155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="944DAEECE2BEDD96E1660E8B9DB1E5CD" ID-CLB-Dataset="121546" ID-DOI="10.11646/zootaxa.5174.5.1" ID-GBIF-Dataset="166971ec-bb80-41bc-9c7b-f6093a0f83e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986555" ID-ZooBank="BDB90B5C-C3DD-464D-AA7F-1635009297A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660327825565" checkinUser="plazi" docAuthor="Moulds, Max & Marshall, David C." docDate="2022" docId="F75887D56F75223FFF2CFB3C03C8F85A" docLanguage="en" docName="zootaxa.5174.5.1.pdf" docOrigin="Zootaxa 5174 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyropsalta Moulds 2012" docType="treatment" docVersion="8" lastPageNumber="497" masterDocId="0B61FFAD6F5B2211FFBBFF96006FFFCF" masterDocTitle="New genera and new species of Western Australian cicadas (Hemiptera: Cicadidae)" masterLastPageNumber="507" masterPageNumber="451" pageNumber="497" updateTime="1699367077226" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
|
<mods:mods id="08F22A3E59A9D44C1E41FD74F35DC01D" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="49F73919D36C6ADCDC25278D7ECF782E">
|
|
<mods:title id="743C9EDBDFB2177C4941E20C9B893101">New genera and new species of Western Australian cicadas (Hemiptera: Cicadidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="5B0042A8823E4AC6C649AB3B865C5BAB" type="personal">
|
|
<mods:role id="A05024C5CFB20D60CA8060392D69615B">
|
|
<mods:roleTerm id="AEB8641F49D3DBB6F62B9A19A5F8C8C7">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="292B91CF2D12A237F6A333F6851835FA">Moulds, Max</mods:namePart>
|
|
<mods:affiliation id="4470B1BF8972DFA7615EF09F443DBAD6">Australian Museum Research Institute, Australian Museum, 1 William St, Sydney, NSW 2010.</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="E39150C9711A3E96359693E90F4DADD4" type="personal">
|
|
<mods:role id="3F1209E0BF266A00466688D2D5CBA3CF">
|
|
<mods:roleTerm id="54E4AD2BF98A7C9669935AF23D1538D4">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="5D6EFD2599172D5DE94BDEFC1DE40B5E">Marshall, David C.</mods:namePart>
|
|
<mods:affiliation id="08189F1692F6C775618BE3408B86AC65">Dept. of Ecology and Evolutionary Biology, University of Connecticut, Storrs, CT 06269, USA. & Current address: Auckland, New Zealand</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource id="50246D8B572C0E50E413A66CFF7E4E29">text</mods:typeOfResource>
|
|
<mods:relatedItem id="50D5D4A397EE6AADF7F12006287CAAAD" type="host">
|
|
<mods:titleInfo id="9E2C414115F0200DCC265B5507E6F23E">
|
|
<mods:title id="4DB0F58D5DE0D584BEEBC2BB7D5A917D">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="0910C657DBBB70152D8D3EB61BE245C7">
|
|
<mods:date id="3A965E2CE8C218065AEC78F628CB45F8">2022</mods:date>
|
|
<mods:detail id="A1BBA44A811D45CA1D1A6960F0503CE1" type="pubDate">
|
|
<mods:number id="644ECEBF1E86DE7BECA58FD4308A71E8">2022-08-12</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="FEF97854CD8FBAFFE276E30C3758C3B2" type="volume">
|
|
<mods:number id="4E29CC16F347704E2B6B89FE743107CD">5174</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="469179A16168D27A7B0AEA82AB14F9B0" type="issue">
|
|
<mods:number id="DAD1AE824743776910EF7538AA04229A">5</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="FA07C24946167167F6D7DBDA5EC40B17" unit="page">
|
|
<mods:start id="893D9C266F919501376B0C8C6EC7962F">451</mods:start>
|
|
<mods:end id="B8071A2B3D9AE0C4F02C61CAFF819D9B">507</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="16C9EB90A2A708DF4B25F4043194C579">journal article</mods:classification>
|
|
<mods:identifier id="6369608F2A47BBBC9F40DF10ED7B054B" type="CLB-Dataset">121546</mods:identifier>
|
|
<mods:identifier id="1771ABB1C197F01130629AFB04B99357" type="DOI">10.11646/zootaxa.5174.5.1</mods:identifier>
|
|
<mods:identifier id="60E1F1AA27CB6FFA4AB23CE956B1B6A4" type="GBIF-Dataset">166971ec-bb80-41bc-9c7b-f6093a0f83e4</mods:identifier>
|
|
<mods:identifier id="55344E1F3715E01BBEB092009C968325" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="DC54A4E92E72B4EC15ED8B9FBC38AADD" type="Zenodo-Dep">6986555</mods:identifier>
|
|
<mods:identifier id="0793B0B4E545E39232A2614D05815E01" type="ZooBank">BDB90B5C-C3DD-464D-AA7F-1635009297A6</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="F75887D56F75223FFF2CFB3C03C8F85A" ID-DOI="http://doi.org/10.5281/zenodo.6987065" ID-GBIF-Taxon="199160331" ID-Zenodo-Dep="6987065" LSID="urn:lsid:plazi:treatment:F75887D56F75223FFF2CFB3C03C8F85A" httpUri="http://treatment.plazi.org/id/F75887D56F75223FFF2CFB3C03C8F85A" lastPageNumber="497" pageId="46" pageNumber="497">
|
|
<subSubSection id="37EB65486F75223FFF2CFB3C01A8FB0B" box="[151,455,1194,1221]" pageId="46" pageNumber="497" type="nomenclature">
|
|
<paragraph id="7F4E36C36F75223FFF2CFB3C01A8FB0B" blockId="46.[151,455,1194,1221]" box="[151,455,1194,1221]" pageId="46" pageNumber="497">
|
|
<heading id="240681AF6F75223FFF2CFB3C01A8FB0B" bold="true" box="[151,455,1194,1221]" fontSize="11" level="1" pageId="46" pageNumber="497" reason="1">
|
|
<taxonomicName id="B8F14D406F75223FFF2CFB3C01A8FB0B" ID-CoL="8HQD7" authority="Moulds, 2012" authorityName="Moulds" authorityYear="2012" box="[151,455,1194,1221]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4D85EAD16F75223FFF2CFB3C01A8FB0B" bold="true" box="[151,455,1194,1221]" pageId="46" pageNumber="497">
|
|
<emphasis id="4D85EAD16F75223FFF2CFB3C0177FB0A" bold="true" box="[151,280,1194,1221]" italics="true" pageId="46" pageNumber="497">Pyropsalta</emphasis>
|
|
<bibRefCitation id="1B604B326F75223FFEA4FB3D01A8FB0B" author="Moulds, M. S." box="[287,455,1194,1221]" pageId="46" pageNumber="497" pagination="1 - 262" refId="ref52056" refString="Moulds, M. S. (2012) A review of the genera of Australian cicadas (Hemiptera: Cicadoidea). Zootaxa, 3287 (1), 1 - 262. https: // doi. org / 10.11646 / zootaxa. 3287.1.1" type="journal article" year="2012">Moulds, 2012</bibRefCitation>
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="37EB65486F75223FFF2CFB650384F99E" pageId="46" pageNumber="497" type="reference_group">
|
|
<paragraph id="7F4E36C36F75223FFF2CFB65022EFA9A" blockId="46.[151,1437,1267,1941]" pageId="46" pageNumber="497">
|
|
<taxonomicName id="B8F14D406F75223FFF2CFB65017EFAC2" ID-CoL="8HQD7" authorityName="Moulds" authorityYear="2012" box="[151,273,1267,1293]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4D85EAD16F75223FFF2CFB65017EFAC2" box="[151,273,1267,1293]" italics="true" pageId="46" pageNumber="497">Pyropsalta</emphasis>
|
|
</taxonomicName>
|
|
is an endemic genus confined to south-western
|
|
<collectingRegion id="BD35F8216F75223FFC9BFB650386FAC2" box="[800,1001,1267,1293]" country="Australia" name="Western Australia" pageId="46" pageNumber="497">Western Australia</collectingRegion>
|
|
.
|
|
<location id="7A2E60186F75223FFC49FB65045AFAC2" LSID="urn:lsid:plazi:treatment:F75887D56F75223FFF2CFB3C03C8F85A:7A2E60186F75223FFC49FB65045AFAC2" box="[1010,1077,1267,1293]" name="When" pageId="46" pageNumber="497" stateProvince="Western Australia">When</location>
|
|
first established it included only the
|
|
<typeStatus id="A04A88616F75223FFF7AFA81009EFAFE" box="[193,241,1303,1329]" pageId="46" pageNumber="497">type</typeStatus>
|
|
species,
|
|
<taxonomicName id="B8F14D406F75223FFEE2FA8102B9FAFE" ID-CoL="4QVBW" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" box="[345,726,1302,1329]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="melete">
|
|
<emphasis id="4D85EAD16F75223FFEE2FA81024CFAFE" box="[345,547,1303,1329]" italics="true" pageId="46" pageNumber="497">Pyropsalta melete</emphasis>
|
|
(
|
|
<bibRefCitation id="1B604B326F75223FFD88FA8102A1FAFE" author="Walker, F." box="[563,718,1302,1329]" pageId="46" pageNumber="497" refId="ref52882" refString="Walker, F. (1850) List of the specimens of homopterous insects in the collection of the British Museum. Part 1. British Museum, London, 260 pp." type="book" year="1850">Walker, 1850</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
, that has remained the only described species in the genus. We now describe three additional species.
|
|
</paragraph>
|
|
<paragraph id="7F4E36C36F75223FFF7CFAC90384F99E" blockId="46.[151,1437,1267,1941]" pageId="46" pageNumber="497">
|
|
These three additional species largely conform to the generic description of
|
|
<bibRefCitation id="1B604B326F75223FFB9AFAC904A6FAB6" author="Moulds, M. S." box="[1057,1225,1375,1401]" pageId="46" pageNumber="497" pagination="1 - 262" refId="ref52056" refString="Moulds, M. S. (2012) A review of the genera of Australian cicadas (Hemiptera: Cicadoidea). Zootaxa, 3287 (1), 1 - 262. https: // doi. org / 10.11646 / zootaxa. 3287.1.1" type="journal article" year="2012">Moulds (2012)</bibRefCitation>
|
|
although there are differences in male timbal morphology and small differences in the male genitalia, the latter not considered significant. The timbals of all three new species have an unusually large timbal cavity that is most extreme in
|
|
<taxonomicName id="B8F14D406F75223FFAA9FA31051AFA0E" ID-CoL="9GDMM" authority="Moulds & Marshall, 2022" authorityName="Moulds & Marshall" authorityYear="2022" box="[1298,1397,1447,1473]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="patula" status="sp. nov.">
|
|
<emphasis id="4D85EAD16F75223FFAA9FA31051AFA0E" box="[1298,1397,1447,1473]" italics="true" pageId="46" pageNumber="497">P. patula</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="4D85EAD16F75223FFAC0FA3100C1FA2A" bold="true" pageId="46" pageNumber="497">
|
|
<taxonomicNameLabel id="56B657AA6F75223FFAC0FA3100C1FA2A" pageId="46" pageNumber="497" rank="species">sp. n.</taxonomicNameLabel>
|
|
</emphasis>
|
|
, each with three long ribs and an expansive anterior membrane largely devoid of structures apart from a short thin rudimentary rib at its dorsal posterior corner, another similar rib tight against the anterior margin and a securitised basal area. This differs from the
|
|
<typeStatus id="A04A88616F75223FFE46F9850242F9E2" box="[509,557,1555,1581]" pageId="46" pageNumber="497">type</typeStatus>
|
|
species that has a timbal cavity of usual proportions, four long ribs, and a small anterior membrane area with one rudimentary short rib at its dorsal anterior.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="37EB65486F75223FFF7CF9CD03C8F85A" pageId="46" pageNumber="497" type="description">
|
|
<paragraph id="7F4E36C36F75223FFF7CF9CD03C8F85A" blockId="46.[151,1437,1267,1941]" pageId="46" pageNumber="497">
|
|
In a molecular study by
|
|
<bibRefCitation id="1B604B326F75223FFE62F9CD02A6F9BA" author="Marshall, D. C. & Hill, K. B. R. & Moulds, M. S. & Vanderpool, D. & Cooley, J. R. & Mohagan, A. B. & Simon, C." box="[473,713,1627,1653]" pageId="46" pageNumber="497" pagination="16 - 34" refId="ref51450" refString="Marshall, D. C., Hill, K. B. R., Moulds, M. S., Vanderpool, D., Cooley, J. R., Mohagan, A. B. & Simon, C. (2016) Inflation of molecular clock rates and dates: molecular phylogenetics, biogeography, and diversification of a global cicada radiation from Australasia (Hemiptera: Cicadidae: Cicadettini). Systematic Biology, 65, 16 - 34. [first issued online 6 November 2015, pp. 1 - 19] https: // doi. org / 10.1093 / sysbio / syv 069" type="journal article" year="2016">
|
|
Marshall
|
|
<emphasis id="4D85EAD16F75223FFDFFF9CD0212F9BA" box="[580,637,1627,1653]" italics="true" pageId="46" pageNumber="497">et al.</emphasis>
|
|
(2016
|
|
</bibRefCitation>
|
|
: fig. 2) species of
|
|
<taxonomicName id="B8F14D406F75223FFC23F9CD047DF9BA" authorityName="Moulds" authorityYear="2012" box="[920,1042,1627,1653]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4D85EAD16F75223FFC23F9CD047DF9BA" box="[920,1042,1627,1653]" italics="true" pageId="46" pageNumber="497">Pyropsalta</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="B8F14D406F75223FFBF2F9CD0531F9BA" authority="Moulds, 2012" authorityName="Moulds" authorityYear="2012" box="[1097,1374,1627,1653]" class="Insecta" family="Cicadidae" genus="Physeema" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4D85EAD16F75223FFBF2F9CD04D4F9BA" box="[1097,1211,1627,1653]" italics="true" pageId="46" pageNumber="497">Physeema</emphasis>
|
|
<bibRefCitation id="1B604B326F75223FFB79F9CD0531F9BA" author="Moulds, M. S." box="[1218,1374,1627,1653]" pageId="46" pageNumber="497" pagination="1 - 262" refId="ref52056" refString="Moulds, M. S. (2012) A review of the genera of Australian cicadas (Hemiptera: Cicadoidea). Zootaxa, 3287 (1), 1 - 262. https: // doi. org / 10.11646 / zootaxa. 3287.1.1" type="journal article" year="2012">Moulds, 2012</bibRefCitation>
|
|
</taxonomicName>
|
|
were found to form a single clade with strong support. However, one of the new species of
|
|
<taxonomicName id="B8F14D406F75223FFBE4F9E904B6F956" authorityName="Moulds" authorityYear="2012" box="[1119,1241,1663,1689]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4D85EAD16F75223FFBE4F9E904B6F956" box="[1119,1241,1663,1689]" italics="true" pageId="46" pageNumber="497">Pyropsalta</emphasis>
|
|
</taxonomicName>
|
|
described below,
|
|
<taxonomicName id="B8F14D406F75223FFF2CF935015CF972" authority="Moulds & Marshall, 2022" authorityName="Moulds & Marshall" authorityYear="2022" box="[151,307,1699,1725]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="rhythmica" status="sp. nov.">
|
|
<emphasis id="4D85EAD16F75223FFF2CF935015CF972" box="[151,307,1699,1725]" italics="true" pageId="46" pageNumber="497">Py. rhythmica</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="4D85EAD16F75223FFE87F9350113F972" bold="true" box="[316,380,1699,1725]" pageId="46" pageNumber="497">
|
|
<taxonomicNameLabel id="56B657AA6F75223FFE87F9350113F972" box="[316,380,1699,1725]" pageId="46" pageNumber="497" rank="species">sp. n.</taxonomicNameLabel>
|
|
</emphasis>
|
|
, was found to be not sister to the
|
|
<typeStatus id="A04A88616F75223FFCBBF935035FF972" box="[768,816,1699,1725]" pageId="46" pageNumber="497">type</typeStatus>
|
|
species of
|
|
<emphasis id="4D85EAD16F75223FFC0AF935045DF972" box="[945,1074,1699,1725]" italics="true" pageId="46" pageNumber="497">
|
|
<taxonomicName id="B8F14D406F75223FFC0AF9350441F972" authorityName="Moulds" authorityYear="2012" box="[945,1070,1699,1725]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">Pyropsalta</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
but separated from it in a sister clade together with
|
|
<taxonomicName id="B8F14D406F75223FFEC8F9510375F92E" authority="(Walker, 1850)" baseAuthorityName="Walker" baseAuthorityYear="1850" box="[371,794,1734,1761]" class="Insecta" family="Cicadidae" genus="Physeema" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="convergens">
|
|
<emphasis id="4D85EAD16F75223FFEC8F9510206F92F" box="[371,617,1735,1761]" italics="true" pageId="46" pageNumber="497">Physeema convergens</emphasis>
|
|
(
|
|
<bibRefCitation id="1B604B326F75223FFDCCF951037EF92E" author="Walker, F." box="[631,785,1734,1761]" pageId="46" pageNumber="497" refId="ref52882" refString="Walker, F. (1850) List of the specimens of homopterous insects in the collection of the British Museum. Part 1. British Museum, London, 260 pp." type="book" year="1850">Walker, 1850</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. This suggests that
|
|
<taxonomicName id="B8F14D406F75223FFC4AF95104E4F92E" authority="Moulds & Marshall, 2022" authorityName="Moulds & Marshall" authorityYear="2022" box="[1009,1163,1735,1761]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="rhythmica" status="sp. nov.">
|
|
<emphasis id="4D85EAD16F75223FFC4AF95104E4F92E" box="[1009,1163,1735,1761]" italics="true" pageId="46" pageNumber="497">Py. rhythmica</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="4D85EAD16F75223FFB29F95104BEF92E" bold="true" box="[1170,1233,1735,1761]" pageId="46" pageNumber="497">
|
|
<taxonomicNameLabel id="56B657AA6F75223FFB29F95104BEF92E" box="[1170,1233,1735,1761]" pageId="46" pageNumber="497" rank="species">sp. n.</taxonomicNameLabel>
|
|
</emphasis>
|
|
(together with the two other species described below) possibly represent a new genus. However, we refrain from taking that action without further evidence, because the support for these relationships in the molecular tree is not entirely conclusive (0.94 Bayesian posterior probabilities, 74 and 69 percent ML bootstrap support). We therefore treat the differences in timbal morphology as indicative of species attributes rather than generic attributes, and because all other attributes are compatible with
|
|
<taxonomicName id="B8F14D406F75223FFE04F8ED0256F85A" authorityName="Moulds" authorityYear="2012" box="[447,569,1915,1941]" class="Insecta" family="Cicadidae" genus="Pyropsalta" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="4D85EAD16F75223FFE04F8ED0256F85A" box="[447,569,1915,1941]" italics="true" pageId="46" pageNumber="497">Pyropsalta</emphasis>
|
|
</taxonomicName>
|
|
with include them in that genus.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |