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<document id="1B2E77D1824ED85F06E809D8379A0CC7" ID-DOI="10.11646/zootaxa.3941.3.1" ID-GBIF-Dataset="67bc3c9d-ec99-4b47-81df-3fca7bd584ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="243860" ID-ZooBank="9907D081-A413-44F1-ABF6-76CC13DC8813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026017375" checkinUser="plazi" docAuthor="Wang, Cheng-Bin &amp; Zhou, Hong-Zhang" docDate="2015" docId="03EC87E6FF97880E9DBF1A40FC1E0FE0" docLanguage="en" docName="zt03941p338.pdf" docOrigin="Zootaxa 3941 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptomaphaginus luoi Wang &amp; Zhou, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="321" masterDocId="FFD5FF9EFF85881A9D281927FFF50E4C" masterDocTitle="Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species" masterLastPageNumber="338" masterPageNumber="301" pageNumber="319" updateTime="1698572131055" updateUser="plazi">
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<mods:title id="C1E4215A59E5F3E3D1AF2FCB41C74EE2">Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species</mods:title>
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<mods:namePart id="F892DC1E55931E5AAB3750B2480A3A5E">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="FDD8640EAE6063F8EFB92E5E3980DC5C">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="CB95629D95FE89DD59AA182143C49943">2015</mods:date>
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<treatment id="03EC87E6FF97880E9DBF1A40FC1E0FE0" ID-DOI="http://doi.org/10.5281/zenodo.5691672" ID-GBIF-Taxon="119606329" ID-Zenodo-Dep="5691672" LSID="urn:lsid:plazi:treatment:03EC87E6FF97880E9DBF1A40FC1E0FE0" httpUri="http://treatment.plazi.org/id/03EC87E6FF97880E9DBF1A40FC1E0FE0" lastPageId="20" lastPageNumber="321" pageId="18" pageNumber="319">
<subSubSection id="C35F657BFF9788089DBF1A40FECC0DEF" pageId="18" pageNumber="319" type="nomenclature">
<paragraph id="8BFA36F0FF9788089DBF1A40FE040DCE" blockId="18.[151,497,871,931]" box="[151,497,871,898]" pageId="18" pageNumber="319">
<heading id="D0B2819CFF9788089DBF1A40FE040DCE" bold="true" box="[151,497,871,898]" fontSize="11" level="1" pageId="18" pageNumber="319" reason="1">
<emphasis id="B931EAE2FF9788089DBF1A40FE040DCE" bold="true" box="[151,497,871,898]" pageId="18" pageNumber="319">
<taxonomicName id="4C454D73FF9788089DBF1A40FE790DCD" ID-CoL="4Q99L" box="[151,396,871,897]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="luoi" status="sp. nov.">
<emphasis id="B931EAE2FF9788089DBF1A40FE790DCD" bold="true" box="[151,396,871,897]" italics="true" pageId="18" pageNumber="319">Ptomaphaginus luoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FF9788089CBB1A4FFE040DCE" box="[403,497,872,898]" pageId="18" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA36F0FF9788089DBF1AADFECC0DEF" blockId="18.[151,497,871,931]" box="[151,313,906,931]" pageId="18" pageNumber="319">
(
<figureCitation id="137E2A75FF9788089DB71AADFF030DEF" box="[159,246,906,931]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Figs. 11</figureCitation>
AK)
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF9788089DBF1AF5FE400A43" pageId="18" pageNumber="319" type="materials_examined">
<paragraph id="8BFA36F0FF9788089DBF1AF5FE400A43" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
<emphasis id="B931EAE2FF9788089DBF1AF5FD7E0DA7" bold="true" box="[151,651,978,1003]" pageId="18" pageNumber="319">
<typeStatus id="54FE8852FF9788089DBF1AF5FF270DA7" box="[151,210,978,1003]" pageId="18" pageNumber="319">Type</typeStatus>
material.
<typeStatus id="54FE8852FF9788089C661AF5FE4A0DA7" box="[334,447,978,1003]" pageId="18" pageNumber="319" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3527660FF9788089CE41AF5FDDE0DA7" box="[460,555,978,1003]" name="China" pageId="18" pageNumber="319">CHINA</collectingCountry>
, Hubei:
</emphasis>
♂, Shennongjia, Jiuchonggangou,
<quantity id="4CBD9B15FF97880899321AF4FB910DA6" box="[1050,1124,979,1003]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="18" pageNumber="319" unit="m" value="870.0">870 m</quantity>
, sifting,
<date id="FFFB1030FF97880899E01AF4FAA60DA7" box="[1224,1363,979,1003]" pageId="18" pageNumber="319" value="1998-07-19">19.VII.1998</date>
, Tian- Hong Luo leg. (IZ-CAS).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF97880E9DEF1D3DFE6B0EFC" lastPageId="20" lastPageNumber="321" pageId="18" pageNumber="319" type="description">
<paragraph id="8BFA36F0FF9788089DEF1D3DFF010AD3" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
<emphasis id="B931EAE2FF9788089DEF1D3DFEAC0A7F" bold="true" box="[199,345,1050,1075]" pageId="18" pageNumber="319">Description.</emphasis>
<emphasis id="B931EAE2FF9788089C481D3DFE6F0A7F" box="[352,410,1050,1075]" italics="true" pageId="18" pageNumber="319">Male</emphasis>
. EBL:
<quantity id="4CBD9B15FF9788089CC51D3CFDA00A7E" box="[493,597,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="2.37" pageId="18" pageNumber="319" unit="mm" value="2.37">2.37 mm</quantity>
. Length of different body parts: HL: AL: PL: ELL = 0.39: 0.61: 0.59:
<quantity id="4CBD9B15FF9788089DBF1D18FF080A1A" box="[151,253,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="18" pageNumber="319" unit="mm" value="1.32">1.32 mm</quantity>
; width: HW: EW: PW: ELW = 0.63: 0.09: 0.97:
<quantity id="4CBD9B15FF9788089E681D18FC530A1A" box="[832,934,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="18" pageNumber="319" unit="mm" value="1.03">1.03 mm</quantity>
. Proportion of antennomeres from base to tip in µm (length × width): 109 × 32, 83 × 39, 47 × 34, 34 × 34, 36 × 41, 24 × 45, 49 × 60, 17 × 61, 46 × 70, 46 × 70, 80 × 74.
</paragraph>
<paragraph id="8BFA36F0FF9788089DEF1D8DFA600B63" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">Habitus elongated oval, relatively convex and lustreless. Moderately pigmented: mostly chestnut brown; mouthparts, basal six antennomeres and apical half of ultimate antennomere, base of pronotum, elytral apex, and tarsi more or less yellowish. Dorsum continually clothed with fine, recumbent and sallow pubescence. Insertions of pubescence on dorsal surfaces of head, pronotum, scutellum, elytra and femora align along transverse striolations.</paragraph>
<paragraph id="8BFA36F0FF9788089DEF1C1DFE850B89" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
Head moderately short and convex, transversely and superficially striolated, interspaces narrower than that on pronotum, HW/HL = 1.64; anterior margin round. Compound eyes slightly reduced, EW/HW = 0.09. Antennae slender (
<figureCitation id="137E2A75FF9788089DD11CA2FEBF0BD2" box="[249,330,1413,1438]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
A), AL/HW = 0.96; 5th slightly longer than 4th; 6th wider than long; 9th similar to 10th, wider than long; 11th pear-like.
</paragraph>
<paragraph id="8BFA36F0FF9788089DEF1CF6FA73087D" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
Pronotum transverse and convex (
<figureCitation id="137E2A75FF9788089F671CF7FD540BA4" box="[591,673,1488,1513]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
D), widest at hind corners, PW/PL = 1.65. Sides regularly curved, narrowing from posterior to anterior; hind corners drawn out and bluntly rounded. Posterior margin with distinct postero-lateral emargination. Surface transversely striolated, SP: 3840, interspaces narrower than that on elytra.
</paragraph>
<paragraph id="8BFA36F0FF9788089DEF1F1BFDC708D1" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
Elytra slender and regularly convex, widest at about basal 1/3, ELL/EW = 1.28. Sides feebly curved, gradually narrowing from base to apices; apices nearly rounded (
<figureCitation id="137E2A75FF9788089E0F1F47FC890834" box="[807,892,1632,1657]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
E). Surface transversely striolated, SE: 6870. Metathoracic wings fully developed.
</paragraph>
<paragraph id="8BFA36F0FF9788089DEF1F8FFF1A0960" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
Prolegs robust, with basal three protarsomeres moderately expanded (
<figureCitation id="137E2A75FF9788089EDE1F8FFBBE088C" box="[1014,1099,1704,1729]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
B): TW/BTW = 1.37. Spinal arrangement on ventral side of protibia as shown in
<figureCitation id="137E2A75FF9788089FF61FEBFCD908A9" box="[734,812,1740,1765]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
C. Profemora wider than protibiae, hairs on ventral side moderately longer and stronger than that of protibiae, but much longer on inner margin. Mesotibiae moderately arcuate.
</paragraph>
<paragraph id="8BFA36F0FF9788089DEF1E1FFEB609D4" blockId="18.[151,1437,978,2017]" pageId="18" pageNumber="319">
Ventrite VII with two large hook-like teeth at posterior margin (
<figureCitation id="137E2A75FF9788089EA71E1FFC2B091C" box="[911,990,1848,1873]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
F). Ventrite VIII widely and moderately emarginate posteriorly (
<figureCitation id="137E2A75FF9788089C8C1E7BFE010939" box="[420,500,1884,1909]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
G). Segment IX: spiculum gastrale very wide, sharply acuminated towards anterior end (
<figureCitation id="137E2A75FF9788089DE61EA7FEEB09D4" box="[206,286,1920,1945]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">Fig. 11</figureCitation>
H).
</paragraph>
<paragraph id="8BFA36F0FF9788099DEF1E83FA9B0F0C" blockId="18.[151,1437,978,2017]" lastBlockId="19.[151,1436,151,320]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
Aedeagus broadly oval in dorsal view (
<figureCitation id="137E2A75FF9788089F561E83FD3A09F1" box="[638,719,1956,1981]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="18" pageNumber="319">
Fig.
<date id="FFFB1030FF9788089F9A1E82FD3A09F1" box="[690,719,1957,1981]" pageId="18" pageNumber="319" value="2011-01">11</date>
</figureCitation>
I): median lobe widest before base and gently narrowed towards middle, then slightly widened again before middle, finally gradually narrowed towards apex; apical expansion distinctly and narrowly stuck out at apex; parameres firmly attached to median lobe, somewhat widened at apical part and more or less hided before middle. Ventral operculum poorly sclerotized, limit inconspicuous, divided in apical half, two lobes asymmetrical, rounded apically at different levels (
<figureCitation id="137E2A75FF9688099ECB19F8FBC30EB4" box="[995,1078,223,248]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="19" pageNumber="320">Fig. 11</figureCitation>
J). In lateral view, median lobe slightly bent ventrad, sharply acuminated to a subacute apex, and three ventrally-oriented setae inserted below apex as well as two just before middle (
<figureCitation id="137E2A75FF9688099F3F1800FD930F0C" box="[535,614,295,320]" captionStart="FIGURE 11" captionStartId="19.[151,250,1918,1940]" captionTargetBox="[175,1411,385,1881]" captionTargetId="figure@19.[151,1436,373,1897]" captionTargetPageId="19" captionText="FIGURE 11. Ptomaphaginus luoi sp. nov. ♂: A, antenna (dorsal view); B, proleg (dorsal view); C, protibia (ventral view); D, pronotum (dorsal view); E, elytral apex (dorsoapical view); F, ventrite VII (ventral view); G, ventrite VIII (ventral view); H, abdominal segment IX (ventral view); I, aedeagus (dorsal view); J, aedeagus (ventral view); K, aedeagus (lateral view). Scales: 0.1 mm." httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="19" pageNumber="320">Fig. 11</figureCitation>
K). Internal stylus moderately wide, multiannulate at about basal 1/2.
</paragraph>
<caption id="DF3A6678FF9688099DBF1E59FF1E09BC" httpUri="https://zenodo.org/record/243871/files/figure.png" pageId="19" pageNumber="320" targetBox="[175,1411,385,1881]" targetPageId="19">
<paragraph id="8BFA36F0FF9688099DBF1E59FF1E09BC" blockId="19.[151,1437,1918,2032]" pageId="19" pageNumber="320">
<emphasis id="B931EAE2FF9688099DBF1E59FD7D09D8" bold="true" box="[151,648,1918,1940]" pageId="19" pageNumber="320">
FIGURE 11.
<taxonomicName id="4C454D73FF9688099C0C1E59FE0209D8" box="[292,503,1918,1940]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="luoi" status="sp. nov.">
<emphasis id="B931EAE2FF9688099C0C1E59FE0209D8" bold="true" box="[292,503,1918,1940]" italics="true" pageId="19" pageNumber="320">Ptomaphaginus luoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FF9688099CD51E59FDBB09D8" box="[509,590,1918,1940]" pageId="19" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
♂: A
</emphasis>
, antenna (dorsal view);
<emphasis id="B931EAE2FF9688099E541E59FC7809D8" bold="true" box="[892,909,1918,1940]" pageId="19" pageNumber="320">B</emphasis>
, proleg (dorsal view);
<emphasis id="B931EAE2FF968809995C1E59FB7309D8" bold="true" box="[1140,1158,1918,1940]" pageId="19" pageNumber="320">C</emphasis>
, protibia (ventral view);
<emphasis id="B931EAE2FF96880998AB1E59FA6009D8" bold="true" box="[1411,1429,1918,1940]" pageId="19" pageNumber="320">D</emphasis>
, pronotum (dorsal view);
<emphasis id="B931EAE2FF9688099CBF1EBAFE5D09FF" bold="true" box="[407,424,1949,1971]" pageId="19" pageNumber="320">E</emphasis>
, elytral apex (dorsoapical view);
<emphasis id="B931EAE2FF9688099E2B1EBAFCE509FF" bold="true" box="[771,784,1949,1971]" pageId="19" pageNumber="320">F</emphasis>
, ventrite VII (ventral view);
<emphasis id="B931EAE2FF96880999141EBAFBB809FF" bold="true" box="[1084,1101,1949,1971]" pageId="19" pageNumber="320">G</emphasis>
, ventrite VIII (ventral view);
<emphasis id="B931EAE2FF96880998AA1EBAFA6309FF" bold="true" box="[1410,1430,1949,1971]" pageId="19" pageNumber="320">H</emphasis>
, abdominal segment IX (ventral view);
<emphasis id="B931EAE2FF9688099F351E9BFDD2099E" bold="true" box="[541,551,1980,2002]" pageId="19" pageNumber="320">I</emphasis>
, aedeagus (dorsal view);
<emphasis id="B931EAE2FF9688099E001E9BFCC0099E" bold="true" box="[808,821,1980,2002]" pageId="19" pageNumber="320">J</emphasis>
, aedeagus (ventral view);
<emphasis id="B931EAE2FF96880999161E9BFBA7099E" bold="true" box="[1086,1106,1980,2002]" pageId="19" pageNumber="320">K</emphasis>
, aedeagus (lateral view). Scales: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BFA36F0FF91880E9DEF19B0FE6B0EFC" blockId="20.[151,1436,151,429]" box="[199,414,151,176]" pageId="20" pageNumber="321">
<emphasis id="B931EAE2FF91880E9DEF19B0FEEE0EFC" box="[199,283,151,176]" italics="true" pageId="20" pageNumber="321">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF91880E9DEF199BFDE50E99" box="[199,528,188,213]" pageId="20" pageNumber="321" type="distribution">
<paragraph id="8BFA36F0FF91880E9DEF199BFDE50E99" blockId="20.[151,1436,151,429]" box="[199,528,188,213]" pageId="20" pageNumber="321">
<emphasis id="B931EAE2FF91880E9DEF199BFE950E99" bold="true" box="[199,352,188,213]" pageId="20" pageNumber="321">Distribution.</emphasis>
<collectingCountry id="F3527660FF91880E9C4F199BFE5F0E99" box="[359,426,188,213]" name="China" pageId="20" pageNumber="321">China</collectingCountry>
(Hubei).
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF91880E9DEF19F8FBDA0F50" pageId="20" pageNumber="321" type="etymology">
<paragraph id="8BFA36F0FF91880E9DEF19F8FBDA0F50" blockId="20.[151,1436,151,429]" pageId="20" pageNumber="321">
<emphasis id="B931EAE2FF91880E9DEF19F8FEBB0EB4" bold="true" box="[199,334,223,248]" pageId="20" pageNumber="321">Etymology.</emphasis>
The specific epithet is dedicated to Dr. Tian-Hong Luo (IZ-CAS, Beijing,
<collectingCountry id="F3527660FF91880E99BF19F8FB2B0EB4" box="[1175,1246,223,248]" name="China" pageId="20" pageNumber="321">China</collectingCountry>
), the collector of this new species and a researcher on forensic coleopterology in our research team.
</paragraph>
</subSubSection>
<subSubSection id="C35F657BFF91880E9DEF1800FC1E0FE0" pageId="20" pageNumber="321" type="discussion">
<paragraph id="8BFA36F0FF91880E9DEF1800FC1E0FE0" blockId="20.[151,1436,151,429]" pageId="20" pageNumber="321">
<emphasis id="B931EAE2FF91880E9DEF1800FECE0F0C" bold="true" box="[199,315,295,320]" pageId="20" pageNumber="321">Remarks.</emphasis>
This new species is sympatric with
<taxonomicName id="4C454D73FF91880E9FFD180EFC610F0C" box="[725,916,295,320]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="shennongensis" status="sp. nov.">
<emphasis id="B931EAE2FF91880E9FFD180EFC610F0C" box="[725,916,295,320]" italics="true" pageId="20" pageNumber="321">P. shennongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025799FF91880E9EB41800FC030F0C" box="[924,1014,295,320]" pageId="20" pageNumber="321" rank="species">
<emphasis id="B931EAE2FF91880E9EB41800FC030F0C" bold="true" box="[924,1014,295,320]" pageId="20" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
, but it seems that the former is much more rare than the latter from specimen quantity. And more, this new species is one of those of the genus
<taxonomicName id="4C454D73FF91880E9DBF1848FEB20FC4" box="[151,327,367,392]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF91880E9DBF1848FEB20FC4" box="[151,327,367,392]" italics="true" pageId="20" pageNumber="321">Ptomaphaginus</emphasis>
</taxonomicName>
that can be identified just from external morphology without aedeagal characters, for its two particularly large and hook-like teeth at the posterior margin of ventrite VII.
</paragraph>
</subSubSection>
</treatment>
</document>