treatments-xml/data/03/EC/87/03EC879BCF408309C19228DE8F42FC12.xml
2024-06-21 12:22:17 +02:00

163 lines
18 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3071CA0929B7F2DA08AF1E8CF6D1EBFF" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza &amp; Reguero, Marcelo A." docDate="2018" docId="03EC879BCF408309C19228DE8F42FC12" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patriarchus diastematus Ameghino 1891" docType="treatment" docVersion="7" lastPageNumber="492" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="492" updateTime="1698526258101" updateUser="ExternalLinkService">
<mods:mods id="3F8E692596B8E46219552F6F2320304D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="05266A7429DBB5F28A9AAC21DA8C77E8">
<mods:title id="60A20EF4EE743B2E61D77A2A3E59C7CF">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
</mods:titleInfo>
<mods:name id="186272D9E589A6232F31D34948AC70A6" type="personal">
<mods:role id="1A6D96F210B087A8952499F8EE9858C4">
<mods:roleTerm id="31A70027EA0674A0ABCCA045C2B451FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23C39EFF128319E50CC08EDEFB23EE0C">Fernández, Mercedes</mods:namePart>
</mods:name>
<mods:name id="C7564B352D164CBF636E18B77F1C0C83" type="personal">
<mods:role id="64C2C1E1724FB225155DADB14950EDCE">
<mods:roleTerm id="9FC9BEB18592EB2EF4D0ED0FFDD6EEB0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B127B3B7A00D16EF797AEB80B543016">Fernicola, Juan Carlos</mods:namePart>
</mods:name>
<mods:name id="41ED9968C29D322E9B1B989310AE5758" type="personal">
<mods:role id="8816406FFE6BA3AAF5EDDB66896A433D">
<mods:roleTerm id="FA51FDFB2E6683D5765CE119F47C0A45">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE12CF5DE8B9EC47F4EC6C42C24FD716">Cerdeño, Esperanza</mods:namePart>
</mods:name>
<mods:name id="799E90F0897F6B5010A5E75758224026" type="personal">
<mods:role id="E141F40D36EC9E5EC28F67EE601287D9">
<mods:roleTerm id="51930631CBC85E2BB4E638E16E6ECAC5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="71E7463B6071095D31CB9450BE3381D7">Reguero, Marcelo A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D5643EE237F19E9A6C7B94C367BFE676">text</mods:typeOfResource>
<mods:relatedItem id="F44F9FC24BCCC3317F5D4AE9FE1FB6F2" type="host">
<mods:titleInfo id="229BD8A78E8E87CFB86B53C8F08B91D2">
<mods:title id="EBC04CF44254B8FDB4BF414366F59ED4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="57654F486DBDCD475B5AA55B91B184B1">
<mods:date id="DD6A37CC8F0A0F4CFE59968A8DA9BAF2">2018</mods:date>
<mods:detail id="3B5711B92A627624C93812160FA21F70" type="pubDate">
<mods:number id="F0408F0EB576BACA29E1B74B0277574B">2018-02-28</mods:number>
</mods:detail>
<mods:detail id="DFA22B87AFF89D891195F4EB2560EB6F" type="volume">
<mods:number id="7EC9AA426DAAFA85C1F4E85080A540F1">4387</mods:number>
</mods:detail>
<mods:detail id="FBC263AB33F540F959B8DBFB384C85DF" type="issue">
<mods:number id="FFE92B9AD2491CAB7BDF89A7DDA56B52">3</mods:number>
</mods:detail>
<mods:extent id="0B9AF1BDC9BFA5BB3B5E0209E1CD0DCB" unit="page">
<mods:start id="EB95A0B86DAE36D1EC9B62CC347DBD45">473</mods:start>
<mods:end id="740C2A8F243C1E419D3951936BC76E6A">498</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="80852357131BBABDBA0688ED0A8F7EE2">journal article</mods:classification>
<mods:identifier id="32FAC52811ED3895DA14A2949CEA1EB4" type="CLB-Dataset">30626</mods:identifier>
<mods:identifier id="8FD4BADEB10B10B4BD40B0656AAC2114" type="DOI">10.11646/zootaxa.4387.3.4</mods:identifier>
<mods:identifier id="DFC6992680A89EC64BBA23E471D02F38" type="GBIF-Dataset">43de8956-0706-4f8d-b5a9-f51c30c276d8</mods:identifier>
<mods:identifier id="B3937FD0898BA048CB4FFA45338157F3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0141F78215D9325E8E88213C9FF4BF3D" type="Zenodo-Dep">1187697</mods:identifier>
<mods:identifier id="9555C2257DABF0B418E48F161656C0E6" type="ZooBank">B53A6B20-5386-4E5A-A00F-A559EDF640DF</mods:identifier>
</mods:mods>
<treatment id="03EC879BCF408309C19228DE8F42FC12" ID-DOI="http://doi.org/10.5281/zenodo.5986709" ID-GBIF-Taxon="141264045" ID-Zenodo-Dep="5986709" LSID="urn:lsid:plazi:treatment:03EC879BCF408309C19228DE8F42FC12" httpUri="http://treatment.plazi.org/id/03EC879BCF408309C19228DE8F42FC12" lastPageNumber="492" pageId="19" pageNumber="492">
<subSubSection id="C35F6506CF408309C19228DE8CCEFF67" box="[151,654,151,178]" pageId="19" pageNumber="492" type="nomenclature">
<paragraph id="8BFA368DCF408309C19228DE8CCEFF67" blockId="19.[151,654,151,178]" box="[151,654,151,178]" pageId="19" pageNumber="492">
<heading id="D0B281E1CF408309C19228DE8CCEFF67" bold="true" box="[151,654,151,178]" fontSize="11" level="1" pageId="19" pageNumber="492" reason="1">
<taxonomicName id="4C454D0ECF408309C19228DE8CCEFF67" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[151,654,151,178]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="diastematus">
<emphasis id="B931EA9FCF408309C19228DE8CCEFF67" bold="true" box="[151,654,151,178]" pageId="19" pageNumber="492">
<emphasis id="B931EA9FCF408309C19228DE8FFDFF64" bold="true" box="[151,445,151,177]" italics="true" pageId="19" pageNumber="492">Patriarchus diastematus</emphasis>
<bibRefCitation id="EFD44B7CCF408309C0C128D18CCEFF67" box="[452,654,152,178]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino, 1891</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F6506CF408309C19228978E85FECE" pageId="19" pageNumber="492" type="materials_examined">
<paragraph id="8BFA368DCF408309C19228978E85FECE" blockId="19.[151,1437,222,967]" pageId="19" pageNumber="492">
<emphasis id="B931EA9FCF408309C19228978F4BFF22" bold="true" box="[151,267,222,247]" pageId="19" pageNumber="492">Holotype.</emphasis>
MACN-A 4044 (
<figureCitation id="137E2A08CF408309C0D028978C77FF22" box="[469,567,222,247]" captionStart="FIGURE 3" captionStartId="15.[151,249,1668,1690]" captionTargetBox="[252,1324,201,1641]" captionTargetId="figure@15.[244,1328,193,1651]" captionTargetPageId="15" captionText="FIGURE 3. Patriarchus palmidens, holotype MACN-A 9651a, mandibular symphysis with right i1p1 and left i12 and the alveolus of i3, in labial (A), occlusal (B) and antero-labial (C) views; reproductions of figures 2 (D), 2a (E) and 2b (F) from Ameghino (1889, plate 15). Patriarchus furculosus, syntype MACN-A 3970, left maxillary fragment with P2M3, in occlusal view (G); syntype MACN-A 3972, right mandibular fragment with p2m3, in occlusal view (H); syntype MACN-A 3977, incomplete skull, dorsal view (I). Patriarchus distortus, holotype MACN-A 9634 to MACN-A 9638: MACN-A 9634, skull, ventral (J)and dorsal (K) views; MACN-A 9636, proximal epiphysis of right femur, in anterior view (L); MACN-A 9637 and MACN-A9638, proximal epiphyses of right tibia and fibula,articulated with the distal epiphysis of right femur, in lateral view (M). Patriarchus rectus, holotype MACN-A 4005, right maxillary fragment (vertically rotated) with alveolus of C and series P1M3, in occlusal view (N); holotype MACN-A 4006, left maxillary fragment (vertically rotated) with M13, in occlusal view (O); holotype MACN-A 4007, mandibular fragment with incomplete symphysis, broken alveoli of left and right i1-2 and complete left i3m3 series, in occlusal view (P). Patriarchus diastematus, holotypeMACN-A 4044, two maxillae with alveoli of CP1 andright P2M3 andleft P2M2, in occlusal view (Q). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187703/files/figure.png" pageId="19" pageNumber="492">Fig. 3 Q</figureCitation>
), two maxillae with alveoli of left and right CP1, right P2M3 and left P2 M2.
</paragraph>
</subSubSection>
<subSubSection id="C35F6506CF408309C1C2296F8D06FDEE" pageId="19" pageNumber="492" type="diagnosis">
<paragraph id="8BFA368DCF408309C1C2296F8B2DFE7E" blockId="19.[151,1437,222,967]" pageId="19" pageNumber="492">
<emphasis id="B931EA9FCF408309C1C2296F8F7AFEEA" bold="true" box="[199,314,294,319]" pageId="19" pageNumber="492">Diagnosis</emphasis>
(
<bibRefCitation id="EFD44B7CCF408309C04B296F8C0DFEEA" box="[334,589,294,319]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891: 293</bibRefCitation>
). “
<emphasis id="B931EA9FCF408309C377296F8ACEFEEA" box="[626,1166,294,319]" italics="true" pageId="19" pageNumber="492">Del mismo tamaño que la especie precedente</emphasis>
[
<taxonomicName id="4C454D0ECF408309C5A5296F8B34FEEB" authorityName="Ameghino" authorityYear="1891" box="[1184,1396,294,319]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="rectus">Patriarchus rectus</taxonomicName>
],
<emphasis id="B931EA9FCF408309C48B296E8B19FE7E" italics="true" pageId="19" pageNumber="492">o apenas algo mayor, de la cual se distingue por el P1 pequeño, cilíndrico y separado del que sigue por un pequeño diastema. Series dentarias superiores un poco más arqueadas. Longitud del P2 al M3, 29 milímetros. Ancho del paladar entre los M3, 18 milímetros; ancho entre los M1, 20,5 milímetros; ancho entre los P2, 13 milímetros</emphasis>
”.
</paragraph>
<paragraph id="8BFA368DCF408309C1C229FF8D06FDEE" blockId="19.[151,1437,222,967]" pageId="19" pageNumber="492">
English translation. Same size as the preceding species [
<taxonomicName id="4C454D0ECF408309C24529FF8A4FFE1B" authorityName="Ameghino" authorityYear="1891" box="[832,1039,438,463]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="rectus">
<emphasis id="B931EA9FCF408309C24529FF8A4FFE1B" box="[832,1039,438,463]" italics="true" pageId="19" pageNumber="492">Patriarchus rectus</emphasis>
</taxonomicName>
] or slightly larger, from which it is distinguished by its small, cylindrical P1, separated from the following premolar by a small diastema. Upper tooth row a little more arched. Length from P2 to M3, 29 millimetres. Width of the palate between M3s, 18 millimetres; between M1s, 20.5 millimetres; between P2s, 13 millimetres.
</paragraph>
</subSubSection>
<subSubSection id="C35F6506CF408309C1C22A0F8F42FC12" pageId="19" pageNumber="492" type="discussion">
<paragraph id="8BFA368DCF408309C1C22A0F8F63FCAB" blockId="19.[151,1437,222,967]" pageId="19" pageNumber="492">
<emphasis id="B931EA9FCF408309C1C22A0F8F0FFD8A" bold="true" box="[199,335,582,607]" pageId="19" pageNumber="492">Comments.</emphasis>
<bibRefCitation id="EFD44B7CCF408309C0532A0F8C5EFD8A" box="[342,542,582,607]" pageId="19" pageNumber="492" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">Ameghino (1889)</bibRefCitation>
described this species without providing any illustration of it. In the Ameghino Collection,
<taxonomicName id="4C454D0ECF408309C01C2A258F8BFD56" authorityName="Ameghino" authorityYear="1891" box="[281,459,618,643]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="diastematus">
<emphasis id="B931EA9FCF408309C01C2A258F8BFD56" box="[281,459,618,643]" italics="true" pageId="19" pageNumber="492">Pa. diastematus</emphasis>
</taxonomicName>
is represented just by one specimen. According to Ameghinos catalogue and to
<bibRefCitation id="EFD44B7CCF408309C4552A228EA3FD72" pageId="19" pageNumber="492" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
, maxilla MACN-A 4044 is the
<typeStatus id="54FE882FCF408309C35D2AC68CC8FD72" box="[600,648,655,679]" pageId="19" pageNumber="492">type</typeStatus>
material. This specimen matches the original description because it presents a slightly larger size (length of P2M3 =
<quantity id="4CBD9B68CF408309C3CF2AFA8D6FFD1E" box="[714,815,691,715]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="19" pageNumber="492" unit="mm" value="28.1">28.1 mm</quantity>
) to that of the
<typeStatus id="54FE882FCF408309C2D02AFB8A77FD1E" box="[981,1079,690,715]" pageId="19" pageNumber="492" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C454D0ECF408309C5592AFD8A8EFD1E" authorityName="Ameghino" authorityYear="1891" box="[1116,1230,692,715]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="rectus">
<emphasis id="B931EA9FCF408309C5592AFD8A8EFD1E" box="[1116,1230,692,715]" italics="true" pageId="19" pageNumber="492">Pa. rectus</emphasis>
</taxonomicName>
(length of P2M3 =
<quantity id="4CBD9B68CF408309C1A82A9F8F52FD3A" box="[173,274,726,751]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="19" pageNumber="492" unit="mm" value="27.5">27.5 mm</quantity>
) and, according to its alveolus, P1 is small and cylindrical and is separated from P2 by a small diastema (~
<quantity id="4CBD9B68CF408309C1B22AB28F51FCC6" box="[183,273,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="492" unit="mm" value="1.4">1.4 mm</quantity>
). Besides, all the measurements (length of P2M3 =
<quantity id="4CBD9B68CF408309C2602AB28D9BFCC7" box="[869,987,763,787]" metricMagnitude="-2" metricUnit="m" metricValue="2.888" pageId="19" pageNumber="492" unit="mm" value="28.88">28.88 mm</quantity>
; width of the palate between both M3 =
<quantity id="4CBD9B68CF408309C1922B568EA8FCE3" box="[151,232,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="492" unit="mm" value="19.0">19 mm</quantity>
; width of the palate between both M1 =
<quantity id="4CBD9B68CF408309C3AE2B568D51FCE3" box="[683,785,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="19" pageNumber="492" unit="mm" value="20.8">20.8 mm</quantity>
; width of the palate between both P2 =
<quantity id="4CBD9B68CF408309C5CF2B568B6FFCE2" box="[1226,1327,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="19" pageNumber="492" unit="mm" value="13.7">13.7 mm</quantity>
) coincide with those provided by
<bibRefCitation id="EFD44B7CCF408309C0B32B0B8CC7FC8E" box="[438,647,834,859]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
. As a result, MACN-A 4044 is confirmed as the
<typeStatus id="54FE882FCF408309C5E62B0B8B05FC8E" box="[1251,1349,834,859]" pageId="19" pageNumber="492" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C454D0ECF408309C4732B0D8F5DFCAA" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="diastematus">
<emphasis id="B931EA9FCF408309C4732B0D8F5DFCAA" italics="true" pageId="19" pageNumber="492">Pa. diastematus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA368DCF408309C1C22BC38F42FC12" blockId="19.[151,1437,222,967]" pageId="19" pageNumber="492">
<materialsCitation id="3B2D3CD0CF408309C1C22BC38F42FC12" ID-GBIF-Occurrence="1828873082" collectionCode="SALMA" location="Santa Cruz Province" pageId="19" pageNumber="492" specimenCount="1" stateProvince="Santa Cruz Province" typeStatus="holotype">
<emphasis id="B931EA9FCF408309C1C22BC38DBEFC76" bold="true" box="[199,1022,906,931]" pageId="19" pageNumber="492">
Chronological and geographical distribution of the
<typeStatus id="54FE882FCF408309C24C2BC38D3DFC76" box="[841,893,906,931]" pageId="19" pageNumber="492">type</typeStatus>
specimen.
</emphasis>
Santacrucian
<collectionCode id="ED54AE48CF408309C5A32BC28B4BFC76" box="[1190,1291,907,931]" pageId="19" pageNumber="492">SALMA</collectionCode>
,
<collectingRegion id="4981F86FCF408309C41E2BC28EBFFC12" country="Argentina" name="Santa Cruz" pageId="19" pageNumber="492">
Santa
<collectingRegion id="4981F86FCF408309C4632BC28EBFFC12" inferredBySuffix="true" pageId="19" pageNumber="492">Cruz Province</collectingRegion>
</collectingRegion>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>