517 lines
72 KiB
XML
517 lines
72 KiB
XML
<document id="2AF6DBD45908B44AB2F2F67FCC48288F" ID-DOI="10.11646/zootaxa.5443.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11045155" ID-ZooBank="173C52E1-C650-4B3A-92AF-AE7AC29E8D2C" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1713857146357" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="03D887EDFF8B815C73E1ACD371C2F677" docLanguage="en" docName="zootaxa.5443.2.4.pdf" docOrigin="Zootaxa 5443 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microedus subtilis Shavrin 2024, comb. n." docType="treatment" docVersion="3" lastPageNumber="221" masterDocId="FFE1FF95FF86814C7376A83D712DF322" masterDocTitle="Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region" masterLastPageNumber="223" masterPageNumber="205" pageNumber="218" updateTime="1713968564068" updateUser="guilherme" zenodo-license-document="CLOSED">
|
||
<mods:mods id="FF412D7BB9940CEC3126ADE33581AEC2" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="789A02D2229B8DDADBA5921E1154680D">
|
||
<mods:title id="A10B71EEB58A93DA8BA54C0EC3717936">Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="372A27B3AC844BFA988E589076AF61BB" type="personal">
|
||
<mods:role id="5DC2075D6808046C886BEDEE28999FC4">
|
||
<mods:roleTerm id="A7D1ECC33662AFAFEDB71ACB857D35BA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8A66373C1C58717A342FEA38A18497A3">Shavrin, Alexey V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1055C871A2C9B44BD4C771C0DBB561BA">text</mods:typeOfResource>
|
||
<mods:relatedItem id="85D68F8EB12D9587A6F3AF8B3FE23CD7" type="host">
|
||
<mods:titleInfo id="B4788E8CC9296D92CBC707EE8CD980B0">
|
||
<mods:title id="320784918BA8D856FDBB03079C7876FC">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="37460D9C4C31EA08E60AA43F4B5E9DFD">
|
||
<mods:date id="6DA414FC9B602B3976EE56BE2659EC0A">2024</mods:date>
|
||
<mods:detail id="2545ED481DC38E783045F34513EAE109" type="pubDate">
|
||
<mods:number id="6F0D3DD4B87AD10FA255D4EEE85A2609">2024-04-22</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A8843AD919639FAE78BC4BC0534C9654" type="volume">
|
||
<mods:number id="21BC4AD4E4B266FCD5C3FD1EDDCED0BB">5443</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A79650C95E2DEF69CC9FEB80C03BDAF5" type="issue">
|
||
<mods:number id="E70F506B387DC2CA575EF0002F8CF43A">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="D06C0667514DE1783B82699761D86F95" unit="page">
|
||
<mods:start id="5B755C45851C7C0FEE8FC731F22D5185">205</mods:start>
|
||
<mods:end id="5013549672003E12B2035ADE7BF577DB">223</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="00FE2976F4C234F5CC9195CA3E9A460B">
|
||
<mods:url id="3858DC9368118663FC8565D15A1F2B48">http://dx.doi.org/10.11646/zootaxa.5443.2.4</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="F053652F31B1F7C4FDA9EEC4BDD5DB22">journal article</mods:classification>
|
||
<mods:identifier id="D44FFDE1F5E2C048432142C0326EBC8D" type="DOI">10.11646/zootaxa.5443.2.4</mods:identifier>
|
||
<mods:identifier id="F109D109DD6A3E15D4020E2F010CE674" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="6CBE6835B6F6EE418FAD654D9916FC50" type="Zenodo-Dep">11045155</mods:identifier>
|
||
<mods:identifier id="9A66CFE660CF76C4FD2502453EEB605F" type="ZooBank">173C52E1-C650-4B3A-92AF-AE7AC29E8D2C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D887EDFF8B815C73E1ACD371C2F677" LSID="urn:lsid:plazi:treatment:03D887EDFF8B815C73E1ACD371C2F677" httpUri="http://treatment.plazi.org/id/03D887EDFF8B815C73E1ACD371C2F677" lastPageId="16" lastPageNumber="221" pageId="13" pageNumber="218">
|
||
<subSubSection id="C36B6570FF8B814173E1ACD373B4F62B" box="[151,665,1262,1289]" pageId="13" pageNumber="218" type="nomenclature">
|
||
<paragraph id="8BCE36FBFF8B814173E1ACD373B4F62B" blockId="13.[151,665,1262,1325]" box="[151,665,1262,1289]" pageId="13" pageNumber="218">
|
||
<heading id="D0868197FF8B814173E1ACD373B4F62B" bold="true" box="[151,665,1262,1289]" fontSize="11" level="1" pageId="13" pageNumber="218" reason="1">
|
||
<emphasis id="B905EAE9FF8B814173E1ACD373B4F62B" bold="true" box="[151,665,1262,1289]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1ACD3730BF62A" authority="(Sharp, 1889)" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[151,550,1262,1289]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis" status="comb. nov.">
|
||
<emphasis id="B905EAE9FF8B814173E1ACD37058F62B" bold="true" box="[151,373,1262,1289]" italics="true" pageId="13" pageNumber="218">Microedus subtilis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE04B0AFF8B814172F3ACD37333F62A" author="Sharp, D." box="[389,542,1262,1288]" pageId="13" pageNumber="218" pagination="463 - 476" refId="ref12891" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2365792FF8B81417158ACD273B4F62B" box="[558,665,1263,1289]" pageId="13" pageNumber="218" rank="species">comb. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6570FF8B814173E1AD2E700AF60F" box="[151,295,1299,1325]" pageId="13" pageNumber="218" type="description">
|
||
<paragraph id="8BCE36FBFF8B814173E1AD2E700AF60F" blockId="13.[151,665,1262,1325]" box="[151,295,1299,1325]" pageId="13" pageNumber="218">
|
||
(
|
||
<figureCitation id="134A2A7EFF8B814173E9AD2E7032F60F" box="[159,287,1299,1325]" captionStart-0="FIGURES 31–33" captionStart-1="FIGURES 34–42" captionStartId-0="14.[151,264,1928,1952]" captionStartId-1="15.[151,264,1844,1868]" captionTargetBox-0="[156,1430,1069,1899]" captionTargetBox-1="[159,1428,189,1812]" captionTargetId-0="figure-379@14.[151,1435,1064,1904]" captionTargetId-1="figure-16@15.[151,1436,181,1820]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 31–33. Lectotype labels (Fig. 31–32) and habitus (Fig. 33) of Microedus subtilis: 31–32—lectotype, with type labels, 33—Japan: Honshu: Mt. Hotaka. Scale bar: 1.0 mm." captionText-1="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11045169" figureDoi-1="http://doi.org/10.5281/zenodo.11045171" httpUri-0="https://zenodo.org/record/11045169/files/figure.png" httpUri-1="https://zenodo.org/record/11045171/files/figure.png" pageId="13" pageNumber="218">Figs 31–42</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6570FF8B814173E1AD6A7427F547" pageId="13" pageNumber="218" type="reference_group">
|
||
<paragraph id="8BCE36FBFF8B814173E1AD6A725CF64D" blockId="13.[151,1290,1367,1637]" box="[151,881,1367,1391]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1AD6A725CF64D" box="[151,881,1367,1391]" pageId="13" pageNumber="218">
|
||
<treatmentCitation id="0AD010EAFF8B814173E1AD6A7309F64D" author="Sharp, D." box="[151,548,1367,1391]" page="471" pageId="13" pageNumber="218" year="1889">
|
||
<taxonomicName id="4C714D78FF8B814173E1AD6A7309F64D" ID-CoL="5VF3V" authority="Sharp, 1889: 471" authorityName="Sharp" authorityPageNumber="471" authorityYear="1889" box="[151,548,1367,1391]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B814173E1AD6A7041F64D" box="[151,364,1367,1391]" italics="true" pageId="13" pageNumber="218">Anthophagus subtilis</emphasis>
|
||
<bibRefCitation id="EFE04B0AFF8B81417205AD6A7309F64D" author="Sharp, D." box="[371,548,1367,1391]" pageId="13" pageNumber="218" pagination="463 - 476" refId="ref12891" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889: 471</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD010EAFF8B81417158AD6A725CF64D" author="Bernhauer, M. & Schubert, K." box="[558,881,1367,1391]" page="81" pageId="13" pageNumber="218" year="1910">
|
||
<bibRefCitation id="EFE04B0AFF8B81417158AD6A725CF64D" author="Bernhauer, M. & Schubert, K." box="[558,881,1367,1391]" pageId="13" pageNumber="218" pagination="1 - 86" refId="ref11818" refString="Bernhauer, M. & Schubert, K. (1910) Staphylinidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 19. Junk, Berlin, pp. 1 - 86. https: // doi. org / 10.1007 / 978 - 94 - 011 - 9697 - 0 _ 1" type="book chapter" year="1910">Bernhauer & Schubert 1910: 81</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1AD4B7214F6AC" blockId="13.[151,1290,1367,1637]" box="[151,825,1398,1422]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1AD4B7214F6AC" box="[151,825,1398,1422]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1AD4B704DF6AC" authority="Scheerpeltz 1933: 1067" box="[151,352,1398,1422]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B814173E1AD4B704DF6AC" box="[151,352,1398,1422]" italics="true" pageId="13" pageNumber="218">Philydrodes subtilis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AD010EAFF8B81417218AD4B7372F6AC" author="Scheerpeltz, O." box="[366,607,1398,1422]" page="1067" pageId="13" pageNumber="218" year="1933">
|
||
<bibRefCitation id="EFE04B0AFF8B81417218AD4B7372F6AC" author="Scheerpeltz, O." box="[366,607,1398,1422]" pageId="13" pageNumber="218" pagination="989 - 1500" refId="ref12854" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz 1933: 1067</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AD010EAFF8B8141711FAD4B7214F6AC" author="Watanabe, Y." box="[617,825,1398,1422]" page="268" pageId="13" pageNumber="218" year="1985">
|
||
<bibRefCitation id="EFE04B0AFF8B8141711FAD4B7214F6AC" author="Watanabe, Y." box="[617,825,1398,1422]" pageId="13" pageNumber="218" pagination="264 - 268" refId="ref13496" refString="Watanabe, Y. (1985) Omaliinae. In: Ueno, S. - L., Kurosawa, Y. & Sato, M. (Ed.), The Coleoptera of Japan in Color 2. Hoikusha, Osaka, pp. 264 - 268, pls. 46 - 47. [in Japanese]" type="book chapter" year="1985">Watanabe 1985: 268</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1ADA97532F6E9" blockId="13.[151,1290,1367,1637]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1ADA975AFF68F" box="[151,1154,1428,1453]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1ADA970E8F68E" authority="Nakane & Sawada 1956: 184" box="[151,453,1428,1453]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis" subGenus="Lioplax">
|
||
<emphasis id="B905EAE9FF8B814173E1ADA9703DF68E" box="[151,272,1428,1452]" italics="true" pageId="13" pageNumber="218">Philydrodes</emphasis>
|
||
(
|
||
<emphasis id="B905EAE9FF8B81417269ADA97041F68E" box="[287,364,1428,1452]" italics="true" pageId="13" pageNumber="218">Lioplax</emphasis>
|
||
)
|
||
<emphasis id="B905EAE9FF8B8141720AADA970E8F68E" box="[380,453,1428,1452]" italics="true" pageId="13" pageNumber="218">subtilis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AD010EAFF8B814172A4ADA973D3F68F" author="Nakane, T. & Sawada, K." box="[466,766,1428,1453]" page="184" pageId="13" pageNumber="218" year="1956">
|
||
<bibRefCitation id="EFE04B0AFF8B814172A4ADA973D3F68F" author="Nakane, T. & Sawada, K." box="[466,766,1428,1453]" pageId="13" pageNumber="218" pagination="179 - 186" refId="ref12719" refString="Nakane, T. & Sawada, K. (1956) On the genus Philydrodes Bernhauer in Japan with descriptions of a new subgenus and several new species. The Scientific Reports of the Saikyo University, Series A, 2, 179 - 186." type="journal article" year="1956">Nakane & Sawada 1956: 184</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD010EAFF8B81417071ADA972EDF68E" author="Adachi, T." box="[775,960,1428,1452]" page="166" pageId="13" pageNumber="218" year="1957">
|
||
<bibRefCitation id="EFE04B0AFF8B81417071ADA972EDF68E" author="Adachi, T." box="[775,960,1428,1452]" pageId="13" pageNumber="218" pagination="166 - 200" refId="ref11792" refString="Adachi, T. (1957) The staphylinid fauna of Japan. The Journal of the Tokyo University, 11, 166 - 200." type="journal article" year="1957">Adachi 1957: 166</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AD010EAFF8B814170BCADA975AFF68F" author="Shibata, Y." box="[970,1154,1428,1453]" page="123" pageId="13" pageNumber="218" year="1976">
|
||
<bibRefCitation id="EFE04B0AFF8B814170BCADA975AFF68F" author="Shibata, Y." box="[970,1154,1428,1453]" pageId="13" pageNumber="218" pagination="71 - 212" refId="ref13284" refString="Shibata, Y. (1976) Provisional check list of the family Staphylinidae of Japan. I. Annual Bulletin of the Nichidai Sanko, 19, 71 - 212." type="journal article" year="1976">Shibata 1976: 123</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1AD8E7532F6E9" box="[151,1055,1459,1483]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1AD8E7052F6E9" ID-CoL="3V748" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[151,383,1459,1483]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B814173E1AD8E7052F6E9" box="[151,383,1459,1483]" italics="true" pageId="13" pageNumber="218">Liophilydrodes subtilis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AD010EAFF8B814172FAAD8E7373F6E9" author="Watanabe, Y." box="[396,606,1459,1483]" page="302" pageId="13" pageNumber="218" year="1990">
|
||
<bibRefCitation id="EFE04B0AFF8B814172FAAD8E7373F6E9" author="Watanabe, Y." box="[396,606,1459,1483]" pageId="13" pageNumber="218" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe 1990: 302</bibRefCitation>
|
||
</treatmentCitation>
|
||
, Shibata
|
||
<emphasis id="B905EAE9FF8B814171CDAD8973CAF6E9" box="[699,743,1459,1483]" italics="true" pageId="13" pageNumber="218">et al</emphasis>
|
||
., 2013: 75,
|
||
<treatmentCitation id="0AD010EAFF8B81417016AD8E7532F6E9" author="Hayashi, Y." box="[864,1055,1459,1483]" page="412" pageId="13" pageNumber="218" year="2022">
|
||
<bibRefCitation id="EFE04B0AFF8B81417016AD8E7532F6E9" author="Hayashi, Y." box="[864,1055,1459,1483]" pageId="13" pageNumber="218" pagination="397 - 427" refId="ref12139" refString="Hayashi, Y. (2022) Key to identify Staphylinidae of Japan (Coleoptera). - 1. Subfamily Omaliinae. Insecta Shiyakeana, a special publication from the Entomological Laboratory, Osaka Museum of Natural History, 2022, 397 - 427. [in Japanese]" type="journal article" year="2022">Hayashi 2022: 412</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1ADEF73C9F6C8" blockId="13.[151,1290,1367,1637]" box="[151,740,1490,1514]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1ADEF73C9F6C8" box="[151,740,1490,1514]" pageId="13" pageNumber="218">
|
||
<treatmentCitation id="0AD010EAFF8B814173E1ADEF73B5F6C8" author="Lafer, G. Sh." box="[151,664,1490,1514]" page="90" pageId="13" pageNumber="218" year="2004">
|
||
<taxonomicName id="4C714D78FF8B814173E1ADEF73B5F6C8" ID-CoL="7HB6Q" authority="Lafer, 2004: 90" authorityName="Lafer" authorityPageNumber="90" authorityYear="2004" box="[151,664,1490,1514]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="subtilis" status="syn. nov." subSpecies="iturupensis">
|
||
<emphasis id="B905EAE9FF8B814173E1ADEF70DAF6CB" box="[151,503,1490,1514]" italics="true" pageId="13" pageNumber="218">Liophilydrodes subtilis iturupensis</emphasis>
|
||
<bibRefCitation id="EFE04B0AFF8B8141728AADEF73B5F6C8" author="Lafer, G. Sh." box="[508,664,1490,1514]" pageId="13" pageNumber="218" pagination="89 - 93" refId="ref12253" refString="Lafer, G. Sh. (2004) [New subspecies Liophilydrodes subtilis iturupensis subsp. n. (Coleoptera, Staphylinidae: Omaliinae) from Kurile Islands]. A. I. Kurentsov's Annual Memorial Meetings, 15, 89 - 93. [in Russian]" type="journal article" year="2004">Lafer, 2004: 90</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
<emphasis id="B905EAE9FF8B814171E9ADEF73C9F6C8" bold="true" box="[671,740,1490,1514]" pageId="13" pageNumber="218">
|
||
<taxonomicNameLabel id="A2365792FF8B814171E9ADEF73C9F6C8" box="[671,740,1490,1514]" pageId="13" pageNumber="218">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1ADCD72A0F52B" blockId="13.[151,1290,1367,1637]" box="[151,909,1520,1545]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1ADCD726CF52B" ID-CoL="4GDNQ" authority="Nakane & K. Sawada, 1956: 184" authorityName="Nakane & K. Sawada" authorityPageNumber="184" authorityYear="1956" box="[151,833,1520,1545]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="troglophila" status="syn. nov." subGenus="Lioplax">
|
||
<emphasis id="B905EAE9FF8B814173E1ADCD703DF52A" box="[151,272,1520,1544]" italics="true" pageId="13" pageNumber="218">Philydrodes</emphasis>
|
||
(
|
||
<emphasis id="B905EAE9FF8B81417269ADCD7041F52A" box="[287,364,1520,1544]" italics="true" pageId="13" pageNumber="218">Lioplax</emphasis>
|
||
)
|
||
<emphasis id="B905EAE9FF8B8141720AADCD70C0F52A" box="[380,493,1520,1544]" italics="true" pageId="13" pageNumber="218">troglophila</emphasis>
|
||
Nakane & K. Sawada, 1956: 184
|
||
</taxonomicName>
|
||
<emphasis id="B905EAE9FF8B8141703EADCC72A0F52B" bold="true" box="[840,909,1521,1545]" pageId="13" pageNumber="218">
|
||
<taxonomicNameLabel id="A2365792FF8B8141703EADCC72A0F52B" box="[840,909,1521,1545]" pageId="13" pageNumber="218">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1AE32754DF505" blockId="13.[151,1290,1367,1637]" box="[151,1120,1551,1575]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1AE32754DF505" box="[151,1120,1551,1575]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1AE3270DAF505" authorityName="Nakane & K.Sawada" authorityYear="1956" box="[151,503,1551,1575]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="troglophilus" subGenus="Lioplax">
|
||
<emphasis id="B905EAE9FF8B814173E1AE32703DF505" box="[151,272,1551,1575]" italics="true" pageId="13" pageNumber="218">Philydrodes</emphasis>
|
||
(
|
||
<emphasis id="B905EAE9FF8B81417269AE327041F505" box="[287,364,1551,1575]" italics="true" pageId="13" pageNumber="218">Lioplax</emphasis>
|
||
)
|
||
<emphasis id="B905EAE9FF8B8141720AAE3270DAF505" box="[380,503,1551,1575]" italics="true" pageId="13" pageNumber="218">troglophilus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AD010EAFF8B81417172AE327201F505" author="Ueno, S. - I. & Watanabe, Y." box="[516,812,1551,1575]" page="321" pageId="13" pageNumber="218" year="1966">
|
||
<bibRefCitation id="EFE04B0AFF8B81417172AE327201F505" author="Ueno, S. - I. & Watanabe, Y." box="[516,812,1551,1575]" pageId="13" pageNumber="218" pagination="321 - 337" refId="ref13377" refString="Ueno, S. - I. & Watanabe, Y. (1966) The subterranean staphylinid beetles of the genus Quedius from Japan. Bulletin of the National Science Museum, 9 (3), 321 - 337." type="journal article" year="1966">Uéno & Watanabe 1966: 321</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AD010EAFF8B81417041AE327525F505" author="Watanabe, Y." box="[823,1032,1551,1575]" page="410" pageId="13" pageNumber="218" year="1982">
|
||
<bibRefCitation id="EFE04B0AFF8B81417041AE327525F505" author="Watanabe, Y." box="[823,1032,1551,1575]" pageId="13" pageNumber="218" pagination="409 - 414" refId="ref13454" refString="Watanabe, Y. (1982) The staphylinid beetles of the Ozegahara moor. Ozegahara. Scientific researches of the Highmoor in central Japan. Japan Society for the Promotion of Science, Tokyo, pp. 409 - 414." type="book chapter" year="1982">Watanabe 1982: 410</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AD010EAFF8B81417765AE32754DF505" author="Watanabe, Y." box="[1043,1120,1551,1575]" page="9" pageId="13" pageNumber="218" year="1996">
|
||
<bibRefCitation id="EFE04B0AFF8B81417765AE32754DF505" author="Watanabe, Y." box="[1043,1120,1551,1575]" pageId="13" pageNumber="218" pagination="8 - 18" refId="ref13692" refString="Watanabe, Y. (1996) Staphylinid Beetles (Coleoptera) Found in Caves and Mines of Japan. Journal of the Speleological Society of Japan, 20, 8 - 18." type="journal article" year="1996">1996: 9</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1AE1373C1F564" blockId="13.[151,1290,1367,1637]" box="[151,748,1582,1606]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1AE1373C1F564" box="[151,748,1582,1606]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1AE13733DF564" authority="Watanabe 1986: 545" box="[151,528,1582,1606]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis" subGenus="Liophilydrodes">
|
||
<emphasis id="B905EAE9FF8B814173E1AE13703DF564" box="[151,272,1582,1606]" italics="true" pageId="13" pageNumber="218">Philydrodes</emphasis>
|
||
(
|
||
<emphasis id="B905EAE9FF8B81417269AE13709AF564" box="[287,439,1582,1606]" italics="true" pageId="13" pageNumber="218">Liophilydrodes</emphasis>
|
||
)
|
||
<emphasis id="B905EAE9FF8B814172B1AE13733DF564" box="[455,528,1582,1606]" italics="true" pageId="13" pageNumber="218">subtilis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AD010EAFF8B8141716BAE1373C1F564" author="Watanabe, Y." box="[541,748,1582,1606]" page="545" pageId="13" pageNumber="218" year="1986">
|
||
<bibRefCitation id="EFE04B0AFF8B8141716BAE1373C1F564" author="Watanabe, Y." box="[541,748,1582,1606]" pageId="13" pageNumber="218" pagination="541 - 547" refId="ref13557" refString="Watanabe, Y. (1986) Staphylinid beetles found along the water edges of mountain streams in Oku-Nikko. In: Editorial Board of Fauna and Flora of Nikko (Ed.), Fauna and Flora of Nikko. Tochinoha-shob. ANt., Kanuma, pp. 541 - 547. [in Japanese]" type="book chapter" year="1986">Watanabe 1986: 545</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173E1AE717427F547" blockId="13.[151,1290,1367,1637]" box="[151,1290,1612,1637]" pageId="13" pageNumber="218">
|
||
<treatmentCitationGroup id="AB6111D5FF8B814173E1AE717427F547" box="[151,1290,1612,1637]" pageId="13" pageNumber="218">
|
||
<taxonomicName id="4C714D78FF8B814173E1AE71709CF546" ID-CoL="6QHBH" authorityName="Nakane & K. Sawada" authorityYear="1956" box="[151,433,1612,1636]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="troglophilus">
|
||
<emphasis id="B905EAE9FF8B814173E1AE71709CF546" box="[151,433,1612,1636]" italics="true" pageId="13" pageNumber="218">Liophilydrodes troglophilus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AD010EAFF8B814172C8AE7173A2F546" author="Watanabe, Y." box="[446,655,1612,1637]" page="306" pageId="13" pageNumber="218" year="1990">
|
||
<bibRefCitation id="EFE04B0AFF8B814172C8AE7173A2F546" author="Watanabe, Y." box="[446,655,1612,1637]" pageId="13" pageNumber="218" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe 1990: 306</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AD010EAFF8B814171ECAE717262F547" author="Zerche, L." box="[666,847,1612,1637]" page="289" pageId="13" pageNumber="218" year="2003">
|
||
<bibRefCitation id="EFE04B0AFF8B814171ECAE717262F547" author="Zerche, L." box="[666,847,1612,1637]" pageId="13" pageNumber="218" pagination="277 - 295" refId="ref13813" refString="Zerche, L. (2003) Arten der Hygrogeus - Gruppe aus dem Fernen Osten Russlands, aus Japan und China (Coleoptera, Staphylinidae, Omaliinae, Anthophagini) (Beitrage zur Kenntnis ostpalaarktischer Insekten; 13). Beitrage zur Entomologie, 53 (2), 277 - 295. https: // doi. org / 10.21248 / contrib. entomol. 53.2.277 - 295" type="journal article" year="2003">Zerche 2003: 289</bibRefCitation>
|
||
</treatmentCitation>
|
||
, Shibata
|
||
<emphasis id="B905EAE9FF8B814170DAAE7072F5F546" box="[940,984,1612,1636]" italics="true" pageId="13" pageNumber="218">et al</emphasis>
|
||
. 2013: 75,
|
||
<treatmentCitation id="0AD010EAFF8B8141773DAE717427F547" author="Hayashi, Y." box="[1099,1290,1612,1637]" page="412" pageId="13" pageNumber="218" year="2022">
|
||
<bibRefCitation id="EFE04B0AFF8B8141773DAE717427F547" author="Hayashi, Y." box="[1099,1290,1612,1637]" pageId="13" pageNumber="218" pagination="397 - 427" refId="ref12139" refString="Hayashi, Y. (2022) Key to identify Staphylinidae of Japan (Coleoptera). - 1. Subfamily Omaliinae. Insecta Shiyakeana, a special publication from the Entomological Laboratory, Osaka Museum of Natural History, 2022, 397 - 427. [in Japanese]" type="journal article" year="2022">Hayashi 2022: 412</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6570FF8B814273E1AEAE757CF3F6" lastPageId="14" lastPageNumber="219" pageId="13" pageNumber="218" type="materials_examined">
|
||
<paragraph id="8BCE36FBFF8B814173E1AEAE7210F443" blockId="13.[151,1437,1682,2033]" pageId="13" pageNumber="218">
|
||
<emphasis id="B905EAE9FF8B814173E1AEAE70E5F58E" bold="true" box="[151,456,1682,1709]" pageId="13" pageNumber="218">Type material examined.</emphasis>
|
||
<materialsCitation id="3B193CA6FF8B814172A5AEAE7210F443" collectionCode="BMNH" collectorName="Sharp" pageId="13" pageNumber="218" specimenCode="NHMUK015009854" specimenCount="1" typeStatus="lectotype">
|
||
<typeStatus id="54CA8859FF8B814172A5AEAE736EF58F" box="[467,579,1683,1709]" pageId="13" pageNumber="218" type="lectotype">Lectotype</typeStatus>
|
||
(
|
||
<emphasis id="B905EAE9FF8B81417121AEAE723AF58E" bold="true" box="[599,791,1682,1709]" pageId="13" pageNumber="218">here designated</emphasis>
|
||
) of
|
||
<taxonomicName id="4C714D78FF8B81417038AEAE75F8F58F" authority="Sharp, 1889" authorityName="Sharp" authorityYear="1889" box="[846,1237,1683,1709]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B81417038AEAE7513F58F" box="[846,1086,1683,1709]" italics="true" pageId="13" pageNumber="218">Anthophagus subtilis</emphasis>
|
||
<bibRefCitation id="EFE04B0AFF8B8141773FAEAE75F8F58F" author="Sharp, D." box="[1097,1237,1683,1709]" pageId="13" pageNumber="218" pagination="463 - 476" refId="ref12891" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
<specimenCount id="9D77FD72FF8B81417796AEAE75D8F58F" box="[1248,1269,1683,1709]" pageId="13" pageNumber="218" type="female">♀</specimenCount>
|
||
(
|
||
<figureCitation id="134A2A7EFF8B8141767EAEAE74BCF58F" box="[1288,1425,1683,1709]" captionStart="FIGURES 31–33" captionStartId="14.[151,264,1928,1952]" captionTargetBox="[156,1430,1069,1899]" captionTargetId="figure-379@14.[151,1435,1064,1904]" captionTargetPageId="14" captionText="FIGURES 31–33. Lectotype labels (Fig. 31–32) and habitus (Fig. 33) of Microedus subtilis: 31–32—lectotype, with type labels, 33—Japan: Honshu: Mt. Hotaka. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11045169" httpUri="https://zenodo.org/record/11045169/files/figure.png" pageId="13" pageNumber="218">Figs 31–32</figureCitation>
|
||
): ‘
|
||
<taxonomicName id="4C714D78FF8B814173EDAE8A701BF5F3" box="[155,310,1719,1745]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">Anthophagus</taxonomicName>
|
||
| subtilis. | Type D.S. | Iwake. Japan’ <handwritten on a card with the specimen, in black India ink>, ‘Type’ <round label with red margin, printed>, ‘Japan [underlined by yellow] | G. Lewis.’ <printed>, ‘
|
||
<collectorName id="2684532DFF8B81417661AEE67470F5D7" box="[1303,1373,1755,1781]" pageId="13" pageNumber="218">Sharp</collectorName>
|
||
Coll. | 1905-313.’ <printed>, ‘
|
||
<specimenCode id="DBD79E80FF8B814172C7AEC3738CF43B" box="[433,673,1790,1817]" pageId="13" pageNumber="218">NHMUK015009854</specimenCode>
|
||
’ <printed label, with BMNH barcode on rigth side of the label>, ‘LECTOTYPE |
|
||
<taxonomicName id="4C714D78FF8B81417223AF1E70C4F41F" box="[341,489,1827,1853]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EAE9FF8B81417223AF1E70C4F41F" box="[341,489,1827,1853]" italics="true" pageId="13" pageNumber="218">Anthophagus</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C714D78FF8B81417288AF1E73F2F41F" authority="Sharp, 1889" authorityName="Sharp" authorityYear="1889" box="[510,735,1827,1853]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B81417288AF1E7362F41F" box="[510,591,1827,1853]" italics="true" pageId="13" pageNumber="218">subtilis</emphasis>
|
||
<bibRefCitation id="EFE04B0AFF8B81417121AF1E73F2F41F" author="Sharp, D." box="[599,735,1827,1853]" pageId="13" pageNumber="218" pagination="463 - 476" refId="ref12891" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
| Shavrin A.V. des. 2024’ <red, printed>, ‘
|
||
<taxonomicName id="4C714D78FF8B814177B6AF1E741AF41F" authorityName="LeConte" authorityYear="1874" box="[1216,1335,1827,1853]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EAE9FF8B814177B6AF1E741AF41F" box="[1216,1335,1827,1853]" italics="true" pageId="13" pageNumber="218">Microedus</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C714D78FF8B8141763DAF1E701CF443" authority="(Sharp, 1889)" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B8141763DAF1E74B1F41F" box="[1355,1436,1827,1853]" italics="true" pageId="13" pageNumber="218">subtilis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE04B0AFF8B814173E9AF7A7004F443" author="Sharp, D." box="[159,297,1863,1889]" pageId="13" pageNumber="218" pagination="463 - 476" refId="ref12891" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
| Shavrin A.V., det. 2024’ <printed> (
|
||
<collectionCode id="ED60AE3EFF8B814171A0AF7A721DF443" box="[726,816,1863,1889]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="13" pageNumber="218">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF8B814173B1AF567427F4D3" blockId="13.[151,1437,1682,2033]" pageId="13" pageNumber="218">
|
||
<typeStatus id="54CA8859FF8B814173B1AF567072F4A7" box="[199,351,1899,1925]" pageId="13" pageNumber="218" type="paralectotype">Paralectotype</typeStatus>
|
||
<specimenCount id="9D77FD72FF8B81417212AF567054F4A7" box="[356,377,1899,1925]" count="1" pageId="13" pageNumber="218" type="female">♀</specimenCount>
|
||
(left apical antennomere, left middle tibia and tarsus and right elytron are absent): ‘
|
||
<taxonomicName id="4C714D78FF8B81417674AF5674B0F4A7" box="[1282,1437,1899,1925]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">Anthophagus</taxonomicName>
|
||
| subtilis. D.S.’ <handwritten on a card with the specimen, in black
|
||
<collectingCountry id="F366766BFF8B814170F9AFB272E4F48B" box="[911,969,1935,1961]" name="India" pageId="13" pageNumber="218">India</collectingCountry>
|
||
ink>, ‘
|
||
<collectingCountry id="F366766BFF8B81417761AFB27572F48B" box="[1047,1119,1935,1961]" name="Japan" pageId="13" pageNumber="218">Japan</collectingCountry>
|
||
. [underlined by yellow] | G. Lewis | 1910-320.’ <printed>, ‘Iwake’ <handrwitten>, ‘
|
||
<taxonomicName id="4C714D78FF8B81417070AF8E72BCF4EF" authorityName="Bernhauer" authorityYear="1929" box="[774,913,1971,1997]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">Philydrodes</taxonomicName>
|
||
[handwritten] |
|
||
<taxonomicName id="4C714D78FF8B81417731AF8E75E8F4EF" authority="Sh." authorityName="Sharp" authorityYear="1889" box="[1095,1221,1971,1997]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">subtilis Sh.</taxonomicName>
|
||
[handwritten] Det. C.Koch’ <printed>, ‘
|
||
<taxonomicName id="4C714D78FF8B814172F4AFEA70D4F4D3" authorityName="LeConte" authorityYear="1874" box="[386,505,2007,2033]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EAE9FF8B814172F4AFEA70D4F4D3" box="[386,505,2007,2033]" italics="true" pageId="13" pageNumber="218">Microedus</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C714D78FF8B8141717AAFEA73D3F4D3" authority="(Sharp, 1889)" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[524,766,2007,2033]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="218" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8B8141717AAFEA7370F4D3" box="[524,605,2007,2033]" italics="true" pageId="13" pageNumber="218">subtilis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE04B0AFF8B8141711AAFEA73DBF4D3" author="Sharp, D." box="[620,758,2007,2033]" pageId="13" pageNumber="218" pagination="463 - 476" refId="ref12891" refString="Sharp, D. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463 - 476. https: // doi. org / 10.1080 / 00222938909460371" type="journal article" year="1889">Sharp, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
| Shavrin A.V., det. 2024’ <printed> (
|
||
<collectionCode id="ED60AE3EFF8B814177D5AFEA75D0F4D3" box="[1187,1277,2007,2033]" pageId="13" pageNumber="218">BMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF88814273B1A8AB757CF3F6" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="219">
|
||
<emphasis id="B905EAE9FF88814273B1A8AB7313F392" bold="true" box="[199,574,150,176]" pageId="14" pageNumber="219">Additional material examined:</emphasis>
|
||
<materialsCitation id="3B193CA6FF888142713FA8AB7560F3F6" collectingDate="2014-08-15" collectionCode="BMNH" collectorName="Honshu B. M. & P. M. Hammond'" country="Japan" elevation="1300" location="Gumma Pref. Mt, Hotaka" pageId="14" pageNumber="219" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="HONSHU">
|
||
<emphasis id="B905EAE9FF888142713FA8AB7208F392" bold="true" box="[585,805,150,176]" pageId="14" pageNumber="219">
|
||
<collectingCountry id="F366766BFF888142713FA8AB738AF392" box="[585,679,150,176]" name="Japan" pageId="14" pageNumber="219">JAPAN</collectingCountry>
|
||
:
|
||
<collectingRegion id="49B5F819FF88814271C1A8AB7236F392" box="[695,795,150,176]" pageId="14" pageNumber="219">HONSHU</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D77FD72FF8881427046A8AA7258F393" box="[816,885,151,177]" count="7" pageId="14" pageNumber="219" type="male">7 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D77FD72FF88814270F3A8AB72E7F393" box="[901,970,150,177]" count="6" pageId="14" pageNumber="219" type="female">6 ♀♀</specimenCount>
|
||
: ‘JAPAN:
|
||
<collectorName id="2684532DFF888142773AA8AB75C5F393" box="[1100,1256,150,177]" pageId="14" pageNumber="219">Honshu B.M.</collectorName>
|
||
1980-492
|
||
<collectorName id="2684532DFF888142761FA8AA7034F3F6" pageId="14" pageNumber="219">P.M. Hammond’</collectorName>
|
||
, ‘
|
||
<location id="8EAE6020FF888142725FA8877361F3F6" LSID="urn:lsid:plazi:treatment:03D887EDFF8B815C73E1ACD371C2F677:8EAE6020FF888142725FA8877361F3F6" box="[297,588,186,213]" country="Japan" name="Gumma Pref. Mt, Hotaka" pageId="14" pageNumber="219" stateProvince="HONSHU">Gumma Pref. Mt, Hotaka</location>
|
||
(foot) ca
|
||
<quantity id="4C899B1EFF88814271CFA8877223F3F7" box="[697,782,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="219" unit="m" value="1300.0">
|
||
<elevation id="005CD1C8FF88814271CFA8877223F3F7" box="[697,782,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="219" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
<date id="FFCF103BFF8881427063A88772A6F3F7" box="[789,907,186,213]" pageId="14" pageNumber="219" value="2014-08-15">
|
||
<collectingDate id="EF8BE9D3FF8881427063A88772A6F3F7" box="[789,907,186,213]" pageId="14" pageNumber="219" value="2014-08-15">14-15.viii.</collectingDate>
|
||
</date>
|
||
[19]80’ (
|
||
<collectionCode id="ED60AE3EFF888142709CA8877569F3F6" box="[1002,1092,186,212]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="14" pageNumber="219">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6570FF88815C73B1A8E37415F1A7" lastPageId="16" lastPageNumber="221" pageId="14" pageNumber="219" type="description">
|
||
<paragraph id="8BCE36FBFF88814273B1A8E370E2F246" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="219">
|
||
<emphasis id="B905EAE9FF88814273B1A8E37058F3DA" bold="true" box="[199,373,222,248]" pageId="14" pageNumber="219">Redescription.</emphasis>
|
||
Measurements (n=15): HW: 0.72–0.87; HL: 0.47–0.63; OL: 0.15–0.21; LT: 0.12–0.14; AL (
|
||
<typeStatus id="54CA8859FF88814273E9A93F7020F23E" box="[159,269,258,284]" pageId="14" pageNumber="219" type="lectotype">lectotype</typeStatus>
|
||
): 2.96; PL: 0.67–0.78; PWmax: 0.90–1.03; PWmin: 0.75–0.87; ESL: 1.02–1.32; EW: 1.30–1.57; MTbL (
|
||
<typeStatus id="54CA8859FF88814273E9A91B7020F262" box="[159,269,294,320]" pageId="14" pageNumber="219" type="lectotype">lectotype</typeStatus>
|
||
): 1.05; MTrL (
|
||
<typeStatus id="54CA8859FF88814272C4A91B730DF262" box="[434,544,294,320]" pageId="14" pageNumber="219" type="lectotype">lectotype</typeStatus>
|
||
): 0.37 (MTrL 1–4: 0.17; MTrL 5: 0.20); AW: 1.35–1.68; AedL: 0.75–0.80; BL: 2.90–5.30 (
|
||
<typeStatus id="54CA8859FF8881427260A97770AEF246" box="[278,387,330,356]" pageId="14" pageNumber="219" type="lectotype">lectotype</typeStatus>
|
||
: 4.70).
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF88814273B1A9537422F1A6" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="219">
|
||
Habitus as in
|
||
<figureCitation id="134A2A7EFF888142721DA95370ECF2AA" box="[363,449,366,392]" captionStart="FIGURES 31–33" captionStartId="14.[151,264,1928,1952]" captionTargetBox="[156,1430,1069,1899]" captionTargetId="figure-379@14.[151,1435,1064,1904]" captionTargetPageId="14" captionText="FIGURES 31–33. Lectotype labels (Fig. 31–32) and habitus (Fig. 33) of Microedus subtilis: 31–32—lectotype, with type labels, 33—Japan: Honshu: Mt. Hotaka. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11045169" httpUri="https://zenodo.org/record/11045169/files/figure.png" pageId="14" pageNumber="219">Fig. 33</figureCitation>
|
||
. Body reddish-brown, usually with darker head and abdomen and paler elytra (some specimens with darker mediobasal portion of elytra;
|
||
<typeStatus id="54CA8859FF8881427194A9AF7264F28E" box="[738,841,402,428]" pageId="14" pageNumber="219" type="lectotype">lectotype</typeStatus>
|
||
and
|
||
<typeStatus id="54CA8859FF8881427008A9AF7538F28E" box="[894,1045,402,428]" pageId="14" pageNumber="219" type="paralectotype">paralectotype</typeStatus>
|
||
somewhat paler); antennae brown, sometimes with paler antennoimeres 1–2; mouthparts and legs yellow-brown to brown; apical maxillary palpomere and tarsi yellowish. Head with very dense isodiametric microreticulation, more transverse in middle of vertex, finer and sometimes indistinct in middle, and coarser on infraorbital portions, mediobasal portion of head with dense and fine isodiametric meshes; neck with dense and coarse isodiametric sculpture; pronotum with very dense and coarse isodiametric microsculpture, distinctly finer and sometimes indistinct in middle; scutellum with very fine transverse or isodiametric meshes; abdominal tergites with very dense and coarse isodiametric microreticulation.
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF88814273B1AAB3722EF086" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="219">Head 1.3–1.5 times as broad as long; medioapical depression slightly or strongly narrowing posteriad toward level of apical margins of eyes; temples 1.2–1.5 times as long as longitudinal length of eyes. Ocelli located distinctly below level of posterior margins of eyes, distance between ocelli 1.3–1.6 times as long as distance between ocellus and posterior margin of eyes. Punctation of head sparse and irregular, sparser and finer (sometimes indistinct) in middle and slightly larger, deeper and denser on infraorbital portions. Antennomeres 4–10 elongate; basal antennomere slightly more than three times as long as broad, 2 slightly narrower and distinctly shorter than basal antennomere, 3–5 distinctly longer than 2, 6–7 slightly longer and indistinctly broader than 5, 8–10 slightly shorter than 7, apical antennomere 1.4–1.7 times as long as 10.</paragraph>
|
||
<paragraph id="8BCE36FBFF88814273B1AB927281F732" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="219">Pronotum 1.3 times as broad as long, 1.1–1.2 times as broad as head, from widest anterior portion slightly or strongly narrowed posteriad, laterobasal margins in front of hind angles sometimes straight and parallel-sided. Punctation dense and fine, sparser and sometimes indistinct in middle.</paragraph>
|
||
<caption id="DF0E6673FF88814273E1AFB573D8F4E6" ID-DOI="http://doi.org/10.5281/zenodo.11045169" ID-Zenodo-Dep="11045169" httpUri="https://zenodo.org/record/11045169/files/figure.png" pageId="14" pageNumber="219" startId="14.[151,264,1928,1952]" targetBox="[156,1430,1069,1899]" targetPageId="14" targetType="figure">
|
||
<paragraph id="8BCE36FBFF88814273E1AFB573D8F4E6" blockId="14.[151,1437,1928,1989]" pageId="14" pageNumber="219">
|
||
<emphasis id="B905EAE9FF88814273E1AFB5707AF482" bold="true" box="[151,343,1928,1952]" pageId="14" pageNumber="219">FIGURES 31–33.</emphasis>
|
||
Lectotype labels (Fig. 31–32) and habitus (Fig. 33) of
|
||
<taxonomicName id="4C714D78FF88814270D3AFB47549F483" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[933,1124,1929,1953]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF88814270D3AFB47549F483" box="[933,1124,1929,1953]" italics="true" pageId="14" pageNumber="219">Microedus subtilis</emphasis>
|
||
</taxonomicName>
|
||
: 31–32—lectotype, with type labels, 33—Japan: Honshu: Mt. Hotaka. Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0E6673FF89814373E1AF097096F495" ID-DOI="http://doi.org/10.5281/zenodo.11045171" ID-Zenodo-Dep="11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="15" pageNumber="220" startId="15.[151,264,1844,1868]" targetBox="[159,1428,189,1812]" targetPageId="15" targetType="figure">
|
||
<paragraph id="8BCE36FBFF89814373E1AF097096F495" blockId="15.[151,1436,1844,1976]" pageId="15" pageNumber="220">
|
||
<emphasis id="B905EAE9FF89814373E1AF097078F46E" bold="true" box="[151,341,1844,1868]" pageId="15" pageNumber="220">FIGURES 34–42.</emphasis>
|
||
Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of
|
||
<taxonomicName id="4C714D78FF8981437647AF0971CDF452" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF8981437647AF0971CDF452" italics="true" pageId="15" pageNumber="220">Microedus subtilis</emphasis>
|
||
</taxonomicName>
|
||
: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCE36FBFF96815C73B1A8AA7540F3F7" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">Elytra 1.1–1.2 times as broad as long, 1.5–1.6 times as long as pronotum. Punctation dense and regular, distinctly larger and deeper than that on pronotum, finer in parascutellar portion and along suture.</paragraph>
|
||
<paragraph id="8BCE36FBFF96815C73B1A8E2751BF28F" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
Male. Apical margin of abdominal tergite VIII truncate (
|
||
<figureCitation id="134A2A7EFF96815C7020A8E27280F3DB" box="[854,941,223,249]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 37</figureCitation>
|
||
). Apical margin of abdominal sternite VIII widely concave (
|
||
<figureCitation id="134A2A7EFF96815C7222A93E7085F23F" box="[340,424,259,285]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 38</figureCitation>
|
||
). Genital segment as in
|
||
<figureCitation id="134A2A7EFF96815C71C4A93E7229F23F" box="[690,772,259,285]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 39</figureCitation>
|
||
. Aedeagus with moderately narrow basal portion, gradually narrowed toward long and narrow median lobe, slightly (
|
||
<figureCitation id="134A2A7EFF96815C7060A91A724AF263" box="[790,871,295,321]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 34</figureCitation>
|
||
) or strongly (
|
||
<figureCitation id="134A2A7EFF96815C7776A91A757CF263" box="[1024,1105,294,321]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 35</figureCitation>
|
||
) narrowed apicad; parameres narrow, distinctly exceeding apex of median lobe, sometimes sligthly broadened apically, with two short apical and two preapical setae; internal sac narrow and long, with two narrow elongate and sclerotized structures in middle (sometimes these structures invisible). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="134A2A7EFF96815C70A9A9AE751CF28F" box="[991,1073,403,429]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 36</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF96815C73B1A98A70A0F2D7" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
Female.Apical margins of abdominal tergite VIII (
|
||
<figureCitation id="134A2A7EFF96815C7183A98A726BF2F3" box="[757,838,439,465]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 40</figureCitation>
|
||
) and sternite VIII (
|
||
<figureCitation id="134A2A7EFF96815C7763A98A7548F2F3" box="[1045,1125,439,465]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 41</figureCitation>
|
||
) somewhat straight. Genital segment as in
|
||
<figureCitation id="134A2A7EFF96815C7241A9E670A4F2D7" box="[311,393,475,501]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Fig. 42</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF96815C73B1A9C37415F1A7" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
<emphasis id="B905EAE9FF96815C73B1A9C3709CF13B" bold="true" box="[199,433,510,537]" pageId="16" pageNumber="221">Comparative notes.</emphasis>
|
||
Based on the shape of the slightly flattened pronotum, the presence of coriaceous ground sculpture on head and pronotum, and the general shape of the aedeagus,
|
||
<taxonomicName id="4C714D78FF96815C70A0AA1E7563F11F" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[982,1102,547,573]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C70A0AA1E7563F11F" box="[982,1102,547,573]" italics="true" pageId="16" pageNumber="221">M. subtilis</emphasis>
|
||
</taxonomicName>
|
||
is similar to the Japanese
|
||
<taxonomicName id="4C714D78FF96815C7608AA1E70E0F143" authority="Watanabe, 1990" authorityName="Watanabe" authorityYear="1990" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="yamanakai">
|
||
<emphasis id="B905EAE9FF96815C7608AA1E703FF143" italics="true" pageId="16" pageNumber="221">M. yamanakai</emphasis>
|
||
<bibRefCitation id="EFE04B0AFF96815C7261AA7A70E0F143" author="Watanabe, Y." box="[279,461,583,609]" pageId="16" pageNumber="221" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
, known from Honshu (
|
||
<bibRefCitation id="EFE04B0AFF96815C71BFAA7A7256F143" author="Watanabe, Y." box="[713,891,583,609]" pageId="16" pageNumber="221" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
|
||
), from which it can be distinguished by the larger eyes, the finer punctation of head and pronotum, and the distinctly narrower median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6570FF96815C73B1AAB371D4F1D3" pageId="16" pageNumber="221" type="distribution">
|
||
<paragraph id="8BCE36FBFF96815C73B1AAB37459F18B" blockId="16.[151,1437,150,1365]" box="[199,1396,654,681]" pageId="16" pageNumber="221">
|
||
<emphasis id="B905EAE9FF96815C73B1AAB3704CF18A" bold="true" box="[199,353,654,680]" pageId="16" pageNumber="221">Distribution.</emphasis>
|
||
<taxonomicName id="4C714D78FF96815C7211AAB2731BF18B" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[359,566,655,681]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C7211AAB2731BF18B" box="[359,566,655,681]" italics="true" pageId="16" pageNumber="221">Microedus subtilis</emphasis>
|
||
</taxonomicName>
|
||
is known from
|
||
<collectingCountry id="F366766BFF96815C7191AAB2720AF18B" box="[743,807,655,681]" name="Japan" pageId="16" pageNumber="221">Japan</collectingCountry>
|
||
(Honshu) and
|
||
<collectingCountry id="F366766BFF96815C70B8AAB27534F18B" box="[974,1049,655,681]" name="Russia" pageId="16" pageNumber="221">Russia</collectingCountry>
|
||
(Iturup Island, Kurile islands).
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF96815C73B1AA8F71D4F1D3" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
<emphasis id="B905EAE9FF96815C73B1AA8F7066F1EE" bold="true" box="[199,331,690,716]" pageId="16" pageNumber="221">Bionomics.</emphasis>
|
||
Some specimens were collected under stones near streams and rivers. Detailed bionomical data are uknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6570FF96815C73B1AAC671C2F677" pageId="16" pageNumber="221" type="discussion">
|
||
<paragraph id="8BCE36FBFF96815C73B1AAC672BAF087" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
<emphasis id="B905EAE9FF96815C73B1AAC67016F037" bold="true" box="[199,315,763,789]" pageId="16" pageNumber="221">Remarks.</emphasis>
|
||
<taxonomicName id="4C714D78FF96815C7230AAC6731AF037" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[326,567,763,789]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C7230AAC6731AF037" box="[326,567,763,789]" italics="true" pageId="16" pageNumber="221">Anthophagus subtilis</emphasis>
|
||
</taxonomicName>
|
||
was originally descibed from “…Iwakisan…” based on “…two mutilated examples”. I studied
|
||
<specimenCount id="9D77FD72FF96815C7208AB227322F01B" box="[382,527,799,825]" count="2" pageId="16" pageNumber="221" type="generic" typeStatus="syntype">two syntypes</specimenCount>
|
||
(females) from BMNH, and one of them, with additional label ‘Type’, I designated here as the
|
||
<typeStatus id="54CA8859FF96815C726BAB7E70A9F07F" box="[285,388,835,861]" pageId="16" pageNumber="221" type="lectotype">lectotype</typeStatus>
|
||
in order to fix the name.
|
||
<bibRefCitation id="EFE04B0AFF96815C71D9AB7E72F5F07F" author="Nakane, T. & Sawada, K." box="[687,984,834,861]" pageId="16" pageNumber="221" pagination="179 - 186" refId="ref12719" refString="Nakane, T. & Sawada, K. (1956) On the genus Philydrodes Bernhauer in Japan with descriptions of a new subgenus and several new species. The Scientific Reports of the Saikyo University, Series A, 2, 179 - 186." type="journal article" year="1956">Nakane & Sawada (1956)</bibRefCitation>
|
||
regarded it as a species of
|
||
<taxonomicName id="4C714D78FF96815C7660AB7E702CF0A3" authorityName="Nakane & K. Sawada" authorityYear="1956" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="subGenus" subGenus="Lioplax">
|
||
<emphasis id="B905EAE9FF96815C7660AB7E74B1F07F" box="[1302,1436,835,861]" italics="true" pageId="16" pageNumber="221">Philydrodes</emphasis>
|
||
(
|
||
<emphasis id="B905EAE9FF96815C73D7AB5A71DAF0A3" box="[161,247,871,897]" italics="true" pageId="16" pageNumber="221">Lioplax</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFE04B0AFF96815C7278AB5A70E2F0A3" author="Watanabe, Y." box="[270,463,871,897]" pageId="16" pageNumber="221" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990)</bibRefCitation>
|
||
redescribed it and transferred it to the genus
|
||
<taxonomicName id="4C714D78FF96815C70A1AB5A75ADF0A3" authorityName="Nakane" authorityYear="1983" box="[983,1152,871,897]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EAE9FF96815C70A1AB5A75ADF0A3" box="[983,1152,871,897]" italics="true" pageId="16" pageNumber="221">Liophilydrodes</emphasis>
|
||
</taxonomicName>
|
||
, and provided additional material from northeastern Honshu and illustrations of the aedeagus.
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF96815C73B1AB9273FDF7E7" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
<taxonomicName id="4C714D78FF96815C73B1AB927366F0EB" authorityName="Nakane & K. Sawada" authorityYear="1956" box="[199,587,943,969]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="troglophila" subGenus="Lioplax">
|
||
<emphasis id="B905EAE9FF96815C73B1AB927060F0EB" box="[199,333,943,969]" italics="true" pageId="16" pageNumber="221">Philydrodes</emphasis>
|
||
(
|
||
<emphasis id="B905EAE9FF96815C7215AB927094F0EB" box="[355,441,943,969]" italics="true" pageId="16" pageNumber="221">Lioplax</emphasis>
|
||
)
|
||
<emphasis id="B905EAE9FF96815C72B8AB927366F0EB" box="[462,587,943,969]" italics="true" pageId="16" pageNumber="221">troglophila</emphasis>
|
||
</taxonomicName>
|
||
was originally described based on
|
||
<specimenCount id="9D77FD72FF96815C7083AB92758DF0EB" box="[1013,1184,943,969]" count="2" pageId="16" pageNumber="221" type="generic">two specimens</specimenCount>
|
||
from “Nippara cave, Okutama,
|
||
<collectingRegion id="49B5F819FF96815C727CABEE707AF0CF" box="[266,343,979,1005]" country="Japan" name="Tokyo" pageId="16" pageNumber="221">Tokyo</collectingRegion>
|
||
”.
|
||
<bibRefCitation id="EFE04B0AFF96815C721CABEE730AF0CF" author="Watanabe, Y." box="[362,551,979,1005]" pageId="16" pageNumber="221" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990)</bibRefCitation>
|
||
redescribed it, transferred it to the genus
|
||
<taxonomicName id="4C714D78FF96815C7098ABEE75BAF0CF" authorityName="Nakane" authorityYear="1983" box="[1006,1175,979,1005]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EAE9FF96815C7098ABEE75BAF0CF" box="[1006,1175,979,1005]" italics="true" pageId="16" pageNumber="221">Liophilydrodes</emphasis>
|
||
</taxonomicName>
|
||
, provided new material from Honshu and illustrations of the aedeagus. Based on the external morphology of the body, and the shape of the aedeagus, this species corresponds with other specimens of
|
||
<taxonomicName id="4C714D78FF96815C7035AC267296F717" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[835,955,1051,1077]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C7035AC267296F717" box="[835,955,1051,1077]" italics="true" pageId="16" pageNumber="221">M. subtilis</emphasis>
|
||
</taxonomicName>
|
||
. I could not find sufficient morphological differences between the
|
||
<typeStatus id="54CA8859FF96815C72D2AC0270F9F77B" box="[420,468,1087,1113]" pageId="16" pageNumber="221">type</typeStatus>
|
||
and additional specimens. The apex of the median lobe is variable and can be narrow or somewhat wide (
|
||
<figureCitation id="134A2A7EFF96815C7223AC5E70F9F75F" box="[341,468,1122,1149]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="16" pageNumber="221">Figs 34–35</figureCitation>
|
||
and Figs
|
||
<quantity id="4C899B1EFF96815C7134AC5E73EFF75F" box="[578,706,1123,1149]" metricMagnitude="4" metricUnit="m" metricValue="1.03483156" pageId="16" pageNumber="221" unit="in" value="407414.0">407, 414 in</quantity>
|
||
<bibRefCitation id="EFE04B0AFF96815C71BEAC5E72B9F75F" author="Watanabe, Y." box="[712,916,1123,1149]" pageId="16" pageNumber="221" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990))</bibRefCitation>
|
||
. It should be noted that the width of the apical portions of the median lobe and the parameres is variable in many species of
|
||
<taxonomicName id="4C714D78FF96815C777CACBA7589F783" box="[1034,1188,1159,1185]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="tribe" tribe="Anthophagini">Anthophagini</taxonomicName>
|
||
(e.g.
|
||
<bibRefCitation id="EFE04B0AFF96815C7790ACBA74BDF783" author="Shavrin, A. V." box="[1254,1424,1159,1185]" pageId="16" pageNumber="221" pagination="451 - 496" refId="ref13084" refString="Shavrin, A. V. (2022 b) The bodemeyeri and the convexicollis groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data fro some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 5213 (5), 451 - 496. https: // doi. org / 10.11646 / zootaxa. 5213.5.1" type="journal article" year="2022">Shavrin 2022b</bibRefCitation>
|
||
). Thus, this species is synonymized with
|
||
<taxonomicName id="4C714D78FF96815C7125AC9673E7F7E7" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[595,714,1195,1221]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="221" phylum="Arthropoda" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C7125AC9673E7F7E7" box="[595,714,1195,1221]" italics="true" pageId="16" pageNumber="221">M. subtilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE36FBFF96815C73B1ACF271C2F677" blockId="16.[151,1437,150,1365]" pageId="16" pageNumber="221">
|
||
<emphasis id="B905EAE9FF96815C73B1ACF27375F7CA" box="[199,600,1231,1257]" italics="true" pageId="16" pageNumber="221">
|
||
<taxonomicName id="4C714D78FF96815C73B1ACF270E6F7CB" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[199,459,1231,1257]" genus="Liophiydrodes" pageId="16" pageNumber="221" rank="species" species="subtilis">Liophiydrodes subtilis</taxonomicName>
|
||
iturupensis
|
||
</emphasis>
|
||
was originally described based on
|
||
<specimenCount id="9D77FD72FF96815C7754ACF27585F7CB" box="[1058,1192,1231,1257]" count="1" pageId="16" pageNumber="221" type="female">one female</specimenCount>
|
||
from Iturup Island (“Konservnaya bay”), Kurile islands. Based on the original description, features of the body of this taxon correspond with morphological features of specimens of
|
||
<taxonomicName id="4C714D78FF96815C71D2AD2A7233F613" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[676,798,1303,1329]" genus="Liophiydrodes" pageId="16" pageNumber="221" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C71D2AD2A7233F613" box="[676,798,1303,1329]" italics="true" pageId="16" pageNumber="221">M. subtilis</emphasis>
|
||
</taxonomicName>
|
||
. Thus,
|
||
<taxonomicName id="4C714D78FF96815C7001AD2A7558F612" authorityName="Lafer" authorityYear="2004" box="[887,1141,1303,1329]" genus="Liophiydrodes" pageId="16" pageNumber="221" rank="subSpecies" species="subtulis" subSpecies="iturupensis">
|
||
<emphasis id="B905EAE9FF96815C7001AD2A7558F612" box="[887,1141,1303,1329]" italics="true" pageId="16" pageNumber="221">L. subtulis iturupensis</emphasis>
|
||
</taxonomicName>
|
||
is synonymyzed with
|
||
<taxonomicName id="4C714D78FF96815C7608AD2A71C5F677" baseAuthorityName="Sharp" baseAuthorityYear="1889" genus="Liophiydrodes" pageId="16" pageNumber="221" rank="species" species="subtilis">
|
||
<emphasis id="B905EAE9FF96815C7608AD2A71C5F677" italics="true" pageId="16" pageNumber="221">M. subtilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |