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<document id="3137C54551662658C2F571DB57368BFB" ID-DOI="10.5852/ejt.2023.864.2083" ID-ISSN="2118-9773" ID-Zenodo-Dep="8011935" ID-ZooBank="4F73AB5F-A4B7-43AD-949E-A40FFEEABF4F" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" approvalRequired="16" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="6" approvalRequired_for_treatments="8" checkinTime="1680182262846" checkinUser="felipe" docAuthor="Vivallo, Felipe" docDate="2023" docId="03CB8782FFE1DA06AD65FEB35C26FD21" docLanguage="en" docName="ejt-864_1-27.pdf" docOrigin="European Journal of Taxonomy 864" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Centrislyngbyei Jensen-Haarup 1908" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFF2FFFAFFEBDA0DAF5DFFBB5E77FFAC" masterDocTitle="The primary types of some species of Centris bees described by European entomologists in the 18 and 20 centuries (Hymenoptera: Apidae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="11" updateTime="1693242320061" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="02FE9E1ED9742E3D95BE1EEA57FA8D52">The primary types of some species of Centris bees described by European entomologists in the 18 and 20 centuries (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="F032E3613915977627569F50B6F483D8">Vivallo, Felipe</mods:namePart>
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<subSubSection id="C378651FFFE1DA07AD65FEB35D8BFE8F" box="[568,1020,264,291]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BDD3694FFE1DA07AD65FEB35D8BFE8F" blockId="10.[568,1020,264,326]" box="[568,1020,264,291]" pageId="10" pageNumber="11">
<heading id="D09581F8FFE1DA07AD65FEB35D8BFE8F" box="[568,1020,264,291]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="4C624D17FFE1DA07AD65FEB35D8BFE8F" authority="Jensen-Haarup, 1908" authorityName="Jensen-Haarup" authorityYear="1908" box="[568,1020,264,291]" family="Apidae" genus="Centrislyngbyei" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" rank="genus">
<emphasis id="B916EA86FFE1DA07AD65FEB35C8BFE8F" bold="true" box="[568,764,264,291]" italics="true" pageId="10" pageNumber="11">Centrislyngbyei</emphasis>
<bibRefCitation id="EFF34B65FFE1DA07AC59FEB25D8BFE8F" author="Jensen-Haarup A. C." box="[772,1020,265,291]" pageId="10" pageNumber="11" pagination="107 - 108" refId="ref11351" refString="Jensen-Haarup A. C. 1908. Two new Argentine species of Apidae. Flora Fauna 8: 107 - 108." type="journal article" year="1908">Jensen-Haarup, 1908</bibRefCitation>
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<paragraph id="8BDD3694FFE1DA07ADB3FE975D32FEEA" blockId="10.[568,1020,264,326]" box="[750,837,300,326]" pageId="10" pageNumber="11">
<figureCitation id="13592A11FFE1DA07ADB3FE975D32FEEA" box="[750,837,300,326]" captionStart="Fig 2" captionStartId="10.[189,230,1963,1989]" captionTargetBox="[189,1399,1303,1895]" captionTargetId="figure-260@10.[189,1399,1300,1895]" captionTargetPageId="10" captionText="Fig 2. Data labels. A. Centriszonalis Dominique, 1898 (lectotype ♀). B. Centris lyngbyei Jensen- Haarup, 1908 (lectotype ♂)." figureDoi="http://doi.org/10.5281/zenodo.8011939" httpUri="https://zenodo.org/record/8011939/files/figure.png" pageId="10" pageNumber="11">Fig. 2B</figureCitation>
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FED55C8DFE24" blockId="10.[189,762,366,392]" box="[189,762,366,392]" pageId="10" pageNumber="11">
<emphasis id="B916EA86FFE1DA07AFE0FED55F09FE24" box="[189,382,366,392]" italics="true" pageId="10" pageNumber="11">Centrislyngbyei</emphasis>
<bibRefCitation id="EFF34B65FFE1DA07AED8FED55C84FE24" author="Jensen-Haarup A. C." box="[389,755,366,392]" pageId="10" pageNumber="11" pagination="107 - 108" refId="ref11351" refString="Jensen-Haarup A. C. 1908. Two new Argentine species of Apidae. Flora Fauna 8: 107 - 108." type="journal article" year="1908">Jensen-Haarup, 1908: 107108</bibRefCitation>
.
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FE145F4EFE65" blockId="10.[189,313,431,457]" box="[189,313,431,457]" pageId="10" pageNumber="11">
<heading id="D09581F8FFE1DA07AFE0FE145F4EFE65" bold="true" box="[189,313,431,457]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B916EA86FFE1DA07AFE0FE145F4EFE65" bold="true" box="[189,313,431,457]" pageId="10" pageNumber="11">
<typeStatus id="54D98836FFE1DA07AFE0FE145E8DFE65" box="[189,250,431,457]" pageId="10" pageNumber="11">Type</typeStatus>
data
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FE655B02FD43" blockId="10.[182,1399,477,751]" pageId="10" pageNumber="11">
To describe this species, Jensen-Haarup mentioned that he studied
<specimenCount id="9D64FD1DFFE1DA07AC96FE665A2AFE5B" box="[971,1117,477,503]" pageId="10" pageNumber="11" type="female">four females</specimenCount>
collected in Chacras de Coria,
<collectingCountry id="F3757604FFE1DA07AE5BFDBA5F0BFDB7" box="[262,380,513,539]" name="Argentina" pageId="10" pageNumber="11">Argentina</collectingCountry>
.However, the characters cited in the description belong to males, not females.The
<typeStatus id="54D98836FFE1DA07AA50FDBA5B01FDB7" box="[1293,1398,513,539]" pageId="10" pageNumber="11">lectotype</typeStatus>
specimen was designated by
<bibRefCitation id="EFF34B65FFE1DA07AD56FD9F5CC2FD92" author="Zanella F. C. V." box="[523,693,548,574]" pageId="10" pageNumber="11" pagination="435 - 488" refId="ref13430" refString="Zanella F. C. V. 2002. Sistematica, filogenia e distribuicao geografica das especies sul-americanas de Centris (Paracentris) Cameron, 1903 e de Centris (Penthemisia) Moure, 1950, incluindo uma analise filogenetica do &quot; grupo Centris &quot; sensu Ayala 1998 (Hymenoptera, Apoidea, Centridini). Revista Brasileira de Entomologia 46 (4): 435 - 488. https: // doi. org / 10.1590 / s 0085 - 56262002000400001" type="journal article" year="2002">Zanella (2002)</bibRefCitation>
and it bears the following data label: [black-rimmed]
<taxonomicName id="4C624D17FFE1DA07AA7FFD9F5F60FDCD" authority="Vivallo, 2023" authorityName="Vivallo" authorityYear="2023" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lyngbyei" status="n. sp.">
<emphasis id="B916EA86FFE1DA07AA7FFD9F5F60FDCD" italics="true" pageId="10" pageNumber="11">Centris lyngbyei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557FDFFE1DA07AE79FDF35F16FDCE" box="[292,353,584,610]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
J.- Hrp [handwriten] Jensen-Haarup det. [printed]\ [black-rimmed] Chacr. de Coria Prov. de
<collectingRegion id="49A6F876FFE1DA07AFE0FDD15F5FFD28" box="[189,296,618,644]" country="Argentina" name="Mendoza" pageId="10" pageNumber="11">Mendoza</collectingRegion>
Rep.
<collectingCountry id="F3757604FFE1DA07AE35FDD15FADFD28" box="[360,474,618,644]" name="Argentina" pageId="10" pageNumber="11">Argentina</collectingCountry>
Jensen-Haarup [handwritten]\ [black-rimmed] Type Coll. J = Hrp. [handwritten]\ [red label] Holo [handwritten]
<collectionCode id="ED73AE51FFE1DA07AD71FD355C03FD04" box="[556,628,654,680]" pageId="10" pageNumber="11">TYPE</collectionCode>
[printed]\ [white label with black discontinuous strokes]
<taxonomicName id="4C624D17FFE1DA07AA7FFD355F79FD67" authorityName="Friese" authorityYear="1900" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis id="B916EA86FFE1DA07AA7FFD355F79FD67" italics="true" pageId="10" pageNumber="11">Centris tricolor</emphasis>
</taxonomicName>
♀ [handwritten] 1907 Friese det. [printed] Fr. [handwritten]\ [pink label]
<typeStatus id="54D98836FFE1DA07AB28FD0A5B6EFD67" box="[1141,1305,689,715]" pageId="10" pageNumber="11">LECTOTYPE</typeStatus>
<taxonomicName id="4C624D17FFE1DA07AA7FFD0A5CAEFD42" authority="Jensen-Haarup, 1908 F. Zanella, 1999" authorityName="Jensen-Haarup, 1908 F. Zanella" authorityYear="1999" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lyngbyei">
<emphasis id="B916EA86FFE1DA07AA7FFD0A5F60FD42" italics="true" pageId="10" pageNumber="11">Centris lyngbyei</emphasis>
<bibRefCitation id="EFF34B65FFE1DA07AE78FD6E5C6FFD42" author="Jensen-Haarup A. C." box="[293,536,724,751]" pageId="10" pageNumber="11" pagination="107 - 108" refId="ref11351" refString="Jensen-Haarup A. C. 1908. Two new Argentine species of Apidae. Flora Fauna 8: 107 - 108." type="journal article" year="1908">Jensen-Haarup, 1908</bibRefCitation>
F. Zanella, 1999
</taxonomicName>
[printed]\ ZMUC 00240257 [printed] (
<collectionCode id="ED73AE51FFE1DA07ABC6FD6F5A80FD42" box="[1179,1271,724,750]" pageId="10" pageNumber="11">NHMD</collectionCode>
) (
<figureCitation id="13592A11FFE1DA07AA52FD6F5B1FFD43" box="[1295,1384,724,751]" captionStart="Fig 2" captionStartId="10.[189,230,1963,1989]" captionTargetBox="[189,1399,1303,1895]" captionTargetId="figure-260@10.[189,1399,1300,1895]" captionTargetPageId="10" captionText="Fig 2. Data labels. A. Centriszonalis Dominique, 1898 (lectotype ♀). B. Centris lyngbyei Jensen- Haarup, 1908 (lectotype ♂)." figureDoi="http://doi.org/10.5281/zenodo.8011939" httpUri="https://zenodo.org/record/8011939/files/figure.png" pageId="10" pageNumber="11">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FCAD5F2AFC83" blockId="10.[189,349,789,816]" box="[189,349,789,816]" pageId="10" pageNumber="11">
<heading id="D09581F8FFE1DA07AFE0FCAD5F2AFC83" bold="true" box="[189,349,789,816]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B916EA86FFE1DA07AFE0FCAD5F2AFC83" bold="true" box="[189,349,789,816]" pageId="10" pageNumber="11">
<typeStatus id="54D98836FFE1DA07AFE0FCAD5E8DFC9C" box="[189,250,790,816]" pageId="10" pageNumber="11">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FCFF5C88FCF2" blockId="10.[189,767,836,862]" box="[189,767,836,862]" pageId="10" pageNumber="11">
<collectingCountry id="F3757604FFE1DA07AFE0FCFF5F40FCF2" box="[189,311,836,862]" name="Argentina" pageId="10" pageNumber="11">Argentina</collectingCountry>
:
<collectingRegion id="49A6F876FFE1DA07AE1CFCFF5C56FCF2" box="[321,545,836,862]" country="Argentina" name="Mendoza" pageId="10" pageNumber="11">Mendoza Province</collectingRegion>
, Chacras de Coria.
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FC3D5C0BFC0F" blockId="10.[189,636,902,931]" box="[189,636,902,931]" pageId="10" pageNumber="11">
<emphasis id="B916EA86FFE1DA07AFE0FC3D5C0BFC0F" bold="true" box="[189,636,902,931]" italics="true" pageId="10" pageNumber="11">Johann Christoph Friedrich Klug</emphasis>
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FC005AA2FBB7" blockId="10.[189,1398,955,1052]" pageId="10" pageNumber="11">
Johann Christoph Friedrich Klug (17751856) was a German entomologist. He taught medicine and entomology at the Universität zu
<collectingRegion id="49A6F876FFE1DA07AD11FC655CE2FC54" box="[588,661,990,1016]" country="Germany" name="Berlin" pageId="10" pageNumber="11">Berlin</collectingRegion>
(currently the Humboldt-Universität zu
<collectingRegion id="49A6F876FFE1DA07AB2AFC655AB5FC55" box="[1143,1218,990,1017]" country="Germany" name="Berlin" pageId="10" pageNumber="11">Berlin</collectingRegion>
) where he also worked as curator of the insect collection (
<bibRefCitation id="EFF34B65FFE1DA07ADEDFBBA5DF3FBB0" author="Gerstaecker C. E. A." box="[688,900,1025,1052]" pageId="10" pageNumber="11" pagination="225 - 237" refId="ref11157" refString="Gerstaecker C. E. A. 1856. Nekrolog. Entomologische Zeitung 17: 225 - 237." type="journal article" year="1856">Gerstaecker 1856</bibRefCitation>
). He died in
<collectingRegion id="49A6F876FFE1DA07AB47FBBA5A11FBB7" box="[1050,1126,1025,1051]" country="Germany" name="Berlin" pageId="10" pageNumber="11">Berlin</collectingRegion>
, aged 80.
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FBF95FDBFBF1" blockId="10.[189,428,1090,1117]" box="[189,428,1090,1117]" pageId="10" pageNumber="11">
<heading id="D09581F8FFE1DA07AFE0FBF95FDBFBF1" bold="true" box="[189,428,1090,1117]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B916EA86FFE1DA07AFE0FBF95FDBFBF1" bold="true" box="[189,428,1090,1117]" pageId="10" pageNumber="11">
Klugs
<taxonomicName id="4C624D17FFE1DA07AE4BFBF85F19FBF1" authorityName="Fabricius" authorityYear="1804" box="[278,366,1091,1117]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA86FFE1DA07AE4BFBF85F19FBF1" bold="true" box="[278,366,1091,1117]" italics="true" pageId="10" pageNumber="11">Centris</emphasis>
</taxonomicName>
bees
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD3694FFE1DA07AFE0FBC95C38FB7E" blockId="10.[189,1398,1137,1234]" pageId="10" pageNumber="11">
In 1810, Klug proposed
<taxonomicName id="4C624D17FFE1DA07AEB8FBCA5C29FB27" authorityName="Klug" authorityYear="1810" box="[485,606,1137,1163]" class="Insecta" family="Apidae" genus="Ptilotopus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA86FFE1DA07AEB8FBCA5C29FB27" box="[485,606,1137,1163]" italics="true" pageId="10" pageNumber="11">Ptilotopus</emphasis>
</taxonomicName>
as a new genus, including in it a single species that he described simultaneously. The species is currently considered valid, but the genus was later downgraded and included as a subgenus of
<taxonomicName id="4C624D17FFE1DA07AEAFFB035C3FFB7E" authorityName="Fabricius" authorityYear="1804" box="[498,584,1208,1234]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA86FFE1DA07AEAFFB035C3FFB7E" box="[498,584,1208,1234]" italics="true" pageId="10" pageNumber="11">Centris</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF1D661CFFE1DA07AFE0F8105C7FF845" ID-DOI="http://doi.org/10.5281/zenodo.8011939" ID-Zenodo-Dep="8011939" httpUri="https://zenodo.org/record/8011939/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,230,1963,1989]" targetBox="[189,1399,1303,1895]" targetPageId="10">
<paragraph id="8BDD3694FFE1DA07AFE0F8105C7FF845" blockId="10.[189,1399,1963,2025]" pageId="10" pageNumber="11">
<emphasis id="B916EA86FFE1DA07AFE0F8105F7FF869" bold="true" box="[189,264,1963,1989]" pageId="10" pageNumber="11">Fig 2.</emphasis>
Data labels.
<emphasis id="B916EA86FFE1DA07AEEDF8105FB2F869" bold="true" box="[432,453,1963,1989]" pageId="10" pageNumber="11">A</emphasis>
.
<taxonomicName id="4C624D17FFE1DA07AE85F8175D1DF869" authority="Dominique, 1898" authorityName="Dominique" authorityYear="1898" box="[472,874,1963,1990]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zonalis">
<emphasis id="B916EA86FFE1DA07AE85F8175CFBF869" box="[472,652,1963,1990]" italics="true" pageId="10" pageNumber="11">Centris zonalis</emphasis>
<bibRefCitation id="EFF34B65FFE1DA07ADC5F8105D1DF869" author="Dominique J." box="[664,874,1963,1989]" pageId="10" pageNumber="11" pagination="57 - 65" refId="ref10986" refString="Dominique J. 1898. Coup d'oeil sur les melliferes sud-americains du Museum de Nantes. Bulletin de la Societe des Sciences naturelles de l'ouest de la France 8 (1): 57 - 65." type="journal article" year="1898">Dominique, 1898</bibRefCitation>
</taxonomicName>
(lectotype ♀).
<emphasis id="B916EA86FFE1DA07AB75F8105A4BF869" bold="true" box="[1064,1084,1963,1989]" pageId="10" pageNumber="11">B</emphasis>
.
<taxonomicName id="4C624D17FFE1DA07AB13F8175B64F869" authorityName="Jensen-Haarup, 1908 F. Zanella" authorityYear="1999" box="[1102,1299,1963,1990]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lyngbyei">
<emphasis id="B916EA86FFE1DA07AB13F8175B64F869" box="[1102,1299,1963,1990]" italics="true" pageId="10" pageNumber="11">Centris lyngbyei</emphasis>
</taxonomicName>
Jensen- Haarup, 1908 (lectotype ♂).
</paragraph>
</caption>
<paragraph id="8BDD3694FFE0DA06AFE0FEB25C26FD21" blockId="11.[189,1399,265,654]" pageId="11" pageNumber="12">
A couple of years before that publication,
<bibRefCitation id="EFF34B65FFE0DA06ADE6FEB25D38FE8F" author="Klug J. C. F." box="[699,847,265,291]" pageId="11" pageNumber="12" pagination="48 - 63" refId="ref11411" refString="Klug J. C. F. 1808. Uber die Geschlechtsverschiedenheit der Piezaten. Andere Halfte der Fabricius'schen Gattungen. Magazin des Gesellschaft naturforschender Freunde 2: 48 - 63." type="journal article" year="1808">Klug (1808)</bibRefCitation>
published an article on sexual dimorphism in
<taxonomicName id="4C624D17FFE0DA06AFE0FE955F29FEE4" authorityName="Linnaeus" authorityYear="1758" box="[189,350,302,328]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
, citing some examples in several genera of wasps, ants and bees. Among his comments on species of
<taxonomicName id="4C624D17FFE0DA06AE3DFEE95FC0FEC0" authorityName="Fabricius" authorityYear="1804" box="[352,439,338,364]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA86FFE0DA06AE3DFEE95FC0FEC0" box="[352,439,338,364]" italics="true" pageId="11" pageNumber="12">Centris</emphasis>
</taxonomicName>
, he cited “
<taxonomicName id="4C624D17FFE0DA06AD64FEEA5CE5FEC7" box="[569,658,337,363]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="byssina">
<emphasis id="B916EA86FFE0DA06AD64FEEA5CE5FEC7" box="[569,658,337,363]" italics="true" pageId="11" pageNumber="12">byssina</emphasis>
</taxonomicName>
”, a species of
<taxonomicName id="4C624D17FFE0DA06AC1DFEE95DD4FEC7" authorityName="Klug" authorityYear="1807" box="[832,931,338,363]" class="Insecta" family="Apidae" genus="Hemisia" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA86FFE0DA06AC1DFEE95DD4FEC7" box="[832,931,338,363]" italics="true" pageId="11" pageNumber="12">Hemisia</emphasis>
</taxonomicName>
that had previously been named by the German entomologist and zoologist Johann Karl Wilhelm Illiger (17751813) but that he never formally described. Klug provided some morphological features that allow the identification of this species and the sex of the specimens he studied, resulting in its formal description. The species remained unnoticed during all these years, until a visit to the
<collectionCode id="ED73AE51FFE0DA06ADC4FE585CAEFE51" box="[665,729,483,509]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="11" pageNumber="12" type="Museum">ZMB</collectionCode>
where I studied the specimens described by Klug, which had previously been recognized as such by another entomologist. The researcher designated one of the specimens as
<typeStatus id="54D98836FFE0DA06AE3BFD905FA1FDE9" box="[358,470,555,581]" pageId="11" pageNumber="12">lectotype</typeStatus>
, but unfortunately, the nomenclatural act was never published. To formalize this action and avoid future taxonomic problems, this designation is published here, acknowledging the previous work carried out by him.
</paragraph>
</subSubSection>
</treatment>
</document>