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Genus 
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<bibRefCitation id="EFE04B0AFF84814E7200A8E27301F3DB" author="LeConte, J. L." box="[374,556,223,249]" pageId="2" pageNumber="207" pagination="265 - 276" refId="ref12303" refString="LeConte, J. L. (1874) [New species]. In: Austin, E. P. (Ed.), Catalogue of the Coleoptera of Mt. Washington, N. H. by E. P. Austin; with descriptions of new species, by J. L. LeConte, M. D. Proceedings of the Boston Society of Natural History, 16, pp. 265 - 276." type="journal article" year="1874">LeConte, 1874</bibRefCitation>
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(
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species: 
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<emphasis id="B905EAE9FF84814E7241A93970F3F23F" box="[311,478,260,285]" italics="true" pageId="2" pageNumber="207">M. austinianus</emphasis>
<bibRefCitation id="EFE04B0AFF84814E7293A93E73A3F23F" author="LeConte, J. L." box="[485,654,259,285]" pageId="2" pageNumber="207" pagination="265 - 276" refId="ref12303" refString="LeConte, J. L. (1874) [New species]. In: Austin, E. P. (Ed.), Catalogue of the Coleoptera of Mt. Washington, N. H. by E. P. Austin; with descriptions of new species, by J. L. LeConte, M. D. Proceedings of the Boston Society of Natural History, 16, pp. 265 - 276." type="journal article" year="1874">LeConte, 1874</bibRefCitation>
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, fixed by monotypy)
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(
<figureCitation id="134A2A7EFF84814E73E9A91A703FF263" box="[159,274,295,321]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–13" captionStart-2="FIGURES 14–17" captionStart-3="FIGURES 18–23" captionStart-4="FIGURES 24–29" captionStartId-0="3.[151,264,1942,1966]" captionStartId-1="7.[151,264,1902,1926]" captionStartId-2="8.[151,264,1176,1200]" captionStartId-3="11.[151,264,1943,1967]" captionStartId-4="12.[151,264,1172,1196]" captionTargetBox-0="[156,1431,347,1909]" captionTargetBox-1="[159,1427,352,1871]" captionTargetBox-2="[159,1428,191,1144]" captionTargetBox-3="[178,1401,196,1911]" captionTargetBox-4="[159,1429,189,1144]" captionTargetId-0="figure-92@3.[151,1435,342,1915]" captionTargetId-1="figure-100@7.[151,1436,345,1879]" captionTargetId-2="figure-18@8.[151,1436,181,1152]" captionTargetId-3="figure-16@11.[170,1417,181,1919]" captionTargetId-4="figure-18@12.[151,1436,181,1149]" captionTargetPageId-0="3" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURES 1–5. Habitus of Microedus: 1—M. austinianus (USA: New Hampshire), 2—M. brevitarsis (holotype), 3—M. kastcheevi (paratype, Kazakhstan: Dzhungar Mts.), 4—A. schilenkovi (Russia: Altai, Korgon Mts.), 5—M. sp. (Kashmir). Scale bar: 1.0 mm." captionText-1="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." captionText-2="FIGURES 14–17. Aedeagus of Microedus austinianus (Figs 14–15: Canada, Quebec, Riviere-a-Claude) and M. brevitarsis (Figs. 16–17: holotype): 14, 16—parameral view, 15, 17—lateral view. Scale bar: 0.1 mm." captionText-3="FIGURES 18–23. Aedeagus of Microedus schilenkovi (Figs 18–19: Russia, Altai, Aygulakskiy Mts.; Fig. 20: Russia, Altai, Teletskoye Lake; Fig. 21: Kazakhstan, Tauchilik River) and A. kastcheevi (Figs. 22–23: paratype, Dzhungar Mts.): 18, 20–22— parameral view, 19, 23—lateral view. Scale bar: 0.1 mm." captionText-4="FIGURES 24–29. Abdominal segments of Microedus schilenkovi: 24—male tergite VIII, 25—male sternite VIII, 26—male genital segment, 27—female tergite VIII, 28—female sternite VIII, 29—female genital segment. Scale bars 0.1 m" figureDoi-0="http://doi.org/10.5281/zenodo.11045157" figureDoi-1="http://doi.org/10.5281/zenodo.11045159" figureDoi-2="http://doi.org/10.5281/zenodo.11045161" figureDoi-3="http://doi.org/10.5281/zenodo.11045163" figureDoi-4="http://doi.org/10.5281/zenodo.11045165" httpUri-0="https://zenodo.org/record/11045157/files/figure.png" httpUri-1="https://zenodo.org/record/11045159/files/figure.png" httpUri-2="https://zenodo.org/record/11045161/files/figure.png" httpUri-3="https://zenodo.org/record/11045163/files/figure.png" httpUri-4="https://zenodo.org/record/11045165/files/figure.png" pageId="2" pageNumber="207">Figs 1–29</figureCitation>
, 
<figureCitation id="134A2A7EFF84814E7268A91A7048F263" box="[286,357,295,321]" captionStart-0="FIGURES 31–33" captionStart-1="FIGURES 34–42" captionStartId-0="14.[151,264,1928,1952]" captionStartId-1="15.[151,264,1844,1868]" captionTargetBox-0="[156,1430,1069,1899]" captionTargetBox-1="[159,1428,189,1812]" captionTargetId-0="figure-379@14.[151,1435,1064,1904]" captionTargetId-1="figure-16@15.[151,1436,181,1820]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 31–33. Lectotype labels (Fig. 31–32) and habitus (Fig. 33) of Microedus subtilis: 31–32—lectotype, with type labels, 33—Japan: Honshu: Mt. Hotaka. Scale bar: 1.0 mm." captionText-1="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11045169" figureDoi-1="http://doi.org/10.5281/zenodo.11045171" httpUri-0="https://zenodo.org/record/11045169/files/figure.png" httpUri-1="https://zenodo.org/record/11045171/files/figure.png" pageId="2" pageNumber="207">31–42</figureCitation>
)
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; 
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[= 
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<bibRefCitation id="EFE04B0AFF84814E77FDA95174BAF2A1" author="Moore, L. &amp; Legner, E. F." box="[1163,1431,363,388]" pageId="2" pageNumber="207" pagination="75 - 77" refId="ref12517" refString="Moore, L. &amp; Legner, E. F. (1972) A new species of Microedus from the Sierra Nevada Mountains (Coleoptera: Staphylinidae). Coleopterists Bulletin, 26 (2), 75 - 77." type="journal article" year="1972">Moore &amp; Legner 1972: 75</bibRefCitation>
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<bibRefCitation id="EFE04B0AFF84814E73B1A9B77002F280" author="Moore, L. &amp; Legner, E. F." box="[199,303,394,418]" pageId="2" pageNumber="207" refId="ref12559" refString="Moore, L. &amp; Legner, E. F. (1975) A catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015. Division of Agricultural Sciences, University of California, Los Angeles, California, 514 pp." type="book" year="1975">1975: 184</bibRefCitation>
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: 435
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<bibRefCitation id="EFE04B0AFF84814E71D5A9B672B5F280" author="Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[675,920,394,418]" pageId="2" pageNumber="207" pagination="1802" refId="ref12766" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="book chapter" year="2000">
Newton 
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. 2000: 340
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<emphasis id="B905EAE9FF84814E73E1A99471C9F2E3" box="[151,228,425,449]" italics="true" pageId="2" pageNumber="207">Lioplax</emphasis>
Nakane &amp; K. Sawada, 1956: 183
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<taxonomicName id="4C714D78FF84814E73E1A9F570DBF2C2" ID-CoL="8VYYB" authority="Nakane, 1983: 148" authorityName="Nakane" authorityPageNumber="148" authorityYear="1983" box="[151,502,456,480]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="genus" status="syn. n.">
<emphasis id="B905EAE9FF84814E73E1A9F57002F2C2" box="[151,303,456,480]" italics="true" pageId="2" pageNumber="207">Liophilydrodes</emphasis>
<bibRefCitation id="EFE04B0AFF84814E7240A9F570DBF2C2" author="Nakane, T." box="[310,502,456,480]" pageId="2" pageNumber="207" pagination="148" refId="ref12681" refString="Nakane, T. (1983) Liophilydrodes. In: Nakane, T., New name for genus Lioplax Nakane et K. Sawada. Fragmenta Coleopterologica, 35 / 37, pp. 148." type="journal article" year="1983">Nakane, 1983: 148</bibRefCitation>
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<emphasis id="B905EAE9FF84814E728AA9F5736FF2C2" bold="true" box="[508,578,456,480]" pageId="2" pageNumber="207">
<taxonomicNameLabel id="A2365792FF84814E728AA9F5736FF2C2" box="[508,578,456,480]" pageId="2" pageNumber="207">syn. n.</taxonomicNameLabel>
</emphasis>
; 
<treatmentCitation id="0AD010EAFF84814E7139A9F57232F2FD" author="Watanabe, Y." box="[591,799,455,480]" page="295" pageId="2" pageNumber="207" year="1990">
<bibRefCitation id="EFE04B0AFF84814E7139A9F57232F2FD" author="Watanabe, Y." box="[591,799,455,480]" pageId="2" pageNumber="207" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe 1990: 295</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="8BCE36FBFF84814E73E1A9DB7313F2DC" blockId="2.[151,1436,363,541]" box="[151,574,486,510]" pageId="2" pageNumber="207">
<treatmentCitationGroup id="AB6111D5FF84814E73E1A9DB7313F2DC" box="[151,574,486,510]" pageId="2" pageNumber="207">
<treatmentCitation id="0AD010EAFF84814E73E1A9DB70DFF2DC" author="Zerche, L." box="[151,498,486,510]" page="107" pageId="2" pageNumber="207" year="1992">
<taxonomicName id="4C714D78FF84814E73E1A9DB70DFF2DC" ID-CoL="8XJH2" authority="Zerche, 1992: 107" authorityName="Zerche" authorityPageNumber="107" authorityYear="1992" box="[151,498,486,510]" class="Insecta" family="Staphylinidae" genus="Altaiodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="genus" status="syn. n.">
<emphasis id="B905EAE9FF84814E73E1A9DB701FF2DC" box="[151,306,486,510]" italics="true" pageId="2" pageNumber="207">Altaiodromicus</emphasis>
<bibRefCitation id="EFE04B0AFF84814E724FA9DB70DFF2DC" author="Zerche, L." box="[313,498,486,510]" pageId="2" pageNumber="207" pagination="105 - 142" refId="ref13769" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992: 107</bibRefCitation>
</taxonomicName>
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<emphasis id="B905EAE9FF84814E728EA9DB7313F2DC" bold="true" box="[504,574,486,510]" pageId="2" pageNumber="207">
<taxonomicNameLabel id="A2365792FF84814E728EA9DB7313F2DC" box="[504,574,486,510]" pageId="2" pageNumber="207">syn. n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCE36FBFF84814E73E1AA38731EF13F" blockId="2.[151,1436,363,541]" box="[151,563,517,541]" pageId="2" pageNumber="207">
For other references see 
<bibRefCitation id="EFE04B0AFF84814E72E6AA387302F13F" author="Herman, L. H." box="[400,559,517,541]" pageId="2" pageNumber="207" pagination="1 - 7" refId="ref12189" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman (2001)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B6570FF84814873E1AA767554F1EF" lastPageId="4" lastPageNumber="209" pageId="2" pageNumber="207" type="description">
<paragraph id="8BCE36FBFF84814E73E1AA767038F03B" blockId="2.[151,1437,587,1982]" pageId="2" pageNumber="207">
<emphasis id="B905EAE9FF84814E73E1AA767068F147" bold="true" box="[151,325,587,613]" pageId="2" pageNumber="207">Redescription.</emphasis>
Head, pronotum and abdomen slightly convex; elytra flattened, slightly or strongly broadened posteriad. Body shiny, with dense coriaceous microsculpture on head and pronotum or without it (
<taxonomicName id="4C714D78FF84814E7795AA4D74A6F1AB" authorityName="Shavrin" authorityYear="2024" baseAuthorityName="Nakane &amp; K. Sawada" baseAuthorityYear="1956" box="[1251,1419,623,649]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="B905EAE9FF84814E7795AA4D74A6F1AB" box="[1251,1419,623,649]" italics="true" pageId="2" pageNumber="207">M. puncticollis</emphasis>
</taxonomicName>
); sometimes elytra subrugosely sculptured (
<taxonomicName id="4C714D78FF84814E71F7AAA973D7F18F" authorityName="Hatch" authorityYear="1957" box="[641,762,660,685]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="rogersi">
<emphasis id="B905EAE9FF84814E71F7AAA973D7F18F" box="[641,762,660,685]" italics="true" pageId="2" pageNumber="207">M. rogersi</emphasis>
</taxonomicName>
); body length varying from 
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. Coloration brown to reddish-brown or black. Head and pronotum with fine and moderately dense punctation; punctation of elytra distinctly denser, larger and deeper than that on pronotum; abdomen with distinct, regular, dense and fine punctation.
</paragraph>
<paragraph id="8BCE36FBFF84814E73B1AB1E70CBF6E7" blockId="2.[151,1437,587,1982]" pageId="2" pageNumber="207">
Head subtrapezoidal, with postocular parts usually widened and relatively long, with distinctly elevated part between bases of antennae and infraorbital portions; supra-antennal protuberances narrow, distinctly elevated, diagonally stretching from base of antennae to about level of apical margins of eyes; medioapical depression deep and wide, slightly or strongly narrowing posteriad toward level of anterior margins of eyes; interocellar depression shallow or well-distinct and moderately deep; anteocellar furrows (grooves in front of ocelli) narrow or very deep and moderately wide, diagonally stretching anteriad toward level of anterior third of eyes. Eyes relatively small, convex. Ocelli convex, small, but well visible. Labrum transverse, with slightly and widely concave apical margin and slightly elongate basolateral projections, with several very long setae in latero-apical and middle portions (
<figureCitation id="134A2A7EFF84814E761CAC2271E9F77F" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="2" pageNumber="207">Figs 6–7</figureCitation>
, and Fig. 
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<bibRefCitation id="EFE04B0AFF84814E720FAC7E7369F77F" author="Watanabe, Y." box="[377,580,1091,1117]" pageId="2" pageNumber="207" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990))</bibRefCitation>
. Mandibles with wide basal part and elongate, strongly curved apical portion; inner margin of each mandible with two elongate teeth (
<figureCitation id="134A2A7EFF84814E7051AC5A72A2F7A3" box="[807,911,1127,1153]" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="2" pageNumber="207">Figs 8–9</figureCitation>
, and Fig. 
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<bibRefCitation id="EFE04B0AFF84814E772AAC5A7406F7A3" author="Watanabe, Y." box="[1116,1323,1127,1153]" pageId="2" pageNumber="207" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990))</bibRefCitation>
. Maxillae relatively wide, with elongate galea and maxillary palpomeres; apical maxillary palpomere somewhat elongate, narrow, distinctly more than twice narrower than markedly widened preceding segment; lacinia moderately short, with several elongate preapical teeth (
<figureCitation id="134A2A7EFF84814E7139ACEE73F9F7CF" box="[591,724,1235,1261]" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="2" pageNumber="207">Figs 10–11</figureCitation>
). Labium narrow; labial palpomere 2 about as long as broad, preapical palpomere distinctly longer than broad, apical palpomere narrow, distinctly longer than preceding segment (
<figureCitation id="134A2A7EFF84814E73E9AD267031F617" box="[159,284,1307,1333]" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="2" pageNumber="207">Figs 12–13</figureCitation>
and Fig. 
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<bibRefCitation id="EFE04B0AFF84814E72B9AD2673B7F617" author="Watanabe, Y." box="[463,666,1307,1333]" pageId="2" pageNumber="207" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990))</bibRefCitation>
; labial palpomere 2 slightly elongate (
<figureCitation id="134A2A7EFF84814E7736AD2675BCF617" box="[1088,1169,1307,1333]" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="2" pageNumber="207">Fig. 12</figureCitation>
) or transverse (
<figureCitation id="134A2A7EFF84814E764AAD2674A2F617" box="[1340,1423,1307,1333]" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="2" pageNumber="207">Fig. 13</figureCitation>
). 
<collectingRegion id="49B5F819FF84814E73E1AD7D71DBF678" box="[151,246,1344,1370]" country="Myanmar" name="Chin" pageId="2" pageNumber="207">Mentum</collectingRegion>
narrow, transverse, as in Fig. 
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<bibRefCitation id="EFE04B0AFF84814E71EDAD027273F67B" author="Watanabe, Y." box="[667,862,1343,1369]" pageId="2" pageNumber="207" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990)</bibRefCitation>
. Gular sutures narrowly (see Fig. 
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<bibRefCitation id="EFE04B0AFF84814E7659AD0271C6F65F" author="Watanabe, Y." pageId="2" pageNumber="207" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990))</bibRefCitation>
or relatively widely separated from each other at level of posterior margins of eyes, strongly divergent posteriad. Antennae long, exceeding middle or apical third length of elytra when reclined, with antennomeres 4–10 slightly or distinctly elongate.
</paragraph>
<paragraph id="8BCE36FBFF84814E73B1ADED7340F5C4" blockId="2.[151,1437,587,1982]" pageId="2" pageNumber="207">
Pronotum convex, distinctly transverse, widest in anterior portion, not or distinctly protruding anteriad (
<taxonomicName id="4C714D78FF84814E7608ADED71CBF52F" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="fenderi">
<emphasis id="B905EAE9FF84814E7608ADED71CBF52F" italics="true" pageId="2" pageNumber="207">M. fenderi</emphasis>
</taxonomicName>
), gradually or strongly narrowed posteriad, narrowly or widely bordered laterally and sometimes distinctly explanate and reflexed (
<taxonomicName id="4C714D78FF84814E72D1AE257336F513" authorityName="Hatch" authorityYear="1957" box="[423,539,1559,1585]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="fenderi">
<emphasis id="B905EAE9FF84814E72D1AE257336F513" box="[423,539,1559,1585]" italics="true" pageId="2" pageNumber="207">M. fenderi</emphasis>
</taxonomicName>
); mediobasal portion sometimes with shallow transverse impression. Prothorax short, strongly transverse, distinctly convex in middle, with long and acute intercoxal process reaching about middle of front coxae. Mesoventrite moderately short, strongly convex, with acute elongate process, reaching apex of relatively long and narrow intercoxal process extending to about middle of mesocoxae. Scutellum large, triangular, without punctation. Metaventrite transverse, convex, with very deep intercoxal cavities and angular intercoxal process extending to mesosternal process.
</paragraph>
<paragraph id="8BCE36FBFF84814E73B1AED2703FF473" blockId="2.[151,1437,587,1982]" pageId="2" pageNumber="207">
Elytra wide, slightly or strongly broadened posteriad, with narrow or wide and distinctly reflexed (
<taxonomicName id="4C714D78FF84814E7668AECD74BEF42B" authorityName="Hatch" authorityYear="1957" box="[1310,1427,1775,1801]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="fenderi">
<emphasis id="B905EAE9FF84814E7668AECD74BEF42B" box="[1310,1427,1775,1801]" italics="true" pageId="2" pageNumber="207">M. fenderi</emphasis>
</taxonomicName>
) lateral margins, elongate or moderately short, reaching apical margin of abdominal tergite III to V. Hind wings fully developed.
</paragraph>
<paragraph id="8BCE36FBFF84814E73B1AF6172EFF49C" blockId="2.[151,1437,587,1982]" pageId="2" pageNumber="207">Legs relatively slender, moderately long, with transverse procoxae and metacoxae, and large suboval mesocoxae; protarsi of males with distinctly dilated protartsomeres 1–4; metatarsus more than twice shorter than metatibiae; apical antennomere slightly shorter than preceding four metatarsomeres.</paragraph>
<paragraph id="8BCE36FBFF85814F73B1A8AA75ADF3F7" blockId="3.[151,1436,151,321]" pageId="3" pageNumber="208">Abdomen slightly or distinctly broader than elytra, with two oval or transverse tomentose spots in the middle of abdominal tergite IV; apical margin of abdominal tergite VII with narrow palisade fringe.</paragraph>
<paragraph id="8BCE36FBFF85814F73B1A8E2754CF263" blockId="3.[151,1436,151,321]" pageId="3" pageNumber="208">
Aedeagus elongate or moderately short (
<taxonomicName id="4C714D78FF85814F71DAA8E27260F3DB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[684,845,223,249]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="kastcheevi" status="sp. n.">
<emphasis id="B905EAE9FF85814F71DAA8E27260F3DB" box="[684,845,223,249]" italics="true" pageId="3" pageNumber="208">M. kastcheevi</emphasis>
</taxonomicName>
<emphasis id="B905EAE9FF85814F702FA8E272B2F3DB" bold="true" box="[857,927,223,249]" pageId="3" pageNumber="208">
<taxonomicNameLabel id="A2365792FF85814F702FA8E272B2F3DB" box="[857,927,223,249]" pageId="3" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C714D78FF85814F70C4A8E27571F3DB" authorityName="Zerche" authorityYear="1992" box="[946,1116,223,249]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="schilenkovi">
<emphasis id="B905EAE9FF85814F70C4A8E27571F3DB" box="[946,1116,223,249]" italics="true" pageId="3" pageNumber="208">M. schilenkovi</emphasis>
</taxonomicName>
), with small or very large phallobase, gradually narrowed apicad; parameres narrow, reaching or exceeding apex of median lobe; internal sac narrow, without or with elongate and sclerotized structures, sometimes with flagellum.
</paragraph>
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<paragraph id="8BCE36FBFF85814F73E1AFAB7037F4D7" blockId="3.[151,1437,1942,2038]" pageId="3" pageNumber="208">
<emphasis id="B905EAE9FF85814F73E1AFAB7012F48C" bold="true" box="[151,319,1942,1966]" pageId="3" pageNumber="208">FIGURES 1–5.</emphasis>
Habitus of 
<taxonomicName id="4C714D78FF85814F72B2AFAB7302F48C" authorityName="LeConte" authorityYear="1874" box="[452,559,1942,1966]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF85814F72B2AFAB7302F48C" box="[452,559,1942,1966]" italics="true" pageId="3" pageNumber="208">Microedus</emphasis>
</taxonomicName>
: 1— 
<taxonomicName id="4C714D78FF85814F7111AFAA722FF48C" authorityName="LeConte" authorityYear="1874" box="[615,770,1943,1966]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="austinianus">
<emphasis id="B905EAE9FF85814F7111AFAA722FF48C" box="[615,770,1943,1966]" italics="true" pageId="3" pageNumber="208">M. austinianus</emphasis>
</taxonomicName>
(USA: New Hampshire), 2— 
<taxonomicName id="4C714D78FF85814F7735AFAA75F8F48C" authorityName="Champion" authorityYear="1925" box="[1091,1237,1942,1966]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF85814F7735AFAA75F8F48C" box="[1091,1237,1942,1966]" italics="true" pageId="3" pageNumber="208">M. brevitarsis</emphasis>
</taxonomicName>
(holotype), 3— 
<taxonomicName id="4C714D78FF85814F76F7AFAA702CF4F0" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="kastcheevi">
<emphasis id="B905EAE9FF85814F76F7AFAA702CF4F0" italics="true" pageId="3" pageNumber="208">M. kastcheevi</emphasis>
</taxonomicName>
(paratype, Kazakhstan: Dzhungar Mts.), 4— 
<taxonomicName id="4C714D78FF85814F71BEAF877278F4F0" authorityName="Zerche" authorityYear="1992" box="[712,853,1978,2002]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="schilenkovi">
<emphasis id="B905EAE9FF85814F71BEAF877278F4F0" box="[712,853,1978,2002]" italics="true" pageId="3" pageNumber="208">A. schilenkovi</emphasis>
</taxonomicName>
(Russia: Altai, Korgon Mts.), 5— 
<emphasis id="B905EAE9FF85814F77DBAF8675E4F4F0" box="[1197,1225,1979,2002]" italics="true" pageId="3" pageNumber="208">M.</emphasis>
sp. (Kashmir). Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BCE36FBFF82814873B1A8AA7304F3F7" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
Female genital segment with narrow and very long gonocoxites, with moderately short and narrow styli, each with long apical setae (
<figureCitation id="134A2A7EFF82814872EFA88670DDF3F7" box="[409,496,187,213]" captionStart="FIGURES 24–29" captionStartId="12.[151,264,1172,1196]" captionTargetBox="[159,1429,189,1144]" captionTargetId="figure-18@12.[151,1436,181,1149]" captionTargetPageId="12" captionText="FIGURES 24–29. Abdominal segments of Microedus schilenkovi: 24—male tergite VIII, 25—male sternite VIII, 26—male genital segment, 27—female tergite VIII, 28—female sternite VIII, 29—female genital segment. Scale bars 0.1 m" figureDoi="http://doi.org/10.5281/zenodo.11045165" httpUri="https://zenodo.org/record/11045165/files/figure.png" pageId="4" pageNumber="209">Figs 29</figureCitation>
, 
<figureCitation id="134A2A7EFF828148728AA8867331F3F7" box="[508,540,187,213]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="4" pageNumber="209">42</figureCitation>
).
</paragraph>
<paragraph id="8BCE36FBFF82814873B1A8E270DBF3DB" blockId="4.[151,1437,151,1977]" box="[199,502,223,249]" pageId="4" pageNumber="209">Immature stages unknown.</paragraph>
<paragraph id="8BCE36FBFF82814873B1A93F7554F1EF" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
<emphasis id="B905EAE9FF82814873B1A93F7083F23F" bold="true" box="[199,430,258,285]" pageId="4" pageNumber="209">Comparative notes.</emphasis>
Based on the general shape of the body and the aedeagus, and the presence of an interocellar depression, the genus 
<taxonomicName id="4C714D78FF82814872FBA91A7329F263" authorityName="LeConte" authorityYear="1874" box="[397,516,295,321]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814872FBA91A7329F263" box="[397,516,295,321]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
is related to genera of the 
<taxonomicName id="4C714D78FF828148705DA91A7288F262" authorityName="Mulsant &amp; Rey" authorityYear="1880" box="[811,933,295,320]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF828148705DA91A7288F262" box="[811,933,295,320]" italics="true" pageId="4" pageNumber="209">Hygrogeus</emphasis>
</taxonomicName>
complex, distributed in the Holarctic Region (
<bibRefCitation id="EFE04B0AFF82814873E9A976701DF247" author="Zerche, L." box="[159,304,331,357]" pageId="4" pageNumber="209" pagination="105 - 142" refId="ref13769" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992</bibRefCitation>
, 
<bibRefCitation id="EFE04B0AFF828148724DA97670FBF247" author="Shavrin, A. V." box="[315,470,331,357]" pageId="4" pageNumber="209" pagination="169 - 186" refId="ref12933" refString="Shavrin, A. V. (2017) Five new species of the genus Philydrodes Bernhauer, 1929 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 4231 (2), 169 - 186. https: // doi. org / 10.11646 / zootaxa. 4231.2.2" type="journal article" year="2017">Shavrin 2017</bibRefCitation>
). Regarding the general shape of the narrow apical segment of the maxillary palpi, the ground sculpture of the forebody, and the shape of the transverse pronotum, 
<taxonomicName id="4C714D78FF828148709EA9527572F2AB" authorityName="LeConte" authorityYear="1874" box="[1000,1119,367,393]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF828148709EA9527572F2AB" box="[1000,1119,367,393]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C714D78FF8281487799A952706BF28F" authority="Tronquet, 1981" authorityName="Tronquet" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487799A95274B0F2AB" box="[1263,1437,367,393]" italics="true" pageId="4" pageNumber="209">Hygrodromicus</emphasis>
Tronquet, 1981
</taxonomicName>
and 
<taxonomicName id="4C714D78FF82814872F4A9AE73FCF28F" authority="Bernhauer, 1929" authorityName="Bernhauer" authorityYear="1929" box="[386,721,403,429]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814872F4A9AE7325F28F" box="[386,520,403,429]" italics="true" pageId="4" pageNumber="209">Philydrodes</emphasis>
Bernhauer, 1929
</taxonomicName>
. It can be distinguished from 
<taxonomicName id="4C714D78FF8281487758A9AE75F1F28F" authorityName="Tronquet" authorityYear="1981" box="[1070,1244,403,429]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487758A9AE75F1F28F" box="[1070,1244,403,429]" italics="true" pageId="4" pageNumber="209">Hygrodromicus</emphasis>
</taxonomicName>
by the narrower apical and somewhat broader preapical segments of the maxillary palpi, longer temples, and the general shape of the aedeagus (basal portion of aedeagus of 
<taxonomicName id="4C714D78FF82814871C3A9E6724EF2D7" authorityName="Tronquet" authorityYear="1981" box="[693,867,475,501]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814871C3A9E6724EF2D7" box="[693,867,475,501]" italics="true" pageId="4" pageNumber="209">Hygrodromicus</emphasis>
</taxonomicName>
usually small and rotated inside the abdomen in lateral position (90°), as in 
<taxonomicName id="4C714D78FF82814872BAA9C2737FF13B" authorityName="Bernhauer" authorityYear="1929" box="[460,594,511,537]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814872BAA9C2737FF13B" box="[460,594,511,537]" italics="true" pageId="4" pageNumber="209">Philydrodes</emphasis>
</taxonomicName>
too). It can be distinguished from 
<taxonomicName id="4C714D78FF82814870ACA9C2754DF13B" authorityName="Bernhauer" authorityYear="1929" box="[986,1120,511,537]" class="Insecta" family="Staphylinidae" genus="Philydrodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814870ACA9C2754DF13B" box="[986,1120,511,537]" italics="true" pageId="4" pageNumber="209">Philydrodes</emphasis>
</taxonomicName>
by the shape of the labrum, broader mandibles, the shorter and broader preapical and longer apical segments of the maxillary palpi, shorter temples, and the significantly broader pronotum. From all genera listed above and other genera of the 
<taxonomicName id="4C714D78FF8281487654AA7A74B1F142" authorityName="Mulsant &amp; Rey" authorityYear="1880" box="[1314,1436,583,608]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487654AA7A74B1F142" box="[1314,1436,583,608]" italics="true" pageId="4" pageNumber="209">Hygrogeus</emphasis>
</taxonomicName>
complex it can be distinguished by the external and internal morphology of the aedeagus. A more detailed analysis of relations of these genera within 
<taxonomicName id="4C714D78FF8281487156AAB273B7F18A" authorityName="Mulsant &amp; Rey" authorityYear="1880" box="[544,666,655,680]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487156AAB273B7F18A" box="[544,666,655,680]" italics="true" pageId="4" pageNumber="209">Hygrogeus</emphasis>
</taxonomicName>
complex will be published in detail in a forthcoming revision of the genus 
<taxonomicName id="4C714D78FF8281487396AA8E7320F1EF" authority="Luze, 1903" authorityName="Luze" authorityYear="1903" box="[224,525,691,717]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487396AA8E70A4F1EF" box="[224,393,691,717]" italics="true" pageId="4" pageNumber="209">Trichodromeus</emphasis>
Luze, 1903
</taxonomicName>
and related genera of the Palaearctic Region (in prep.).
</paragraph>
</subSubSection>
<subSubSection id="C36B6570FF82814873B1AAEB7590F0A3" pageId="4" pageNumber="209" type="distribution">
<paragraph id="8BCE36FBFF82814873B1AAEB70A8F01B" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
<emphasis id="B905EAE9FF82814873B1AAEB704CF1D2" bold="true" box="[199,353,726,752]" pageId="4" pageNumber="209">Distribution.</emphasis>
Species of the genus 
<taxonomicName id="4C714D78FF828148712BAAEA73F9F1D3" authorityName="LeConte" authorityYear="1874" box="[605,724,727,753]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF828148712BAAEA73F9F1D3" box="[605,724,727,753]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
are distributed in the Nearctic (
<collectingCountry id="F366766BFF8281487737AAEA75B4F1D3" box="[1089,1177,727,753]" name="Canada" pageId="4" pageNumber="209">Canada</collectingCountry>
, 
<collectingCountry id="F366766BFF82814877D0AAEA75CDF1D3" box="[1190,1248,727,753]" name="United States of America" pageId="4" pageNumber="209">USA</collectingCountry>
) and Palaearctic regions (
<collectingCountry id="F366766BFF828148738DAAC6706AF037" box="[251,327,763,789]" name="Russia" pageId="4" pageNumber="209">Russia</collectingCountry>
(
<collectingRegion id="49B5F819FF828148722FAAC670B8F037" box="[345,405,763,789]" country="Russia" name="Altay" pageId="4" pageNumber="209">Altai</collectingRegion>
, Russian Far East), 
<collectingCountry id="F366766BFF82814871F6AAC6722EF037" box="[640,771,763,789]" name="Kazakhstan" pageId="4" pageNumber="209">Kazakhstan</collectingCountry>
(Altai and Dzhungar mts.), 
<collectingCountry id="F366766BFF828148773DAAC675A8F037" box="[1099,1157,763,789]" name="India" pageId="4" pageNumber="209">India</collectingCountry>
(Himalayas), and 
<collectingCountry id="F366766BFF828148762AAAC674B1F037" box="[1372,1436,763,789]" name="Japan" pageId="4" pageNumber="209">Japan</collectingCountry>
(
<collectingRegion id="49B5F819FF82814873E9AB22703FF01B" box="[159,274,799,825]" country="Japan" name="Hokkaido" pageId="4" pageNumber="209">Hokkaido</collectingRegion>
, Honshu).
</paragraph>
<paragraph id="8BCE36FBFF82814873B1AB7F7590F0A3" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
<emphasis id="B905EAE9FF82814873B1AB7F7066F07E" bold="true" box="[199,331,834,860]" pageId="4" pageNumber="209">Bionomics.</emphasis>
Species of 
<taxonomicName id="4C714D78FF82814872B8AB7E7368F07F" authorityName="LeConte" authorityYear="1874" box="[462,581,835,861]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814872B8AB7E7368F07F" box="[462,581,835,861]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
inhabit sand or gravel banks of rivers, streams and lakes, in moss and litter, and can be found frequently in the mountains (
<bibRefCitation id="EFE04B0AFF82814871D7AB5A7205F0A3" author="Hatch, M. H." box="[673,808,870,897]" pageId="4" pageNumber="209" pagination="1 - 384" refId="ref12098" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, 16 (2), i - ix + 1 - 384." type="journal article" year="1957">Hatch 1957</bibRefCitation>
, 
<bibRefCitation id="EFE04B0AFF8281487045AB5A7520F0A3" author="Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[819,1037,871,897]" pageId="4" pageNumber="209" pagination="1802" refId="ref12766" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="book chapter" year="2000">
Newton 
<emphasis id="B905EAE9FF82814870E5AB5A72E9F0A3" box="[915,964,871,897]" italics="true" pageId="4" pageNumber="209">et al</emphasis>
. 2000
</bibRefCitation>
, present study).
</paragraph>
</subSubSection>
<subSubSection id="C36B6570FF82814873B1ABB673B9F49B" pageId="4" pageNumber="209" type="discussion">
<paragraph id="8BCE36FBFF82814873B1ABB67375F573" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
<emphasis id="B905EAE9FF82814873B1ABB67016F087" bold="true" box="[199,315,907,933]" pageId="4" pageNumber="209">Remarks.</emphasis>
The genus 
<taxonomicName id="4C714D78FF82814872CCABB6734BF087" authorityName="Zerche" authorityYear="1992" box="[442,614,907,933]" class="Insecta" family="Staphylinidae" genus="Altaiodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814872CCABB6734BF087" box="[442,614,907,933]" italics="true" pageId="4" pageNumber="209">Altaiodromicus</emphasis>
</taxonomicName>
was originally described based on a single species (
<taxonomicName id="4C714D78FF82814877D3ABB671FDF0EB" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="schilenkovi">
<emphasis id="B905EAE9FF82814877D3ABB6746CF087" box="[1189,1345,907,933]" italics="true" pageId="4" pageNumber="209">A. schilenkovi</emphasis>
<bibRefCitation id="EFE04B0AFF828148763EABB671FDF0EB" author="Zerche, L." pageId="4" pageNumber="209" pagination="105 - 142" refId="ref13769" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
</taxonomicName>
) from 
<collectingRegion id="49B5F819FF8281487254AB927077F0EB" box="[290,346,943,969]" country="Russia" name="Altay" pageId="4" pageNumber="209">Altai</collectingRegion>
Mts., 
<collectingCountry id="F366766BFF82814872DEAB9270DBF0EB" box="[424,502,943,969]" name="Russia" pageId="4" pageNumber="209">Russia</collectingCountry>
. Based on the shape of the body (
<figureCitation id="134A2A7EFF82814870FFAB9272FCF0EB" box="[905,977,943,969]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1942,1966]" captionTargetBox="[156,1431,347,1909]" captionTargetId="figure-92@3.[151,1435,342,1915]" captionTargetPageId="3" captionText="FIGURES 1–5. Habitus of Microedus: 1—M. austinianus (USA: New Hampshire), 2—M. brevitarsis (holotype), 3—M. kastcheevi (paratype, Kazakhstan: Dzhungar Mts.), 4—A. schilenkovi (Russia: Altai, Korgon Mts.), 5—M. sp. (Kashmir). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11045157" httpUri="https://zenodo.org/record/11045157/files/figure.png" pageId="4" pageNumber="209">Fig. 4</figureCitation>
), it was compared with 
<taxonomicName id="4C714D78FF8281487799AB9274B0F0EB" authorityName="Tronquet" authorityYear="1981" box="[1263,1437,943,969]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487799AB9274B0F0EB" box="[1263,1437,943,969]" italics="true" pageId="4" pageNumber="209">Hygrodromicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C714D78FF82814873BFABEE7364F0CF" authority="Redtenbacher, 1857" authorityName="Redtenbacher" authorityYear="1857" box="[201,585,978,1005]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814873BFABEE7072F0CF" box="[201,351,979,1005]" italics="true" pageId="4" pageNumber="209">Geodromicus</emphasis>
Redtenbacher, 1857
</taxonomicName>
and distinguished by the narrow apical maxillary palpomere and different morphology of the aedeagus. During the study of 
<taxonomicName id="4C714D78FF82814871A2ABCA7264F733" box="[724,841,1015,1041]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="223" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
material collected by Dr. Kastcheev, I found a new, similar species from 
<collectingCountry id="F366766BFF8281487209AC267328F717" box="[383,517,1051,1077]" name="Kazakhstan" pageId="4" pageNumber="209">Kazakhstan</collectingCountry>
. This new species can be clearly distinguished from 
<taxonomicName id="4C714D78FF828148773AAC2675C4F717" authorityName="Zerche" authorityYear="1992" box="[1100,1257,1051,1077]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="schilenkovi">
<emphasis id="B905EAE9FF828148773AAC2675C4F717" box="[1100,1257,1051,1077]" italics="true" pageId="4" pageNumber="209">A. schilenkovi</emphasis>
</taxonomicName>
by the presence of sclerotized elongate structures within the internal sac (
<figureCitation id="134A2A7EFF8281487071AC02727BF77B" box="[775,854,1087,1113]" captionStart="FIGURES 18–23" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[178,1401,196,1911]" captionTargetId="figure-16@11.[170,1417,181,1919]" captionTargetPageId="11" captionText="FIGURES 18–23. Aedeagus of Microedus schilenkovi (Figs 18–19: Russia, Altai, Aygulakskiy Mts.; Fig. 20: Russia, Altai, Teletskoye Lake; Fig. 21: Kazakhstan, Tauchilik River) and A. kastcheevi (Figs. 22–23: paratype, Dzhungar Mts.): 18, 20–22— parameral view, 19, 23—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045163" httpUri="https://zenodo.org/record/11045163/files/figure.png" pageId="4" pageNumber="209">Fig. 22</figureCitation>
) and some other morphological features (see below). Besides that, I studied the 
<typeStatus id="54CA8859FF82814872B4AC5E7309F75F" box="[450,548,1123,1149]" pageId="4" pageNumber="209" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C714D78FF828148713CAC5E750CF75F" authority="Champion, 1925" authorityName="Champion" authorityYear="1925" box="[586,1057,1122,1149]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF828148713CAC5E7273F75F" box="[586,862,1123,1149]" italics="true" pageId="4" pageNumber="209">Geodromicus brevitarsis</emphasis>
<bibRefCitation id="EFE04B0AFF8281487010AC5E750CF75F" author="Champion, G. C." box="[870,1057,1122,1149]" pageId="4" pageNumber="209" pagination="101 - 112" refId="ref11916" refString="Champion, G. C. (1925) Some Indian [and Tibetan] Coleoptera (17). The Entomologist's Monthly Magazine, 61, 101 - 112." type="journal article" year="1925">Champion, 1925</bibRefCitation>
</taxonomicName>
from northern 
<collectingCountry id="F366766BFF82814877BBAC5E742AF75F" box="[1229,1287,1123,1149]" name="India" pageId="4" pageNumber="209">India</collectingCountry>
(
<figureCitation id="134A2A7EFF8281487660AC5E7471F75F" box="[1302,1372,1123,1149]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1942,1966]" captionTargetBox="[156,1431,347,1909]" captionTargetId="figure-92@3.[151,1435,342,1915]" captionTargetPageId="3" captionText="FIGURES 1–5. Habitus of Microedus: 1—M. austinianus (USA: New Hampshire), 2—M. brevitarsis (holotype), 3—M. kastcheevi (paratype, Kazakhstan: Dzhungar Mts.), 4—A. schilenkovi (Russia: Altai, Korgon Mts.), 5—M. sp. (Kashmir). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11045157" httpUri="https://zenodo.org/record/11045157/files/figure.png" pageId="4" pageNumber="209">Fig. 2</figureCitation>
). The apical maxillary palpomere of this species is also narrow, and based on the slightly elongate antennomeres 4–10, 
<taxonomicName id="4C714D78FF82814873E1AC96701FF7E7" authorityName="Champion" authorityYear="1925" box="[151,306,1195,1221]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF82814873E1AC96701FF7E7" box="[151,306,1195,1221]" italics="true" pageId="4" pageNumber="209">G. brevitarsis</emphasis>
</taxonomicName>
is somewhat similar to the two species listed above, but differs by the smaller and narrower body, with markedly narrower pronotum, and different morphology of the aedeagus. Based on the external morphological features, one unnamed species from Kashmir is also similar to 
<taxonomicName id="4C714D78FF828148703FACCE72CCF62F" authorityName="Champion" authorityYear="1925" box="[841,993,1267,1293]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF828148703FACCE72CCF62F" box="[841,993,1267,1293]" italics="true" pageId="4" pageNumber="209">G. brevitarsis</emphasis>
</taxonomicName>
(see below). Several years ago, with the assistance of the late A. Smetana, I received several Nearctic species of 
<taxonomicName id="4C714D78FF82814870B6AD2A751AF613" authorityName="LeConte" authorityYear="1874" box="[960,1079,1303,1329]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814870B6AD2A751AF613" box="[960,1079,1303,1329]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F366766BFF828148770AAD2A75FDF613" box="[1148,1232,1303,1329]" name="Canada" pageId="4" pageNumber="209">Canada</collectingCountry>
and 
<collectingCountry id="F366766BFF8281487671AD2A746EF613" box="[1287,1347,1303,1329]" name="United States of America" pageId="4" pageNumber="209">USA</collectingCountry>
. During the study of mouthparts and aedeagi of these species, I found that they are similar to species of 
<taxonomicName id="4C714D78FF82814877C7AD067470F677" authorityName="Zerche" authorityYear="1992" box="[1201,1373,1339,1365]" class="Insecta" family="Staphylinidae" genus="Altaiodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814877C7AD067470F677" box="[1201,1373,1339,1365]" italics="true" pageId="4" pageNumber="209">Altaiodromicus</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A7EFF828148761CAD0671FCF65B" captionStart="FIGURES 6–13" captionStartId="7.[151,264,1902,1926]" captionTargetBox="[159,1427,352,1871]" captionTargetId="figure-100@7.[151,1436,345,1879]" captionTargetPageId="7" captionText="FIGURES 6–13. Mouthparts (dorsal view) of M. austinianus (Figs. 6, 8, 10, 12) and M. schilenkovi (Figs. 7, 9, 11, 13): 6–7— labrum, 8–9—mandibles (left and right), 10–11—maxilla (right), 12–13—labium. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045159" httpUri="https://zenodo.org/record/11045159/files/figure.png" pageId="4" pageNumber="209">Figs 6–13</figureCitation>
) and 
<taxonomicName id="4C714D78FF8281487266AD627087F65B" authorityName="Champion" authorityYear="1925" box="[272,426,1375,1401]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF8281487266AD627087F65B" box="[272,426,1375,1401]" italics="true" pageId="4" pageNumber="209">G. brevitarsis</emphasis>
</taxonomicName>
. The general shapes of the body (
<figureCitation id="134A2A7EFF8281487051AD627245F65B" box="[807,872,1375,1401]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1942,1966]" captionTargetBox="[156,1431,347,1909]" captionTargetId="figure-92@3.[151,1435,342,1915]" captionTargetPageId="3" captionText="FIGURES 1–5. Habitus of Microedus: 1—M. austinianus (USA: New Hampshire), 2—M. brevitarsis (holotype), 3—M. kastcheevi (paratype, Kazakhstan: Dzhungar Mts.), 4—A. schilenkovi (Russia: Altai, Korgon Mts.), 5—M. sp. (Kashmir). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11045157" httpUri="https://zenodo.org/record/11045157/files/figure.png" pageId="4" pageNumber="209">Fig. 1</figureCitation>
) and the aedeagus (
<figureCitation id="134A2A7EFF828148773DAD6275B1F65B" box="[1099,1180,1375,1401]" captionStart="FIGURES 14–17" captionStartId="8.[151,264,1176,1200]" captionTargetBox="[159,1428,191,1144]" captionTargetId="figure-18@8.[151,1436,181,1152]" captionTargetPageId="8" captionText="FIGURES 14–17. Aedeagus of Microedus austinianus (Figs 14–15: Canada, Quebec, Riviere-a-Claude) and M. brevitarsis (Figs. 16–17: holotype): 14, 16—parameral view, 15, 17—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045161" httpUri="https://zenodo.org/record/11045161/files/figure.png" pageId="4" pageNumber="209">Fig. 14</figureCitation>
) of the type species of 
<taxonomicName id="4C714D78FF82814873E1ADBE7023F6BF" authorityName="LeConte" authorityYear="1874" box="[151,270,1411,1437]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814873E1ADBE7023F6BF" box="[151,270,1411,1437]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C714D78FF8281487269ADBE70E8F6BE" authorityName="LeConte" authorityYear="1874" box="[287,453,1411,1436]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="austinianus">
<emphasis id="B905EAE9FF8281487269ADBE70E8F6BE" box="[287,453,1411,1436]" italics="true" pageId="4" pageNumber="209">M. austinianus</emphasis>
</taxonomicName>
) are somewhat similar to these of 
<taxonomicName id="4C714D78FF828148703CADBE72CBF6BF" authorityName="Champion" authorityYear="1925" box="[842,998,1411,1437]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF828148703CADBE72CBF6BF" box="[842,998,1411,1437]" italics="true" pageId="4" pageNumber="209">M. brevitarsis</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A7EFF8281487080ADBE7564F6BF" box="[1014,1097,1411,1437]" captionStart="FIGURES 14–17" captionStartId="8.[151,264,1176,1200]" captionTargetBox="[159,1428,191,1144]" captionTargetId="figure-18@8.[151,1436,181,1152]" captionTargetPageId="8" captionText="FIGURES 14–17. Aedeagus of Microedus austinianus (Figs 14–15: Canada, Quebec, Riviere-a-Claude) and M. brevitarsis (Figs. 16–17: holotype): 14, 16—parameral view, 15, 17—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045161" httpUri="https://zenodo.org/record/11045161/files/figure.png" pageId="4" pageNumber="209">Fig. 16</figureCitation>
). The shapes of antennomeres 4–10 of 
<taxonomicName id="4C714D78FF8281487387AD9A70BBF6E2" authorityName="LeConte" authorityYear="1874" box="[241,406,1447,1472]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="austinianus">
<emphasis id="B905EAE9FF8281487387AD9A70BBF6E2" box="[241,406,1447,1472]" italics="true" pageId="4" pageNumber="209">M. austinianus</emphasis>
</taxonomicName>
are also similar to these in all Asian species listed above. Other Nearctic species of 
<taxonomicName id="4C714D78FF8281487650AD9A74B0F6E3" authorityName="LeConte" authorityYear="1874" box="[1318,1437,1447,1473]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487650AD9A74B0F6E3" box="[1318,1437,1447,1473]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
studied by me (
<taxonomicName id="4C714D78FF8281487232ADF67366F6C7" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[324,587,1482,1509]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="fenderi">
<emphasis id="B905EAE9FF8281487232ADF67076F6C6" box="[324,347,1483,1508]" italics="true" pageId="4" pageNumber="209">M</emphasis>
. 
<emphasis id="B905EAE9FF828148721FADF67095F6C7" box="[361,440,1483,1509]" italics="true" pageId="4" pageNumber="209">fenderi</emphasis>
<bibRefCitation id="EFE04B0AFF82814872C9ADF67366F6C7" author="Hatch, M. H." box="[447,587,1482,1509]" pageId="4" pageNumber="209" pagination="1 - 384" refId="ref12098" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, 16 (2), i - ix + 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C714D78FF8281487120ADF672E1F6C7" authority="(Mannerheim, 1843)" baseAuthorityName="Mannerheim" baseAuthorityYear="1843" box="[598,972,1483,1509]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B905EAE9FF8281487120ADF67340F6C6" box="[598,621,1483,1508]" italics="true" pageId="4" pageNumber="209">M</emphasis>
. 
<emphasis id="B905EAE9FF828148710DADF673F0F6C7" box="[635,733,1483,1509]" italics="true" pageId="4" pageNumber="209">laticollis</emphasis>
(
<bibRefCitation id="EFE04B0AFF828148719AADF672EEF6C7" author="Mannerheim, C. G. von" box="[748,963,1483,1509]" pageId="4" pageNumber="209" pagination="175 - 314" refId="ref12384" refString="Mannerheim, C. G. von (1843) Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16 (2), 175 - 314. https: // doi. org / 10.5962 / bhl. title. 37833" type="journal article" year="1843">Mannerheim, 1843</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C714D78FF8281487774ADF67435F6C7" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[1026,1304,1482,1509]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="porterae">
<emphasis id="B905EAE9FF8281487774ADF67534F6C6" box="[1026,1049,1483,1508]" italics="true" pageId="4" pageNumber="209">M</emphasis>
. 
<emphasis id="B905EAE9FF8281487751ADF675AAF6C6" box="[1063,1159,1483,1508]" italics="true" pageId="4" pageNumber="209">porterae</emphasis>
<bibRefCitation id="EFE04B0AFF82814877F8ADF67435F6C7" author="Hatch, M. H." box="[1166,1304,1482,1509]" pageId="4" pageNumber="209" pagination="1 - 384" refId="ref12098" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, 16 (2), i - ix + 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
) have more elongate antennomeres 4–10. Besides that, all these species have similar ground sculpture of the forebody and shape of the female genital segment (
<figureCitation id="134A2A7EFF8281487281AE2E7361F50F" box="[503,588,1555,1581]" captionStart="FIGURES 24–29" captionStartId="12.[151,264,1172,1196]" captionTargetBox="[159,1429,189,1144]" captionTargetId="figure-18@12.[151,1436,181,1149]" captionTargetPageId="12" captionText="FIGURES 24–29. Abdominal segments of Microedus schilenkovi: 24—male tergite VIII, 25—male sternite VIII, 26—male genital segment, 27—female tergite VIII, 28—female sternite VIII, 29—female genital segment. Scale bars 0.1 m" figureDoi="http://doi.org/10.5281/zenodo.11045165" httpUri="https://zenodo.org/record/11045165/files/figure.png" pageId="4" pageNumber="209">Fig. 29</figureCitation>
). Thus, I decided to synonymize the genus 
<taxonomicName id="4C714D78FF828148774EAE2E75C9F50F" authorityName="Zerche" authorityYear="1992" box="[1080,1252,1555,1581]" class="Insecta" family="Staphylinidae" genus="Altaiodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF828148774EAE2E75C9F50F" box="[1080,1252,1555,1581]" italics="true" pageId="4" pageNumber="209">Altaiodromicus</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C714D78FF8281487650AE2E74B0F50F" authorityName="LeConte" authorityYear="1874" box="[1318,1437,1555,1581]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487650AE2E74B0F50F" box="[1318,1437,1555,1581]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
and transfer 
<taxonomicName id="4C714D78FF8281487255AE0A7091F573" authorityName="Champion" authorityYear="1925" box="[291,444,1591,1617]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="brevitarsis">
<emphasis id="B905EAE9FF8281487255AE0A7091F573" box="[291,444,1591,1617]" italics="true" pageId="4" pageNumber="209">G. brevitarsis</emphasis>
</taxonomicName>
to this genus.
</paragraph>
<paragraph id="8BCE36FBFF82814873B1AE66739EF40B" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
Species of 
<taxonomicName id="4C714D78FF8281487249AE6670C5F557" authorityName="Nakane" authorityYear="1983" box="[319,488,1627,1653]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487249AE6670C5F557" box="[319,488,1627,1653]" italics="true" pageId="4" pageNumber="209">Liophilydrodes</emphasis>
</taxonomicName>
, known from 
<collectingCountry id="F366766BFF82814871F7AE6673ECF557" box="[641,705,1627,1653]" name="Japan" pageId="4" pageNumber="209">Japan</collectingCountry>
and Far Eastern 
<collectingCountry id="F366766BFF828148700EAE6672EEF557" box="[888,963,1627,1653]" name="Russia" pageId="4" pageNumber="209">Russia</collectingCountry>
have a narrow apical segment of maxillary palpi, similar general shape of the body and mouthparts (see Figs 
<quantity id="4C899B1EFF82814870D2AE42750AF5BB" box="[932,1063,1663,1689]" metricMagnitude="1" metricUnit="m" metricValue="1.00965" metricValueMax="1.01346" metricValueMin="1.00584" pageId="4" pageNumber="209" unit="in" value="397.5" valueMax="399.0" valueMin="396.0">396–399 in</quantity>
<bibRefCitation id="EFE04B0AFF8281487747AE4275D0F5BB" author="Watanabe, Y." box="[1073,1277,1663,1689]" pageId="4" pageNumber="209" pagination="59 - 391" refId="ref13615" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990))</bibRefCitation>
as in species from Middle Asia and the Nearctic Region. All species of 
<taxonomicName id="4C714D78FF8281487054AE9E72E6F59F" authorityName="Nakane" authorityYear="1983" box="[802,971,1699,1725]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487054AE9E72E6F59F" box="[802,971,1699,1725]" italics="true" pageId="4" pageNumber="209">Liophilydrodes</emphasis>
</taxonomicName>
have elongate antennomeres 4–10. Some species have similar shapes of aedeagi as those from Middle Asia, with the presence of elongate structures within the internal sac (
<figureCitation id="134A2A7EFF828148723AAED670B3F427" box="[332,414,1771,1797]" captionStart="FIGURES 34–42" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[159,1428,189,1812]" captionTargetId="figure-16@15.[151,1436,181,1820]" captionTargetPageId="15" captionText="FIGURES 34–42. Aedeagus (Figs. 34–36: Japan: Honshu, Mt. Hotaka) and abdominal segments (Figs. 37–42) of Microedus subtilis: 34—aedeagus, parameral view, 35—aedeagus, apex of median lobe, 36—aedeagus, lateral view, 37—male tergite VIII, 38—male sternite VIII, 39—male genital segment, 40—female tergite VIII, 41—female sternite VIII, 42—female genital segment. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11045171" httpUri="https://zenodo.org/record/11045171/files/figure.png" pageId="4" pageNumber="209">Fig. 34</figureCitation>
), sometimes with flagellum as in 
<figureCitation id="134A2A7EFF8281487066AED672BEF427" box="[784,915,1771,1797]" captionStart-0="FIGURES 18–23" captionStart-1="FIGURES 24–29" captionStartId-0="11.[151,264,1943,1967]" captionStartId-1="12.[151,264,1172,1196]" captionTargetBox-0="[178,1401,196,1911]" captionTargetBox-1="[159,1429,189,1144]" captionTargetId-0="figure-16@11.[170,1417,181,1919]" captionTargetId-1="figure-18@12.[151,1436,181,1149]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 18–23. Aedeagus of Microedus schilenkovi (Figs 18–19: Russia, Altai, Aygulakskiy Mts.; Fig. 20: Russia, Altai, Teletskoye Lake; Fig. 21: Kazakhstan, Tauchilik River) and A. kastcheevi (Figs. 22–23: paratype, Dzhungar Mts.): 18, 20–22— parameral view, 19, 23—lateral view. Scale bar: 0.1 mm." captionText-1="FIGURES 24–29. Abdominal segments of Microedus schilenkovi: 24—male tergite VIII, 25—male sternite VIII, 26—male genital segment, 27—female tergite VIII, 28—female sternite VIII, 29—female genital segment. Scale bars 0.1 m" figureDoi-0="http://doi.org/10.5281/zenodo.11045163" figureDoi-1="http://doi.org/10.5281/zenodo.11045165" httpUri-0="https://zenodo.org/record/11045163/files/figure.png" httpUri-1="https://zenodo.org/record/11045165/files/figure.png" pageId="4" pageNumber="209">
Figs. 
<quantity id="4C899B1EFF828148703BAED672BEF427" box="[845,915,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.604000000000001" metricValueMin="5.3340000000000005" pageId="4" pageNumber="209" unit="in" value="23.5" valueMax="26.0" valueMin="21.0">21–26</quantity>
</figureCitation>
in 
<bibRefCitation id="EFE04B0AFF82814870C2AED675C8F427" author="Shavrin, A. V. &amp; Makarov, K. V." box="[948,1253,1771,1797]" pageId="4" pageNumber="209" pagination="36 - 53" refId="ref13153" refString="Shavrin, A. V. &amp; Makarov, K. V. (2019) Contribution to the knowledge of the fauna of rove beetles of the subfamily Omaliinae MacLeay, 1825 (Coleoptera: Staphylinidae) of Kunashir Island, Kurile Islands. Russian Entomological Journal, 28 (1), 36 - 53. https: // doi. org / 10.15298 / rusentj. 28.1.06" type="journal article" year="2019">Shavrin &amp; Makarov (2019)</bibRefCitation>
. Thus, the genus 
<taxonomicName id="4C714D78FF82814873E1AF32706DF40B" authorityName="Nakane" authorityYear="1983" box="[151,320,1807,1833]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814873E1AF32706DF40B" box="[151,320,1807,1833]" italics="true" pageId="4" pageNumber="209">Liophilydrodes</emphasis>
</taxonomicName>
is synonymized with 
<taxonomicName id="4C714D78FF8281487143AF327381F40B" authorityName="LeConte" authorityYear="1874" box="[565,684,1807,1833]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF8281487143AF327381F40B" box="[565,684,1807,1833]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCE36FBFF82814873B1AF0E73B9F49B" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="209">
The Palaearctic species of 
<taxonomicName id="4C714D78FF828148728AAF0E735EF46F" authorityName="LeConte" authorityYear="1874" box="[508,627,1843,1869]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF828148728AAF0E735EF46F" box="[508,627,1843,1869]" italics="true" pageId="4" pageNumber="209">Microedus</emphasis>
</taxonomicName>
vary in proportions of the antennomeres, the shape of the interocellar foveae, the pronotum and the aedeagus as in some other taxa of the 
<taxonomicName id="4C714D78FF82814870B5AF6A7510F452" authorityName="Mulsant &amp; Rey" authorityYear="1880" box="[963,1085,1879,1904]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAE9FF82814870B5AF6A7510F452" box="[963,1085,1879,1904]" italics="true" pageId="4" pageNumber="209">Hygrogeus</emphasis>
</taxonomicName>
group (e.g. 
<bibRefCitation id="EFE04B0AFF82814877A7AF6A74B5F453" author="Shavrin, A. V." box="[1233,1432,1879,1905]" pageId="4" pageNumber="209" pagination="451 - 496" refId="ref13084" refString="Shavrin, A. V. (2022 b) The bodemeyeri and the convexicollis groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data fro some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae). Zootaxa, 5213 (5), 451 - 496. https: // doi. org / 10.11646 / zootaxa. 5213.5.1" type="journal article" year="2022">Shavrin (2022b))</bibRefCitation>
. Species of the Nearctic Region are in need of a comprehensive taxonomic revision due to significant morphological differences between species (e.g. 
<taxonomicName id="4C714D78FF8281487179AFA273AEF49B" authorityName="Hatch" authorityYear="1957" box="[527,643,1951,1977]" class="Insecta" family="Staphylinidae" genus="Microedus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="fenderi">
<emphasis id="B905EAE9FF8281487179AFA273AEF49B" box="[527,643,1951,1977]" italics="true" pageId="4" pageNumber="209">M. fenderi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>