249 lines
40 KiB
XML
249 lines
40 KiB
XML
<document id="C9F16444D3920253CF7F007430369763" ID-CLB-Dataset="276818" ID-DOI="10.11646/zootaxa.5178.3.5" ID-GBIF-Dataset="941b1506-c7c8-4ea0-95e3-054be82c7fd8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7026260" ID-ZooBank="911C1854-08AD-4FF9-8374-3197753AB1C8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661535791432" checkinUser="plazi" docAuthor="Mu, Yannan & Zhang, Feng" docDate="2022" docId="A535F669FFCBC4231CD2E6E9FE2B3E53" docLanguage="en" docName="zootaxa.5178.3.5.pdf" docOrigin="Zootaxa 5178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lingulatus Mu & Zhang 2022, gen. nov." docType="treatment" docVersion="5" lastPageNumber="267" masterDocId="590C8E11FFCAC4211C45E717FFDD3F12" masterDocTitle="Lingulatus gen. nov., a new genus with description of three new species and one new combination (Araneae: Phrurolithidae)" masterLastPageNumber="277" masterPageNumber="265" pageNumber="266" updateTime="1699377397059" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B0B01728C3FAEA5AA17F1967AF14BBEB">Lingulatus gen. nov., a new genus with description of three new species and one new combination (Araneae: Phrurolithidae)</mods:title>
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<mods:name id="3685AAC1F5D84300737EAA5BFDFB5998" type="personal">
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<mods:roleTerm id="85765831B609D09A8177F080758A01EE">Author</mods:roleTerm>
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<mods:namePart id="30F729BB9454B894D396239AF1883C31">Mu, Yannan</mods:namePart>
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<mods:nameIdentifier id="B49DDF89C7536A5971E8DE6ABC30500C" type="ORCID">0000-0002-2504-673X</mods:nameIdentifier>
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<mods:affiliation id="E14484936A8E1BB86C1171F74A695AF4">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, P. R. China & 15188605531 @ 163. com; https: // orcid. org / 0000 - 0002 - 2504 - 673 X</mods:affiliation>
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<mods:namePart id="AFD742BBB928965296DADD703BA9AC0C">Zhang, Feng</mods:namePart>
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<mods:nameIdentifier id="127B8C143F4B56256A6552CCAE7E3106" type="ORCID">0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="F51B54A9D27FED38807C25CA2DAC12C9">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, P. R. China & dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031</mods:affiliation>
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<mods:title id="060299FD66B72C9DCBCD4E98DED62A1E">Zootaxa</mods:title>
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<mods:part id="C1749CC933DC72E3969E09A89E1F80BA">
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<mods:date id="B35A43454B3E7DFB303F05779FBFD734">2022</mods:date>
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<mods:number id="9B0A792655D001806D04F9BD2EF01DC9">2022-08-26</mods:number>
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<mods:number id="D61B4288642061D2B6DD2F0F7D85DA4A">5178</mods:number>
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<treatment id="A535F669FFCBC4231CD2E6E9FE2B3E53" ID-DOI="http://doi.org/10.5281/zenodo.7026142" ID-GBIF-Taxon="217309006" ID-Zenodo-Dep="7026142" LSID="urn:lsid:plazi:treatment:A535F669FFCBC4231CD2E6E9FE2B3E53" httpUri="http://treatment.plazi.org/id/A535F669FFCBC4231CD2E6E9FE2B3E53" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266">
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<subSubSection id="658614F4FFCBC4201CD2E6E9FDD93D0A" box="[151,516,510,537]" pageId="1" pageNumber="266" type="nomenclature">
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<paragraph id="2D23477FFFCBC4201CD2E6E9FDD93D0A" blockId="1.[151,516,510,537]" box="[151,516,510,537]" pageId="1" pageNumber="266">
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<heading id="766BF013FFCBC4201CD2E6E9FDD93D0A" bold="true" box="[151,516,510,537]" fontSize="11" level="1" pageId="1" pageNumber="266" reason="1">
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<emphasis id="1FE89B6DFFCBC4201CD2E6E9FDD93D0A" bold="true" box="[151,516,510,537]" pageId="1" pageNumber="266">
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<taxonomicName id="EA9C3CFCFFCBC4201CD2E6E9FEC63D0B" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[151,283,510,537]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1FE89B6DFFCBC4201CD2E6E9FEC63D0B" bold="true" box="[151,283,510,537]" italics="true" pageId="1" pageNumber="266">Lingulatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="04DB2616FFCBC4201D66E6E8FE4D3D0B" box="[291,400,511,537]" pageId="1" pageNumber="266" rank="genus">gen. nov.</taxonomicNameLabel>
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(fflfla)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="658614F4FFCBC4201CD2E550FC313D97" pageId="1" pageNumber="266" type="etymology">
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<paragraph id="2D23477FFFCBC4201CD2E550FC313D97" blockId="1.[151,1437,583,2013]" pageId="1" pageNumber="266">
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<emphasis id="1FE89B6DFFCBC4201CD2E550FEC33D73" bold="true" box="[151,286,583,609]" pageId="1" pageNumber="266">Etymology.</emphasis>
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The generic name “
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<taxonomicName id="EA9C3CFCFFCBC4201E45E550FDA53D73" authorityName="Mu & Zhang" authorityYear="2022" box="[512,632,583,609]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="genus">
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<emphasis id="1FE89B6DFFCBC4201E45E550FDA53D73" box="[512,632,583,609]" italics="true" pageId="1" pageNumber="266">Lingulatus</emphasis>
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</taxonomicName>
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” is a Latin adjective meaning “tongue-shaped”, referring to the tongueshaped tegular sheet originating from the tegulum. The gender is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="658614F4FFCBC4201C82E599FA4E3B27" pageId="1" pageNumber="266" type="diagnosis">
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<paragraph id="2D23477FFFCBC4201C82E599FA4E3B27" blockId="1.[151,1437,583,2013]" pageId="1" pageNumber="266">
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<emphasis id="1FE89B6DFFCBC4201C82E599FE9F3DBA" bold="true" box="[199,322,654,680]" pageId="1" pageNumber="266">Diagnosis.</emphasis>
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This new genus can be distinguished from all other Asian phrurolithid genera by the tegular sheet originating from the tegulum, located near the base of the embolus, combined with the tibia with a ventral tegular apophysis and retrodorsal tibial apophysis; the new genus can also be distinguished from all other phrurolithid genera by the round ventral tibial apophysis or regular, tapering, pointed apophysis located at the middle of the ventral surface of the tibia. This new genus is similar to
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<taxonomicName id="EA9C3CFCFFCBC4201F4AE408FBA83C2B" authority="Wunderlich, 1992" authorityName="Wunderlich" authorityYear="1992" box="[783,1141,799,825]" class="Arachnida" family="Phrurolithidae" genus="Liophrurillus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="genus">
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<emphasis id="1FE89B6DFFCBC4201F4AE408FC793C2B" box="[783,932,799,825]" italics="true" pageId="1" pageNumber="266">Liophrurillus</emphasis>
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Wunderlich, 1992
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</taxonomicName>
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by having a tegular sheet, but can be distinguished by: 1) embolus thin, acicular-shaped or flake-shaped (vs extremely stout, hook-shaped, compare
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<figureCitation id="B5A75BFAFFCBC4201CBAE470FE843C93" box="[255,345,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1608,1632]" captionTargetBox="[158,1426,191,1577]" captionTargetId="figure-22@4.[151,1436,181,1584]" captionTargetPageId="4" captionText="FIGURE 3. Lingulatus brevis sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view.Abbreviations:E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TA—tegular apophysis; TS—tegular sheet; vTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7026266" httpUri="https://zenodo.org/record/7026266/files/figure.png" pageId="1" pageNumber="266">Figs 3C</figureCitation>
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,
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<figureCitation id="B5A75BFAFFCBC4201D23E470FE543C93" box="[358,393,871,897]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[198,1399,182,1906]" captionTargetId="figure-22@8.[179,1408,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Lingulatus longulus sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA— retrolateral tibial apophysis; TA—tegular apophysis; TS—tegular sheet; vTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7026272" httpUri="https://zenodo.org/record/7026272/files/figure.png" pageId="1" pageNumber="266">6B</figureCitation>
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,
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<figureCitation id="B5A75BFAFFCBC4201DD0E470FE683C93" box="[405,437,871,897]" captionStart="FIGURE 8" captionStartId="10.[151,250,1826,1850]" captionTargetBox="[155,1436,741,1795]" captionTargetId="figure-276@10.[151,1436,741,1803]" captionTargetPageId="10" captionText="FIGURE 8. Lingulatus pingbian sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TS—tegular sheet; vTA—ventral tibial apophysis; B—bursa; CD—copulatory duct; CO—copulatory opening; CT—connecting tube; GA—glandular appendages; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.7026283" httpUri="https://zenodo.org/record/7026283/files/figure.png" pageId="1" pageNumber="266">8B</figureCitation>
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with fig.
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<bibRefCitation id="490D3A8EFFCBC4201E26E470FC883C93" author="Wunderlich, J." box="[611,853,871,897]" pageId="1" pageNumber="266" pagination="1 - 246" refId="ref6014" refString="Wunderlich, J. (2012) Fifteen papers on extant and fossil spiders (Araneae) Beitrage zur Araneologie, 7, 1 - 246." type="journal article" year="2012">Wunderlich 2012: 22</bibRefCitation>
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); 2) tegular sheet medium or small in size (vs thick, compare
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<figureCitation id="B5A75BFAFFCBC4201D45E49CFE863CB7" box="[256,347,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1608,1632]" captionTargetBox="[158,1426,191,1577]" captionTargetId="figure-22@4.[151,1436,181,1584]" captionTargetPageId="4" captionText="FIGURE 3. Lingulatus brevis sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view.Abbreviations:E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TA—tegular apophysis; TS—tegular sheet; vTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7026266" httpUri="https://zenodo.org/record/7026266/files/figure.png" pageId="1" pageNumber="266">Figs 3C</figureCitation>
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,
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<figureCitation id="B5A75BFAFFCBC4201D2DE49CFE503CB7" box="[360,397,907,933]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[198,1399,182,1906]" captionTargetId="figure-22@8.[179,1408,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Lingulatus longulus sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA— retrolateral tibial apophysis; TA—tegular apophysis; TS—tegular sheet; vTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7026272" httpUri="https://zenodo.org/record/7026272/files/figure.png" pageId="1" pageNumber="266">6A</figureCitation>
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,
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<figureCitation id="B5A75BFAFFCBC4201DDCE49CFE673CB7" box="[409,442,907,933]" captionStart="FIGURE 8" captionStartId="10.[151,250,1826,1850]" captionTargetBox="[155,1436,741,1795]" captionTargetId="figure-276@10.[151,1436,741,1803]" captionTargetPageId="10" captionText="FIGURE 8. Lingulatus pingbian sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TS—tegular sheet; vTA—ventral tibial apophysis; B—bursa; CD—copulatory duct; CO—copulatory opening; CT—connecting tube; GA—glandular appendages; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.7026283" httpUri="https://zenodo.org/record/7026283/files/figure.png" pageId="1" pageNumber="266">8B</figureCitation>
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with fig.
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); 3) femoral apophysis well-developed (vs a small triangular protrusion, compare
|
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<figureCitation id="B5A75BFAFFCBC4201DB0E4B8FD8C3CDB" box="[501,593,943,969]" captionStart="FIGURE 3" captionStartId="4.[151,250,1608,1632]" captionTargetBox="[158,1426,191,1577]" captionTargetId="figure-22@4.[151,1436,181,1584]" captionTargetPageId="4" captionText="FIGURE 3. Lingulatus brevis sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view.Abbreviations:E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TA—tegular apophysis; TS—tegular sheet; vTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7026266" httpUri="https://zenodo.org/record/7026266/files/figure.png" pageId="1" pageNumber="266">Figs 3A</figureCitation>
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,
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<figureCitation id="B5A75BFAFFCBC4201E1BE4B8FD5E3CDB" box="[606,643,943,969]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[198,1399,182,1906]" captionTargetId="figure-22@8.[179,1408,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Lingulatus longulus sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA— retrolateral tibial apophysis; TA—tegular apophysis; TS—tegular sheet; vTA—ventral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.7026272" httpUri="https://zenodo.org/record/7026272/files/figure.png" pageId="1" pageNumber="266">6A</figureCitation>
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,
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<figureCitation id="B5A75BFAFFCBC4201ECAE4B8FD6D3CDB" box="[655,688,943,969]" captionStart="FIGURE 8" captionStartId="10.[151,250,1826,1850]" captionTargetBox="[155,1436,741,1795]" captionTargetId="figure-276@10.[151,1436,741,1803]" captionTargetPageId="10" captionText="FIGURE 8. Lingulatus pingbian sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TS—tegular sheet; vTA—ventral tibial apophysis; B—bursa; CD—copulatory duct; CO—copulatory opening; CT—connecting tube; GA—glandular appendages; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.7026283" httpUri="https://zenodo.org/record/7026283/files/figure.png" pageId="1" pageNumber="266">8A</figureCitation>
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with fig.
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<bibRefCitation id="490D3A8EFFCBC4201F1AE4B8FB8C3CDB" author="Wunderlich, J." box="[863,1105,943,969]" pageId="1" pageNumber="266" pagination="1 - 246" refId="ref6014" refString="Wunderlich, J. (2012) Fifteen papers on extant and fossil spiders (Araneae) Beitrage zur Araneologie, 7, 1 - 246." type="journal article" year="2012">Wunderlich 2012: 22</bibRefCitation>
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); 4) the thin copulatory ducts and connecting tubes (vs thick and snail-shaped, compare
|
||
<figureCitation id="B5A75BFAFFCBC4201F69E4C4FC743CFF" box="[812,937,979,1005]" captionStart="FIGURE 4" captionStartId="5.[151,250,1111,1135]" captionTargetBox="[162,1434,530,1076]" captionTargetId="figure-178@5.[151,1436,525,1088]" captionTargetPageId="5" captionText="FIGURE 4. Lingulatus brevis sp. nov.: A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: B—bursa; CD— copulatory duct; CO—copulatory opening; CT—connecting tube; FD—fertilization duct; GA—glandular appendage; S— spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7026268" httpUri="https://zenodo.org/record/7026268/files/figure.png" pageId="1" pageNumber="266">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="B5A75BFAFFCBC4201FF3E4C4FC2F3CFF" box="[950,1010,979,1005]" captionStart="FIGURE 8" captionStartId="10.[151,250,1826,1850]" captionTargetBox="[155,1436,741,1795]" captionTargetId="figure-276@10.[151,1436,741,1803]" captionTargetPageId="10" captionText="FIGURE 8. Lingulatus pingbian sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TS—tegular sheet; vTA—ventral tibial apophysis; B—bursa; CD—copulatory duct; CO—copulatory opening; CT—connecting tube; GA—glandular appendages; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.7026283" httpUri="https://zenodo.org/record/7026283/files/figure.png" pageId="1" pageNumber="266">8E–F</figureCitation>
|
||
with figs
|
||
<quantity id="EA64EA9AFFCBC4201823E4C4FB173CFF" box="[1126,1226,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="7.874" metricValueMin="7.62" pageId="1" pageNumber="266" unit="in" value="30.5" valueMax="31.0" valueMin="30.0">30–31 in</quantity>
|
||
<bibRefCitation id="490D3A8EFFCBC4201894E4C4FF6A3B03" author="Wunderlich, J." pageId="1" pageNumber="266" pagination="1 - 246" refId="ref6014" refString="Wunderlich, J. (2012) Fifteen papers on extant and fossil spiders (Araneae) Beitrage zur Araneologie, 7, 1 - 246." type="journal article" year="2012">Wunderlich 2012: 22</bibRefCitation>
|
||
); 5) the presence of glandular appendages and bursa (vs absent, compare
|
||
<figureCitation id="B5A75BFAFFCBC4201FACE4E0FB9F3B03" box="[1001,1090,1015,1041]" captionStart="FIGURE 4" captionStartId="5.[151,250,1111,1135]" captionTargetBox="[162,1434,530,1076]" captionTargetId="figure-178@5.[151,1436,525,1088]" captionTargetPageId="5" captionText="FIGURE 4. Lingulatus brevis sp. nov.: A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: B—bursa; CD— copulatory duct; CO—copulatory opening; CT—connecting tube; FD—fertilization duct; GA—glandular appendage; S— spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7026268" httpUri="https://zenodo.org/record/7026268/files/figure.png" pageId="1" pageNumber="266">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="B5A75BFAFFCBC420180BE4E0FBB63B03" box="[1102,1131,1015,1041]" captionStart="FIGURE 8" captionStartId="10.[151,250,1826,1850]" captionTargetBox="[155,1436,741,1795]" captionTargetId="figure-276@10.[151,1436,741,1803]" captionTargetPageId="10" captionText="FIGURE 8. Lingulatus pingbian sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TS—tegular sheet; vTA—ventral tibial apophysis; B—bursa; CD—copulatory duct; CO—copulatory opening; CT—connecting tube; GA—glandular appendages; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.7026283" httpUri="https://zenodo.org/record/7026283/files/figure.png" pageId="1" pageNumber="266">8F</figureCitation>
|
||
with figs
|
||
<quantity id="EA64EA9AFFCBC420189EE4E0FACE3B03" box="[1243,1299,1015,1041]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="1" pageNumber="266" unit="in" value="31.0">31 in</quantity>
|
||
<bibRefCitation id="490D3A8EFFCBC420195CE4E0FF233B27" author="Wunderlich, J." pageId="1" pageNumber="266" pagination="1 - 246" refId="ref6014" refString="Wunderlich, J. (2012) Fifteen papers on extant and fossil spiders (Araneae) Beitrage zur Araneologie, 7, 1 - 246." type="journal article" year="2012">Wunderlich 2012: 22</bibRefCitation>
|
||
); 6) copulatory openings small (vs large, compare
|
||
<figureCitation id="B5A75BFAFFCBC4201F70E30CFC4C3B27" box="[821,913,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[151,250,1111,1135]" captionTargetBox="[162,1434,530,1076]" captionTargetId="figure-178@5.[151,1436,525,1088]" captionTargetPageId="5" captionText="FIGURE 4. Lingulatus brevis sp. nov.: A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: B—bursa; CD— copulatory duct; CO—copulatory opening; CT—connecting tube; FD—fertilization duct; GA—glandular appendage; S— spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7026268" httpUri="https://zenodo.org/record/7026268/files/figure.png" pageId="1" pageNumber="266">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="B5A75BFAFFCBC4201FD9E30CFC613B27" box="[924,956,1051,1077]" captionStart="FIGURE 8" captionStartId="10.[151,250,1826,1850]" captionTargetBox="[155,1436,741,1795]" captionTargetId="figure-276@10.[151,1436,741,1803]" captionTargetPageId="10" captionText="FIGURE 8. Lingulatus pingbian sp. nov.: A. Male left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; FA—femoral apophysis; rDTA—retrodorsal tibial apophysis; RTA—retrolateral tibial apophysis; TS—tegular sheet; vTA—ventral tibial apophysis; B—bursa; CD—copulatory duct; CO—copulatory opening; CT—connecting tube; GA—glandular appendages; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.7026283" httpUri="https://zenodo.org/record/7026283/files/figure.png" pageId="1" pageNumber="266">8E</figureCitation>
|
||
with figs 30,
|
||
<quantity id="EA64EA9AFFCBC4201813E30CFB523B27" box="[1110,1167,1051,1077]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="1" pageNumber="266" unit="in" value="32.0">32 in</quantity>
|
||
<bibRefCitation id="490D3A8EFFCBC42018D3E30CFA5B3B27" author="Wunderlich, J." box="[1174,1414,1051,1077]" pageId="1" pageNumber="266" pagination="1 - 246" refId="ref6014" refString="Wunderlich, J. (2012) Fifteen papers on extant and fossil spiders (Araneae) Beitrage zur Araneologie, 7, 1 - 246." type="journal article" year="2012">Wunderlich 2012: 22</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="658614F4FFCBC4201C82E329FCCD3817" pageId="1" pageNumber="266" type="description">
|
||
<paragraph id="2D23477FFFCBC4201C82E329FBF0393F" blockId="1.[151,1437,583,2013]" pageId="1" pageNumber="266">
|
||
<emphasis id="1FE89B6DFFCBC4201C82E329FE843B4A" bold="true" box="[199,345,1086,1112]" pageId="1" pageNumber="266">Description.</emphasis>
|
||
Small spiders, total length
|
||
<quantity id="EA64EA9AFFCBC4201EEDE328FC893B4B" box="[680,852,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="3.295" metricValueMax="3.71" metricValueMin="2.88" pageId="1" pageNumber="266" unit="mm" value="3.295" valueMax="3.71" valueMin="2.88">2.88–3.71 mm</quantity>
|
||
. Carapace with nearly round thoracic region and narrowed cephalic region, smooth, yellow to blackish-brown, widest at coxae II, highest near fovea. Cervical groove inconspicuous. Fovea longitudinal, dark-red. Eight eyes in two rows, PER slightly wider than AER, AER recurved in dorsal view, PER slightly recurved in dorsal view. Eyes moderate in size, AME almost same size as ALE, larger than PME and PLE. Chelicerae yellow, promargin with three well-separated teeth and retromargin with four denticles close together (
|
||
<figureCitation id="B5A75BFAFFCBC4201E63E3E4FD403A1F" box="[550,669,1267,1293]" captionStart="FIGURE 1" captionStartId="2.[151,250,1901,1925]" captionTargetBox="[155,1436,531,1877]" captionTargetId="figure-105@2.[151,1436,525,1877]" captionTargetPageId="2" captionText="FIGURE 1. Left chelicerae: A. Lingulatus pingbian sp. nov., retromarginal view (male holotype); B. Same, retromarginal view (female paratype); C. Same, promarginal view (female paratype); D. L. longulus sp. nov., retromarginal view (male holotype); E. L. brevis sp. nov., retromarginal view (male holotype); F. Same, retromarginal view (female paratype)." figureDoi="http://doi.org/10.5281/zenodo.7026262" httpUri="https://zenodo.org/record/7026262/files/figure.png" pageId="1" pageNumber="266">Fig. 1E, F</figureCitation>
|
||
), with one long and one short spine anteriorly. Endites longer than wide, labium wider than long. Sternum yellow to black, smooth, without pattern, longer than wide, shield-shaped; precoxal triangles present, intercoxal sclerites absent. Legs yellow to dark brown; all femora with one to three dorsal spines, femur I with four prolateral spines distally, femur II with two or three prolateral spines distally; tibia I usually with six to seven proventral spines and six to eight retroventral spines, tibia II usually with six or seven proventral spines and five or six retroventral spines; metatarsus I usually with four pairs of ventral spines, metatarsus II usually with four proventral spines and three retroventral spines; posterior legs lack spines; metatarsi III and IV with distal preening brush. Leg formula: 1423. Abdomen oval, dorsum black, with dorsal scutum in males; venter yellow; colulus absent; transverse row of white hairs present just anterior of anal tubercle.
|
||
</paragraph>
|
||
<paragraph id="2D23477FFFCBC4201C82E120FCCD3817" blockId="1.[151,1437,583,2013]" pageId="1" pageNumber="266">Male palp: femur with well-developed femoral apophysis located at middle of ventral surface; tibia with round or modified ventral tibial apophysis, medium-sized tooth-shaped retrolateral apophysis, and large retrodorsal apophysis; embolus acicular or flake-shaped; tegulum oval, tegular apophysis present or absent, tegular sheet originating near base of embolus on tegulum; conductor absent. Female epigyne: epigynal plate weakly sclerotized; copulatory openings small and far apart; spermathecae small, located posteriorly; glandular appendages digit- or mastoid-shaped; a pair of long and oval bursa anteriorly.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="658614F4FFCBC4201C82E018FC2B3863" pageId="1" pageNumber="266" type="type_taxon">
|
||
<paragraph id="2D23477FFFCBC4201C82E018FCE5383B" blockId="1.[151,1437,583,2013]" box="[199,824,1806,1833]" pageId="1" pageNumber="266">
|
||
<emphasis id="1FE89B6DFFCBC4201C82E018FEB8383A" bold="true" box="[199,357,1806,1833]" pageId="1" pageNumber="266">
|
||
<typeStatus id="F227F9DDFFCBC4201C82E018FEDF383B" box="[199,258,1807,1833]" pageId="1" pageNumber="266">Type</typeStatus>
|
||
species:
|
||
</emphasis>
|
||
<taxonomicName id="EA9C3CFCFFCBC4201D29E018FD93383B" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[364,590,1807,1833]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="pingbian" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFCBC4201D29E018FD93383B" box="[364,590,1807,1833]" italics="true" pageId="1" pageNumber="266">Lingulatus pingbian</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201E10E018FD73383B" bold="true" box="[597,686,1807,1833]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201E10E018FD73383B" box="[597,686,1807,1833]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="1FE89B6DFFCBC4201EFAE018FCF23834" bold="true" box="[703,815,1807,1830]" pageId="1" pageNumber="266">屏ṁfflfl</emphasis>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="2D23477FFFCBC4201C82E025FC2B3863" blockId="1.[151,1437,583,2013]" pageId="1" pageNumber="266">
|
||
<emphasis id="1FE89B6DFFCBC4201C82E025FEBA385E" bold="true" box="[199,359,1842,1868]" pageId="1" pageNumber="266">Composition.</emphasis>
|
||
Four species, including three new species described here:
|
||
<taxonomicName id="EA9C3CFCFFCBC4201FABE024FB93385F" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[1006,1102,1843,1869]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFCBC4201FABE024FB93385F" box="[1006,1102,1843,1869]" italics="true" pageId="1" pageNumber="266">L. brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201811E024FB70385F" bold="true" box="[1108,1197,1843,1869]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201811E024FB70385F" box="[1108,1197,1843,1869]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EA9C3CFCFFCBC42018FFE024FAEB385F" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[1210,1334,1843,1869]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="longulus" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFCBC42018FFE024FAEB385F" box="[1210,1334,1843,1869]" italics="true" pageId="1" pageNumber="266">L. longulus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201978E024FA48385F" bold="true" box="[1341,1429,1843,1869]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201978E024FA48385F" box="[1341,1429,1843,1869]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EA9C3CFCFFCBC4201CD2E040FEC43863" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[151,281,1879,1905]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="pingbian" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFCBC4201CD2E040FEC43863" box="[151,281,1879,1905]" italics="true" pageId="1" pageNumber="266">L. pingbian</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201D5AE040FEA43863" bold="true" box="[287,377,1879,1905]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201D5AE040FEA43863" box="[287,377,1879,1905]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EA9C3CFCFFCBC4201DEAE040FCB53863" ID-CoL="6TCLW" authority="(Jager & Wunderlich, 2012)" authorityName="Mu & Zhang" authorityYear="2022" baseAuthorityName="Jager & Wunderlich" baseAuthorityYear="2012" box="[431,872,1879,1905]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="christae" status="comb. nov.">
|
||
<emphasis id="1FE89B6DFFCBC4201DEAE040FDFA3863" box="[431,551,1879,1905]" italics="true" pageId="1" pageNumber="266">L. christae</emphasis>
|
||
(
|
||
<bibRefCitation id="490D3A8EFFCBC4201E73E040FCBC3863" author="Jager, P. & Wunderlich, J." box="[566,865,1879,1905]" pageId="1" pageNumber="266" pagination="251 - 271" refId="ref5582" refString="Jager, P. & Wunderlich, J. (2012) Seven new species of the spider genus Otacilia Thorell 1897 (Araneae: Corinnidae) from China, Laos and Thailand. Beitrage zur Araneologie, 7, 251 - 271 + 352 - 357." type="journal article" year="2012">Jäger & Wunderlich, 2012</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201F2AE040FC323863" bold="true" box="[879,1007,1879,1905]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201F2AE040FC323863" box="[879,1007,1879,1905]" pageId="1" pageNumber="266" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="658614F4FFCBC4231C82E06CFD8E3E0F" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266" type="discussion">
|
||
<paragraph id="2D23477FFFCBC4231C82E06CFD8E3E0F" blockId="1.[151,1437,583,2013]" lastBlockId="2.[151,1436,150,321]" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266">
|
||
<emphasis id="1FE89B6DFFCBC4201C82E06CFEE63887" bold="true" box="[199,315,1915,1941]" pageId="1" pageNumber="266">Remarks.</emphasis>
|
||
Members of this genus can be divided into two species groups. Firstly, the
|
||
<emphasis id="1FE89B6DFFCBC4201886E06CFAED3887" box="[1219,1328,1915,1941]" italics="true" pageId="1" pageNumber="266">pingbian-</emphasis>
|
||
group (
|
||
<taxonomicName id="EA9C3CFCFFCBC42019C3E06CFF2A38AB" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="longulus" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFCBC42019C3E06CFF2A38AB" italics="true" pageId="1" pageNumber="266">L. longulus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201CBBE088FE8A38AB" bold="true" box="[254,343,1951,1977]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201CBBE088FE8A38AB" box="[254,343,1951,1977]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EA9C3CFCFFCBC4201DC8E088FDD338AB" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[397,526,1951,1977]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="pingbian" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFCBC4201DC8E088FDD338AB" box="[397,526,1951,1977]" italics="true" pageId="1" pageNumber="266">L. pingbian</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFCBC4201E51E088FDB038AB" bold="true" box="[532,621,1951,1977]" pageId="1" pageNumber="266">
|
||
<taxonomicNameLabel id="04DB2616FFCBC4201E51E088FDB038AB" box="[532,621,1951,1977]" pageId="1" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) is characterized by: 1) DS is narrower than the abdomen, and more than half the length of the abdomen; 2) embolus flattened; 3) vTA with a small tip directed towards the tegulum; 4) TS longer than half the length of the tegulum; and 5) COs located posteriorly. Secondly, the
|
||
<emphasis id="1FE89B6DFFC8C4231830E780FB1F3FA3" box="[1141,1218,151,177]" italics="true" pageId="2" pageNumber="267">brevis-</emphasis>
|
||
group (
|
||
<taxonomicName id="EA9C3CFCFFC8C4231956E780FAA93FA3" authority="Mu & Zhang, 2022" authorityName="Mu & Zhang" authorityYear="2022" box="[1299,1396,151,177]" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="267" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="1FE89B6DFFC8C4231956E780FAA93FA3" box="[1299,1396,151,177]" italics="true" pageId="2" pageNumber="267">L. brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFC8C423193EE780FF153FC7" bold="true" pageId="2" pageNumber="267">
|
||
<taxonomicNameLabel id="04DB2616FFC8C423193EE780FF153FC7" pageId="2" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="EA9C3CFCFFC8C4231D47E7ACFD1E3FC7" authority="(Jager & Wunderlich, 2012)" authorityName="Mu & Zhang" authorityYear="2022" baseAuthorityName="Jager & Wunderlich" baseAuthorityYear="2012" box="[258,707,186,213]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="267" phylum="Arthropoda" rank="species" species="christae" status="comb. nov.">
|
||
<emphasis id="1FE89B6DFFC8C4231D47E7ACFEA63FC7" box="[258,379,187,213]" italics="true" pageId="2" pageNumber="267">L. christae</emphasis>
|
||
<emphasis id="1FE89B6DFFC8C4231DC1E7ADFE503FC6" bold="true" box="[388,397,186,212]" pageId="2" pageNumber="267">(</emphasis>
|
||
<bibRefCitation id="490D3A8EFFC8C4231DC8E7ACFD663FC7" author="Jager, P. & Wunderlich, J." box="[397,699,187,213]" pageId="2" pageNumber="267" pagination="251 - 271" refId="ref5582" refString="Jager, P. & Wunderlich, J. (2012) Seven new species of the spider genus Otacilia Thorell 1897 (Araneae: Corinnidae) from China, Laos and Thailand. Beitrage zur Araneologie, 7, 251 - 271 + 352 - 357." type="journal article" year="2012">Jäger & Wunderlich, 2012</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="1FE89B6DFFC8C4231E8EE7ACFCEE3FC7" bold="true" box="[715,819,187,213]" pageId="2" pageNumber="267">
|
||
<taxonomicNameLabel id="04DB2616FFC8C4231E8EE7ACFCEE3FC7" box="[715,819,187,213]" pageId="2" pageNumber="267" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) is characterized by: 1) DS large, almost covering the entire abdomen; 2) embolus acicular; 3) VTA round, without a small tip; 4) TS shorter than half the length of the tegulum; and 5) COs located anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="658614F4FFC8C4231C82E631FE2B3E53" box="[199,502,294,321]" pageId="2" pageNumber="267" type="distribution">
|
||
<paragraph id="2D23477FFFC8C4231C82E631FE2B3E53" blockId="2.[151,1436,150,321]" box="[199,502,294,321]" pageId="2" pageNumber="267">
|
||
<emphasis id="1FE89B6DFFC8C4231C82E631FEBC3E52" bold="true" box="[199,353,294,320]" pageId="2" pageNumber="267">Distribution.</emphasis>
|
||
<collectingCountry id="558B07EFFFC8C4231D22E630FE703E53" box="[359,429,295,321]" name="China" pageId="2" pageNumber="267">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="558B07EFFFC8C4231DFDE630FE2F3E53" box="[440,498,295,321]" name="Laos" pageId="2" pageNumber="267">Laos</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |