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<document id="9B655F09F654E33749FB657E09D926EF" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. &amp; Hughes, Nigel C." docDate="2022" docId="03CB87846E781D25FF3AC59B4B4AF68F" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Illaenus caecoides Reed, Complete 1917" docType="treatment" docVersion="6" lastPageNumber="346" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reeds Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="346" updateTime="1699356965331" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B84A690B165F37C0996FD02AC0E3DCCF">Revision of F. R. C. Reeds Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:namePart id="875BBC24029F98056AF4191FEA0D8EBA">Fortey, Richard A.</mods:namePart>
<mods:affiliation id="C071D4DCA8CC15E6703BE2A9B893DD21">Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="6B038E3F59C8C677E0C2204DCC963093">Wernette, Shelly J.</mods:namePart>
<mods:affiliation id="8AB3785A7203EA7EAAF5FAD7B817AB6B">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. &amp; Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<mods:namePart id="27FE0FB69C71EF9EC3681869C5E97999">Hughes, Nigel C.</mods:namePart>
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<treatment id="03CB87846E781D25FF3AC59B4B4AF68F" ID-DOI="http://doi.org/10.5281/zenodo.6810380" ID-GBIF-Taxon="197214797" ID-Zenodo-Dep="6810380" LSID="urn:lsid:plazi:treatment:03CB87846E781D25FF3AC59B4B4AF68F" httpUri="http://treatment.plazi.org/id/03CB87846E781D25FF3AC59B4B4AF68F" lastPageNumber="346" pageId="45" pageNumber="346">
<subSubSection id="C37865196E781D25FF3AC59B481DF4B8" box="[151,518,474,501]" pageId="45" pageNumber="346" type="nomenclature">
<paragraph id="8BDD36926E781D25FF3AC59B481DF4B8" blockId="45.[151,518,474,537]" box="[151,518,474,501]" pageId="45" pageNumber="346">
<heading id="D09581FE6E781D25FF3AC59B481DF4B8" bold="true" box="[151,518,474,501]" fontSize="11" level="1" pageId="45" pageNumber="346" reason="1">
<taxonomicName id="4C624D116E781D25FF3AC59B481DF4B8" authority="Reed, 1917" authorityName="Reed, Complete" authorityYear="1917" box="[151,518,474,501]" class="Trilobita" family="Illaenidae" genus="Illaenus" kingdom="Animalia" order="Corynexochida" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="caecoides">
<emphasis id="B916EA806E781D25FF3AC59B481DF4B8" bold="true" box="[151,518,474,501]" pageId="45" pageNumber="346">
<emphasis id="B916EA806E781D25FF3AC59B4B6DF4B9" bold="true" box="[151,374,474,501]" italics="true" pageId="45" pageNumber="346">Illaenus caecoides</emphasis>
<bibRefCitation id="EFF34B636E781D25FED0C59A481DF4B8" author="Reed, F. R. C." box="[381,518,474,501]" pageId="45" pageNumber="346" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37865196E781D25FF3AC5BE4955F717" pageId="45" pageNumber="346" type="description">
<paragraph id="8BDD36926E781D25FF3AC5BE4AE7F755" blockId="45.[151,518,474,537]" box="[151,252,510,537]" pageId="45" pageNumber="346">
<figureCitation id="13592A176E781D25FF3AC5BE4AE7F755" box="[151,252,510,537]" captionStart="FIGURE 15" captionStartId="48.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-29@48.[151,1436,181,1531]" captionTargetPageId="48" captionText="FIGURE 15. (14) Metopolichas? sp. 1 (Reed) from the Li-Lu and Kunlein formations (Katian), northern Shan State. 1. Hypostome (original of Reed 1915, pl. 7, fig. 15), GSI 11546, Mân-ngai. 2. Hypostome (original of Reed 1915, pl. 7, fig. 16), GSI 11547, Lilu. 3. Partial cranidium (original of Reed 1915, pl. 7, fig. 14), GSI 11545, Tâ-Pangtawng. 4a, b. Immature cephalon (original of Reed 1915, pl. 7, fig. 17), GSI 11548, Makmongshai, original and retrodeformation. (5) Illaenus caecoides Reed, Complete specimen (refigured from Reed 1917, pl. 7, fig. 9), GSI 11898, Shihtien Formation (Darriwilian), Banpo Village, Shidian County, Baoshan Prefecture, western Yunnan, China. (68) Phorocephala mansuyi, Cranidium (refigured and original from Reed 1917, pl. 8, fig. 7), GSI 11848, dorsal and lateral views, Shihtien Formation (Darriwilian), Pu-piao, Baoshan Prefecture, western Yunnan, China. (9) Metopolichas sp. 2 (Reed), Cephalon (original of Reed 1917, pl. 8, fig. 9), GSI 11910, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (10) Stygina? sp. (Reed), Cranidium (original of Reed 1915, pl. 7, fig. 13), GSI 11544, Hwe Mawng Beds (uppermost Katian), Mong-Ha, northern part of Shan State, Myanmar. Scale bars = 2 mm for 1, 10; = 1 mm for 2, 4, 69; = 5 mm for 3, 5." figureDoi="http://doi.org/10.5281/zenodo.6810334" httpUri="https://zenodo.org/record/6810334/files/figure.png" pageId="45" pageNumber="346">Fig. 15.5</figureCitation>
</paragraph>
<paragraph id="8BDD36926E781D25FF3AC6024955F717" blockId="45.[151,846,579,603]" box="[151,846,579,603]" pageId="45" pageNumber="346">
1917
<taxonomicName id="4C624D116E781D25FF5AC6024BA8F717" authority="Fortey &amp; Wernette &amp; Hughes, 2022" authorityName="Fortey &amp; Wernette &amp; Hughes" authorityYear="2022" box="[247,435,579,603]" class="Trilobita" family="Illaenidae" genus="Illaenus" kingdom="Animalia" order="Corynexochida" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="caecoides" status="sp. nov.">
<emphasis id="B916EA806E781D25FF5AC6024BA8F717" box="[247,435,579,603]" italics="true" pageId="45" pageNumber="346">Illaenus caecoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557FB6E781D25FE14C6054810F717" box="[441,523,580,603]" pageId="45" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
; Reed, p. 456, pl. 7, figs 910.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E781D25FF3AC6C8499CF7A0" pageId="45" pageNumber="346" type="materials_examined">
<paragraph id="8BDD36926E781D25FF3AC6C8499CF7A0" blockId="45.[151,1436,649,964]" pageId="45" pageNumber="346">
<emphasis id="B916EA806E781D25FF3AC6C84B1DF7EF" bold="true" box="[151,262,649,675]" pageId="45" pageNumber="346">Material.</emphasis>
<typeStatus id="54D988306E781D25FEA2C6CB4B64F7E8" box="[271,383,650,676]" pageId="45" pageNumber="346" type="lectotype">Lectotype</typeStatus>
(selected herein): dorsal exoskeleton,
<figureCitation id="13592A176E781D25FC99C6C84981F7E8" box="[820,922,649,676]" captionStart="FIGURE 15" captionStartId="48.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-29@48.[151,1436,181,1531]" captionTargetPageId="48" captionText="FIGURE 15. (14) Metopolichas? sp. 1 (Reed) from the Li-Lu and Kunlein formations (Katian), northern Shan State. 1. Hypostome (original of Reed 1915, pl. 7, fig. 15), GSI 11546, Mân-ngai. 2. Hypostome (original of Reed 1915, pl. 7, fig. 16), GSI 11547, Lilu. 3. Partial cranidium (original of Reed 1915, pl. 7, fig. 14), GSI 11545, Tâ-Pangtawng. 4a, b. Immature cephalon (original of Reed 1915, pl. 7, fig. 17), GSI 11548, Makmongshai, original and retrodeformation. (5) Illaenus caecoides Reed, Complete specimen (refigured from Reed 1917, pl. 7, fig. 9), GSI 11898, Shihtien Formation (Darriwilian), Banpo Village, Shidian County, Baoshan Prefecture, western Yunnan, China. (68) Phorocephala mansuyi, Cranidium (refigured and original from Reed 1917, pl. 8, fig. 7), GSI 11848, dorsal and lateral views, Shihtien Formation (Darriwilian), Pu-piao, Baoshan Prefecture, western Yunnan, China. (9) Metopolichas sp. 2 (Reed), Cephalon (original of Reed 1917, pl. 8, fig. 9), GSI 11910, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (10) Stygina? sp. (Reed), Cranidium (original of Reed 1915, pl. 7, fig. 13), GSI 11544, Hwe Mawng Beds (uppermost Katian), Mong-Ha, northern part of Shan State, Myanmar. Scale bars = 2 mm for 1, 10; = 1 mm for 2, 4, 69; = 5 mm for 3, 5." figureDoi="http://doi.org/10.5281/zenodo.6810334" httpUri="https://zenodo.org/record/6810334/files/figure.png" pageId="45" pageNumber="346">Fig. 15.5</figureCitation>
(
<bibRefCitation id="EFF34B636E781D25FC07C6C84E34F7EF" author="Reed, F. R. C." box="[938,1071,649,675]" pageId="45" pageNumber="346" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
, pl. 7, fig. 9),
<collectionCode id="ED73AE576E781D25FB7EC6C84F1BF7EF" box="[1235,1280,649,675]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="45" pageNumber="346">GSI</collectionCode>
11898. Additional type material: pygidium (
<bibRefCitation id="EFF34B636E781D25FE5AC6EC4861F78B" author="Reed, F. R. C." box="[503,634,685,711]" pageId="45" pageNumber="346" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
, pl. 7, fig. 10),
<collectionCode id="ED73AE576E781D25FC8FC6EC4954F78B" box="[802,847,685,711]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="45" pageNumber="346">GSI</collectionCode>
11899. Both specimens from the Shihtien Formation (Darriwilian) at Banpo Village in Shidian County, western
<collectingRegion id="49A6F8706E781D25FC84C6934999F7A0" box="[809,898,722,748]" country="China" name="Yunnan" pageId="45" pageNumber="346">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E781D25FF6AC6B44B4AF68F" pageId="45" pageNumber="346" type="discussion">
<paragraph id="8BDD36926E781D25FF6AC6B44B4AF68F" blockId="45.[151,1436,649,964]" pageId="45" pageNumber="346">
<emphasis id="B916EA806E781D25FF6AC6B44B20F643" bold="true" box="[199,315,757,783]" pageId="45" pageNumber="346">Remarks.</emphasis>
We were unable to make a satisfactory cast of the
<typeStatus id="54D988306E781D25FC29C6B449F5F643" box="[900,1006,757,783]" pageId="45" pageNumber="346" type="lectotype">lectotype</typeStatus>
, and Reeds original illustration is reproduced here. Although the dorsal exoskeleton is complete it is much abraded.
<bibRefCitation id="EFF34B636E781D25FB9DC7584EACF67F" author="Reed, F. R. C." box="[1072,1207,793,819]" pageId="45" pageNumber="346" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed (1917</bibRefCitation>
, p. 46) inferred that this was a blind species of “
<taxonomicName id="4C624D116E781D25FE79C77C4835F61B" box="[468,558,829,855]" class="Trilobita" family="Illaenidae" genus="Illaenus" kingdom="Animalia" order="Corynexochida" pageId="45" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E781D25FE79C77C4835F61B" box="[468,558,829,855]" italics="true" pageId="45" pageNumber="346">Illaenus</emphasis>
</taxonomicName>
” but we regard it as more probable that the abrasion of the dorsal surface has removed evidence of the facial sutures and the visual surfaces from a more typical illaenid. If this is so there are very few possible specific characters, and the species will not be recognisable until more collections are made from the type locality.
</paragraph>
</subSubSection>
</treatment>
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