,1591,1616]" italics="true" pageId="11" pageNumber="12">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EEB14FF90FFFDFE682D60FE59F9C2" box="[388,447,1591,1617]" pageId="11" pageNumber="12" sensu="stricto">s. str.</taxonomicNameLabel>
(
<bibRefCitation id="EFF8F78CFF90FFFDFE212D60FD47F9C1" author="Jaloszynski, P." box="[461,673,1591,1618]" pageId="11" pageNumber="12" pagination="255 - 270" refId="ref25908" refString="Jaloszynski, P. (2021 b) Taxonomy of &quot; Euconnus complex &quot;. Part XXIII. Status of Napochus Thomson, Pycnophus Casey, and Filonapochus Franz revisited (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5026 (2), 255 - 270. https: // doi. org / 10.11646 / zootaxa. 5026.2.6" type="journal article" year="2021">Jałoszyński 2021b</bibRefCitation>
), the placement of 
<taxonomicName id="4C69F1FEFF90FFFDFC9A2D60FC0DF9C2" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[886,1003,1591,1617]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF90FFFDFC9A2D60FC0DF9C2" box="[886,1003,1591,1617]" italics="true" pageId="11" pageNumber="12">
E. 
<emphasis id="B91D566FFF90FFFDFC792D60FC0DF9C2" box="[917,1003,1591,1617]" pageId="11" pageNumber="12">fustiger</emphasis>
</emphasis>
</taxonomicName>
in 
<taxonomicName id="4C69F1FEFF90FFFDFBE22D60FB9AF9C3" box="[1038,1148,1591,1616]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF90FFFDFBE22D60FB9AF9C3" box="[1038,1148,1591,1616]" italics="true" pageId="11" pageNumber="12">Euconnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EEB14FF90FFFDFB682D60FB59F9C2" box="[1156,1215,1591,1617]" pageId="11" pageNumber="12" sensu="stricto">s. str.</taxonomicNameLabel>
can be confirmed.
</paragraph>
<paragraph id="8BD68A7DFF90FFFDFF2B2D0CFD90F90A" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">
This is the commonest of mainland Japanese 
<taxonomicName id="4C69F1FEFF90FFFDFD3F2D0CFCA7F9E7" box="[723,833,1627,1652]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B91D566FFF90FFFDFD3F2D0CFCA7F9E7" box="[723,833,1627,1652]" italics="true" pageId="11" pageNumber="12">Euconnus</emphasis>
</taxonomicName>
(s. str.) species; it inhabits leaf litter in lowland and mountainous deciduous and mixed forests.
</paragraph>
<paragraph id="8BD68A7DFF90FFFDFF2B2DF4FD3EF82A" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">
Efforts should be made to obtain non-distorted preparations (by avoiding maceration in alkali!), and descriptions of new species belonging to this complex should be supported by a profound understanding of geographical variation and character variability. I have seen many Japanese specimens of 
<taxonomicName id="4C69F1FEFF90FFFDFC6F2DBCFC17F897" box="[899,1009,1771,1796]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B91D566FFF90FFFDFC6F2DBCFC17F897" box="[899,1009,1771,1796]" italics="true" pageId="11" pageNumber="12">Euconnus</emphasis>
</taxonomicName>
which markedly differ in body length from 
<taxonomicName id="4C69F1FEFF90FFFDFF3A2C58FEABF8BA" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[214,333,1807,1833]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF90FFFDFF3A2C58FEABF8BA" box="[214,333,1807,1833]" italics="true" pageId="11" pageNumber="12">
E. 
<emphasis id="B91D566FFF90FFFDFF1B2C58FEABF8BA" box="[247,333,1807,1833]" pageId="11" pageNumber="12">fustiger</emphasis>
</emphasis>
</taxonomicName>
(being either much smaller or much larger), showing the same form of aedeagus. Unfortunately, many such species have already been described (see comments in further parts of this paper), based on single or few specimens with strongly distorted aedeagi, which makes it impossible to unambiguously identify some (or all) of them. The taxonomy of the Japanese ‘ 
<taxonomicName id="4C69F1FEFF90FFFDFDD12C2CFCEEF806" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[573,776,1915,1941]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF90FFFDFDD12C2CFCEEF806" box="[573,776,1915,1941]" italics="true" pageId="11" pageNumber="12">
Euconnus 
<emphasis id="B91D566FFF90FFFDFD5E2C2CFCEEF806" box="[690,776,1915,1941]" pageId="11" pageNumber="12">fustiger</emphasis>
</emphasis>
</taxonomicName>
group’ seems to have reached the level of chaos that makes future studies extremely difficult, if not impossible.
</paragraph>
</subSubSection>
</treatment>
</document>