185 lines
19 KiB
XML
185 lines
19 KiB
XML
<document id="57D7712AC09DC89CF2124EF1C90F795F" ID-DOI="10.11646/zootaxa.3608.2.1" ID-GBIF-Dataset="dc20b97c-cdf4-467d-8ce9-98e912fe060c" ID-ISSN="1175-5326" ID-Zenodo-Dep="223235" ID-ZooBank="0E56C847-5694-4315-9A51-B4F207CA4F91" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460569345297" checkinUser="plazi" docAuthor="Criscione, Francesco & Köhler, Frank" docDate="2013" docId="03D887BF001A2503929B15097594FBBE" docLanguage="en" docName="zt03608p115.pdf" docOrigin="Zootaxa 3608 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Australocosmica pallida Criscione & Köhler, 2013, n. sp." docType="treatment" docVersion="9" lastPageNumber="112" masterDocId="FFE1FFC700112508920C14147605FFF2" masterDocTitle="Six new species of Australocosmica Köhler, 2011 from the Kimberley islands, Western Australia (Mollusca: Pulmonata: Camaenidae)" masterLastPageNumber="115" masterPageNumber="101" pageNumber="112" updateTime="1698321793467" updateUser="plazi">
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<mods:mods id="7897FB87D79D7C6E04D3CB7873117857" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8464A9136724AC05928E8BE60B747FDC">
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<mods:title id="5180A946AE3890AD957ED167FD56FCBD">Six new species of Australocosmica Köhler, 2011 from the Kimberley islands, Western Australia (Mollusca: Pulmonata: Camaenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FC04000BE9D5370EF46974CD1CE8C5F3" type="personal">
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<mods:role id="38797D48D64901E8F6510FA1909B771C">
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<mods:roleTerm id="CB96FE0A650BE8F65188C8452326F9EF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="12E53F32E7861D14659B74069B5417D4">Criscione, Francesco</mods:namePart>
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</mods:name>
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<mods:name id="6B81F1A3D26EFD16E4CE6E648E081336" type="personal">
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<mods:role id="0D2EF05548A590E64F2278EF9DE0F649">
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<mods:roleTerm id="A18270D3CFB912CC2629337B43527B45">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="77DCC6F8EB80690EAE13409007503059">Köhler, Frank</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B856DD07E171D43A65B7F65791ADBFDE">text</mods:typeOfResource>
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<mods:titleInfo id="2A2D437216961CF8B2F733101FD85F52">
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<mods:title id="A71AB9AE10500DEA1DDBB5874A170071">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1AFD32A03242619E3D1513534D157FF3">
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<mods:date id="81E6636A830C25296FED9B4853C1FABA">2013</mods:date>
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<mods:detail id="3F9B5E4BC03F593A99486D37B7BDA3A4" type="volume">
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<mods:number id="BF135BE62B7858F971DE9C7B6F6A148C">3608</mods:number>
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</mods:detail>
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<mods:detail id="0EB92A7DB98F29A69C44598CDF11A894" type="issue">
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<mods:number id="2ED7565789C0D3A5EBB0D90127B22881">2</mods:number>
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<mods:start id="FE102CBB08748DE8C4C0183E7D4891EA">101</mods:start>
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<mods:end id="A2F643A72739FC97ED11CD27368A84B9">115</mods:end>
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<mods:classification id="FAA81E7E0ECC82D375CE819DD09AD4B9">journal article</mods:classification>
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<mods:identifier id="E4E207EEFBE0005415D159FEDACF3835" type="DOI">10.11646/zootaxa.3608.2.1</mods:identifier>
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<mods:identifier id="FF4E45E67C6E022B4391B410D839661E" type="GBIF-Dataset">dc20b97c-cdf4-467d-8ce9-98e912fe060c</mods:identifier>
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<mods:identifier id="819DD46F05395C3AA3A9E5B1F60B32F0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="03887D3FD39C821C8EE5059A38F206BC" type="Zenodo-Dep">223235</mods:identifier>
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<mods:identifier id="9D6599AF4EC92466502512D41D365D47" type="ZooBank">0E56C847-5694-4315-9A51-B4F207CA4F91</mods:identifier>
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</mods:mods>
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<treatment id="03D887BF001A2503929B15097594FBBE" ID-DOI="http://doi.org/10.5281/zenodo.6153553" ID-GBIF-Taxon="119558298" ID-Zenodo-Dep="6153553" LSID="urn:lsid:plazi:treatment:03D887BF001A2503929B15097594FBBE" httpUri="http://treatment.plazi.org/id/03D887BF001A2503929B15097594FBBE" lastPageNumber="112" pageId="11" pageNumber="112">
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<subSubSection id="C36B6522001A2503929B1509776BFEAA" pageId="11" pageNumber="112" type="nomenclature">
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<paragraph id="8BCE36A9001A2503929B15097400FEC5" blockId="11.[151,517,285,344]" box="[151,517,285,311]" pageId="11" pageNumber="112">
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<heading id="D08681C5001A2503929B15097400FEC5" bold="true" box="[151,517,285,311]" fontSize="11" level="1" pageId="11" pageNumber="112" reason="1">
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<emphasis id="B905EABB001A2503929B15097400FEC5" bold="true" box="[151,517,285,311]" pageId="11" pageNumber="112">
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<taxonomicName id="4C714D2A001A2503929B150977BEFEC5" ID-CoL="5VXPW" box="[151,443,285,311]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="pallida" status="sp. nov.">
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<emphasis id="B905EABB001A2503929B150977BEFEC5" bold="true" box="[151,443,285,311]" italics="true" pageId="11" pageNumber="112">Australocosmica pallida</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23657C0001A250393CE15097400FEC5" box="[450,517,285,311]" pageId="11" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BCE36A9001A2503929B152B776BFEAA" blockId="11.[151,517,285,344]" box="[151,366,319,344]" pageId="11" pageNumber="112">
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<figureCitation id="134A2A2C001A2503929B152B76DEFEA5" box="[151,219,319,343]" captionStart="FIGURE 3" captionStartId="5.[151,250,1564,1587]" captionTargetBox="[326,1260,203,1521]" captionTargetId="figure@5.[313,1279,195,1537]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Australocosmica species. A. A. bernoulliensis n. sp. (paratype FMNH 219104). B. A. buffonensis n. sp. (paratype WAM S 83007). C. A. crassicostata n. sp. (paratype FMNH 219181) D. A. nana n. sp. (paratype WAM S 83039). E. A. pallida n. sp. (paratype WAM S 83006). F. A. rotunda n. sp. (paratype WAM S 49237). Scale bar = 10 mm." httpUri="https://zenodo.org/record/223238/files/figure.png" pageId="11" pageNumber="112">Figs 3</figureCitation>
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E, 5C–D, 7A
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6522001A2503929B159C7461FD85" pageId="11" pageNumber="112" type="materials_examined">
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<paragraph id="8BCE36A9001A2503929B159C72D2FE36" blockId="11.[151,1436,391,1100]" pageId="11" pageNumber="112">
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<bibRef id="52D1A25F001A2503929B159C72D2FE36" pageId="11" pageNumber="112" type="journal article">
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||
<title id="F975DD9F001A2503929B159C744BFE36" jopScore="1.8751719" pageId="11" pageNumber="112">
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||
<emphasis id="B905EABB001A2503929B159C76FBFE52" box="[151,254,392,416]" italics="true" pageId="11" pageNumber="112">
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||
<typeStatus id="54CA880B001A2503929B159C76FBFE52" box="[151,254,392,416]" pageId="11" pageNumber="112" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
.
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<collectingCountry id="F3667639001A250393021593777DFE6D" box="[270,376,391,415]" name="Australia" pageId="11" pageNumber="112">Australia</collectingCountry>
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, WA, Kimberley, Bonaparte Archipelago, Brunswick Bay: Sta. KC-100, small island SW of Lucas Island, -15.231, 124.494 (coll. V
|
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</title>
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.
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<author id="53CCDE4C001A2503905515B974D9FE36" box="[601,732,428,452]" fns="I" npo="LF" pageId="11" pageNumber="112">Kessner, A.</author>
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<journalOrPublisher id="51B03C65001A250390EF15B87575FE36" box="[739,880,428,452]" jopScore="1.9258522" pageId="11" pageNumber="112">Longbottom</journalOrPublisher>
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,
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<part id="FFD59C34001A2503917715B87592FE36" box="[891,919,428,452]" pageId="11" pageNumber="112" type="volume">24</part>
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July
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<year id="FFD977FA001A250391D815B87208FE36" box="[980,1037,428,452]" pageId="11" pageNumber="112">1988</year>
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) (WAM
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<accessionNumber id="9422AB4A001A2503967D15B872CEFE36" box="[1137,1227,428,452]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S84056" pageId="11" pageNumber="112" type="EnaNcbi">
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S
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<pagination id="B2B8C85D001A2503967315B872CEFE36" box="[1151,1227,428,452]" pageId="11" pageNumber="112" type="pagination">84056</pagination>
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||
</accessionNumber>
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||
).
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</bibRef>
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</paragraph>
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<paragraph id="8BCE36A9001A250392CB15C57461FD85" blockId="11.[151,1436,391,1100]" pageId="11" pageNumber="112">
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||
<bibRef id="52D1A25F001A250392CB15C57461FD85" pageId="11" pageNumber="112" type="journal article">
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||
<title id="F975DD9F001A250392CB15C5773CFE1A" box="[199,313,465,488]" jopScore="0.039930373" pageId="11" pageNumber="112">
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||
<emphasis id="B905EABB001A250392CB15C5773CFE1A" box="[199,313,465,488]" italics="true" pageId="11" pageNumber="112">
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<typeStatus id="54CA880B001A250392CB15C5773CFE1A" box="[199,313,465,488]" pageId="11" pageNumber="112" type="paratype">Paratypes</typeStatus>
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</emphasis>
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</title>
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.
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<author id="53CCDE4C001A2503934B15DB77E9FE1A" box="[327,492,463,488]" fns="I" npo="LF" pageId="11" pageNumber="112">
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<collectingCountry id="F3667639001A2503934B15DB77B5FE15" box="[327,432,463,487]" name="Australia" pageId="11" pageNumber="112">Australia</collectingCountry>
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, WA
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</author>
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,
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<journalOrPublisher id="51B03C65001A250393FB15DB730DFDA6" jopScore="2.8350565" pageId="11" pageNumber="112">
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Kimberley, Bonaparte Archipelago, Brunswick Bay: same data as for
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<typeStatus id="54CA880B001A2503970315DB7371FE15" box="[1295,1396,463,487]" pageId="11" pageNumber="112" type="holotype">holotype</typeStatus>
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, 14 wet (WAM
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<accessionNumber id="9422AB4A001A2503932015E07782FDFE" box="[300,391,500,524]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83006" pageId="11" pageNumber="112" type="EnaNcbi">S83006</accessionNumber>
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); 16 wet (FMNH 219401); Sta. KC-099, small island SW of Lucas Island, -15.231, 124.483(coll. V. Kessner, A. Longbottom, 24 July 1988), 38 dry (WAM
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<accessionNumber id="9422AB4A001A250391D716037230FDDD" box="[987,1077,535,559]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83004" pageId="11" pageNumber="112" type="EnaNcbi">S83004</accessionNumber>
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); 38 dry (FMNH 219396); Sta. KC099, small island SW of Lucas Island, -15.231, 124.483 (coll. V. Kessner, A. Longbottom, 24 July
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</journalOrPublisher>
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<year id="FFD977FA001A250397031628734FFDA6" box="[1295,1354,572,596]" pageId="11" pageNumber="112">1988</year>
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),
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<part id="FFD59C34001A2503975216287369FDA6" box="[1374,1388,572,596]" pageId="11" pageNumber="112" type="volume">6</part>
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wet (WAM
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<accessionNumber id="9422AB4A001A250392E0164B7743FD85" box="[236,326,607,631]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83005" pageId="11" pageNumber="112" type="EnaNcbi">S83005</accessionNumber>
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); 6 wet (FMNH
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<pagination id="B2B8C85D001A250393F3164B745DFD85" box="[511,600,607,631]" pageId="11" pageNumber="112" type="pagination">219397</pagination>
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).
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</bibRef>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6522001A250392CB169076ECFD4D" pageId="11" pageNumber="112" type="etymology">
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<paragraph id="8BCE36A9001A250392CB169076ECFD4D" blockId="11.[151,1436,391,1100]" pageId="11" pageNumber="112">
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<emphasis id="B905EABB001A250392CB16907738FD6E" box="[199,317,644,668]" italics="true" pageId="11" pageNumber="112">Etymology</emphasis>
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. In reference to the pale shell colour, derived from “
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<emphasis id="B905EABB001A2503919E169075E9FD6E" box="[914,1004,644,668]" italics="true" pageId="11" pageNumber="112">pallidus</emphasis>
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” (Latin = pale); adjective of feminine gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6522001A250392CB16D97502FC6A" pageId="11" pageNumber="112" type="description">
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<paragraph id="8BCE36A9001A250392CB16D97502FC6A" blockId="11.[151,1436,391,1100]" pageId="11" pageNumber="112">
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<emphasis id="B905EABB001A250392CB16D9774EFD16" box="[199,331,717,740]" italics="true" pageId="11" pageNumber="112">Description</emphasis>
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. Shell (
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<figureCitation id="134A2A2C001A250393A316D877FDFD16" box="[431,504,716,740]" captionStart="FIGURE 3" captionStartId="5.[151,250,1564,1587]" captionTargetBox="[326,1260,203,1521]" captionTargetId="figure@5.[313,1279,195,1537]" captionTargetPageId="5" captionText="FIGURE 3. Shells of Australocosmica species. A. A. bernoulliensis n. sp. (paratype FMNH 219104). B. A. buffonensis n. sp. (paratype WAM S 83007). C. A. crassicostata n. sp. (paratype FMNH 219181) D. A. nana n. sp. (paratype WAM S 83039). E. A. pallida n. sp. (paratype WAM S 83006). F. A. rotunda n. sp. (paratype WAM S 49237). Scale bar = 10 mm." httpUri="https://zenodo.org/record/223238/files/figure.png" pageId="11" pageNumber="112">Fig. 3</figureCitation>
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E; 5C–D). Large, elevated and broadly conical. Whorls with slightly angulate periphery, moderately shouldered underneath suture. Axial ribs weak. Colour horn. Genitalia (
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<figureCitation id="134A2A2C001A250396E416FB732AFCF5" box="[1256,1327,751,775]" captionStart="FIGURE 7" captionStartId="10.[151,250,956,979]" captionTargetBox="[156,1429,178,930]" captionTargetId="figure@10.[151,1436,171,939]" captionTargetPageId="10" captionText="FIGURE 7. Penial anatomy of Australocosmica species. A. A. pallida n. sp. (holotype WAM S 84056). B. A. rotunda n. sp. (holotype WAM S 84061). Scale bar = 2 mm." httpUri="https://zenodo.org/record/223242/files/figure.png" pageId="11" pageNumber="112">Fig. 7</figureCitation>
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A). Penis almost half as long as anterior part of oviduct (= vagina and free oviduct). Inner penial wall supporting complex sculpture, posteriorly with three large pads bearing transverse folds and tubercles, anteriorly with longitudinal folds extending into atrium. Vagina almost half as long as penis and free oviduct. Spermoviduct twice as long as free oviduct and about 1.5 times longer than albumen gland.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6522001A250392CB17B07594FBBE" pageId="11" pageNumber="112" type="discussion">
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<paragraph id="8BCE36A9001A250392CB17B07594FBBE" blockId="11.[151,1436,391,1100]" pageId="11" pageNumber="112">
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<emphasis id="B905EABB001A250392CB17B07735FC4E" box="[199,304,932,956]" italics="true" pageId="11" pageNumber="112">Remarks.</emphasis>
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Shell colour lighter than other congeners. On average shell larger than other congeners except
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<taxonomicName id="4C714D2A001A2503978817B076F9FC12" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="augustae">
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<emphasis id="B905EABB001A2503978817B076F9FC12" italics="true" pageId="11" pageNumber="112">A. augustae</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C714D2A001A2503930117DC778DFC12" box="[269,392,968,992]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="rotunda">
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<emphasis id="B905EABB001A2503930117DC778DFC12" box="[269,392,968,992]" italics="true" pageId="11" pageNumber="112">A. rotunda</emphasis>
|
||
</taxonomicName>
|
||
,
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<taxonomicName id="4C714D2A001A2503939517DC7492FC12" box="[409,663,968,992]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="sanctumpatriciusae">
|
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<emphasis id="B905EABB001A2503939517DC7492FC12" box="[409,663,968,992]" italics="true" pageId="11" pageNumber="112">A. sanctumpatriciusae</emphasis>
|
||
</taxonomicName>
|
||
; more elevated than other congeners except
|
||
<taxonomicName id="4C714D2A001A250396AF17DC732EFC12" box="[1187,1323,968,992]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="augustae">
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<emphasis id="B905EABB001A250396AF17DC732EFC12" box="[1187,1323,968,992]" italics="true" pageId="11" pageNumber="112">A. augustae</emphasis>
|
||
</taxonomicName>
|
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; with the largest number of whorls. Sculpture finer than in
|
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<taxonomicName id="4C714D2A001A250390E017F87573FBF6" box="[748,886,1004,1028]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="augustae">
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<emphasis id="B905EABB001A250390E017F87573FBF6" box="[748,886,1004,1028]" italics="true" pageId="11" pageNumber="112">A. augustae</emphasis>
|
||
</taxonomicName>
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,
|
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<taxonomicName id="4C714D2A001A2503918517F87247FBF6" box="[905,1090,1004,1028]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="crassicostata">
|
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<emphasis id="B905EABB001A2503918517F87247FBF6" box="[905,1090,1004,1028]" italics="true" pageId="11" pageNumber="112">A. crassicostata</emphasis>
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</taxonomicName>
|
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,
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<taxonomicName id="4C714D2A001A2503965917F872B7FBF6" box="[1109,1202,1004,1028]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="nana">
|
||
<emphasis id="B905EABB001A2503965917F872B7FBF6" box="[1109,1202,1004,1028]" italics="true" pageId="11" pageNumber="112">A. nana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C714D2A001A250396CA17F87346FBF6" box="[1222,1347,1004,1028]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="rotunda">
|
||
<emphasis id="B905EABB001A250396CA17F87346FBF6" box="[1222,1347,1004,1028]" italics="true" pageId="11" pageNumber="112">A. rotunda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C714D2A001A2503978817F87777FBDA" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="sanctumpatriciusae">
|
||
<emphasis id="B905EABB001A2503978817F87777FBDA" italics="true" pageId="11" pageNumber="112">A. sanctumpatriciusae</emphasis>
|
||
</taxonomicName>
|
||
. Inner penial wall sculpture similar to that of
|
||
<taxonomicName id="4C714D2A001A25039176100475D7FBDA" box="[890,978,1040,1064]" class="Gastropoda" family="Camaenidae" genus="Australocosmica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="112" phylum="Mollusca" rank="species" species="nana">
|
||
<emphasis id="B905EABB001A25039176100475D7FBDA" box="[890,978,1040,1064]" italics="true" pageId="11" pageNumber="112">A. nana</emphasis>
|
||
</taxonomicName>
|
||
but with less developed transverse folds on lateral pads.This is Solem’s (1991) manuscript species ‘NSP 76’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |