583 lines
73 KiB
XML
583 lines
73 KiB
XML
<document id="E0B2C9BE60D35076F2F746E973EE1893" ID-CLB-Dataset="54709" ID-DOI="10.5852/ejt.2023.875.2147" ID-GBIF-Dataset="3351d966-cbeb-42a8-ac49-627b5fc261c7" ID-ISSN="2118-9773" ID-Zenodo-Dep="8083773" ID-ZooBank="32FA008C-B35D-483C-9DBE-1DCCD0868FAC" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687788869828" checkinUser="felipe" docAuthor="Fasbender, Andrew" docDate="2023" docId="03D887BE817936252A03FE4544250CC8" docLanguage="en" docName="ejt-875_159-202.pdf" docOrigin="European Journal of Taxonomy 875" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ceratoculicoides borkenti Fasbender 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="174" masterDocId="FFE1FFC68175362A284DFF9741030970" masterDocTitle="Revision of the New World Ceratoculicoides Wirth & Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)" masterLastPageNumber="202" masterPageNumber="159" pageNumber="171" updateTime="1687807085140" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B93B7ED5AB79D6E698B90E19825F1245">Revision of the New World Ceratoculicoides Wirth & Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)</mods:title>
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<mods:namePart id="F7CFE4BE9770ADC667857CCA5E4F4BA4">Fasbender, Andrew</mods:namePart>
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<mods:affiliation id="BC0D2DF50A48D277180380CA84184A14">Rhithron Associates Inc., 33 Fort Missoula Road, Missoula, MT 59847, USA.</mods:affiliation>
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<mods:nameIdentifier id="F5AD5D4D8E11D0B147B2C3B1E11ABFE6" type="email">afasbender@rhithron.com</mods:nameIdentifier>
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<mods:title id="ADC4DCF18FF7E2868782376BB2BFB2E9">European Journal of Taxonomy</mods:title>
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<mods:part id="F1F550C52A4574608B7DB8F36CA00A68">
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<mods:date id="262252E38CCF4F70B6D47B078AF14C5A">2023</mods:date>
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<mods:number id="EC309D640B9C7659FB0EE161FDE46EC6">2023-06-26</mods:number>
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<treatment id="03D887BE817936252A03FE4544250CC8" ID-DOI="http://doi.org/10.5281/zenodo.8083795" ID-Zenodo-Dep="8083795" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8" httpUri="http://treatment.plazi.org/id/03D887BE817936252A03FE4544250CC8" lastPageId="15" lastPageNumber="174" pageId="12" pageNumber="171">
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<subSubSection id="C36B6523817936262A03FE4542E5089D" box="[590,998,466,493]" pageId="12" pageNumber="171" type="nomenclature">
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<paragraph id="8BCE36A8817936262A03FE4542E5089D" blockId="12.[354,1233,466,563]" box="[590,998,466,493]" pageId="12" pageNumber="171">
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<heading id="D08681C4817936262A03FE4542E5089D" box="[590,998,466,493]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="171" reason="2">
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<taxonomicName id="4C714D2B817936262A03FE454287089D" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[590,900,466,493]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="borkenti" status="sp. nov.">
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<emphasis id="B905EABA817936262A03FE454287089D" bold="true" box="[590,900,466,493]" italics="true" pageId="12" pageNumber="171">Ceratoculicoidesborkenti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23657C1817936262BC6FE4442E5089D" box="[907,998,467,493]" pageId="12" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B652381793626292FFE62429A0B43" pageId="12" pageNumber="171" type="description">
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<paragraph id="8BCE36A881793626292FFE6245D20B60" blockId="12.[354,1233,466,563]" box="[354,1233,501,528]" pageId="12" pageNumber="171">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD70C7D817936262AC9FE6245D20B60" box="[644,1233,501,528]" pageId="12" pageNumber="171">545F2DF7-08BE-4ACD-B8D8-B18BC4F57A7D</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BCE36A8817936262AD6FD8E429A0B43" blockId="12.[354,1233,466,563]" box="[667,921,536,563]" pageId="12" pageNumber="171">
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<figureCitation id="134A2A2D817936262AD6FD8E43F00B43" box="[667,755,536,563]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[190,1398,1040,1891]" captionTargetId="figure-160@10.[190,1398,1040,1891]" captionTargetPageId="10" captionText="Fig. 5. Male heads II. Ceratoculicoidesmoravicus group. a. C. sp. M1. b. C.moravicusKnoz, 1987. c. C.propinquus sp. nov. d. C.grogani sp. nov. e. C.borkenti sp. nov. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083789" httpUri="https://zenodo.org/record/8083789/files/figure.png" pageId="12" pageNumber="171">Figs 5e</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A2D817936262AB2FD8E421E0B43" box="[767,797,537,563]" captionStart="Fig" captionStartId="12.[189,230,1963,1989]" captionTargetBox="[196,1390,1017,1903]" captionTargetId="figure-171@12.[196,1390,1017,1903]" captionTargetPageId="12" captionText="Fig. 7. Female heads II.Ceratoculicoidesmoravicus group.a. C.aliciae Huerta & Borkent, 2005. b. C. aliciae antenna. c. C. borkenti sp. nov. d. C.grogani sp. nov. e. C.moravicusKnoz, 1987. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083793" httpUri="https://zenodo.org/record/8083793/files/figure.png" pageId="12" pageNumber="171">7c</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A2D817936262B64FD8E42610B43" box="[809,866,537,563]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="12" pageNumber="171">9d–f</figureCitation>
|
||
,
|
||
<figureCitation id="134A2A2D817936262B23FD8E429A0B43" box="[878,921,537,563]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta & Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="12" pageNumber="171">11b</figureCitation>
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B65238179362728F0FDCD40DB08E1" lastPageId="13" lastPageNumber="172" pageId="12" pageNumber="171" type="diagnosis">
|
||
<paragraph id="8BCE36A88179362628F0FDCD40350B04" blockId="12.[189,310,602,628]" box="[189,310,602,628]" pageId="12" pageNumber="171">
|
||
<heading id="D08681C48179362628F0FDCD40350B04" bold="true" box="[189,310,602,628]" fontSize="11" level="3" pageId="12" pageNumber="171" reason="3">
|
||
<emphasis id="B905EABA8179362628F0FDCD40350B04" bold="true" box="[189,310,602,628]" pageId="12" pageNumber="171">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88179362628A7FD1E40290BD3" blockId="12.[189,1399,649,781]" box="[234,298,649,675]" pageId="12" pageNumber="171">
|
||
<emphasis id="B905EABA8179362628A7FD1E40290BD3" bold="true" box="[234,298,649,675]" pageId="12" pageNumber="171">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88179362628F0FD3B44580A7C" blockId="12.[189,1399,649,781]" pageId="12" pageNumber="171">
|
||
<taxonomicName id="4C714D2B8179362628F0FD3B40EF0BB6" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[189,492,684,710]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="borkenti" status="sp. nov.">
|
||
<emphasis id="B905EABA8179362628F0FD3B40EF0BB6" box="[189,492,684,710]" italics="true" pageId="12" pageNumber="171">Ceratoculicoides borkenti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C18179362629BAFD3B43570BB6" box="[503,596,684,710]" pageId="12" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from congeners by the following combination of characters: femora and tibiae brown; apices of parameres acute, tapering distally; aedeagus lateral margins an evenly concave arc, nearly entire medial portion of aedeagus hyaline, expanding anteriorly (
|
||
<figureCitation id="134A2A2D817936262CB3FD65444E0A7C" box="[1278,1357,754,780]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="12" pageNumber="171">Fig. 9f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0E66208179362628F0F83C44520E99" ID-DOI="http://doi.org/10.5281/zenodo.8083793" ID-Zenodo-Dep="8083793" httpUri="https://zenodo.org/record/8083793/files/figure.png" pageId="12" pageNumber="171" startId="12.[189,230,1963,1989]" targetBox="[196,1390,1017,1903]" targetPageId="12">
|
||
<paragraph id="8BCE36A88179362628F0F83C44520E99" blockId="12.[189,1396,1963,2025]" pageId="12" pageNumber="171">
|
||
<emphasis id="B905EABA8179362628F0F83C40060EB5" bold="true" box="[189,261,1963,1989]" pageId="12" pageNumber="171">Fig. 7.</emphasis>
|
||
Female heads II.
|
||
<emphasis id="B905EABA81793626298DF83C43FA0EB5" box="[448,761,1963,1989]" italics="true" pageId="12" pageNumber="171">Ceratoculicoidesmoravicus</emphasis>
|
||
group.
|
||
<emphasis id="B905EABA817936262B05F83C42550EB5" bold="true" box="[840,854,1963,1989]" pageId="12" pageNumber="171">a</emphasis>
|
||
.
|
||
<taxonomicName id="4C714D2B817936262B29F83B45DC0EB6" authority="Huerta & Borkent, 2005" authorityName="Huerta & Borkent" authorityYear="2005" box="[868,1247,1963,1990]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="aliciae">
|
||
<emphasis id="B905EABA817936262B29F83B42CC0EB5" box="[868,975,1963,1990]" italics="true" pageId="12" pageNumber="171">C.aliciae</emphasis>
|
||
<bibRefCitation id="EFE04B59817936262B9EF83B45DC0EB6" author="Huerta H. & Borkent A." box="[979,1247,1963,1990]" pageId="12" pageNumber="171" pagination="111 - 119" refId="ref23309" refString="Huerta H. & Borkent A. 2005. A new species and first record of Ceratoculicoides Wirth & Ratanaworabhan from the Neotropical region and new species and records of Brachypogon Kieffer from Mexico (Diptera: Ceratopogonidae). Folia Entomologica Mexicana 44: 111 - 119." type="journal article" year="2005">Huerta & Borkent, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B905EABA817936262CAAF83C45F40EB5" bold="true" box="[1255,1271,1963,1989]" pageId="12" pageNumber="171">b</emphasis>
|
||
.
|
||
<taxonomicName id="4C714D2B817936262D48F83B44770EB5" authorityName="Huerta & Borkent" authorityYear="2005" box="[1285,1396,1963,1990]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="aliciae">
|
||
<emphasis id="B905EABA817936262D48F83B44770EB5" box="[1285,1396,1963,1990]" italics="true" pageId="12" pageNumber="171">C. aliciae</emphasis>
|
||
</taxonomicName>
|
||
antenna.
|
||
<emphasis id="B905EABA81793626296CF859402E0E98" bold="true" box="[289,301,1998,2024]" pageId="12" pageNumber="171">c</emphasis>
|
||
.
|
||
<taxonomicName id="4C714D2B817936262976F85840BC0E98" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[315,447,1998,2025]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="borkenti" status="sp. nov.">
|
||
<emphasis id="B905EABA817936262976F85840BC0E98" box="[315,447,1998,2025]" italics="true" pageId="12" pageNumber="171">C. borkenti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C181793626298BF858431F0E99" box="[454,540,1999,2025]" pageId="12" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B905EABA817936262A6CF85943320E98" bold="true" box="[545,561,1998,2024]" pageId="12" pageNumber="171">d</emphasis>
|
||
.
|
||
<taxonomicName id="4C714D2B817936262A72F85843B50E98" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[575,694,1999,2025]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="grogani" status="sp. nov.">
|
||
<emphasis id="B905EABA817936262A72F85843B50E98" box="[575,694,1999,2025]" italics="true" pageId="12" pageNumber="171">C.grogani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C1817936262AF0F85842100E99" box="[701,787,1999,2025]" pageId="12" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B905EABA817936262B54F85942260E98" bold="true" box="[793,805,1998,2024]" pageId="12" pageNumber="171">e</emphasis>
|
||
.
|
||
<taxonomicName id="4C714D2B817936262B7EF858455B0E98" authority="Knoz, 1987" authorityName="Knoz" authorityYear="1987" box="[819,1112,1998,2025]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="171" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B905EABA817936262B7EF85842CC0E98" box="[819,975,1999,2025]" italics="true" pageId="12" pageNumber="171">C.moravicus</emphasis>
|
||
<bibRefCitation id="EFE04B59817936262B83F858455B0E98" author="Knoz J." box="[974,1112,1998,2025]" pageId="12" pageNumber="171" pagination="388 - 392" refId="ref23379" refString="Knoz J. 1987. Description of a new species of Ceratoculicoides (Diptera, Ceratopogonidae) from Czechoslovakia and a review of the genus. Acta Entomologica Bohemoslovaca 84: 388 - 392." type="journal article" year="1987">Knoz, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
. Scale bars = 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCE36A88178362728A7FE9E40460853" blockId="13.[189,1398,265,401]" box="[234,325,265,291]" pageId="13" pageNumber="172">
|
||
<emphasis id="B905EABA8178362728A7FE9E40460853" bold="true" box="[234,325,265,291]" pageId="13" pageNumber="172">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0FEBA40DB08E1" blockId="13.[189,1398,265,401]" pageId="13" pageNumber="172">
|
||
Only species of
|
||
<taxonomicName id="4C714D2B817836272933FEBA43450837" authorityName=", Wirth & Ratanaworabhan" authorityYear="1971" box="[382,582,301,327]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EABA817836272933FEBA43450837" box="[382,582,301,327]" italics="true" pageId="13" pageNumber="172">Ceratoculicoides</emphasis>
|
||
</taxonomicName>
|
||
with the following combination of characters: FR 1.41–1.47; femora and tibiae brown, wing length
|
||
<quantity id="4C899B4D817836272A62FEC543A8081C" box="[559,683,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="13" pageNumber="172" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
|
||
; 2 major spermathecae, largest 52–67; medial margin of 9
|
||
<superScript id="7C049BE0817836272D24FEC64475082F" attach="none" box="[1385,1398,337,351]" fontSize="6" pageId="13" pageNumber="172">th</superScript>
|
||
sternite deeply concave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B65238178362728F0FE2E42C70B58" pageId="13" pageNumber="172" type="etymology">
|
||
<paragraph id="8BCE36A88178362728F0FE2E404708A3" blockId="13.[189,324,441,467]" box="[189,324,441,467]" pageId="13" pageNumber="172">
|
||
<heading id="D08681C48178362728F0FE2E404708A3" bold="true" box="[189,324,441,467]" fontSize="11" level="3" pageId="13" pageNumber="172" reason="3">
|
||
<emphasis id="B905EABA8178362728F0FE2E404708A3" bold="true" box="[189,324,441,467]" pageId="13" pageNumber="172">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0FE7E42C70B58" blockId="13.[189,1399,489,552]" pageId="13" pageNumber="172">
|
||
This species is named in honor of Art Borkent for encouraging my study of
|
||
<taxonomicName id="4C714D2B817836272C7CFE7E45FA0B73" authorityName="Newman" authorityYear="1834" box="[1073,1273,489,515]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="172" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
|
||
, including suggesting this project after I collected a specimen of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B65238178362728F0FDC745330DA1" pageId="13" pageNumber="172" type="materials_examined">
|
||
<paragraph id="8BCE36A88178362728F0FDC740A90B1A" blockId="13.[189,426,592,618]" box="[189,426,592,618]" pageId="13" pageNumber="172">
|
||
<heading id="D08681C48178362728F0FDC740A90B1A" bold="true" box="[189,426,592,618]" fontSize="11" level="3" pageId="13" pageNumber="172" reason="3">
|
||
<emphasis id="B905EABA8178362728F0FDC740A90B1A" bold="true" box="[189,426,592,618]" pageId="13" pageNumber="172">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B193CF58178362728A7FD1644110BCF" collectingDate="1987-05-01" collectionCode="CNCI" collectorName="A. Borkent" country="United States of America" county="Santa Cruz Co." location="Nogales" pageId="13" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="Arizona" typeStatus="holotype">
|
||
<paragraph id="8BCE36A88178362728A7FD16405F0BEB" blockId="13.[189,1298,641,704]" box="[234,348,641,667]" pageId="13" pageNumber="172">
|
||
<emphasis id="B905EABA8178362728A7FD16405F0BEB" bold="true" box="[234,348,641,667]" pageId="13" pageNumber="172">
|
||
<typeStatus id="54CA880A8178362728A7FD16405F0BEB" box="[234,348,641,667]" pageId="13" pageNumber="172">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0FD3244110BCF" blockId="13.[189,1298,641,704]" box="[189,1298,677,704]" pageId="13" pageNumber="172">
|
||
<collectingCountry id="F36676388178362728F0FD3241F50BCF" box="[189,246,677,703]" name="United States of America" pageId="13" pageNumber="172">USA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D77FD21817836272942FD31402B0BB0" box="[271,296,678,704]" pageId="13" pageNumber="172" type="male">♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B5F84A81783627297EFD3240960BCF" box="[307,405,677,703]" country="United States of America" name="Arizona" pageId="13" pageNumber="172">Arizona</collectingRegion>
|
||
,
|
||
<collectingCounty id="62AF4E248178362729EDFD3243560BCF" box="[416,597,677,703]" pageId="13" pageNumber="172">Santa Cruz Co.</collectingCounty>
|
||
,
|
||
<locationDeviation id="9A7C2F8A817836272A2CFD3243EE0BCF" bearing="270.0" box="[609,749,677,704]" distance="24000.0" pageId="13" pageNumber="172">
|
||
<quantity id="4C899B4D817836272A2CFD3243A80BCF" box="[609,683,677,703]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="172" unit="km" value="24.0">24 km</quantity>
|
||
W of
|
||
</locationDeviation>
|
||
<location id="8EAE6073817836272AB9FD32425B0BCF" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272AB9FD32425B0BCF" box="[756,856,677,703]" country="United States of America" county="Santa Cruz Co." name="Nogales" pageId="13" pageNumber="172" stateProvince="Arizona">Nogales</location>
|
||
;
|
||
<date id="FFCF1068817836272B29FD3242F60BCF" box="[868,1013,677,704]" pageId="13" pageNumber="172" value="1987-05-01">
|
||
<collectingDate id="EF8BE980817836272B29FD3242F60BCF" box="[868,1013,677,704]" pageId="13" pageNumber="172" value="1987-05-01">1 May 1987</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E817836272C4DFD3245810BCF" box="[1024,1154,677,703]" pageId="13" pageNumber="172">A. Borkent</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D817836272C89FD32440D0BCF" box="[1220,1294,677,703]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="13" pageNumber="172">CNCI</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B193CF58178362728A7FD7F429C0A57" collectionCode="CNCI" country="United States of America" location="USA" pageId="13" pageNumber="172" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
|
||
<paragraph id="8BCE36A88178362728A7FD7F40640A72" blockId="13.[189,927,744,807]" box="[234,359,744,770]" pageId="13" pageNumber="172">
|
||
<emphasis id="B905EABA8178362728A7FD7F40640A72" bold="true" box="[234,359,744,770]" pageId="13" pageNumber="172">
|
||
<typeStatus id="54CA880A8178362728A7FD7F40640A72" box="[234,359,744,770]" pageId="13" pageNumber="172">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0FC9A429C0A57" blockId="13.[189,927,744,807]" box="[189,927,780,807]" pageId="13" pageNumber="172">
|
||
<collectingCountry id="F36676388178362728F0FC9A41F50A57" box="[189,246,781,807]" name="United States of America" pageId="13" pageNumber="172">USA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D77FD21817836272942FC9A403E0A57" box="[271,317,781,807]" pageId="13" pageNumber="172" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D77FD21817836272907FC9A408E0A57" box="[330,397,781,807]" pageId="13" pageNumber="172" type="female">3 ♀♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED60AE6D817836272B1CFC9A42980A57" box="[849,923,781,807]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="13" pageNumber="172">CNCI</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BCE36A88178362728A7FCD840A50A19" blockId="13.[189,1380,847,911]" box="[234,422,847,873]" pageId="13" pageNumber="172">
|
||
<emphasis id="B905EABA8178362728A7FCD840A50A19" bold="true" box="[234,422,847,873]" pageId="13" pageNumber="172">Other material</emphasis>
|
||
</paragraph>
|
||
<materialsCitation id="3B193CF58178362728F0FCE341F60AA1" collectingDate="1987-04-29" collectingDateMax="1991-07-08" collectingDateMin="1987-04-29" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" county="Santa Cruz Co." location="Lake Cowichan" pageId="13" pageNumber="172" specimenCount="4" specimenCount-male="4" stateProvince="British Columbia">
|
||
<paragraph id="8BCE36A88178362728F0FCE344670AFE" blockId="13.[189,1380,847,911]" box="[189,1380,884,911]" pageId="13" pageNumber="172">
|
||
<collectingCountry id="F36676388178362728F0FCE3403A0AFE" box="[189,313,884,910]" name="Canada" pageId="13" pageNumber="172">CANADA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D77FD2181783627291FFCE340820AFF" box="[338,385,884,911]" pageId="13" pageNumber="172" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B5F84A8178362729C0FCE343580AFE" box="[397,603,884,910]" country="Canada" name="British Columbia" pageId="13" pageNumber="172">British Columbia</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A7C2F8A817836272A2AFCE343F10AFE" bearing="270.0" box="[615,754,884,911]" distance="25000.0" pageId="13" pageNumber="172">
|
||
<quantity id="4C899B4D817836272A2AFCE343B20AFE" box="[615,689,884,911]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="172" unit="km" value="25.0">25 km</quantity>
|
||
W of
|
||
</locationDeviation>
|
||
<location id="8EAE6073817836272AB7FCE342B40AFE" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272AB7FCE342B40AFE" box="[762,951,884,910]" country="Canada" county="Santa Cruz Co." name="Lake Cowichan" pageId="13" pageNumber="172" stateProvince="British Columbia">Lake Cowichan</location>
|
||
;
|
||
<date id="FFCF1068817836272B8FFCE345440AFE" box="[962,1095,884,910]" pageId="13" pageNumber="172" value="1991-07-08">
|
||
<collectingDate id="EF8BE980817836272B8FFCE345440AFE" box="[962,1095,884,910]" pageId="13" pageNumber="172" value="1991-07-08">8 Jul. 1991</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E817836272C1FFCE345D60AFE" box="[1106,1237,884,910]" pageId="13" pageNumber="172">A. Borkent</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D817836272D5BFCE344630AFE" box="[1302,1376,884,910]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="13" pageNumber="172">CNCI</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0FC2045330DA1" blockId="13.[189,1399,951,1234]" pageId="13" pageNumber="172">
|
||
<collectingCountry id="F36676388178362728F0FC2041F60AA1" box="[189,245,951,977]" name="United States of America" pageId="13" pageNumber="172">USA</collectingCountry>
|
||
•
|
||
<materialsCitation id="3B193CF581783627295DFC2044670AA1" box="[272,1380,951,977]" collectingDate="1987-04-29" collectingDateMax="1991-07-08" collectingDateMin="1987-04-29" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" county="Santa Cruz Co." location="Lake Cowichan" pageId="13" pageNumber="172" specimenCount="4" specimenCount-male="4" stateProvince="British Columbia">
|
||
<specimenCount id="9D77FD2181783627295DFC2040550AA1" box="[272,342,951,977]" pageId="13" pageNumber="172" type="male">3 ♂♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B5F84A81783627292CFC2040C20AA1" box="[353,449,951,977]" country="United States of America" name="Arizona" pageId="13" pageNumber="172">Arizona</collectingRegion>
|
||
,
|
||
<collectingCounty id="62AF4E24817836272980FC2043820AA1" box="[461,641,951,977]" pageId="13" pageNumber="172">Santa Cruz Co.</collectingCounty>
|
||
,
|
||
<locationDeviation id="9A7C2F8A817836272AC3FC2042280AA1" bearing="225.0" box="[654,811,951,977]" distance="11000.0" pageId="13" pageNumber="172">
|
||
<quantity id="4C899B4D817836272AC3FC2043D40AA1" box="[654,727,951,977]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="172" unit="km" value="11.0">11 km</quantity>
|
||
SW of
|
||
</locationDeviation>
|
||
<location id="8EAE6073817836272B7EFC2042AB0AA1" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272B7EFC2042AB0AA1" box="[819,936,951,977]" country="Canada" county="Santa Cruz Co." name="Patagonia" pageId="13" pageNumber="172" stateProvince="British Columbia">Patagonia</location>
|
||
;
|
||
<date id="FFCF1068817836272BF8FC2045530AA1" box="[949,1104,951,977]" pageId="13" pageNumber="172" value="1987-04-29">
|
||
<collectingDate id="EF8BE980817836272BF8FC2045530AA1" box="[949,1104,951,977]" pageId="13" pageNumber="172" value="1987-04-29">29 Apr. 1987</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E817836272C16FC2045DF0AA1" box="[1115,1244,951,977]" pageId="13" pageNumber="172">A. Borkent</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D817836272D52FC2044670AA1" box="[1311,1380,951,977]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="13" pageNumber="172">CNCI</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B193CF58178362728F0FC4B42FA0A86" box="[189,1017,987,1014]" collectingDate="1987-04-23" collectionCode="CNCI" location="Portal" pageId="13" pageNumber="172" specimenCount="2" specimenCount-male="2">
|
||
<specimenCount id="9D77FD218178362728F0FC4B40020A86" box="[189,257,988,1014]" pageId="13" pageNumber="172" type="male">2 ♂♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<location id="8EAE6073817836272AF2FC4C42040A85" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272AF2FC4C42040A85" box="[703,775,987,1013]" name="Portal" pageId="13" pageNumber="172">Portal</location>
|
||
;
|
||
<date id="FFCF1068817836272B5FFC4B42AA0A86" box="[786,937,988,1014]" pageId="13" pageNumber="172" value="1987-04-23">
|
||
<collectingDate id="EF8BE980817836272B5FFC4B42AA0A86" box="[786,937,988,1014]" pageId="13" pageNumber="172" value="1987-04-23">23 Apr. 1987</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode id="ED60AE6D817836272BF9FC4B42FA0A86" box="[948,1017,988,1014]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="13" pageNumber="172">CNCI</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B193CF5817836272C42FC4B42A90D6A" collectingDate="1987-05-08" collectionCode="CNCI" county="Yavapai Co." location="Sedona" pageId="13" pageNumber="172" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D77FD21817836272C42FC4B45380A86" box="[1039,1083,988,1014]" pageId="13" pageNumber="172" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectingCounty id="62AF4E24817836272970FB9740CE0D6A" box="[317,461,1024,1050]" pageId="13" pageNumber="172">Yavapai Co.</collectingCounty>
|
||
,
|
||
<locationDeviation id="9A7C2F8A817836272994FB97435F0D6A" bearing="180.0" box="[473,604,1024,1050]" distance="19000.0" pageId="13" pageNumber="172">
|
||
<quantity id="4C899B4D817836272994FB9743200D6A" box="[473,547,1024,1050]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="172" unit="km" value="19.0">19 km</quantity>
|
||
S of
|
||
</locationDeviation>
|
||
<location id="8EAE6073817836272A2EFB9743B80D6A" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272A2EFB9743B80D6A" box="[611,699,1024,1050]" county="Yavapai Co." name="Sedona" pageId="13" pageNumber="172">Sedona</location>
|
||
;
|
||
<date id="FFCF1068817836272A85FB97425A0D6A" box="[712,857,1024,1051]" pageId="13" pageNumber="172" value="1987-05-08">
|
||
<collectingDate id="EF8BE980817836272A85FB97425A0D6A" box="[712,857,1024,1051]" pageId="13" pageNumber="172" value="1987-05-08">8 May 1987</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode id="ED60AE6D817836272B28FB9742A90D6A" box="[869,938,1024,1050]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="13" pageNumber="172">CNCI</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B193CF5817836272B89FB97444B0D4F" collectingDate="1970-04-05" collectionCode="USNM" collectorName="L. Lapre" county="Riverside Co." location="Philip L. Boyd Deep Canyon Desert Research Center" municipality="Philip L. Boyd Deep Canyon Desert Research Center" pageId="13" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="California">
|
||
<specimenCount id="9D77FD21817836272B89FB9742F00D6B" box="[964,1011,1024,1051]" pageId="13" pageNumber="172" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B5F84A817836272C4CFB97457B0D6A" box="[1025,1144,1024,1050]" country="United States of America" name="California" pageId="13" pageNumber="172">California</collectingRegion>
|
||
,
|
||
<collectingCounty id="62AF4E24817836272CC9FB9744260D6A" box="[1156,1317,1024,1050]" pageId="13" pageNumber="172">Riverside Co.</collectingCounty>
|
||
,
|
||
<location id="8EAE6073817836272D7FFB9743DA0D4F" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272D7FFB9743DA0D4F" county="Riverside Co." municipality="Philip L. Boyd Deep Canyon Desert Research Center" name="Philip L. Boyd Deep Canyon Desert Research Center" pageId="13" pageNumber="172" stateProvince="California">Philip L. Boyd Deep Canyon Desert Research Center</location>
|
||
,
|
||
<location id="8EAE6073817836272AA8FBB242AE0D4F" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272AA8FBB242AE0D4F" box="[741,941,1061,1087]" county="Riverside Co." municipality="Philip L. Boyd Deep Canyon Desert Research Center" name="Horsethief Creek" pageId="13" pageNumber="172" stateProvince="California">Horsethief Creek</location>
|
||
;
|
||
<date id="FFCF1068817836272BF4FBB345470D4F" box="[953,1092,1060,1087]" pageId="13" pageNumber="172" value="1970-04-05">
|
||
<collectingDate id="EF8BE980817836272BF4FBB345470D4F" box="[953,1092,1060,1087]" pageId="13" pageNumber="172" value="1970-04-05">5 Apr. 1970</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E817836272C1DFBB245B00D4F" box="[1104,1203,1061,1087]" pageId="13" pageNumber="172">L. Lapre</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D817836272CB9FBB2444B0D4F" box="[1268,1352,1061,1087]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="172" type="Museum">USNM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B193CF5817836272D25FBB2442B0D13" collectingDate="1955-04-03" collectionCode="USNM" collectorName="W. R. Mason" location="Thousand Palms" municipality="Thousand Palms" pageId="13" pageNumber="172" specimenCount="2" specimenCount-male="2">
|
||
<specimenCount id="9D77FD21817836272D25FBB241EF0D14" pageId="13" pageNumber="172" type="male">2 ♂♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectingMunicipality id="6BAAACD2817836272AE5FBDE42690D13" box="[680,874,1097,1123]" pageId="13" pageNumber="172">Thousand Palms</collectingMunicipality>
|
||
;
|
||
<date id="FFCF1068817836272B39FBDE42FE0D14" box="[884,1021,1097,1124]" pageId="13" pageNumber="172" value="1955-04-03">
|
||
<collectingDate id="EF8BE980817836272B39FBDE42FE0D14" box="[884,1021,1097,1124]" pageId="13" pageNumber="172" value="1955-04-03">3 Apr. 1955</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E817836272C4AFBDD45950D14" box="[1031,1174,1098,1124]" pageId="13" pageNumber="172">W.R. Mason</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D817836272C99FBDE442B0D13" box="[1236,1320,1097,1123]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="172" type="Museum">USNM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B193CF5817836272D09FBDE431C0DDD" collectingDate="2017-07-15" collectionCode="AFPC" collectorName="A. Fasbender" county="Idaho Co." latitude="46.5149" location="Imnamatnoon Creek" longLatPrecision="6" longitude="-114.7635" municipality="Clearwater National Forest" pageId="13" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="Idaho">
|
||
<specimenCount id="9D77FD21817836272D09FBDE44710D14" box="[1348,1394,1097,1124]" pageId="13" pageNumber="172" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B5F84A8178362728F0FBF940020DF8" box="[189,257,1134,1160]" country="United States of America" name="Idaho" pageId="13" pageNumber="172">Idaho</collectingRegion>
|
||
,
|
||
<collectingCounty id="62AF4E24817836272942FBF940850DF8" box="[271,390,1134,1160]" pageId="13" pageNumber="172">Idaho Co.</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BAAACD28178362729D9FBF943D70DF8" box="[404,724,1134,1160]" pageId="13" pageNumber="172">Clearwater National Forest</collectingMunicipality>
|
||
,
|
||
<location id="8EAE6073817836272AAFFBF942D70DF8" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272AAFFBF942D70DF8" box="[738,980,1134,1160]" county="Idaho Co." latitude="46.5149" longLatPrecision="6" longitude="-114.7635" municipality="Clearwater National Forest" name="Imnamatnoon Creek" pageId="13" pageNumber="172" stateProvince="Idaho">Imnamatnoon Creek</location>
|
||
;
|
||
<geoCoordinate id="EE45506F817836272BAFFBF9456E0DF8" box="[994,1133,1134,1161]" degrees="46.5149" direction="north" orientation="latitude" pageId="13" pageNumber="172" precision="5" value="46.5149">46.5149° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE45506F817836272C36FBF9441B0DF8" box="[1147,1304,1134,1161]" degrees="114.7635" direction="west" orientation="longitude" pageId="13" pageNumber="172" precision="5" value="-114.7635">114.7635° W</geoCoordinate>
|
||
;
|
||
<date id="FFCF1068817836272D6AFBF941F90DDD" pageId="13" pageNumber="172" value="2017-07-15">
|
||
<collectingDate id="EF8BE980817836272D6AFBF941F90DDD" pageId="13" pageNumber="172" value="2017-07-15">15 Jul. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E81783627294EFB0440980DDC" box="[259,411,1170,1197]" pageId="13" pageNumber="172">A. Fasbender</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D81783627299AFB04431C0DDD" box="[471,543,1171,1197]" pageId="13" pageNumber="172">AFPC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B193CF5817836272A7BFB0445330DA1" collectingDate="2021-06-26" collectionCode="AFPC" collectorName="A. Fasbender" county="Missoula Co." latitude="46.7125" location="Lolo National Forest" longLatPrecision="6" longitude="-114.5324" pageId="13" pageNumber="172" specimenCount="1" specimenCount-male="1" stateProvince="Montana">
|
||
<specimenCount id="9D77FD21817836272A7BFB0443610DDD" box="[566,610,1171,1197]" pageId="13" pageNumber="172" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49B5F84A817836272A23FB0443D40DDD" box="[622,727,1171,1197]" country="United States of America" name="Montana" pageId="13" pageNumber="172">Montana</collectingRegion>
|
||
,
|
||
<collectingCounty id="62AF4E24817836272AAFFB05427F0DDD" box="[738,892,1170,1197]" pageId="13" pageNumber="172">Missoula Co.</collectingCounty>
|
||
,
|
||
<location id="8EAE6073817836272BCAFB0545740DDD" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272BCAFB0545740DDD" box="[903,1143,1170,1197]" county="Missoula Co." latitude="46.7125" longLatPrecision="6" longitude="-114.5324" name="Lolo National Forest" pageId="13" pageNumber="172" stateProvince="Montana">Lolo National Forest</location>
|
||
,
|
||
<location id="8EAE6073817836272CCCFB0444750DDC" LSID="urn:lsid:plazi:treatment:03D887BE817936252A03FE4544250CC8:8EAE6073817836272CCCFB0444750DDC" box="[1153,1398,1170,1197]" county="Missoula Co." latitude="46.7125" longLatPrecision="6" longitude="-114.5324" name="East Fork Lolo Creek" pageId="13" pageNumber="172" stateProvince="Montana">East Fork Lolo Creek</location>
|
||
at
|
||
<locationDeviation id="9A7C2F8A817836272895FB2040270DA1" box="[216,292,1207,1233]" pageId="13" pageNumber="172">US 12</locationDeviation>
|
||
;
|
||
<geoCoordinate id="EE45506F81783627297DFB2040BB0DA2" box="[304,440,1207,1234]" degrees="46.7125" direction="north" orientation="latitude" pageId="13" pageNumber="172" precision="5" value="46.7125">46.7125° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE45506F81783627298EFB20435E0DA2" box="[451,605,1207,1234]" degrees="114.5324" direction="west" orientation="longitude" pageId="13" pageNumber="172" precision="5" value="-114.5324">114.5324° W</geoCoordinate>
|
||
;
|
||
<date id="FFCF1068817836272A25FB2043FC0DA1" box="[616,767,1207,1234]" pageId="13" pageNumber="172" value="2021-06-26">
|
||
<collectingDate id="EF8BE980817836272A25FB2043FC0DA1" box="[616,767,1207,1234]" pageId="13" pageNumber="172" value="2021-06-26">26 Jun. 2021</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2684537E817836272B44FB2042A10DA1" box="[777,930,1207,1233]" pageId="13" pageNumber="172">A. Fasbender</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED60AE6D817836272BADFB20452F0DA1" box="[992,1068,1207,1233]" pageId="13" pageNumber="172">AFPC</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="C36B65238178362528F0FB6D40B50B62" lastPageId="15" lastPageNumber="174" pageId="13" pageNumber="172" type="description">
|
||
<paragraph id="8BCE36A88178362728F0FB6D404D0C64" blockId="13.[189,334,1274,1300]" box="[189,334,1274,1300]" pageId="13" pageNumber="172">
|
||
<heading id="D08681C48178362728F0FB6D404D0C64" bold="true" box="[189,334,1274,1300]" fontSize="11" level="3" pageId="13" pageNumber="172" reason="3">
|
||
<emphasis id="B905EABA8178362728F0FB6D404D0C64" bold="true" box="[189,334,1274,1300]" pageId="13" pageNumber="172">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728A7FABD40290C34" blockId="13.[189,1399,1322,1532]" box="[234,298,1322,1348]" pageId="13" pageNumber="172">
|
||
<emphasis id="B905EABA8178362728A7FABD40290C34" bold="true" box="[234,298,1322,1348]" pageId="13" pageNumber="172">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0FAD840DB0C8C" blockId="13.[189,1399,1322,1532]" pageId="13" pageNumber="172">
|
||
<smallCapsWord id="8D28A0748178362728F0FAD8406D0C17" baselines="1378,1378" box="[189,366,1359,1385]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="13" pageNumber="172">MEASUREMENTS</smallCapsWord>
|
||
(n = 5). Head width (n = 3) 293–305(305); flagellomeres 34–48(45), 22–28(26), 23– 29(25), 22–25(24), 20–27(24), 24–27(25), 25–27(25), 23–30(27), 21–29(25), 21–32(25), 65–86(68), 53–64(56), 55–65(58); AR 0.65–0.76(0.69); FR (n = 1) 1.49; wing length 0.95–1.0(0.97) mm; wing width
|
||
<quantity id="4C899B4D817836272945FA2B40760CA7" box="[264,373,1468,1495]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="172" unit="mm" value="0.35">0.35 mm</quantity>
|
||
; costal ratio (n = 1) 0.50; GCR 2.13–2.62(2.42); GSR 0.82–0.89(0.86); aedeagus ratio (n = 3) 3.33–3.67(3.65).
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0F9B345FB0F12" blockId="13.[189,1398,1572,1635]" pageId="13" pageNumber="172">
|
||
<smallCapsWord id="8D28A0748178362728F0F9B3401A0F4C" baselines="1591,1591" box="[189,281,1572,1598]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="13" pageNumber="172">THORAX</smallCapsWord>
|
||
. Dorsocentral punctations inconspicuous, present only at posteriormost portion of dorsocentral setae near scutellum, may be absent in some specimens. Legs with femora and tibiae brown.
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728F0F91C40420E48" blockId="13.[189,1399,1675,1848]" pageId="13" pageNumber="172">
|
||
<smallCapsWord id="8D28A0748178362728F0F91C40350FD4" baselines="1694,1695" box="[189,310,1675,1701]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="13" pageNumber="172">GENITALIA</smallCapsWord>
|
||
(
|
||
<figureCitation id="134A2A2D817836272901F91C40CB0FD5" box="[332,456,1675,1701]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="13" pageNumber="172">Fig. 9d–f</figureCitation>
|
||
). Distal portion of parameres tapering gradually to acute apex. Aedeagus lateral margins concave, smoothly rounded, apical point posterolaterally directed, triangular with adjacent lateral obtuse spur-like posterolateral point without accessory spines, posterior margin emarginate, hyaline, hyaline medial incision extensive, margins parallel to lateral margins and basal arch.
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362728A7F8F640460E0B" blockId="13.[189,1399,1889,2025]" box="[234,325,1889,1915]" pageId="13" pageNumber="172">
|
||
<emphasis id="B905EABA8178362728A7F8F640460E0B" bold="true" box="[234,325,1889,1915]" pageId="13" pageNumber="172">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A88178362528F0F81142F70837" blockId="13.[189,1399,1889,2025]" lastBlockId="15.[189,1399,265,327]" lastPageId="15" lastPageNumber="174" pageId="13" pageNumber="172">
|
||
<smallCapsWord id="8D28A0748178362728F0F811406D0EEE" baselines="1945,1945" box="[189,366,1926,1952]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="13" pageNumber="172">MEASUREMENTS</smallCapsWord>
|
||
(n = 4). Head width (n = 3) 253–278; flagellomeres (n = 3) 26–28(27), 15–17(15), 18–20(20), 20–28(21), 21–25(24), 17–24(24), 24–27(25), 25–31(30), 39–41(40, 35–43(42), 40–46(40), 30–40(38), 41–50(43); AR 1.08–1.15; FR 1.41–1.47; wing length 0.97–1.19(1.13) mm; wing width 0.40–0.52(0.45) mm; costal ratio (n = 1) 0.55; spermathecal length 52–67(61); spermathecal width 37–49(43); spermathecal neck 9–16(14); spermatheca/neck ratio 0.22.
|
||
</paragraph>
|
||
<caption id="DF0E6620817B362428F0F96C45400E99" ID-DOI="http://doi.org/10.5281/zenodo.8083798" ID-Zenodo-Dep="8083798" httpUri="https://zenodo.org/record/8083798/files/figure.png" pageId="14" pageNumber="173" startId="14.[189,232,1787,1813]" targetBox="[259,1269,265,1731]" targetPageId="14">
|
||
<paragraph id="8BCE36A8817B362428F0F96C45400E99" blockId="14.[189,1399,1787,2025]" pageId="14" pageNumber="173">
|
||
<emphasis id="B905EABA817B362428F0F96C40130E65" bold="true" box="[189,272,1787,1813]" pageId="14" pageNumber="173">Fig. 8.</emphasis>
|
||
Male genitalia I.
|
||
<taxonomicName id="4C714D2B817B362429BEF96C42360E65" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[499,821,1787,1813]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="173" phylum="Arthropoda" rank="species" species="confusus" status="sp. nov.">
|
||
<emphasis id="B905EABA817B362429BEF96C42360E65" box="[499,821,1787,1813]" italics="true" pageId="14" pageNumber="173">Ceratoculicoides confusus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C1817B36242B77F96B42990E65" box="[826,922,1788,1814]" pageId="14" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B905EABA817B36242BE2F96C42BD0E65" bold="true" box="[943,958,1787,1813]" pageId="14" pageNumber="173">a</emphasis>
|
||
. Genitalia, aedeagus and parameres removed.
|
||
<emphasis id="B905EABA817B36242978F88940460E48" bold="true" box="[309,325,1822,1848]" pageId="14" pageNumber="173">b</emphasis>
|
||
. Parameres.
|
||
<emphasis id="B905EABA817B362429AEF88940F30E48" bold="true" box="[483,496,1822,1848]" pageId="14" pageNumber="173">c</emphasis>
|
||
. Aedeagus.
|
||
<taxonomicName id="4C714D2B817B36242ACBF88842120E48" authorityName="Fasbender" authorityYear="2023" box="[646,785,1822,1849]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="173" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="B905EABA817B36242ACBF88842120E48" box="[646,785,1822,1849]" italics="true" pageId="14" pageNumber="173">C.confusus</emphasis>
|
||
</taxonomicName>
|
||
(Colombia):
|
||
<emphasis id="B905EABA817B36242BF8F88942C60E48" bold="true" box="[949,965,1822,1848]" pageId="14" pageNumber="173">d</emphasis>
|
||
. Genitalia, aedeagus and parameres removed.
|
||
<emphasis id="B905EABA817B3624297DF8D5403E0E2C" bold="true" box="[304,317,1858,1884]" pageId="14" pageNumber="173">e</emphasis>
|
||
. Parameres.
|
||
<emphasis id="B905EABA817B3624299DF8D640D90E2B" bold="true" box="[464,474,1857,1883]" pageId="14" pageNumber="173">f</emphasis>
|
||
. Aedeagus.
|
||
<taxonomicName id="4C714D2B817B36242A2EF8D543EC0E2C" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[611,751,1858,1884]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="173" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov">
|
||
<emphasis id="B905EABA817B36242A2EF8D543EC0E2C" box="[611,751,1858,1884]" italics="true" pageId="14" pageNumber="173">C. pacificus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C1817B36242AB8F8D5424B0E2C" box="[757,840,1858,1884]" pageId="14" pageNumber="173" rank="species">sp. nov</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B905EABA817B36242B1BF8D542660E2C" bold="true" box="[854,869,1858,1884]" pageId="14" pageNumber="173">g</emphasis>
|
||
. Genitalia, aedeagus and parameres removed.
|
||
<emphasis id="B905EABA817B362428F0F8F241CE0E0F" bold="true" box="[189,205,1893,1919]" pageId="14" pageNumber="173">h</emphasis>
|
||
. Parameres.
|
||
<emphasis id="B905EABA817B3624292BF8F3406D0E0E" bold="true" box="[358,366,1892,1918]" pageId="14" pageNumber="173">i</emphasis>
|
||
. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCE36A8817A362528F0FEF843C808F8" blockId="15.[189,715,366,393]" box="[189,715,366,393]" pageId="15" pageNumber="174">
|
||
<smallCapsWord id="8D28A074817A362528F0FEF8401A08F7" baselines="386,386" box="[189,281,367,393]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="15" pageNumber="174">THORAX</smallCapsWord>
|
||
. Legs with femora and tibiae brown.
|
||
</paragraph>
|
||
<paragraph id="8BCE36A8817A362528F0FE2640B50B62" blockId="15.[189,1398,432,530]" pageId="15" pageNumber="174">
|
||
<smallCapsWord id="8D28A074817A362528F0FE26403108B9" baselines="452,452" box="[189,306,433,459]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="15" pageNumber="174">GENITALIA</smallCapsWord>
|
||
(
|
||
<figureCitation id="134A2A2D817A36252901FE2740BA08BA" box="[332,441,432,459]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta & Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="15" pageNumber="174">Fig. 11b</figureCitation>
|
||
). 9
|
||
<superScript id="7C049BE0817A362529A8FE2740F108CE" attach="left" box="[485,498,432,446]" fontSize="6" pageId="15" pageNumber="174">th</superScript>
|
||
sternite anterior branch acutely pointed, apices of each half touching medially or not; posterior branch evenly curving towards apex, spiniform, tip acute to rounded. 2 major spermathecae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6523817A362528F0FDAE45830BF3" box="[189,1152,569,643]" pageId="15" pageNumber="174" type="distribution">
|
||
<paragraph id="8BCE36A8817A362528F0FDAE40550B23" blockId="15.[189,342,569,595]" box="[189,342,569,595]" pageId="15" pageNumber="174">
|
||
<heading id="D08681C4817A362528F0FDAE40550B23" bold="true" box="[189,342,569,595]" fontSize="11" level="3" pageId="15" pageNumber="174" reason="3">
|
||
<emphasis id="B905EABA817A362528F0FDAE40550B23" bold="true" box="[189,342,569,595]" pageId="15" pageNumber="174">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A8817A362528F0FDFE45830BF3" blockId="15.[189,1152,617,643]" box="[189,1152,617,643]" pageId="15" pageNumber="174">
|
||
<collectingRegion id="49B5F84A817A362528F0FDFE408B0BF3" box="[189,392,617,643]" country="Canada" name="British Columbia" pageId="15" pageNumber="174">British Columbia</collectingRegion>
|
||
(
|
||
<collectingCountry id="F3667638817A362529D5FDFE40F40BF3" box="[408,503,617,643]" name="Canada" pageId="15" pageNumber="174">Canada</collectingCountry>
|
||
);
|
||
<collectingRegion id="49B5F84A817A36252A44FDFE43680BF3" box="[521,619,617,643]" country="United States of America" name="Arizona" pageId="15" pageNumber="174">Arizona</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B5F84A817A36252A3BFDFE43F30BF3" box="[630,752,617,643]" country="United States of America" name="California" pageId="15" pageNumber="174">California</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B5F84A817A36252AB6FDFE42420BF3" box="[763,833,617,643]" country="United States of America" name="Idaho" pageId="15" pageNumber="174">Idaho</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B5F84A817A36252B00FDFE42B60BF3" box="[845,949,617,643]" country="United States of America" name="Montana" pageId="15" pageNumber="174">Montana</collectingRegion>
|
||
(
|
||
<collectingCountry id="F3667638817A36252B88FDFE45000BF3" box="[965,1027,617,643]" name="United States of America" pageId="15" pageNumber="174">USA</collectingCountry>
|
||
) (
|
||
<figureCitation id="134A2A2D817A36252C51FDFE456D0BF3" box="[1052,1134,617,643]" captionStart="Fig" captionStartId="38.[189,232,1822,1848]" captionTargetBox="[283,1385,608,1735]" captionTargetId="figure-200@38.[203,1385,608,1735]" captionTargetPageId="38" captionText="Fig. 13. Distribution. a. Western North America. Solid star =Ceratoculicoides aliciae Huerta & Borkent, 2005; solid circles = C.borkenti sp. nov.; open triangles = C.grogani sp. nov.; open star = C. longipennis (Wirth, 1952); open square = C. pacificus sp. nov.; solid triangle = C.propinquus sp. nov.; solid cross C. sp. M1. b. Eastern North America. Solid circles = C. confusus sp. nov.; open squares = C.virginianus (Wirth, 1951). c. Central and northern South America. Solid circle = C.confusus; open squares = C. sp. F1; open triangles = C. sp. F2; solid triangle = C. sp. F3." figureDoi="http://doi.org/10.5281/zenodo.8083814" httpUri="https://zenodo.org/record/8083814/files/figure.png" pageId="15" pageNumber="174">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6523817A362528F0FD3C44250CC8" pageId="15" pageNumber="174" type="discussion">
|
||
<paragraph id="8BCE36A8817A362528F0FD3C402C0BB5" blockId="15.[189,303,683,709]" box="[189,303,683,709]" pageId="15" pageNumber="174">
|
||
<heading id="D08681C4817A362528F0FD3C402C0BB5" bold="true" box="[189,303,683,709]" fontSize="11" level="3" pageId="15" pageNumber="174" reason="3">
|
||
<emphasis id="B905EABA817A362528F0FD3C402C0BB5" bold="true" box="[189,303,683,709]" pageId="15" pageNumber="174">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCE36A8817A362528F0FD4D40E10AD6" blockId="15.[188,1399,730,934]" pageId="15" pageNumber="174">
|
||
<emphasis id="B905EABA817A362528F0FD4D40ED0B84" box="[189,494,730,756]" italics="true" pageId="15" pageNumber="174">Ceratoculicoidesborkenti</emphasis>
|
||
<taxonomicNameLabel id="A23657C1817A362529B8FD4D434D0B84" box="[501,590,730,756]" pageId="15" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is one of the most distinctive species of
|
||
<taxonomicName id="4C714D2B817A36252C69FD4D45EF0B84" authorityName=", Wirth & Ratanaworabhan" authorityYear="1971" box="[1060,1260,730,756]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EABA817A36252C69FD4D45EF0B84" box="[1060,1260,730,756]" italics="true" pageId="15" pageNumber="174">Ceratoculicoides</emphasis>
|
||
</taxonomicName>
|
||
in the adult male life stage. As member of the
|
||
<taxonomicName id="4C714D2B817A36252A1AFD6943F00A67" authorityName="Knoz" authorityYear="1987" box="[599,755,766,792]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="moravicus">
|
||
<emphasis id="B905EABA817A36252A1AFD6943F00A67" box="[599,755,766,792]" italics="true" pageId="15" pageNumber="174">C.moravicus</emphasis>
|
||
</taxonomicName>
|
||
group, the lateral margins of the aedeagus are heavily sclerotized, deeply concave, and taper in a smooth arc to a narrow apex (
|
||
<figureCitation id="134A2A2D817A36252C41FCB6455C0A4B" box="[1036,1119,801,827]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="15" pageNumber="174">Fig. 9f</figureCitation>
|
||
). The median portion of the aedeagus is so hyaline as to be nearly transparent, and unlike any other
|
||
<taxonomicName id="4C714D2B817A36252C6BFCD245ED0A2F" authorityName=", Wirth & Ratanaworabhan" authorityYear="1971" box="[1062,1262,837,863]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905EABA817A36252C6BFCD245ED0A2F" box="[1062,1262,837,863]" italics="true" pageId="15" pageNumber="174">Ceratoculicoides</emphasis>
|
||
</taxonomicName>
|
||
the hyaline incision expands above the basal arch subtrapezoidally, leaving only the lateral and anterior margins of the aedeagus sclerotized.
|
||
</paragraph>
|
||
<paragraph id="8BCE36A8817A362528F0FC5944250CC8" blockId="15.[187,1399,974,1464]" pageId="15" pageNumber="174">
|
||
The females of
|
||
<taxonomicName id="4C714D2B817A3625293AFC5940FB0A98" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[375,504,974,1000]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="borkenti" status="sp. nov.">
|
||
<emphasis id="B905EABA817A3625293AFC5940FB0A98" box="[375,504,974,1000]" italics="true" pageId="15" pageNumber="174">C.borkenti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C1817A36252A4CFC5843580A99" box="[513,603,975,1001]" pageId="15" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="134A2A2D817A36252A21FC5943CD0A98" box="[620,718,974,1000]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta & Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="15" pageNumber="174">Fig. 11b</figureCitation>
|
||
) fall into a group of several species with the length of the largest spermathecae <70 μm. Of the described species, it can be differentiated from
|
||
<taxonomicName id="4C714D2B817A36252CDCFC6544150D7B" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[1169,1302,1010,1036]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="grogani" status="sp. nov.">
|
||
<emphasis id="B905EABA817A36252CDCFC6544150D7B" box="[1169,1302,1010,1036]" italics="true" pageId="15" pageNumber="174">C. grogani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C1817A36252D50FC6544740D7C" box="[1309,1399,1010,1036]" pageId="15" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the medial margin of the 9
|
||
<superScript id="7C049BE0817A36252A42FB82431F0D53" attach="left" box="[527,540,1045,1059]" fontSize="6" pageId="15" pageNumber="174">th</superScript>
|
||
sternite being deeply concave and rounded (vs nearly straight and sinuous,
|
||
<figureCitation id="134A2A2D817A362528F0FBAE401A0D23" box="[189,281,1081,1107]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta & Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="15" pageNumber="174">Fig. 11e</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C714D2B817A3625291BFBAE40FA0D23" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[342,505,1081,1107]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="virginianus">
|
||
<emphasis id="B905EABA817A3625291BFBAE40FA0D23" box="[342,505,1081,1107]" italics="true" pageId="15" pageNumber="174">C.virginianus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134A2A2D817A36252A4BFBAE43670D23" box="[518,612,1081,1107]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="15" pageNumber="174">Fig. 12c</figureCitation>
|
||
) by the 9
|
||
<superScript id="7C049BE0817A36252A81FBAF43DA0D36" attach="left" box="[716,729,1080,1094]" fontSize="6" pageId="15" pageNumber="174">th</superScript>
|
||
sternite’s acute or triangular anterior branch (vs rounded or truncate) and evenly curved posteromedially directed spiniform posterior branch (base of branch directed posteriorly, curving medially in apical half in
|
||
<taxonomicName id="4C714D2B817A36252AA8FB1642880DEA" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[741,907,1153,1179]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="virginianus">
|
||
<emphasis id="B905EABA817A36252AA8FB1642880DEA" box="[741,907,1153,1179]" italics="true" pageId="15" pageNumber="174">C. virginianus</emphasis>
|
||
</taxonomicName>
|
||
, usually tapering only in apical portion). However, there are a number of unassociated female specimens which display a similar morphology, including a species from
|
||
<collectingCountry id="F3667638817A36252997FB5F434F0D92" box="[474,588,1224,1250]" name="Colombia" pageId="15" pageNumber="174">Colombia</collectingCountry>
|
||
(
|
||
<emphasis id="B905EABA817A36252A11FB5F43750D92" box="[604,630,1224,1250]" italics="true" pageId="15" pageNumber="174">C.</emphasis>
|
||
sp. F1,
|
||
<figureCitation id="134A2A2D817A36252A9DFB5F422D0D92" box="[720,814,1224,1250]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="15" pageNumber="174">Fig. 12c</figureCitation>
|
||
) and another from
|
||
<collectingCountry id="F3667638817A36252C45FB5F45860D92" box="[1032,1157,1224,1250]" name="Costa Rica" pageId="15" pageNumber="174">Costa Rica</collectingCountry>
|
||
(
|
||
<emphasis id="B905EABA817A36252CD9FB5F45AC0D92" box="[1172,1199,1224,1250]" italics="true" pageId="15" pageNumber="174">C.</emphasis>
|
||
sp. F3,
|
||
<figureCitation id="134A2A2D817A36252D45FB5F446A0D92" box="[1288,1385,1224,1250]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="15" pageNumber="174">Fig. 12e</figureCitation>
|
||
). The Costa Rican
|
||
<emphasis id="B905EABA817A362529CEFB7B409D0C76" box="[387,414,1260,1286]" italics="true" pageId="15" pageNumber="174">C.</emphasis>
|
||
sp F3 differs in having the base of the posterior branch directed posteriorly, vs evenly curving along the whole length in
|
||
<taxonomicName id="4C714D2B817A36252A74FA8743B90C59" authorityName="Fasbender" authorityYear="2023" box="[569,698,1295,1322]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="borkenti">
|
||
<emphasis id="B905EABA817A36252A74FA8743B90C59" box="[569,698,1295,1322]" italics="true" pageId="15" pageNumber="174">C.borkenti</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<emphasis id="B905EABA817A36252AA2FA87420A0C5A" box="[751,777,1296,1322]" italics="true" pageId="15" pageNumber="174">C.</emphasis>
|
||
sp. F1 from
|
||
<collectingCountry id="F3667638817A36252BDAFA98450A0C59" box="[919,1033,1295,1321]" name="Colombia" pageId="15" pageNumber="174">Colombia</collectingCountry>
|
||
cannot be reliably differentiated by the 9
|
||
<superScript id="7C049BE0817A3625295AFAA540270C30" attach="left" box="[279,292,1330,1344]" fontSize="6" pageId="15" pageNumber="174">th</superScript>
|
||
sternite.
|
||
<taxonomicName id="4C714D2B817A362529C4FAA443080C3D" authorityName="Fasbender" authorityYear="2023" box="[393,523,1331,1357]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="borkenti">
|
||
<emphasis id="B905EABA817A362529C4FAA443080C3D" box="[393,523,1331,1357]" italics="true" pageId="15" pageNumber="174">C.borkenti</emphasis>
|
||
</taxonomicName>
|
||
has a lower flagellum ratio (FR <1.5) than either of these provisional species. As there is only a limited sample of reliably associated females, and there are
|
||
<specimenCount id="9D77FD21817A36252C18FAC045C20C00" box="[1109,1217,1366,1393]" pageId="15" pageNumber="174" type="male">two male</specimenCount>
|
||
morphospecies without a female association sharing the range of
|
||
<taxonomicName id="4C714D2B817A36252AB7FAEC427F0CE4" authorityName="Fasbender" authorityYear="2023" box="[762,892,1402,1429]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="borkenti">
|
||
<emphasis id="B905EABA817A36252AB7FAEC427F0CE4" box="[762,892,1402,1429]" italics="true" pageId="15" pageNumber="174">C.borkenti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C714D2B817A36252BC0FAEC45330CE4" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[909,1072,1403,1429]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="174" phylum="Arthropoda" rank="species" species="propinquus" status="sp. nov.">
|
||
<emphasis id="B905EABA817A36252BC0FAEC45330CE4" box="[909,1072,1403,1429]" italics="true" pageId="15" pageNumber="174">C.propinquus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23657C1817A36252C75FAEC45920CE5" box="[1080,1169,1403,1429]" pageId="15" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<emphasis id="B905EABA817A36252C84FAEC45E70CE5" box="[1225,1252,1403,1429]" italics="true" pageId="15" pageNumber="174">C.</emphasis>
|
||
sp. M1), the characters cited above may prove non-diagnostic upon examination of a broader range of material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |