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<document id="E1B0CE1A05F9DEC51D56AE388DB96C0A" ID-CLB-Dataset="9477" ID-DOI="10.11646/zootaxa.4885.4.8" ID-GBIF-Dataset="8cd9d74c-2aac-43b8-be04-c649f9300b1d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296987" ID-ZooBank="16AC9A8F-D326-42DC-91A2-029E2CF1A0ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606685483353" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03EC87C09F4AF30D20D8F8AAFC0FFC49" docLanguage="en" docName="zootaxa.4885.4.8.pdf" docOrigin="Zootaxa 4885 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discheramocephalus bisulcatus Darby 2013" docType="treatment" docVersion="8" lastPageNumber="595" masterDocId="FFD5FFB89F49F309204FFFB2FFDEFF81" masterDocTitle="Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae)" masterLastPageNumber="595" masterPageNumber="591" pageNumber="594" updateTime="1698846858831" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2964720C1183ED96A049A9259E0930B4">Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae)</mods:title>
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<mods:number id="BBA6C51ADD9248678B6E71315119E049">2020-11-27</mods:number>
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<mods:classification id="33A4D838F63112424613D4EE05B5BC9C">journal article</mods:classification>
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<treatment id="03EC87C09F4AF30D20D8F8AAFC0FFC49" ID-DOI="http://doi.org/10.5281/zenodo.4324253" ID-GBIF-Taxon="170599101" ID-Zenodo-Dep="4324253" LSID="urn:lsid:plazi:treatment:03EC87C09F4AF30D20D8F8AAFC0FFC49" httpUri="http://treatment.plazi.org/id/03EC87C09F4AF30D20D8F8AAFC0FFC49" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594">
<subSubSection id="C35F655D9F4AF30A20D8F8AAFDB0F8B2" box="[151,622,1816,1843]" pageId="3" pageNumber="594" type="nomenclature">
<paragraph id="8BFA36D69F4AF30A20D8F8AAFDB0F8B2" blockId="3.[151,622,1816,1879]" box="[151,622,1816,1843]" pageId="3" pageNumber="594">
<heading id="D0B281BA9F4AF30A20D8F8AAFDB0F8B2" bold="true" box="[151,622,1816,1843]" fontSize="11" level="1" pageId="3" pageNumber="594" reason="1">
<taxonomicName id="4C454D559F4AF30A20D8F8AAFDB0F8B2" ID-CoL="36SC5" authority="Darby" authorityName="Darby" authorityYear="2013" box="[151,622,1816,1843]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="bisulcatus">
<emphasis id="B931EAC49F4AF30A20D8F8AAFDB0F8B2" bold="true" box="[151,622,1816,1843]" pageId="3" pageNumber="594">
<emphasis id="B931EAC49F4AF30A20D8F8AAFDC9F8B2" bold="true" box="[151,535,1816,1843]" italics="true" pageId="3" pageNumber="594">Discheramocephalus bisulcatus</emphasis>
Darby
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F655D9F4AF30A20D8F88EFEA9F8D6" box="[151,375,1852,1879]" pageId="3" pageNumber="594" type="description">
<paragraph id="8BFA36D69F4AF30A20D8F88EFEA9F8D6" blockId="3.[151,622,1816,1879]" box="[151,375,1852,1879]" pageId="3" pageNumber="594">
(
<figureCitation id="137E2A539F4AF30A20EFF88EFEDEF8D6" box="[160,256,1852,1879]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="3" pageNumber="594">Fig. 1–3</figureCitation>
,
<figureCitation id="137E2A539F4AF30A2143F88FFEC2F8D6" box="[268,284,1853,1879]" captionStart="FIGURES 7–9" captionStartId="2.[151,264,1967,1991]" captionTargetBox="[166,1421,1562,1926]" captionTargetId="figure-566@2.[151,1436,1528,1943]" captionTargetPageId="2" captionText="FIGURES 7–9. Abdominal internal ‘cavities’ of segment VIII. Discheramocephalus vasilii Darby, male (7); D. bisulcatus Darby, male (8) and female (9)." figureDoi="http://doi.org/10.5281/zenodo.4296991" httpUri="https://zenodo.org/record/4296991/files/figure.png" pageId="3" pageNumber="594">7</figureCitation>
,
<figureCitation id="137E2A539F4AF30A2167F88FFEB1F8D6" box="[296,367,1853,1879]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="3" pageNumber="594">10–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F655D9F4AF30A20D8F8C1FC45F80C" box="[151,923,1907,1933]" pageId="3" pageNumber="594" type="reference_group">
<paragraph id="8BFA36D69F4AF30A20D8F8C1FC45F80C" blockId="3.[151,923,1907,1933]" box="[151,923,1907,1933]" pageId="3" pageNumber="594">
<treatmentCitationGroup id="AB5511F89F4AF30A20D8F8C1FC45F80C" box="[151,923,1907,1933]" pageId="3" pageNumber="594">
<treatmentCitation id="0AE410C79F4AF30A20D8F8C1FD16F80C" ID-DOI="http://doi.org/10.5281/zenodo.6154587" ID-Zenodo-Dep="6154587" LSID="urn:lsid:zoobank.org:act:16EC7F5F-21AC-4F5C-B000-16F2A863EB74" author="Darby, M." box="[151,712,1907,1933]" httpUri="http://treatment.plazi.org/id/187B87854057091F3BE6FEEAFC9EF346" page="241" pageId="3" pageNumber="594" year="2013">
<taxonomicName id="4C454D559F4AF30A20D8F8C1FD16F80C" ID-CoL="36SC5" authority="Darby, 2013: 241" authorityName="Darby" authorityPageNumber="241" authorityYear="2013" box="[151,712,1907,1933]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="bisulcatus">
<emphasis id="B931EAC49F4AF30A20D8F8C1FE23F80C" box="[151,509,1907,1933]" italics="true" pageId="3" pageNumber="594">Discheramocephalus bisulcatus</emphasis>
<bibRefCitation id="EFD44B279F4AF30A224CF8C1FD16F80C" author="Darby, M." box="[515,712,1907,1933]" pageId="3" pageNumber="594" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby, 2013: 241</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<figureCitation id="137E2A539F4AF30A229BF8C1FCCBF80C" box="[724,789,1907,1933]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="3" pageNumber="594">Fig. 1</figureCitation>
, 19, 21, 27.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35F655D9F4AF30D20D8F809FBC0FD4D" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594" type="materials_examined">
<paragraph id="8BFA36D69F4AF30A20D8F809FC6DF878" blockId="3.[151,1436,1979,2042]" pageId="3" pageNumber="594">
<emphasis id="B931EAC49F4AF30A20D8F809FE83F854" bold="true" box="[151,349,1979,2005]" pageId="3" pageNumber="594">Material studied</emphasis>
: 1 ♁,
<materialsCitation id="3B2D3C8B9F4AF30A21EBF809FC71F878" ID-GBIF-Occurrence="2985804302" collectingDate="2019-12-04" collectionCode="MNHW" collectorName="P. Jaloszynski" country="Madagascar" elevation="950" latitude="-21.26" location="Ranomafana National Park" longLatPrecision="758" longitude="47.43" pageId="3" pageNumber="594" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D43FD5F9F4AF30A21EBF809FE3AF857" box="[420,484,1979,2006]" pageId="3" pageNumber="594" type="female">2 ♀♀</specimenCount>
,
<collectingCountry id="F35276469F4AF30A21BEF809FDA4F854" box="[497,634,1979,2005]" name="Madagascar" pageId="3" pageNumber="594">Madagascar</collectingCountry>
,
<location id="8E9A600D9F4AF30A22CAF809FC69F854" LSID="urn:lsid:plazi:treatment:03EC87C09F4AF30D20D8F8AAFC0FFC49:8E9A600D9F4AF30A22CAF809FC69F854" box="[645,951,1979,2005]" country="Madagascar" latitude="-21.26" longLatPrecision="758" longitude="47.43" name="Ranomafana National Park" pageId="3" pageNumber="594">Ranomafana National Park</location>
,
<geoCoordinate id="EE7150119F4AF30A238DF809FBCDF854" box="[962,1043,1979,2005]" degrees="21.26" direction="south" orientation="latitude" pageId="3" pageNumber="594" precision="555" value="-21.26">21.26S</geoCoordinate>
,
<geoCoordinate id="EE7150119F4AF30A2450F809FBACF854" box="[1055,1138,1979,2005]" degrees="47.43" direction="east" orientation="longitude" pageId="3" pageNumber="594" precision="555" value="47.43">47.43E</geoCoordinate>
,
<quantity id="4CBD9B339F4AF30A2432F809FAD2F857" box="[1149,1292,1979,2006]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="594" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
<elevation id="0068D1E59F4AF30A2432F809FAD2F857" box="[1149,1292,1979,2006]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="594" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</elevation>
</quantity>
,
<date id="FFFB10169F4AF30A2558F809FA46F854" box="[1303,1432,1979,2005]" pageId="3" pageNumber="594" value="2019-12-04">
<collectingDate id="EFBFE9FE9F4AF30A2558F809FA46F854" box="[1303,1432,1979,2005]" pageId="3" pageNumber="594" value="2019-12-04">12.04.2019</collectingDate>
</date>
, sifted from leaf litter in secondary forest, leg.
<collectorName id="26B053009F4AF30A22D5F852FCEBF878" box="[666,821,2015,2042]" pageId="3" pageNumber="594">P. Jałoszyński</collectorName>
(
<collectionCode id="ED54AE139F4AF30A230BF86DFC78F878" box="[836,934,2015,2041]" pageId="3" pageNumber="594">MNHW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFA36D69F4DF30D2088FF24FD16FE09" blockId="4.[151,1437,150,968]" pageId="4" pageNumber="595">
<emphasis id="B931EAC49F4DF30D2088FF24FE68FF31" bold="true" box="[199,438,150,176]" pageId="4" pageNumber="595">Emended diagnosis.</emphasis>
Head with biarcuate (broadly M-shaped) frontal groove, lacking well-defined paired impressions; frons in front of groove strongly microsculptured; pronotum with two pairs of longitudinal sulci reaching anterior pronotal margin, inner pair gradually broadening posterad, pronotum also with two pairs of oval antebasal impressions between sulci; proximal flagellomeres more than 3 times as long as broad; ‘cavities’ of abdominal segment VIII in male and female narrower than 1/4 width of segment; posterior margin of sternite IX in male and female similar, with broad median area devoid of denticles; aedeagus stout, in dorsal view with rounded sides, in lateral view abruptly bent ventrad at obtuse angle.
</paragraph>
<paragraph id="8BFA36D69F4DF30D2088FE20FAC3FDBD" blockId="4.[151,1437,150,968]" pageId="4" pageNumber="595">
<emphasis id="B931EAC49F4DF30D2088FE20FDC8FE2D" bold="true" box="[199,534,402,428]" pageId="4" pageNumber="595">Supplementary description.</emphasis>
Male (
<figureCitation id="137E2A539F4DF30D2221FE20FD0FFE2C" box="[622,721,402,429]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="4" pageNumber="595">Fig. 4–5</figureCitation>
) not differing externally from female; antennae (
<figureCitation id="137E2A539F4DF30D255FFE20FA89FE2C" box="[1296,1367,402,429]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="4" pageNumber="595">Fig. 6</figureCitation>
) with conspicuously elongate antennomeres 3–9, each 3 times or more longer than broad; abdominal segment VIII in both sexes with pair of small lateral internal ‘cavities’, each narrower than 1/4 of width of segment, in male (
<figureCitation id="137E2A539F4DF30D251EFE68FA4DFE75" box="[1361,1427,474,500]" captionStart="FIGURES 7–9" captionStartId="2.[151,264,1967,1991]" captionTargetBox="[166,1421,1562,1926]" captionTargetId="figure-566@2.[151,1436,1528,1943]" captionTargetPageId="2" captionText="FIGURES 7–9. Abdominal internal ‘cavities’ of segment VIII. Discheramocephalus vasilii Darby, male (7); D. bisulcatus Darby, male (8) and female (9)." figureDoi="http://doi.org/10.5281/zenodo.4296991" httpUri="https://zenodo.org/record/4296991/files/figure.png" pageId="4" pageNumber="595">Fig. 8</figureCitation>
) ‘cavities’ appear slightly smaller than in female (
<figureCitation id="137E2A539F4DF30D22F3FE4CFCDFFD99" box="[700,769,510,536]" captionStart="FIGURES 7–9" captionStartId="2.[151,264,1967,1991]" captionTargetBox="[166,1421,1562,1926]" captionTargetId="figure-566@2.[151,1436,1528,1943]" captionTargetPageId="2" captionText="FIGURES 7–9. Abdominal internal ‘cavities’ of segment VIII. Discheramocephalus vasilii Darby, male (7); D. bisulcatus Darby, male (8) and female (9)." figureDoi="http://doi.org/10.5281/zenodo.4296991" httpUri="https://zenodo.org/record/4296991/files/figure.png" pageId="4" pageNumber="595">Fig. 9</figureCitation>
); posterior margin of sternite IX in both sexes (
<figureCitation id="137E2A539F4DF30D2557FE4CFA4AFD99" box="[1304,1428,510,536]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="4" pageNumber="595">Fig. 13–14</figureCitation>
) of similar shape, with lateral groups of 2–3 large denticles and arcuate median region lacking denticles.
</paragraph>
<paragraph id="8BFA36D69F4DF30D2088FDF4FCBDFD28" blockId="4.[151,1437,150,968]" pageId="4" pageNumber="595">
Aedeagus (
<figureCitation id="137E2A539F4DF30D2109FDF4FE1CFDE0" box="[326,450,582,609]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="4" pageNumber="595">Fig. 15–17</figureCitation>
) relatively stout and short, less than 0.2 times as long as body, in dorsal view (
<figureCitation id="137E2A539F4DF30D250DFDF4FA4AFDE0" box="[1346,1428,582,609]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="4" pageNumber="595">Fig. 15</figureCitation>
) with evenly rounded sides, apex (
<figureCitation id="137E2A539F4DF30D2240FDD8FDBFFD04" box="[527,609,618,645]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="4" pageNumber="595">Fig. 16</figureCitation>
) subtrapezoidal, in lateral view distinctly bent ventrad by an obtuse angle (
<figureCitation id="137E2A539F4DF30D20EFFD3CFF2DFD29" box="[160,243,654,680]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="4" pageNumber="595">Fig. 17</figureCitation>
). Endophallus poorly visible (also with phase contrast).
</paragraph>
<paragraph id="8BFA36D69F4DF30D2088FD00FBC0FD4D" blockId="4.[151,1437,150,968]" box="[199,1054,690,716]" pageId="4" pageNumber="595">
Spermatheca (
<figureCitation id="137E2A539F4DF30D2128FD00FE66FD4D" box="[359,440,690,716]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="4" pageNumber="595">Fig. 18</figureCitation>
) simple, lacking loops and coils, with elongate funnel.
</paragraph>
</subSubSection>
<subSubSection id="C35F655D9F4DF30D2088FD64FC0FFC49" pageId="4" pageNumber="595" type="discussion">
<paragraph id="8BFA36D69F4DF30D2088FD64FEA6FCB9" blockId="4.[151,1437,150,968]" pageId="4" pageNumber="595">
<emphasis id="B931EAC49F4DF30D2088FD64FEE5FD71" bold="true" box="[199,315,726,752]" pageId="4" pageNumber="595">Remarks.</emphasis>
Males of this species were not known so far; they are externally indistinguishable from females. The latter were adequately described by
<bibRefCitation id="EFD44B279F4DF30D2260FD48FD14FC95" author="Darby, M." box="[559,714,762,788]" pageId="4" pageNumber="595" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby (2013)</bibRefCitation>
, except for the details of abdominal structures, which are added in the present study.
</paragraph>
<paragraph id="8BFA36D69F4DF30D2088FCF0FC0FFC49" blockId="4.[151,1437,150,968]" pageId="4" pageNumber="595">
<taxonomicName id="4C454D559F4DF30D2088FCF0FDF3FCDD" authorityName="Darby" authorityYear="2013" box="[199,557,834,860]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="bisulcatus">
<emphasis id="B931EAC49F4DF30D2088FCF0FDF3FCDD" box="[199,557,834,860]" italics="true" pageId="4" pageNumber="595">Discheramocephalus bisulcatus</emphasis>
</taxonomicName>
clearly differs from its sympatric Malagasy relative
<taxonomicName id="4C454D559F4DF30D2437FCF1FB00FCDD" authorityName="in Darby" authorityYear="2013" box="[1144,1246,834,860]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="vasilii">
<emphasis id="B931EAC49F4DF30D2437FCF1FB00FCDD" box="[1144,1246,834,860]" italics="true" pageId="4" pageNumber="595">D. vasilii</emphasis>
</taxonomicName>
in cephalic, pronotal and genital characters while both species have a similar shape of the posterior margin of the sternite IX. This latter structure, i.e. the median portion of the posterior margin lacking denticles and flanked by groups of large teeth, differentiates the Malagasy species from those known from other regions.
</paragraph>
</subSubSection>
</treatment>
</document>
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