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<document id="F14A161EF1EC9E375042BDAB804182B7" ID-CLB-Dataset="22330" ID-DOI="10.11646/zootaxa.4772.1.5" ID-GBIF-Dataset="b14b0d0d-cf67-4895-965c-12792c622b21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3814079" ID-ZooBank="B497198A-08CD-4A21-AE04-14390499853B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588834845818" checkinUser="plazi" docAuthor="Polhemus, Dan A." docDate="2020" docId="03FE00115B64FFC8FF38FA6CFED709D7" docLanguage="en" docName="zootaxa.4772.1.5.pdf" docOrigin="Zootaxa 4772 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enithares arfak Polhemus 2020, new species" docType="treatment" docVersion="8" lastPageNumber="158" masterDocId="FFC778695B7DFFD2FFAFFFC8FFF90E21" masterDocTitle="Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist" masterLastPageNumber="182" masterPageNumber="132" pageNumber="157" updateTime="1698838866946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F8516223437F39D4FC79C1CBBECC8905" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0719EB0DE880BDC910436F58A369F19">
<mods:title id="F5672C9F281C7739E76517F296D74092">Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist</mods:title>
</mods:titleInfo>
<mods:name id="1E690BC4BDA671B6E93ECC7ED84301EC" type="personal">
<mods:role id="156293D61DEA338FAA619AFEE12C1A93">
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<mods:namePart id="E23F382A9FC720A0E75513D9E06B6D94">Polhemus, Dan A.</mods:namePart>
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<mods:title id="B3E365444DFEE1B36BA2AC9212297DD6">Zootaxa</mods:title>
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<mods:part id="98E01733C36E93D3C0D524C372B3B9E6">
<mods:date id="7E62A3BC1A46BD80F2D1E776424E174D">2020</mods:date>
<mods:detail id="BB41DAE89185A1358AA4E47476D10D76" type="pubDate">
<mods:number id="B91B253697E56D79E04677EDEC61258B">2020-05-07</mods:number>
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<mods:number id="417A40932638C52DB1CDBF385292DA4E">4772</mods:number>
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<mods:identifier id="25914632570CBF077D21E2ECB5000C0D" type="DOI">10.11646/zootaxa.4772.1.5</mods:identifier>
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<treatment id="03FE00115B64FFC8FF38FA6CFED709D7" ID-DOI="http://doi.org/10.5281/zenodo.3815488" ID-GBIF-Taxon="163975559" ID-Zenodo-Dep="3815488" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7" httpUri="http://treatment.plazi.org/id/03FE00115B64FFC8FF38FA6CFED709D7" lastPageId="26" lastPageNumber="158" pageId="25" pageNumber="157">
<subSubSection id="C34DE28C5B64FFCBFF38FA6CFE0B0B9F" box="[151,498,1444,1471]" pageId="25" pageNumber="157" type="nomenclature">
<paragraph id="8BE8B1075B64FFCBFF38FA6CFE0B0B9F" blockId="25.[151,498,1444,1507]" box="[151,498,1444,1471]" pageId="25" pageNumber="157">
<heading id="D0A0066B5B64FFCBFF38FA6CFE0B0B9F" bold="true" box="[151,498,1444,1471]" fontSize="11" level="1" pageId="25" pageNumber="157" reason="1">
<emphasis id="B9236D155B64FFCBFF38FA6CFE0B0B9F" bold="true" box="[151,498,1444,1471]" pageId="25" pageNumber="157">
<taxonomicName id="4C57CA845B64FFCBFF38FA6CFEA00B9E" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[151,345,1444,1471]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="157" phylum="Arthropoda" rank="species" species="arfak" status="sp. nov.">
<emphasis id="B9236D155B64FFCBFF38FA6CFEA00B9E" bold="true" box="[151,345,1444,1471]" italics="true" pageId="25" pageNumber="157">Enithares arfak</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210D06E5B64FFCBFECFFA6CFE0B0B9F" box="[352,498,1444,1470]" pageId="25" pageNumber="157" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B64FFCBFF38FA00FE580BC2" box="[151,417,1479,1507]" pageId="25" pageNumber="157" type="description">
<paragraph id="8BE8B1075B64FFCBFF38FA00FE580BC2" blockId="25.[151,498,1444,1507]" box="[151,417,1479,1507]" pageId="25" pageNumber="157">
(
<figureCitation id="136CAD825B64FFCBFF0FFA00FF090BC3" box="[160,240,1480,1506]" captionStart="FIGURES 27" captionStartId="3.[151,264,1014,1039]" captionTargetBox="[151,1436,159,999]" captionTargetId="figure@3.[151,1436,159,1000]" captionTargetPageId="3" captionText="FIGURES 27. Enithares species, metaxiphus. 2. E. arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 3. E. bosavi n. sp.; holotype specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 4. E. insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 5. E. kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 6. E. orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Reinduk, CL 7036. 7. E. papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157." figureDoi="http://doi.org/10.5281/zenodo.3814083" httpUri="https://zenodo.org/record/3814083/files/figure.png" pageId="25" pageNumber="157">Figs. 2</figureCitation>
,
<figureCitation id="136CAD825B64FFCBFF53FA00FEBF0BC3" box="[252,326,1480,1506]" captionStart="FIGURES 4148" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 4148. Enithares species in the E. megalops group, dorsal habitus photographs. 4142. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 4344. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 4546. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 4748. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="25" pageNumber="157">47, 48</figureCitation>
,
<figureCitation id="136CAD825B64FFCBFEFEFA0FFE960BC3" box="[337,367,1479,1506]" captionStart="FIGURES 4954" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 4954. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="25" pageNumber="157">51</figureCitation>
,
<figureCitation id="136CAD825B64FFCBFED3FA00FE630BC2" box="[380,410,1480,1507]" captionStart="FIGURE 66" captionStartId="38.[151,250,1007,1032]" captionTargetBox="[161,1429,190,974]" captionTargetId="figure@38.[151,1436,181,982]" captionTargetPageId="38" captionText="FIGURE 66. Distribution of species in the E. megalops group on New Guinea. Triangles = E. megalops; circles = E. paramegalops Lansbury; pentagons = E. arfak." figureDoi="http://doi.org/10.5281/zenodo.3814899" httpUri="https://zenodo.org/record/3814899/files/figure.png" pageId="25" pageNumber="157">66</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B64FFCBFF38F9C3FE0C0964" pageId="25" pageNumber="157" type="materials_examined">
<paragraph id="8BE8B1075B64FFCBFF38F9C3FE0C0964" blockId="25.[151,1437,1546,2041]" pageId="25" pageNumber="157">
<emphasis id="B9236D155B64FFCBFF38F9C3FE430804" bold="true" box="[151,442,1546,1573]" pageId="25" pageNumber="157">Type material examined</emphasis>
.
<materialsCitation id="3B3FBB5A5B64FFCBFE67F9C3FC5D084C" ID-GBIF-Occurrence="2610672499" collectingDate="1991-10-18" collectionCode="A, BPBM" collectorName="Polhemus &amp; J. T. Polhemus" country="Indonesia" county="New" elevation="150" latitude="-1.1116667" location="Warkomi" longLatPrecision="21" longitude="134.08444" municipality="Mupi River" pageId="25" pageNumber="157" specimenCount="150" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="54EC0FA55B64FFCBFE67F9C3FDCD0804" box="[456,564,1547,1573]" pageId="25" pageNumber="157" type="holotype">Holotype</typeStatus>
, male (dissected):
<collectingCountry id="F340F1975B64FFCBFCA9F9C3FC5A0804" box="[774,931,1547,1573]" name="Indonesia" pageId="25" pageNumber="157">INDONESIA</collectingCountry>
,
<collectingRegion id="49937FE55B64FFCBFC02F9C2FB880804" box="[941,1137,1546,1573]" country="Indonesia" name="Papua Barat" pageId="25" pageNumber="157">
<emphasis id="B9236D155B64FFCBFC02F9C2FB900804" bold="true" box="[941,1129,1546,1573]" pageId="25" pageNumber="157">Irian Jaya Prov</emphasis>
.
</collectingRegion>
[
<collectingRegion id="49937FE55B64FFCBFBD1F9C3FAF40804" box="[1150,1293,1546,1573]" country="Indonesia" name="Papua Barat" pageId="25" pageNumber="157">Papua Prov.</collectingRegion>
],
<collectingCounty id="6289C98B5B64FFCBFAB1F9C3FAAA0804" box="[1310,1363,1547,1573]" pageId="25" pageNumber="157">New</collectingCounty>
Guin- ea,
<collectingMunicipality id="6B8C2B7D5B64FFCBFF12F9E7FEB90868" box="[189,320,1583,1609]" pageId="25" pageNumber="157">Mupi River</collectingMunicipality>
above
<location id="8E88E7DC5B64FFCBFE20F9E7FE020868" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7:8E88E7DC5B64FFCBFE20F9E7FE020868" box="[399,507,1583,1609]" country="Indonesia" county="New" latitude="-1.1116667" longLatPrecision="21" longitude="134.08444" municipality="Mupi River" name="Warkomi" pageId="25" pageNumber="157" stateProvince="Papua Barat">Warkomi</location>
,
<location id="8E88E7DC5B64FFCBFDACF9E7FD3E0868" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7:8E88E7DC5B64FFCBFDACF9E7FD3E0868" box="[515,711,1583,1609]" country="Indonesia" county="New" latitude="-1.1116667" longLatPrecision="21" longitude="134.08444" municipality="Mupi River" name="Arfak Mountains" pageId="25" pageNumber="157" stateProvince="Papua Barat">Arfak Mountains</location>
,
<quantity id="4CAF1CE25B64FFCBFD7EF9E7FCE20868" box="[721,795,1583,1609]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="157" unit="km" value="38.0">38 km</quantity>
. S. of
<location id="8E88E7DC5B64FFCBFCF0F9E7FC1D0868" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7:8E88E7DC5B64FFCBFCF0F9E7FC1D0868" box="[863,996,1583,1609]" country="Indonesia" county="New" latitude="-1.1116667" longLatPrecision="21" longitude="134.08444" municipality="Mupi River" name="Manokwari" pageId="25" pageNumber="157" stateProvince="Papua Barat">Manokwari</location>
,
<specimenCount id="9D517A8E5B64FFCBFC40F9E6FBC70868" box="[1007,1086,1582,1609]" pageId="25" pageNumber="157" type="generic">
<quantity id="4CAF1CE25B64FFCBFC40F9E6FBC30868" box="[1007,1082,1582,1609]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="157" unit="m" value="150.0">
<elevation id="007A56345B64FFCBFC40F9E6FBC30868" box="[1007,1082,1582,1609]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="157" unit="m" value="150.0">150 m</elevation>
</quantity>
.
</specimenCount>
,
<date id="FFE997C75B64FFCBFBE6F9E7FAF10868" box="[1097,1288,1583,1609]" pageId="25" pageNumber="157" value="1991-10-18">
<collectingDate id="EFAD6E2F5B64FFCBFBE6F9E7FAF10868" box="[1097,1288,1583,1609]" pageId="25" pageNumber="157" value="1991-10-18">18 October 1991</collectingDate>
</date>
,
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,
<geoCoordinate id="EE63D7C05B64FFCBFF38F99AFED5084C" box="[151,300,1618,1645]" degrees="134" direction="east" minutes="05" orientation="longitude" pageId="25" pageNumber="157" precision="15" seconds="04" value="134.08444">134°0504E</geoCoordinate>
, CL 2646, D.
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.
<collectorName id="26A2D4D15B64FFCBFE43F99BFDA5084C" box="[492,604,1619,1645]" pageId="25" pageNumber="157">Polhemus</collectorName>
and
<collectorName id="26A2D4D15B64FFCBFD3DF99BFCC1084C" box="[658,824,1619,1645]" pageId="25" pageNumber="157">J. T. Polhemus</collectorName>
(
<collectionCode id="ED4629C25B64FFCBFCE8F99BFC65084C" box="[839,924,1619,1645]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="25" pageNumber="157" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3FBB5A5B64FFCBFC01F99CFBCE08F8" ID-GBIF-Occurrence="2610672524" collectingDate="1991-10-19" collectionCode="USNM, A" collectorName="Polhemus &amp; J. T. Polhemus" country="Indonesia" county="New" elevation="140" latitude="-0.9113889" location="Arfak Mountains" longLatPrecision="21" longitude="133.79417" municipality="Aimasi Hulu" pageId="25" pageNumber="157" specimenCount="163" specimenCount-female="12" specimenCount-male="11" stateProvince="Papua Barat" typeStatus="paratype">
<emphasis id="B9236D155B64FFCBFC01F99CFBD8084C" box="[942,1057,1620,1645]" italics="true" pageId="25" pageNumber="157">
<typeStatus id="54EC0FA55B64FFCBFC01F99CFBD8084C" box="[942,1057,1620,1645]" pageId="25" pageNumber="157" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F340F1975B64FFCBFB9FF99BFB34084C" box="[1072,1229,1619,1645]" name="Indonesia" pageId="25" pageNumber="157">INDONESIA</collectingCountry>
,
<collectingRegion id="49937FE55B64FFCBFB77F99AFA65084C" box="[1240,1436,1618,1645]" country="Indonesia" name="Papua Barat" pageId="25" pageNumber="157">
<emphasis id="B9236D155B64FFCBFB77F99AFA6C084C" bold="true" box="[1240,1429,1618,1645]" pageId="25" pageNumber="157">Irian Jaya Prov</emphasis>
.
</collectingRegion>
[
<collectingRegion id="49937FE55B64FFCBFF32F9BFFEC908B0" box="[157,304,1654,1681]" country="Indonesia" name="Papua Barat" pageId="25" pageNumber="157">Papua Prov.</collectingRegion>
],
<collectingCounty id="6289C98B5B64FFCBFEEBF9BFFE8008B0" box="[324,377,1655,1681]" pageId="25" pageNumber="157">New</collectingCounty>
Guinea:
<specimenCount id="9D517A8E5B64FFCBFE49F9BFFDB908B0" box="[486,576,1655,1681]" pageId="25" pageNumber="157" type="male">9 males</specimenCount>
(1 dissected),
<specimenCount id="9D517A8E5B64FFCBFD42F9BFFCA608B0" box="[749,863,1655,1681]" pageId="25" pageNumber="157" type="female">7 females</specimenCount>
, same data as holotype (
<collectionCode id="ED4629C25B64FFCBFBD0F9BFFB2E08B0" box="[1151,1239,1655,1681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="157" type="Museum">USNM</collectionCode>
);
<specimenCount id="9D517A8E5B64FFCBFB42F9BFFABE08B0" box="[1261,1351,1655,1681]" pageId="25" pageNumber="157" type="male">2 males</specimenCount>
(1 dis- sected),
<specimenCount id="9D517A8E5B64FFCBFF56F952FE940894" box="[249,365,1690,1717]" pageId="25" pageNumber="157" type="female">5 females</specimenCount>
, small rocky stream at
<collectingMunicipality id="6B8C2B7D5B64FFCBFD2EF953FCE10894" box="[641,792,1691,1717]" pageId="25" pageNumber="157">Aimasi Hulu</collectingMunicipality>
,
<location id="8E88E7DC5B64FFCBFC89F953FC170894" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7:8E88E7DC5B64FFCBFC89F953FC170894" box="[806,1006,1691,1717]" country="Indonesia" county="New" latitude="-0.9113889" longLatPrecision="21" longitude="133.79417" municipality="Aimasi Hulu" name="Arfak Mountains" pageId="25" pageNumber="157" stateProvince="Papua Barat">Arfak Mountains</location>
,
<quantity id="4CAF1CE25B64FFCBFC52F952FBB50894" box="[1021,1100,1690,1717]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="157" unit="km" value="65.0">65 km</quantity>
.
<locationDeviation id="9A5AA8255B64FFCBFBF6F953FACD0894" box="[1113,1332,1691,1717]" location="Manokwari" pageId="25" pageNumber="157">SW of Manokwari</locationDeviation>
,
<specimenCount id="9D517A8E5B64FFCBFAECF953FA6E0894" box="[1347,1431,1691,1717]" pageId="25" pageNumber="157" type="generic">
<quantity id="4CAF1CE25B64FFCBFAECF953FA6A0894" box="[1347,1427,1691,1717]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="25" pageNumber="157" unit="m" value="140.0">
<elevation id="007A56345B64FFCBFAECF953FA6A0894" box="[1347,1427,1691,1717]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="25" pageNumber="157" unit="m" value="140.0">140 m</elevation>
</quantity>
.
</specimenCount>
,
<geoCoordinate id="EE63D7C05B64FFCBFF38F976FEF608F8" box="[151,271,1726,1753]" degrees="0" direction="south" minutes="54" orientation="latitude" pageId="25" pageNumber="157" precision="15" seconds="41" value="-0.9113889">0°5441S</geoCoordinate>
,
<geoCoordinate id="EE63D7C05B64FFCBFEB2F976FE4B08F8" box="[285,434,1726,1753]" degrees="133" direction="east" minutes="47" orientation="longitude" pageId="25" pageNumber="157" precision="15" seconds="39" value="133.79417">133°4739E</geoCoordinate>
, water temp. 25° C.,
<date id="FFE997C75B64FFCBFD1FF977FC8F08F8" box="[688,886,1727,1753]" pageId="25" pageNumber="157" value="1991-10-19">
<collectingDate id="EFAD6E2F5B64FFCBFD1FF977FC8F08F8" box="[688,886,1727,1753]" pageId="25" pageNumber="157" value="1991-10-19">19 October 1991</collectingDate>
</date>
, CL 2649, D.
<collectionCode id="ED4629C25B64FFCBFBB0F977FBCE08F8" box="[1055,1079,1727,1753]" pageId="25" pageNumber="157">A</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3FBB5A5B64FFCBFBEAF977FE5908DC" ID-GBIF-Occurrence="2610672378" collectionCode="BPBM, USNM, LIPI" collectorName="Polhemus &amp; J. T. Polhemus" country="Indonesia" pageId="25" pageNumber="157" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
<collectorName id="26A2D4D15B64FFCBFBEAF977FB4C08F8" box="[1093,1205,1727,1753]" pageId="25" pageNumber="157">Polhemus</collectorName>
and
<collectorName id="26A2D4D15B64FFCBFB5EF977FA6508F8" box="[1265,1436,1727,1753]" pageId="25" pageNumber="157">J. T. Polhemus</collectorName>
(
<collectionCode id="ED4629C25B64FFCBFF30F92BFF0B08DC" box="[159,242,1763,1789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="25" pageNumber="157" type="Museum">BPBM</collectionCode>
,
<collectionCode id="ED4629C25B64FFCBFF51F92BFEAB08DC" box="[254,338,1763,1789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="157" type="Museum">USNM</collectionCode>
,
<collectionCode id="ED4629C25B64FFCBFEF0F92BFE6108DC" box="[351,408,1763,1789]" pageId="25" pageNumber="157">LIPI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3FBB5A5B64FFCBFE03F92BFEF20964" ID-GBIF-Occurrence="2610672598" collectingDate="1991-10-19" collectionCode="A" country="Indonesia" county="Arfak Mtns." elevation="215" latitude="-0.9586111" location="Prafi River" longLatPrecision="21" longitude="133.88777" municipality="Warmari" pageId="25" pageNumber="157" specimenCount="5" specimenCount-female="5" stateProvince="Papua Barat" typeStatus="paratype">
<emphasis id="B9236D155B64FFCBFE03F92BFD3208DC" box="[428,715,1763,1789]" italics="true" pageId="25" pageNumber="157">Other material examined</emphasis>
(not
<typeStatus id="54EC0FA55B64FFCBFCA6F92BFC8508DC" box="[777,892,1763,1789]" pageId="25" pageNumber="157" type="paratype">paratypes</typeStatus>
):
<specimenCount id="9D517A8E5B64FFCBFC20F92AFBF808DC" box="[911,1025,1762,1789]" pageId="25" pageNumber="157" type="female">5 females</specimenCount>
,
<location id="8E88E7DC5B64FFCBFBA2F92BFB7508DC" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7:8E88E7DC5B64FFCBFBA2F92BFB7508DC" box="[1037,1164,1763,1789]" country="Indonesia" county="Arfak Mtns." latitude="-0.9586111" longLatPrecision="21" longitude="133.88777" municipality="Warmari" name="Prafi River" pageId="25" pageNumber="157" stateProvince="Papua Barat">Prafi River</location>
at
<collectingMunicipality id="6B8C2B7D5B64FFCBFB1FF92BFAEC08DC" box="[1200,1301,1763,1789]" pageId="25" pageNumber="157">Warmari</collectingMunicipality>
,
<locationDeviation id="9A5AA8255B64FFCBFA8DF92BFEC20900" location="Manokwari" pageId="25" pageNumber="157">
<quantity id="4CAF1CE25B64FFCBFA8DF92BFA9308DC" box="[1314,1386,1763,1789]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="25" pageNumber="157" unit="km" value="48.0">48 km</quantity>
SW of Manokwari
</locationDeviation>
,
<collectingCounty id="6289C98B5B64FFCBFEEAF8CFFE2B0900" box="[325,466,1799,1825]" pageId="25" pageNumber="157">Arfak Mtns.</collectingCounty>
,
<quantity id="4CAF1CE25B64FFCBFE72F8CEFDDE0900" box="[477,551,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="25" pageNumber="157" unit="m" value="215.0">
<elevation id="007A56345B64FFCBFE72F8CEFDDE0900" box="[477,551,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="25" pageNumber="157" unit="m" value="215.0">215 m</elevation>
</quantity>
,
<geoCoordinate id="EE63D7C05B64FFCBFD9DF8CEFD530900" box="[562,682,1798,1825]" degrees="0" direction="south" minutes="57" orientation="latitude" pageId="25" pageNumber="157" precision="15" seconds="31" value="-0.9586111">0°5731S</geoCoordinate>
,
<geoCoordinate id="EE63D7C05B64FFCBFD1AF8CEFCB30900" box="[693,842,1798,1825]" degrees="133" direction="east" minutes="53" orientation="longitude" pageId="25" pageNumber="157" precision="15" seconds="16" value="133.88777">133°5316E</geoCoordinate>
, water temp. 26.5° C., CL 2652,
<date id="FFE997C75B64FFCBFB11F8CFFA860900" box="[1214,1407,1799,1825]" pageId="25" pageNumber="157" value="1991-10-19">
<collectingDate id="EFAD6E2F5B64FFCBFB11F8CFFA860900" box="[1214,1407,1799,1825]" pageId="25" pageNumber="157" value="1991-10-19">19 October 1991</collectingDate>
</date>
, J. T. &amp; D.
<collectionCode id="ED4629C25B64FFCBFF5CF8E3FEF20964" box="[243,267,1835,1861]" pageId="25" pageNumber="157">A</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3FBB5A5B64FFCBFEBAF8E3FE080964" ID-GBIF-Occurrence="2610672416" box="[277,497,1835,1861]" collectionCode="BPBM" country="Indonesia" location="Polhemus" pageId="25" pageNumber="157" specimenCount="1" stateProvince="Papua Barat">
<location id="8E88E7DC5B64FFCBFEBAF8E3FE7C0964" LSID="urn:lsid:plazi:treatment:03FE00115B64FFC8FF38FA6CFED709D7:8E88E7DC5B64FFCBFEBAF8E3FE7C0964" box="[277,389,1835,1861]" country="Indonesia" name="Polhemus" pageId="25" pageNumber="157" stateProvince="Papua Barat">Polhemus</location>
(
<collectionCode id="ED4629C25B64FFCBFE3BF8E3FE100964" box="[404,489,1835,1861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="25" pageNumber="157" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B64FFC8FF68F886FB360A2C" lastPageId="26" lastPageNumber="158" pageId="25" pageNumber="157" type="description">
<paragraph id="8BE8B1075B64FFCBFF68F886FC160948" blockId="25.[151,1437,1546,2041]" box="[199,1007,1870,1897]" pageId="25" pageNumber="157">
<emphasis id="B9236D155B64FFCBFF68F886FEAA0948" bold="true" box="[199,339,1870,1897]" pageId="25" pageNumber="157">Description</emphasis>
.
<emphasis id="B9236D155B64FFCBFECEF887FE670948" bold="true" box="[353,414,1871,1897]" pageId="25" pageNumber="157">Male</emphasis>
: Length
<quantity id="4CAF1CE25B64FFCBFDAAF886FD940948" box="[517,621,1870,1897]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="25" pageNumber="157" unit="mm" value="8.5">8.50 mm</quantity>
, width across pronotum
<quantity id="4CAF1CE25B64FFCBFC2CF887FC130948" box="[899,1002,1871,1897]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="25" pageNumber="157" unit="mm" value="3.1">3.10 mm</quantity>
.
</paragraph>
<paragraph id="8BE8B1075B64FFC8FF68F8BBFEC50F44" blockId="25.[151,1437,1546,2041]" lastBlockId="26.[151,1437,153,2038]" lastPageId="26" lastPageNumber="158" pageId="25" pageNumber="157">
<emphasis id="B9236D155B64FFCBFF68F8BBFEBA09AC" box="[199,323,1907,1933]" italics="true" pageId="25" pageNumber="157">Coloration</emphasis>
: Ground color brown to blackish, with varying extent of pale marking on scutellum and hemelytra (
<figureCitation id="136CAD825B64FFCBFF0FF85FFF0C0990" box="[160,245,1943,1969]" captionStart="FIGURES 4148" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 4148. Enithares species in the E. megalops group, dorsal habitus photographs. 4142. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 4344. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 4546. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 4748. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="25" pageNumber="157">Fig. 47</figureCitation>
). Vertex and frons creamy white to yellowish; frons bearing a pair (1+1) of small roughly circular brown patches near anterior apex, and another roughly triangular brown patch above base of labrum; lower section of frons adjacent to labrum sometimes suffused with pale blue; labrum dark brown, sometimes dark yellow centrally. Pronotum with anterior half between pronotal foveae creamy white to tan; posterior half brown to black; propleura creamy white. Scutellum creamy yellowish-white, suffused to varying degrees with dark brown to black at anterolateral angles and centrally. Hemelytra brown to black, with outer half of clavus, outer half of corium, and sometimes inner margin of clavus bordering scutellum white; wing membrane uniform fumate brown, translucent. Legs yellowish brown, anterior edges of all femora margined with dark brown to black. Venter brown, with coxal plates greyish.
</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68FEA7FC750C1D" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
<emphasis id="B9236D155B67FFC8FF68FEA7FE470FA8" box="[199,446,367,393]" italics="true" pageId="26" pageNumber="158">Structural characters</emphasis>
: Head broadly rounded anteriorly when viewed dorsally. Head length 1.10; greatest width 2.50, equal to 0.80 pronotal width; anterior width of vertex 0.70, equal to 0.63× head length. Synthlipsis 0.50, about 0.71 anterior width of vertex and clearly shorter than pronotum. Pronotal length along midline 0.95, humeral width 3.10, lateral margins convex, posterior margin weakly sinuate. Dorsal margin of pronotal fovea directed caudad behind eyes (
<figureCitation id="136CAD825B67FFC8FE49FE36FDC30C39" box="[486,570,510,536]" captionStart="FIGURES 4148" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 4148. Enithares species in the E. megalops group, dorsal habitus photographs. 4142. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 4344. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 4546. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 4748. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="26" pageNumber="158">Fig. 47</figureCitation>
). Nodal furrow curved weakly cephalad, removed about 2.0× its length from membranal suture, length 0.50, distance to membranal suture 0.90.
</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68FD8EFAAF0CEA" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">Front and hind legs typical for genus, lacking unusual modifications. Middle trochanter rounded. Middle femur with single large subapical tooth, bordered basally by about 12 small black pegs, distally by about 4 similar pegs. Lengths of leg segments as follows: fore femurtibiatarsal 1tarsal 2 = 1.60/1.50/0.50/0.25; middle fe-murtibiatarsal 1tarsal 2 = 1.90/1.65/0.55/0.50; hind femurtibiatarsal 1tarsal 2 = 3.00/2.50/1.10/0.80.</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68FD1DFBA60CCE" blockId="26.[151,1437,153,2038]" box="[199,1119,725,751]" pageId="26" pageNumber="158">
Ventral abdomen with metaxyphus triangular, slightly concave, tip acute (
<figureCitation id="136CAD825B67FFC8FBA4FD1DFBAB0CCE" box="[1035,1106,725,751]" captionStart="FIGURES 27" captionStartId="3.[151,264,1014,1039]" captionTargetBox="[151,1436,159,999]" captionTargetId="figure@3.[151,1436,159,1000]" captionTargetPageId="3" captionText="FIGURES 27. Enithares species, metaxiphus. 2. E. arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 3. E. bosavi n. sp.; holotype specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 4. E. insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 5. E. kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 6. E. orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Reinduk, CL 7036. 7. E. papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157." figureDoi="http://doi.org/10.5281/zenodo.3814083" httpUri="https://zenodo.org/record/3814083/files/figure.png" pageId="26" pageNumber="158">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68FD30FE730DE4" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
<emphasis id="B9236D155B67FFC8FF68FD30FE880D33" box="[199,369,760,786]" italics="true" pageId="26" pageNumber="158">Male genitalia</emphasis>
when viewed laterally (
<figureCitation id="136CAD825B67FFC8FD3EFD30FD1E0D32" box="[657,743,760,787]" captionStart="FIGURES 4954" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 4954. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="26" pageNumber="158">Fig. 51</figureCitation>
) with proctiger truncate; posterior lobe of pygophore with posteroventral angle obtuse, bearing a prominent setal tuft, lobe of proctiger erect, posteriorly setiferous, apex broadly rounded, not notched on dorsal margin; paramere small, subovate, not even reaching half distance to apex of posterior lobe; lateral arm of basal plate strongly bent, roughly L-shaped, dorsal margin broadly curving, apex turned slightly upward; aedeagus semicircular, with numerous transverse folds, bearing serrate row of dark teeth along ventral margin.
</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68FC07FB360A2C" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
<emphasis id="B9236D155B67FFC8FF68FC07FEE60DC8" bold="true" box="[199,287,975,1001]" pageId="26" pageNumber="158">Female</emphasis>
: Length
<quantity id="4CAF1CE25B67FFC8FE27FC07FE0C0DC8" box="[392,501,975,1001]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="26" pageNumber="158" unit="mm" value="8.2">8.20 mm</quantity>
.; width across pronotum 3.00 mm. Similar to male in general structure and coloration, but with pale coloration on pronotum, scutellum and hemelytra more extensive (
<figureCitation id="136CAD825B67FFC8FBC2FC3AFB3B0A2C" box="[1133,1218,1010,1037]" captionStart="FIGURES 4148" captionStartId="27.[151,264,1822,1847]" captionTargetBox="[169,1417,181,1797]" captionTargetId="figure@27.[169,1417,181,1797]" captionTargetPageId="27" captionText="FIGURES 4148. Enithares species in the E. megalops group, dorsal habitus photographs. 4142. Enithares megalops Lansbury, male and female; male specimen from Indonesia, West Papua Province, Kasim, CL 2620, female paratype specimen from Indonesia, West Papua Prov., Bomberai Peninsula. 4344. Enithares paramegalops Lansbury, male and female; male specimen from Papua New Guinea, Gulf Province, Kopi, CL 7000, female paratype specimen from Indonesia, Papua Prov., Enarotali. 4546. Enithares kasim n. sp., male and female; specimens from Indonesia, West Papua Province, Kasim, CL 2630. 4748. Enithares arfak n. sp., male and female; specimens from Indonesia, West Papua Province, Mupi River, CL 2646." figureDoi="http://doi.org/10.5281/zenodo.3814101" httpUri="https://zenodo.org/record/3814101/files/figure.png" pageId="26" pageNumber="158">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B67FFC8FF68FBDEFE310A75" pageId="26" pageNumber="158" type="etymology">
<paragraph id="8BE8B1075B67FFC8FF68FBDEFE310A75" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
<emphasis id="B9236D155B67FFC8FF68FBDEFEB00A11" bold="true" box="[199,329,1046,1072]" pageId="26" pageNumber="158">Etymology</emphasis>
. The name “arfak” is a noun in apposition, and refers to the Arfak Mountains
<typeStatus id="54EC0FA55B67FFC8FB66FBDFFB030A10" box="[1225,1274,1047,1073]" pageId="26" pageNumber="158">type</typeStatus>
locality in far northwestern New
<collectingCountry id="F340F1975B67FFC8FEC0FBF2FE3D0A75" box="[367,452,1082,1108]" name="Guinea-Bissau" pageId="26" pageNumber="158">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B67FFC8FF68FB95FED709D7" pageId="26" pageNumber="158" type="discussion">
<paragraph id="8BE8B1075B67FFC8FF68FB95FAAE0B53" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
<emphasis id="B9236D155B67FFC8FF68FB95FEBF0A59" bold="true" box="[199,326,1117,1144]" pageId="26" pageNumber="158">Discussion</emphasis>
.
<taxonomicName id="4C57CA845B67FFC8FEFBFB96FDFD0A59" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[340,516,1118,1144]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak" status="sp. nov.">
<emphasis id="B9236D155B67FFC8FEFBFB96FDFD0A59" box="[340,516,1118,1144]" italics="true" pageId="26" pageNumber="158">Enithares arfak</emphasis>
</taxonomicName>
<emphasis id="B9236D155B67FFC8FDA5FB96FDB00A59" bold="true" box="[522,585,1118,1144]" pageId="26" pageNumber="158">
<taxonomicNameLabel id="A210D06E5B67FFC8FDA5FB96FDB00A59" box="[522,585,1118,1144]" pageId="26" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C57CA845B67FFC8FDF7FB97FD400A59" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[600,697,1118,1144]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="kasim" status="sp. nov.">
<emphasis id="B9236D155B67FFC8FDF7FB97FD400A59" box="[600,697,1118,1144]" italics="true" pageId="26" pageNumber="158">E. kasim</emphasis>
</taxonomicName>
<emphasis id="B9236D155B67FFC8FD10FB96FD070A59" bold="true" box="[703,766,1118,1144]" pageId="26" pageNumber="158">
<taxonomicNameLabel id="A210D06E5B67FFC8FD10FB96FD070A59" box="[703,766,1118,1144]" pageId="26" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C57CA845B67FFC8FCA3FB97FBF00A59" authority="Lansbury" authorityName="Lansbury" authorityYear="1968" box="[780,1033,1118,1144]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="B9236D155B67FFC8FCA3FB97FC6E0A59" box="[780,919,1118,1144]" italics="true" pageId="26" pageNumber="158">E. megalops</emphasis>
Lansbury
</taxonomicName>
and
<taxonomicName id="4C57CA845B67FFC8FB90FB97FA8B0A59" authority="Lansbury" authorityName="Lansbury" authorityYear="1968" box="[1087,1394,1118,1144]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="paramegalops">
<emphasis id="B9236D155B67FFC8FB90FB97FAF90A59" box="[1087,1280,1118,1144]" italics="true" pageId="26" pageNumber="158">E. paramegalops</emphasis>
Lansbury
</taxonomicName>
appear to be a closely knit group of sister species, all of which share certain common male genitalic character states, including a highly reduced paramere; a rounded or angular apex on the PL, without a dorsal notch; and a relatively simple morphology of the LABP, with the apex not expanded or acuminate (
<figureCitation id="136CAD825B67FFC8FC56FB01FB7D0AC2" box="[1017,1156,1224,1251]" captionStart="FIGURES 4954" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 4954. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="26" pageNumber="158">Figs. 4951</figureCitation>
). The genitalic morphology of
<taxonomicName id="4C57CA845B67FFC8FF49FB25FEB80B26" authorityName="Polhemus" authorityYear="2020" box="[230,321,1261,1287]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FF49FB25FEB80B26" box="[230,321,1261,1287]" italics="true" pageId="26" pageNumber="158">E. arfak</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C57CA845B67FFC8FD8FFB25FD520B26" authorityName="Lansbury" authorityYear="1968" box="[544,683,1261,1287]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="B9236D155B67FFC8FD8FFB25FD520B26" box="[544,683,1261,1287]" italics="true" pageId="26" pageNumber="158">E. megalops</emphasis>
</taxonomicName>
, but differs in the somewhat more broadly rounded apex of the PL, the slightly larger paramere, and the different shape of the LABP, which is more strongly bent and L-shaped in
<taxonomicName id="4C57CA845B67FFC8FA2BFAD9FF2A0B6F" authorityName="Polhemus" authorityYear="2020" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FA2BFAD9FF2A0B6F" italics="true" pageId="26" pageNumber="158">E. arfak</emphasis>
</taxonomicName>
and has the apex slightly upturned in lateral view (compare
<figureCitation id="136CAD825B67FFC8FCD6FAFCFBF90B6F" box="[889,1024,1331,1358]" captionStart="FIGURES 4954" captionStartId="31.[151,264,1639,1664]" captionTargetBox="[151,1435,181,1614]" captionTargetId="figure@31.[151,1436,181,1614]" captionTargetPageId="31" captionText="FIGURES 4954. Enithares species in the E. megalops group, male genitalic complex, left lateral view. 49. Enithares megalops Lansbury; specimen from Indonesia, West Papua Prov., Kasim, CL 2620. 50. Enithares paramegalops Lansbury; specimen from Papua New Guinea, Gulf Prov., Kopi, CL 7000, 51. Enithares arfak n. sp.; specimen from Indonesia, West Papua Prov., Mupi River, CL 2646. 52. Enithares kasim n. sp.; specimen from Indonesia, West Papua Prov., Kasim, CL 2630. 53. Enithares nigra Lansbury, specimen from Papua New Guinea, Southern Highlands Prov., Moro, CL 7024. 54. Enithares vulgaris Lansbury n. sp.; specimen from Indonesia, Papua Prov., Biak Island, CL 2643." figureDoi="http://doi.org/10.5281/zenodo.3814103" httpUri="https://zenodo.org/record/3814103/files/figure.png" pageId="26" pageNumber="158">Figs. 49, 51</figureCitation>
).
<taxonomicName id="4C57CA845B67FFC8FBBBFAFCFB3A0B6F" authorityName="Polhemus" authorityYear="2020" box="[1044,1219,1332,1358]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FBBBFAFCFB3A0B6F" box="[1044,1219,1332,1358]" italics="true" pageId="26" pageNumber="158">Enithares arfak</emphasis>
</taxonomicName>
is also a somewhat larger and stouter species, with body lengths ranging from
<quantity id="4CAF1CE25B67FFC8FC9DFA9FFC470B53" box="[818,958,1367,1394]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="8.9" metricValueMin="8.5" pageId="26" pageNumber="158" unit="mm" value="8.7" valueMax="8.9" valueMin="8.5">8.58.9 mm</quantity>
, versus 7.0
<quantity id="4CAF1CE25B67FFC8FBE5FA90FB590B53" box="[1098,1184,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="26" pageNumber="158" unit="mm" value="7.7">7.7 mm</quantity>
in
<taxonomicName id="4C57CA845B67FFC8FB6BFA91FAA90B53" authorityName="Lansbury" authorityYear="1968" box="[1220,1360,1368,1394]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="B9236D155B67FFC8FB6BFA91FAA90B53" box="[1220,1360,1368,1394]" italics="true" pageId="26" pageNumber="158">E. megalops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68FAB3FACB08DA" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
<emphasis id="B9236D155B67FFC8FF68FAB3FE730BB7" bold="true" box="[199,394,1403,1430]" pageId="26" pageNumber="158">Ecological notes</emphasis>
.
<taxonomicName id="4C57CA845B67FFC8FE37FAB4FDB00BB7" authorityName="Polhemus" authorityYear="2020" box="[408,585,1404,1430]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FE37FAB4FDB00BB7" box="[408,585,1404,1430]" italics="true" pageId="26" pageNumber="158">Enithares arfak</emphasis>
</taxonomicName>
belongs to a set of four putatively related species that have partially segregated but also partially overlapping ranges in the Vogelkop Peninsula of western New
<collectingCountry id="F340F1975B67FFC8FBC3FA57FB380B98" box="[1132,1217,1439,1465]" name="Guinea-Bissau" pageId="26" pageNumber="158">Guinea</collectingCountry>
. Within this grouping,
<taxonomicName id="4C57CA845B67FFC8FF67FA0CFE700BFC" authorityName="Lansbury" authorityYear="1968" box="[200,393,1475,1501]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="paramegalops">
<emphasis id="B9236D155B67FFC8FF67FA0CFE700BFC" box="[200,393,1475,1501]" italics="true" pageId="26" pageNumber="158">E. paramegalops</emphasis>
</taxonomicName>
as currently interpreted has an extensive range, occurring from the
<collectingRegion id="49937FE55B67FFC8FBD5FA0BFB2D0BFC" box="[1146,1236,1475,1501]" country="Indonesia" name="Sulawesi Utara" pageId="26" pageNumber="158">Celebes</collectingRegion>
eastward through the
<collectingCountry id="F340F1975B67FFC8FF6EFA2FFED60820" box="[193,303,1511,1537]" name="Indonesia" pageId="26" pageNumber="158">Moluccas</collectingCountry>
to the Papuan
<collectingRegion id="49937FE55B67FFC8FE75FA2FFDF70820" box="[474,526,1511,1537]" country="Papua New Guinea" name="Gulf" pageId="26" pageNumber="158">Gulf</collectingRegion>
lowlands of
<collectingCountry id="F340F1975B67FFC8FD0DFA2FFC870820" box="[674,894,1511,1537]" name="Papua New Guinea" pageId="26" pageNumber="158">Papua New Guinea</collectingCountry>
(
<figureCitation id="136CAD825B67FFC8FC22FA2FFC1A0823" box="[909,995,1511,1538]" captionStart="FIGURE 66" captionStartId="38.[151,250,1007,1032]" captionTargetBox="[161,1429,190,974]" captionTargetId="figure@38.[151,1436,181,982]" captionTargetPageId="38" captionText="FIGURE 66. Distribution of species in the E. megalops group on New Guinea. Triangles = E. megalops; circles = E. paramegalops Lansbury; pentagons = E. arfak." figureDoi="http://doi.org/10.5281/zenodo.3814899" httpUri="https://zenodo.org/record/3814899/files/figure.png" pageId="26" pageNumber="158">Fig. 66</figureCitation>
). By contrast,
<taxonomicName id="4C57CA845B67FFC8FB27FA20FB1D0820" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[1160,1252,1511,1537]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak" status="sp. nov.">
<emphasis id="B9236D155B67FFC8FB27FA20FB1D0820" box="[1160,1252,1511,1537]" italics="true" pageId="26" pageNumber="158">E. arfak</emphasis>
</taxonomicName>
<emphasis id="B9236D155B67FFC8FB44FA2FFAD20820" bold="true" box="[1259,1323,1511,1537]" pageId="26" pageNumber="158">
<taxonomicNameLabel id="A210D06E5B67FFC8FB44FA2FFAD20820" box="[1259,1323,1511,1537]" pageId="26" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C57CA845B67FFC8FA95FA20FA650820" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[1338,1436,1511,1537]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="kasim" status="sp. nov.">
<emphasis id="B9236D155B67FFC8FA95FA20FA650820" box="[1338,1436,1511,1537]" italics="true" pageId="26" pageNumber="158">E. kasim</emphasis>
</taxonomicName>
<emphasis id="B9236D155B67FFC8FF38F9C3FF2E0804" bold="true" box="[151,215,1547,1573]" pageId="26" pageNumber="158">
<taxonomicNameLabel id="A210D06E5B67FFC8FF38F9C3FF2E0804" box="[151,215,1547,1573]" pageId="26" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C57CA845B67FFC8FEB8F9C3FDEC0805" authority="Lansbury" authorityName="Lansbury" authorityYear="1968" box="[279,533,1546,1572]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="B9236D155B67FFC8FEB8F9C3FE5B0805" box="[279,418,1546,1572]" italics="true" pageId="26" pageNumber="158">E. megalops</emphasis>
Lansbury
</taxonomicName>
have more limited ranges in western New
<collectingCountry id="F340F1975B67FFC8FC50F9C2FBA80805" box="[1023,1105,1546,1572]" name="Guinea-Bissau" pageId="26" pageNumber="158">Guinea</collectingCountry>
and the
<collectingCountry id="F340F1975B67FFC8FB1CF9C2FADC0805" box="[1203,1317,1546,1572]" name="Indonesia" pageId="26" pageNumber="158">Moluccas</collectingCountry>
.
<taxonomicName id="4C57CA845B67FFC8FA80F9C2FEFA0869" authorityName="Lansbury" authorityYear="1968" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="B9236D155B67FFC8FA80F9C2FEFA0869" italics="true" pageId="26" pageNumber="158">Enithares megalops</emphasis>
</taxonomicName>
occurs from
<collectingCountry id="F340F1975B67FFC8FE39F9E6FE120869" box="[406,491,1582,1608]" name="Indonesia" pageId="26" pageNumber="158">Ambon</collectingCountry>
to the Lengguru Fold Belt of the New
<collectingCountry id="F340F1975B67FFC8FC34F9E6FC140869" box="[923,1005,1582,1608]" name="Guinea-Bissau" pageId="26" pageNumber="158">Guinea</collectingCountry>
“birds neck” (
<figureCitation id="136CAD825B67FFC8FB38F9E6FB120868" box="[1175,1259,1582,1609]" captionStart="FIGURE 66" captionStartId="38.[151,250,1007,1032]" captionTargetBox="[161,1429,190,974]" captionTargetId="figure@38.[151,1436,181,982]" captionTargetPageId="38" captionText="FIGURE 66. Distribution of species in the E. megalops group on New Guinea. Triangles = E. megalops; circles = E. paramegalops Lansbury; pentagons = E. arfak." figureDoi="http://doi.org/10.5281/zenodo.3814899" httpUri="https://zenodo.org/record/3814899/files/figure.png" pageId="26" pageNumber="158">Fig. 66</figureCitation>
);
<taxonomicName id="4C57CA845B67FFC8FB50F9E7FA990869" authorityName="Polhemus" authorityYear="2020" box="[1279,1376,1582,1608]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="kasim">
<emphasis id="B9236D155B67FFC8FB50F9E7FA990869" box="[1279,1376,1582,1608]" italics="true" pageId="26" pageNumber="158">E. kasim</emphasis>
</taxonomicName>
(
<figureCitation id="136CAD825B67FFC8FAC1F9E6FF4D084C" captionStart="FIGURE 67" captionStartId="38.[151,250,1916,1941]" captionTargetBox="[158,1430,1097,1882]" captionTargetId="figure@38.[151,1436,1090,1890]" captionTargetPageId="38" captionText="FIGURE 67. Distribution of species in the E. megalops group on New Guinea. Square = E. kasim n. sp.; circles = E. nigra Lansbury; triangles = E. alexis Lansbury; pentagons = E. vulgaris Lansbury." figureDoi="http://doi.org/10.5281/zenodo.3814901" httpUri="https://zenodo.org/record/3814901/files/figure.png" pageId="26" pageNumber="158">Fig. 67</figureCitation>
) has been taken only at the western end of the Vogelkop; and
<taxonomicName id="4C57CA845B67FFC8FCDAF99BFC28084D" authorityName="Polhemus" authorityYear="2020" box="[885,977,1618,1644]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FCDAF99BFC28084D" box="[885,977,1618,1644]" italics="true" pageId="26" pageNumber="158">E. arfak</emphasis>
</taxonomicName>
has so far been collected only in the Arfak Mountains of the eastern Vogelkop (
<figureCitation id="136CAD825B67FFC8FDCEF9BEFD4F08B0" box="[609,694,1654,1681]" captionStart="FIGURE 66" captionStartId="38.[151,250,1007,1032]" captionTargetBox="[161,1429,190,974]" captionTargetId="figure@38.[151,1436,181,982]" captionTargetPageId="38" captionText="FIGURE 66. Distribution of species in the E. megalops group on New Guinea. Triangles = E. megalops; circles = E. paramegalops Lansbury; pentagons = E. arfak." figureDoi="http://doi.org/10.5281/zenodo.3814899" httpUri="https://zenodo.org/record/3814899/files/figure.png" pageId="26" pageNumber="158">Fig. 66</figureCitation>
). These patterns of putative endemism, though intriguing, should be considered provisional, however, since too little collecting has been done to establish the precise geographic ranges of these species with any confidence. Based on current records, however,
<taxonomicName id="4C57CA845B67FFC8FB9EF976FB7508F6" authorityName="Polhemus" authorityYear="2020" box="[1073,1164,1725,1751]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FB9EF976FB7508F6" box="[1073,1164,1725,1751]" italics="true" pageId="26" pageNumber="158">E. arfak</emphasis>
</taxonomicName>
appears confined to the Vogelkop Highlands areas of freshwater endemism (Area 5) as defined by D.
<bibRefCitation id="EFC6CCF65B67FFC8FBA8F929FAD708DA" author="Polhemus, D. A. &amp; Allen, G. A." box="[1031,1326,1761,1787]" pageId="26" pageNumber="158" pagination="207 - 245" refId="ref39028" refString="Polhemus, D. A. &amp; Allen, G. A. (2007) Freshwater biogeography of Papua. In: Marshall, A. J. &amp; Beehler, B. M. (Eds.), Ecology of Papua. Part 1. Periplus Editions (HK) Ltd., Singapore, pp. 207 - 245." type="book chapter" year="2007">Polhemus &amp; Allen (2007)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE8B1075B67FFC8FF68F8CDFED709D7" blockId="26.[151,1437,153,2038]" pageId="26" pageNumber="158">
The Mupi River at the
<typeStatus id="54EC0FA55B67FFC8FE73F8CDFDF4093E" box="[476,525,1797,1823]" pageId="26" pageNumber="158">type</typeStatus>
locality was a fast, clear, powerful river coming out of the Arfak Mountains through a bed of boulders, rocks and gravel. The stream is known to the local people for its cold waters. The heavily scoured main channel was relatively poor habitat for aquatic Heteroptera, but quiet, spring-fed side channels harbored a variety of species, including
<taxonomicName id="4C57CA845B67FFC8FDF2F8B8FCF409AB" authorityName="Polhemus" authorityYear="2020" box="[605,781,1904,1930]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="158" phylum="Arthropoda" rank="species" species="arfak">
<emphasis id="B9236D155B67FFC8FDF2F8B8FCF409AB" box="[605,781,1904,1930]" italics="true" pageId="26" pageNumber="158">Enithares arfak</emphasis>
</taxonomicName>
. This species was also taken deeper into the Arfak Mountains from a small, clear stream issuing from primary rain forest adjacent to oil palm plantations at Aimasi Hulu. The habitat at this latter site consisted of shaded pools and small riffles in a bed of rocks and sand, with water depth of
<quantity id="4CAF1CE25B67FFC8FF19F812FED309D7" box="[182,298,2010,2038]" metricMagnitude="-1" metricUnit="m" metricValue="3.15" metricValueMax="4.5" metricValueMin="1.8" pageId="26" pageNumber="158" unit="cm" value="31.5" valueMax="45.0" valueMin="18.0">1845 cm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>