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<document id="1233E64C7BE556343B923BA667C9CCFF" ID-DOI="10.11646/zootaxa.3539.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10096759" ID-ZooBank="0564EF02-FCF9-49A8-BDAC-F432876C16B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629821597637" checkinUser="felipe" docAuthor="Ramos, Magno S. &amp; Azevedo, Celso O." docDate="2012" docId="03D887D08A0AFFB71399FDF6FB2AFCDB" docLanguage="en" docName="zt03539p080.pdf" docOrigin="Zootaxa 3539 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3539.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eupsenella wanamara Ramos &amp; Azevedo, sp. nov." docType="treatment" docVersion="3" lastPageNumber="59" masterDocId="FFE1FFA88A33FF8D130EFFB1FFFCFFEC" masterDocTitle="Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="58" updateTime="1699581041301" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DA0BD0BA1FA50EE2BA7F939A95AAC22D">Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)</mods:title>
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<mods:name id="EA5E85FC860D7E6A47492DA16538E459" type="personal">
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<mods:namePart id="23A6C09686AA3D54532216B05B7D34CA">Ramos, Magno S.</mods:namePart>
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<mods:namePart id="82D24BE13D46BAC2F63F477EE173812E">Azevedo, Celso O.</mods:namePart>
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<treatment id="03D887D08A0AFFB71399FDF6FB2AFCDB" LSID="urn:lsid:plazi:treatment:03D887D08A0AFFB71399FDF6FB2AFCDB" httpUri="http://treatment.plazi.org/id/03D887D08A0AFFB71399FDF6FB2AFCDB" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<subSubSection id="C36B654D8A0AFFB41399FDF6FCFCFD8D" box="[151,768,583,609]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BCE36C68A0AFFB41399FDF6FCFCFD8D" blockId="57.[151,768,583,642]" box="[151,768,583,609]" pageId="57" pageNumber="58">
<heading id="D08681AA8A0AFFB41399FDF6FCFCFD8D" bold="true" box="[151,768,583,609]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="B905EAD48A0AFFB41399FDF6FCFCFD8D" bold="true" box="[151,768,583,609]" pageId="57" pageNumber="58">
<taxonomicName id="4C714D458A0AFFB41399FDF6FD6AFD8D" authority="Ramos &amp; Azevedo" authorityName="Ramos &amp; Azevedo" box="[151,662,583,609]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="wanamara" status="sp. nov.">
<emphasis id="B905EAD48A0AFFB41399FDF6FE5AFD8D" bold="true" box="[151,422,583,609]" italics="true" pageId="57" pageNumber="58">Eupsenella wanamara</emphasis>
Ramos &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A23657AF8A0AFFB411ADFDF6FCFCFD8D" box="[675,768,583,609]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A0AFFB71399FDDBFE66FEF0" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph id="8BCE36C68A0AFFB41399FDDBFEA3FD6E" blockId="57.[151,768,583,642]" box="[151,351,618,642]" pageId="57" pageNumber="58">
(
<figureCitation id="134A2A438A0AFFB41391FDDBFF09FD6E" box="[159,245,618,642]" captionStart="FIGURES 3947" captionStartId="65.[151,264,1445,1468]" captionTargetBox="[151,1436,193,1424]" captionTargetId="figure-71@65.[151,1436,193,1424]" captionTargetPageId="65" captionText="FIGURES 3947. Head in dorsal view. 39. Eupsenella kokatha sp. nov. 40. E. larrakia sp. nov. 41. E. maya sp. nov. 42. E. malgana sp. nov. 43. E. nanda sp. nov. 44. E. nangatara sp. nov. 45. E. pangkala sp. nov. 46. E. wanamara sp. nov. 47. E. agilis. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096769" httpUri="https://zenodo.org/record/10096769/files/figure.png" pageId="57" pageNumber="58">Figs 46</figureCitation>
,
<figureCitation id="134A2A438A0AFFB4120FFDDBFEE3FD6E" box="[257,287,618,642]" captionStart="FIGURES 8492" captionStartId="70.[151,264,1931,1954]" captionTargetBox="[161,1425,181,1910]" captionTargetId="figure-72@70.[161,1425,181,1910]" captionTargetPageId="70" captionText="FIGURES 8492. Mesosoma in dorsal view. 84. Eupsenella kokatha sp. nov. 85. E. larrakia sp. nov. 86. E. maya sp. nov. 87." figureDoi="http://doi.org/10.5281/zenodo.10096773" httpUri="https://zenodo.org/record/10096773/files/figure.png" pageId="57" pageNumber="58">91</figureCitation>
,
<figureCitation id="134A2A438A0AFFB41225FDDBFEAAFD6E" box="[299,342,618,642]" captionStart="FIGURES 133137" captionStartId="75.[151,264,1228,1251]" captionTargetBox="[151,1436,193,1207]" captionTargetId="figure-50@75.[151,1436,193,1207]" captionTargetPageId="75" captionText="FIGURES 133137. Right forewing. 133. Eupsenella nanda sp. nov. 134. E. nangatara sp. nov. 135. E. s pangkala sp. nov. 136. E. wanamara sp. nov. 137. E. agilis. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096783" httpUri="https://zenodo.org/record/10096783/files/figure.png" pageId="57" pageNumber="58">136</figureCitation>
)
</paragraph>
<paragraph id="8BCE36C68A0AFFB41399FD03FC36FD26" blockId="57.[151,1436,690,1939]" box="[151,970,690,715]" pageId="57" pageNumber="58">
<emphasis id="B905EAD48A0AFFB41399FD03FED4FD27" bold="true" box="[151,296,690,715]" inLineHeading="true" pageId="57" pageNumber="58" reason="1">Description.</emphasis>
FEMALE (
<typeStatus id="54CA88648A0AFFB412A0FD03FDEBFD26" box="[430,535,690,714]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
). Body length
<quantity id="4C899B238A0AFFB411B7FD03FCDCFD26" box="[697,800,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="6.58" pageId="57" pageNumber="58" unit="mm" value="6.58">6.58 mm</quantity>
. LFW
<quantity id="4C899B238A0AFFB41063FD03FC3AFD26" box="[877,966,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="57" pageNumber="58" unit="mm" value="4.1">4.1 mm</quantity>
.
</paragraph>
<paragraph id="8BCE36C68A0AFFB413C9FD67FB79FCB6" blockId="57.[151,1436,690,1939]" pageId="57" pageNumber="58">
<emphasis id="B905EAD48A0AFFB413C9FD67FEB5FD02" box="[199,329,726,750]" italics="true" pageId="57" pageNumber="58">Coloration.</emphasis>
Head dark castaneous almost black; scape, pedicel and flagellum castaneous; mandible castaneous with teeth dark castaneous; palpi light castaneous; mesosoma dark castaneous almost black; tegula castaneous; wing venation light castaneous; procoxa dark castaneous; mesocoxa, metacoxa, trochanters and profemur castaneous; mesofemur, metafemur and tarsi light castaneous; metasoma dark castaneous.
</paragraph>
<paragraph id="8BCE36C68A0AFFB413C9FCD7FDF8FABE" blockId="57.[151,1436,690,1939]" pageId="57" pageNumber="58">
<emphasis id="B905EAD48A0AFFB413C9FCD7FEF9FC92" box="[199,261,870,894]" italics="true" pageId="57" pageNumber="58">Head</emphasis>
(
<figureCitation id="134A2A438A0AFFB41213FCD7FE86FC92" box="[285,378,870,894]" captionStart="FIGURES 3947" captionStartId="65.[151,264,1445,1468]" captionTargetBox="[151,1436,193,1424]" captionTargetId="figure-71@65.[151,1436,193,1424]" captionTargetPageId="65" captionText="FIGURES 3947. Head in dorsal view. 39. Eupsenella kokatha sp. nov. 40. E. larrakia sp. nov. 41. E. maya sp. nov. 42. E. malgana sp. nov. 43. E. nanda sp. nov. 44. E. nangatara sp. nov. 45. E. pangkala sp. nov. 46. E. wanamara sp. nov. 47. E. agilis. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096769" httpUri="https://zenodo.org/record/10096769/files/figure.png" pageId="57" pageNumber="58">Fig. 46</figureCitation>
). Head wider than long, subrectangular in profile; gena weakly concave in profile. Mandible evenly wide, lowermost tooth larger than upper ones, lower tooth curved outward, lower margin strongly setose. Clypeus coriaceous, median clypeal lobe subangulate, median concavity of median clypeal lobe absent, 0.06 LH, well marked off from frons, median carina absent. Malar space conspicuous, 0.31 mandibular base. Antenna
<quantity id="4C899B238A0AFFB412DDFC47FDD3FBE2" box="[467,559,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="57" pageNumber="58" unit="mm" value="1.9">1.9 mm</quantity>
; first five antennal sectionsantennomeres in ratio of about 11:11:11:11:11; flagellomere 1 1.22 longer than wide, flagellar pubescence dense, with many outstanding erect setae. Region between torulus and eye slightly concave. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.14 LH. Eye protuberant, 0.58 LH, with sparse hairs, located more laterally, lower margin without strong depression. Frons strongly coriaceous-punctate. WH 1.28 LH. WF 0.57 WH. WF 1.3 HE. OOL 0.88 WOT. VOL 0.5 HE. Distance from posterior ocellus to vertex crest 0.25 DAO. Ocelli large, DAO 0.11 WF, ocellar triangle not compact, distance between posterior ocelli 2.4 DAO, anterior ocellus not surpassing imaginary top eye line, posterior ocellus almost reaching vertex. Vertex concave, long setae, 0.20 LH, temple profile slightly convergent in dorsal view, with corner angulate, not projected postero-laterad. Hipostomal carina arched, low.
</paragraph>
<paragraph id="8BCE36C68A0AFFB413C9FAEEFF0DF8CA" blockId="57.[151,1436,690,1939]" pageId="57" pageNumber="58">
<emphasis id="B905EAD48A0AFFB413C9FAEEFEC3FA9A" box="[199,319,1375,1398]" italics="true" pageId="57" pageNumber="58">Mesosoma</emphasis>
(
<figureCitation id="134A2A438A0AFFB4125BFAEFFE53FA9A" box="[341,431,1374,1398]" captionStart="FIGURES 8492" captionStartId="70.[151,264,1931,1954]" captionTargetBox="[161,1425,181,1910]" captionTargetId="figure-72@70.[161,1425,181,1910]" captionTargetPageId="70" captionText="FIGURES 8492. Mesosoma in dorsal view. 84. Eupsenella kokatha sp. nov. 85. E. larrakia sp. nov. 86. E. maya sp. nov. 87." figureDoi="http://doi.org/10.5281/zenodo.10096773" httpUri="https://zenodo.org/record/10096773/files/figure.png" pageId="57" pageNumber="58">Fig. 91</figureCitation>
). Pronotal disc coriaceous-punctate, 0.42 as long as wide, anterior corner rounded, posterior margin concave medially, pronotal disc without short longitudinal furrow. Parapsidal furrow wholly conspicuous. Notauli convergent posterad. Mesoscutum 0.75 length pronotal disc. Scutellum coriaceous, 0.74 length pronotal disc, almost flat in profile, posterior margin convex medially, scutellar pit elliptical, narrowed, distant each other 11.0 its own diameter. Dorsal axillar surface with fovea subrectangular. Metanotum with median region almost flat, coriaceous-puncticulate, lower than scutellum in profile, with fovea elliptical laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.18 length propodeal disc, incomplete posterad; discal carina 0.05 length propodeal disc, converging posterad, slightly arched; space between median and discal carina almost polished and smooth. Mesopleuron with subtegular furrow oval; mesepimeral suture almost inconspicuous posterad, mesopleural epicoxal lobe inconspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression heart-shaped; median suture evident.
</paragraph>
<paragraph id="8BCE36C68A0AFFB413C9F882FA67F87E" blockId="57.[151,1436,690,1939]" pageId="57" pageNumber="58">
<emphasis id="B905EAD48A0AFFB413C9F882FEEFF8A6" box="[199,275,1843,1866]" italics="true" pageId="57" pageNumber="58">Wings.</emphasis>
Macropterous. Forewing (
<figureCitation id="134A2A438A0AFFB41133F883FD61F8A6" box="[573,669,1842,1866]" captionStart="FIGURES 133137" captionStartId="75.[151,264,1228,1251]" captionTargetBox="[151,1436,193,1207]" captionTargetId="figure-50@75.[151,1436,193,1207]" captionTargetPageId="75" captionText="FIGURES 133137. Right forewing. 133. Eupsenella nanda sp. nov. 134. E. nangatara sp. nov. 135. E. s pangkala sp. nov. 136. E. wanamara sp. nov. 137. E. agilis. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096783" httpUri="https://zenodo.org/record/10096783/files/figure.png" pageId="57" pageNumber="58">Fig. 136</figureCitation>
) with 1M cell subpentagonal; 1M and 1R1 cells not aligned basally; C cell widening distally;1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs
<emphasis id="B905EAD48A0AFFB41780F8E7FB60F882" box="[1166,1180,1878,1902]" italics="true" pageId="57" pageNumber="58">b</emphasis>
convex; 2R1 cell 0.15 LFW; Distal margin of Rs
<emphasis id="B905EAD48A0AFFB412CDF8CAFE33F87E" box="[451,463,1915,1938]" italics="true" pageId="57" pageNumber="58">c</emphasis>
convex; r-rs vein inclined; R1
<emphasis id="B905EAD48A0AFFB41023F8CAFCC7F87F" box="[813,827,1915,1939]" italics="true" pageId="57" pageNumber="58">b</emphasis>
entirely on anterior margin of forewing; Cu
<emphasis id="B905EAD48A0AFFB41638F8CAFAB8F87F" box="[1334,1348,1915,1939]" italics="true" pageId="57" pageNumber="58">b</emphasis>
absent;
</paragraph>
<paragraph id="8BCE36C68A09FFB713C9FF28FC87FF38" blockId="58.[151,1436,151,824]" pageId="58" pageNumber="59">
<emphasis id="B905EAD48A09FFB713C9FF28FEFFFF5C" box="[199,259,153,176]" italics="true" pageId="58" pageNumber="59">Legs.</emphasis>
Profemur very swollen, 0.65 as wide as long. Metacoxa with median region 0.5 as wide as long. Metacoxal projection in dorsal region conspicuous, apex rounded.
</paragraph>
<paragraph id="8BCE36C68A09FFB713C9FF50FC6BFF1B" blockId="58.[151,1436,151,824]" box="[199,919,223,248]" pageId="58" pageNumber="59">
<emphasis id="B905EAD48A09FFB713C9FF50FEBFFF14" box="[199,323,225,248]" italics="true" pageId="58" pageNumber="59">Metasoma.</emphasis>
0.3 as wide as long, ventral carina of petiole present.
</paragraph>
<paragraph id="8BCE36C68A09FFB713C9FEB5FE66FEF0" blockId="58.[151,1436,151,824]" box="[199,410,260,284]" pageId="58" pageNumber="59">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A09FFB713C9FE96FC50FE88" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8BCE36C68A09FFB713C9FE96FC50FE88" blockId="58.[151,1436,151,824]" pageId="58" pageNumber="59">
<emphasis id="B905EAD48A09FFB713C9FE96FE4EFEAC" bold="true" box="[199,434,295,320]" inLineHeading="true" pageId="58" pageNumber="59" reason="1">Material examined.</emphasis>
<materialsCitation id="3B193C9B8A09FFB712B5FE99FC50FE88" collectingDate="1980-05-09" collectionCode="ANIC" collectorName="I. D. Naumann &amp; J. C. Cardale" country="Australia" location="AUSTRALIA" pageId="58" pageNumber="59" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B905EAD48A09FFB712B5FE99FDDDFEAC" box="[443,545,296,320]" italics="true" pageId="58" pageNumber="59">
<typeStatus id="54CA88648A09FFB712B5FE99FDDDFEAC" box="[443,545,296,320]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D77FD4F8A09FFB71125FE99FDBCFED3" box="[555,576,296,319]" pageId="58" pageNumber="59" type="female"></specimenCount>
.
<collectingCountry id="F36676568A09FFB7115EFE96FD0DFED3" box="[592,753,295,319]" name="Australia" pageId="58" pageNumber="59">AUSTRALIA</collectingCountry>
, Q[ueensland], (near School),
<date id="FFCF10068A09FFB71741FE96FB20FED3" box="[1103,1244,295,320]" pageId="58" pageNumber="59" value="1980-05-09">
<collectingDate id="EF8BE9EE8A09FFB71741FE96FB20FED3" box="[1103,1244,295,320]" pageId="58" pageNumber="59" value="1980-05-09">9 May 1980</collectingDate>
</date>
,
<collectingMethod id="52304ED18A09FFB717E4FE96FA67FED3" box="[1258,1435,295,319]" pageId="58" pageNumber="59">yellow pan trap</collectingMethod>
(ethanediol), transferred to alcohol,
<collectorName id="268453108A09FFB71127FEFDFD38FE8F" box="[553,708,332,355]" pageId="58" pageNumber="59">I.D.Naumann</collectorName>
,
<collectorName id="268453108A09FFB711DEFEFDFCA9FE88" box="[720,853,332,356]" pageId="58" pageNumber="59">J.C.Cardale</collectorName>
,
<collectionCode id="ED60AE038A09FFB7106FFEFDFC55FE88" box="[865,937,332,356]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="58" pageNumber="59">ANIC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A09FFB713C9FEDEFD8FFE6B" box="[199,627,367,392]" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8BCE36C68A09FFB713C9FEDEFD8FFE6B" blockId="58.[151,1436,151,824]" box="[199,627,367,392]" pageId="58" pageNumber="59">
<emphasis id="B905EAD48A09FFB713C9FEDEFE9CFE64" bold="true" box="[199,352,367,392]" inLineHeading="true" pageId="58" pageNumber="59" reason="1">Distribution.</emphasis>
<collectingCountry id="F36676568A09FFB71269FEDEFE32FE6B" box="[359,462,367,391]" name="Australia" pageId="58" pageNumber="59">Australia</collectingCountry>
(
<collectingRegion id="49B5F8248A09FFB712D3FEDEFD94FE6B" box="[477,616,367,391]" country="Australia" name="Queensland" pageId="58" pageNumber="59">Queensland</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A09FFB713C9FE25FB2AFCDB" pageId="58" pageNumber="59" type="discussion">
<paragraph id="8BCE36C68A09FFB713C9FE25FEF8FDD0" blockId="58.[151,1436,151,824]" pageId="58" pageNumber="59">
<emphasis id="B905EAD48A09FFB713C9FE25FEC7FE40" bold="true" box="[199,315,404,428]" inLineHeading="true" pageId="58" pageNumber="59" reason="1">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C714D458A09FFB71164FE24FD1EFE40" authorityName="Terayama" authorityYear="2004" box="[618,738,404,428]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ceciliae">
<emphasis id="B905EAD48A09FFB71164FE24FD1EFE40" box="[618,738,404,428]" italics="true" pageId="58" pageNumber="59">E. ceciliae</emphasis>
</taxonomicName>
by having the vertex crest concave, the median clypeal lobe subangulate to slightly rounded, median carina of clypeus absent, eyes protuberant, but it differs from it by having the median clypeal lobe clearly shorter, the eyes large, the median carina of propodeal disc very short, whereas
<taxonomicName id="4C714D458A09FFB7168AFE6CFF12FDF4" authorityName="Terayama" authorityYear="2004" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ceciliae">
<emphasis id="B905EAD48A09FFB7168AFE6CFF12FDF4" italics="true" pageId="58" pageNumber="59">E. ceciliae</emphasis>
</taxonomicName>
has the median clypeal lobe broad and slightly long, the eyes large, and the median carina of propodeal disc long.
</paragraph>
<paragraph id="8BCE36C68A09FFB713C9FDF9FB2AFCDB" blockId="58.[151,1436,151,824]" pageId="58" pageNumber="59">
It is also similar to
<taxonomicName id="4C714D458A09FFB712AAFDF8FDC1FD8C" box="[420,573,585,608]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="wanamara" status="sp. nov.">
<emphasis id="B905EAD48A09FFB712AAFDF8FDC1FD8C" box="[420,573,585,608]" italics="true" pageId="58" pageNumber="59">E. wanamara</emphasis>
</taxonomicName>
<emphasis id="B905EAD48A09FFB7114BFDF9FD5CFD8C" bold="true" box="[581,672,584,608]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="A23657AF8A09FFB7114BFDF9FD5CFD8C" box="[581,672,584,608]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
because both species present clypeus with the median clypeal lobe without median carina, the vertex crest concave, the eyes small and protuberant. However this species has the area between the anterior margin of eyes and demarkation of median clypeal lobe short, the median clypeal lobe subangulate, short, well-defined in relation to frons and region between the anterior margin of eye and antennal torulus not angulate, whereas
<taxonomicName id="4C714D458A09FFB712F1FD68FD60FD1C" box="[511,668,729,752]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="wanamara" status="sp. nov.">
<emphasis id="B905EAD48A09FFB712F1FD68FD60FD1C" box="[511,668,729,752]" italics="true" pageId="58" pageNumber="59">E. wanamara</emphasis>
</taxonomicName>
<emphasis id="B905EAD48A09FFB711A4FD69FCF5FD1C" bold="true" box="[682,777,728,752]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="A23657AF8A09FFB711A4FD69FCF5FD1C" box="[682,777,728,752]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the area between the anterior margin of eyes and demarkation of median clypeal lobe long, the median clypeal lobe broad, the median clypeal lobe ill-defined in relation to frons and the region between the anterior margin of eye and antennal torulus angulate.
</paragraph>
</subSubSection>
</treatment>
</document>