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<document id="5109E13FD321BBA2F7B294BC2133BB67" ID-DOI="10.5281/zenodo.4504254" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504254" ID-ZooBank="9F76CF88-A3DD-4F0E-B348-EEB9558DBBC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465889004" checkinUser="carolina" docAuthor="Ng, Peter K. L., Schubart, Christoph D. &amp; Lukhaup, Christian" docDate="2015" docId="03CD4D1DD025FFDBF747FF13FC7AFD14" docLanguage="en" docName="RafflesBZool.63.3-13.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Geosesarma hagen Ng &amp; Schubart &amp; Lukhaup 2015, new species" docType="treatment" docUuid="F2A209A9-49AA-412C-8161-E00530FA8045" docUuidSource="ZooBank" docVersion="6" lastPageNumber="11" masterDocId="FFF43565D02DFFD3F654FFAEFFC0FFC1" masterDocTitle="New species of “ vampire crabs ” (Geosesarma De Man, 1892) from central Java, Indonesia, and the identity of Sesarma (Geosesarma) nodulifera De Man, 1892 (Crustacea, Brachyura, Thoracotremata, Sesarmidae)" masterLastPageNumber="13" masterPageNumber="3" pageNumber="11" updateTime="1698866089694" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="4429F2FC9534D07974F7C04E2233F9B7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66C0A699563E3D342C1FA0E8BEBF2BDF">
<mods:title id="342167DF776209248A1CB713525ED45E">New species of “ vampire crabs ” (Geosesarma De Man, 1892) from central Java, Indonesia, and the identity of Sesarma (Geosesarma) nodulifera De Man, 1892 (Crustacea, Brachyura, Thoracotremata, Sesarmidae)</mods:title>
</mods:titleInfo>
<mods:name id="1AEA4D68520B73AD967E77C182EBD315" type="personal">
<mods:role id="57CA5403BC2FC8404BF7FC8648810ED6">
<mods:roleTerm id="A3AD886BF2985BA9E94A7D68B644B9CA">Author</mods:roleTerm>
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<mods:namePart id="1448926948478CB3406CB9A67AD859CE">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="A55D7B0A0352127E54B0FF236C69F23E" type="personal">
<mods:role id="A03F7590C0D01B080C2C1242399028E1">
<mods:roleTerm id="7E54E92B92B0BCDB53C3D26A858C03E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2AEF7BDCB9C06C7D87D198C4B78EB24">Schubart, Christoph D.</mods:namePart>
</mods:name>
<mods:name id="7E004426162F89256419FF476194EB1B" type="personal">
<mods:role id="4F70C8DF06348C16FDFA5A975AC4D448">
<mods:roleTerm id="3C2525E4CBA8469B2CE71729D1C8B5E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="467DEB2710B5529F1941B5B65EEB5F31">Lukhaup, Christian</mods:namePart>
</mods:name>
<mods:typeOfResource id="85560B2054F14A39DA8D1913161606AE">text</mods:typeOfResource>
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<mods:title id="C63520D616B555D8C194B02EB494543D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="7A59208BC4919A652C0F79D32C948C74">
<mods:date id="8E50B5BBDAA64B2E058AFB374B0AC8F6">2015</mods:date>
<mods:detail id="2452CC791013AE7EFCB30FC5F5568D35" type="pubDate">
<mods:number id="60F94E5AEFE9E9B68C2B54AF6F831C77">2015-01-16</mods:number>
</mods:detail>
<mods:detail id="12FE31E2195491FF01FC34181AE80B16" type="volume">
<mods:number id="ABD71ACB4A65985C96ED20E3373DE7F8">63</mods:number>
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<mods:start id="A4D8342C9A23D5807078F84E9A74CB6A">3</mods:start>
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<mods:classification id="B85F7BA80CFC16B672FFF331B5CB7EB3">journal article</mods:classification>
<mods:identifier id="747C675576C345AE3804696FA36C2A1A" type="DOI">10.5281/zenodo.4504254</mods:identifier>
<mods:identifier id="121990280C1059C9A098521870C3E46B" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="482DA66F3E54DB818134BE23D2E08CA1" type="Zenodo-Dep">4504254</mods:identifier>
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<treatment id="03CD4D1DD025FFDBF747FF13FC7AFD14" ID-DOI="http://doi.org/10.5281/zenodo.4504561" ID-Zenodo-Dep="4504561" LSID="urn:lsid:zoobank.org:act:F2A209A9-49AA-412C-8161-E00530FA8045" httpUri="http://treatment.plazi.org/id/03CD4D1DD025FFDBF747FF13FC7AFD14" lastPageNumber="11" pageId="8" pageNumber="11">
<subSubSection id="C37EAF80D025FFDBF747FF13FDBCFF14" box="[275,636,188,213]" pageId="8" pageNumber="11" type="nomenclature">
<paragraph id="8BDBFC0BD025FFDBF747FF13FDBCFF14" blockId="8.[225,686,188,245]" box="[275,636,188,213]" pageId="8" pageNumber="11">
<heading id="D0934B67D025FFDBF747FF13FDBCFF14" bold="true" box="[275,636,188,213]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="11" reason="2">
<emphasis id="B9102019D025FFDBF747FF13FDBCFF14" bold="true" box="[275,636,188,213]" pageId="8" pageNumber="11">
<taxonomicName id="4C648788D025FFDBF747FF13FE27FF15" ID-CoL="6KBGZ" authority="Ng &amp; Schubart &amp; Lukhaup, 2015" authorityName="Ng &amp; Schubart &amp; Lukhaup" authorityYear="2015" box="[275,487,188,213]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="11" phylum="Arthropoda" rank="species" species="hagen" status="sp. nov.">
<emphasis id="B9102019D025FFDBF747FF13FE27FF15" bold="true" box="[275,487,188,213]" italics="true" pageId="8" pageNumber="11">Geosesarma hagen</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2239D62D025FFDBF7A2FF13FDBCFF14" box="[502,636,189,213]" pageId="8" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37EAF80D025FFDBF6B5FF73FD6EFF34" box="[225,686,221,245]" pageId="8" pageNumber="11" type="description">
<paragraph id="8BDBFC0BD025FFDBF6B5FF73FD6EFF34" blockId="8.[225,686,188,245]" box="[225,686,221,245]" pageId="8" pageNumber="11">
(
<figureCitation id="135FE08ED025FFDBF6BDFF73FEABFF35" box="[233,363,221,245]" captionStart="Fig" captionStartId="4.[136,171,1121,1143]" captionTargetBox="[136,1474,189,1106]" captionTargetId="figure-8@4.[136,1474,189,1108]" captionTargetPageId="4" captionText="Fig. 3. A, B, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; C, D, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, C, overall view; B, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504260" httpUri="https://zenodo.org/record/4504260/files/figure.png" pageId="8" pageNumber="11">Figs. 3C, D</figureCitation>
,
<figureCitation id="135FE08ED025FFDBF72CFF73FE45FF34" box="[376,389,221,245]" captionStart="Fig" captionStartId="4.[136,171,2034,2056]" captionTargetBox="[305,1305,1212,2019]" captionTargetId="figure-56@4.[297,1312,1203,2022]" captionTargetPageId="4" captionText="Fig. 4. AG, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; HM, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, H, left third maxilliped; B, I, male abdominal somites 36 and telson; CG, JM, left G1 (drawn from different angles). All structures denuded. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4504262" httpUri="https://zenodo.org/record/4504262/files/figure.png" pageId="8" pageNumber="11">4</figureCitation>
HM, 5E, G, 6DF, 7E, F)
</paragraph>
</subSubSection>
<subSubSection id="C37EAF80D025FFDBF6DCFEB3FD5CFD74" pageId="8" pageNumber="11" type="materials_examined">
<paragraph id="8BDBFC0BD025FFDBF6DCFEB3FD5CFD74" blockId="8.[136,775,285,693]" pageId="8" pageNumber="11">
<emphasis id="B9102019D025FFDBF6DCFEB3FEB3FEF4" bold="true" box="[136,371,285,309]" pageId="8" pageNumber="11">Material examined.</emphasis>
<materialsCitation id="3B0CF656D025FFDBF72AFEB3FE3BFE54" collectingDate="2011-10-05" collectionCode="MZB" collectorName="C. Lukhaup" country="Indonesia" latitude="-7.4638886" location="Cilacap town" longLatPrecision="21" longitude="108.83778" municipality="Cilacap Regency" pageId="8" pageNumber="11" specimenCode="MZB 4062" specimenCount="1" stateProvince="Jawa Tengah" typeStatus="holotype">
<typeStatus id="54DF42A9D025FFDBF72AFEB3FE2AFEF4" box="[382,490,285,309]" pageId="8" pageNumber="11" type="holotype">Holotype</typeStatus>
: male (13.9 ×
<quantity id="4C9C51EED025FFDBF4CCFEB3FD3FFEF4" box="[664,767,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="8" pageNumber="11" unit="mm" value="13.3">13.3 mm</quantity>
) (
<specimenCode id="DBC25470D025FFDBF6DBFE93FEC9FE94" box="[143,265,317,341]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="11" type="Museum">MZB 4062</specimenCode>
), station R6051, north of
<location id="8EBBAAD0D025FFDBF44AFE93FD6DFE95" LSID="urn:lsid:plazi:treatment:03CD4D1DD025FFDBF747FF13FC7AFD14:8EBBAAD0D025FFDBF44AFE93FD6DFE95" box="[542,685,317,341]" country="Indonesia" latitude="-7.4638886" longLatPrecision="21" longitude="108.83778" municipality="Cilacap Regency" name="Cilacap town" pageId="8" pageNumber="11" stateProvince="Jawa Tengah">Cilacap town</location>
,
<collectingMunicipality id="6BBF6671D025FFDBF4E2FE93FF28FEB4" pageId="8" pageNumber="11">Cilacap Regency</collectingMunicipality>
,
<collectingRegion id="49A032E9D025FFDBF6A7FEF3FEB4FEB5" box="[243,372,349,373]" country="Indonesia" name="Jawa Tengah" pageId="8" pageNumber="11">central Java</collectingRegion>
,
<geoCoordinate id="EE509ACCD025FFDBF72BFEF3FE33FEB4" box="[383,499,349,373]" degrees="7" direction="south" minutes="27" orientation="latitude" pageId="8" pageNumber="11" precision="15" seconds="50" value="-7.4638886">7°2750”S</geoCoordinate>
,
<geoCoordinate id="EE509ACCD025FFDBF7AAFEF3FD4DFEB4" box="[510,653,349,373]" degrees="108" direction="east" minutes="50" orientation="longitude" pageId="8" pageNumber="11" precision="15" seconds="16" value="108.83778">108°5016”E</geoCoordinate>
,
<collectingCountry id="F373BC9BD025FFDBF4CCFEF3FCC2FEB4" box="[664,770,349,373]" name="Indonesia" pageId="8" pageNumber="11">Indonesia</collectingCountry>
, coll.
<collectorName id="269199DDD025FFDBF6EBFED3FE82FE54" box="[191,322,381,405]" pageId="8" pageNumber="11">C. Lukhaup</collectorName>
,
<date id="FFDADACBD025FFDBF71AFED3FE3BFE54" box="[334,507,381,405]" pageId="8" pageNumber="11" value="2011-10-05">
<collectingDate id="EF9E2323D025FFDBF71AFED3FE3BFE54" box="[334,507,381,405]" pageId="8" pageNumber="11" value="2011-10-05">5 October 2011</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="54DF42A9D025FFDBF452FED3FDB2FE54" box="[518,626,381,405]" pageId="8" pageNumber="11" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D623782D025FFDBF429FED3FD07FE54" box="[637,711,381,405]" pageId="8" pageNumber="11" type="male">1 male</specimenCount>
(13.2 ×
<quantity id="4C9C51EED025FFDBF6CBFE33FEC2FE74" box="[159,258,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="11" unit="mm" value="12.2">12.2 mm</quantity>
),
<materialsCitation id="3B0CF656D025FFDBF742FE33FE41FE14" collectingDate="2011-10-05" collectionCode="ZRC" collectorName="C. Lukhaup" country="Indonesia" latitude="-7.4638886" location="Cilacap town" longLatPrecision="21" longitude="108.83778" municipality="Cilacap Regency" pageId="8" pageNumber="11" specimenCode="ZRC 2014.0269" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="9D623782D025FFDBF742FE33FEB4FE74" box="[278,372,413,437]" pageId="8" pageNumber="11" type="female">1 female</specimenCount>
(14.5 ×
<quantity id="4C9C51EED025FFDBF785FE33FDF2FE74" box="[465,562,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="8" pageNumber="11" unit="mm" value="13.4">13.4 mm</quantity>
) (
<specimenCode id="DBC25470D025FFDBF41DFE33FD3CFE74" box="[585,764,413,437]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="11">ZRC 2014.0269</specimenCode>
), same data as holotype
</materialsCitation>
;
<specimenCount id="9D623782D025FFDBF7C4FE13FE27FE14" box="[400,487,445,469]" pageId="8" pageNumber="11" type="male">2 males</specimenCount>
(larger 12.5 ×
<quantity id="4C9C51EED025FFDBF4C0FE13FD3BFE14" box="[660,763,445,469]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="11" unit="mm" value="12.2">12.2 mm</quantity>
),
<materialsCitation id="3B0CF656D025FFDBF6DCFE73FD64FE34" accessionNumber="SMF 46287" box="[136,676,477,501]" collectionCode="SMF" country="Indonesia" pageId="8" pageNumber="11" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="9D623782D025FFDBF6DCFE73FF32FE34" box="[136,242,477,501]" pageId="8" pageNumber="11" type="female">2 females</specimenCount>
(larger 11.6 ×
<quantity id="4C9C51EED025FFDBF7CFFE73FE3EFE34" box="[411,510,477,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="8" pageNumber="11" unit="mm" value="10.9">10.9 mm</quantity>
) (
<collectionCode id="ED7564CED025FFDBF442FE73FD8DFE34" box="[534,589,477,501]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="11">SMF</collectionCode>
46287)
</materialsCitation>
,
<specimenCount id="9D623782D025FFDBF4E6FE73FCC7FE34" box="[690,775,477,501]" pageId="8" pageNumber="11" type="male">2 males</specimenCount>
(larger 12.2 ×
<quantity id="4C9C51EED025FFDBF771FE53FE48FDD4" box="[293,392,509,533]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="8" pageNumber="11" unit="mm" value="11.6">11.6 mm</quantity>
),
<materialsCitation id="3B0CF656D025FFDBF7C8FE53FD9BFDF4" collectingDate="2011-10-05" collectionCode="ZRC" collectorName="C. Lukhaup" country="Indonesia" latitude="-7.4638886" location="Cilacap town" longLatPrecision="21" longitude="108.83778" municipality="Cilacap Regency" pageId="8" pageNumber="11" specimenCode="ZRC 2014.0268" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="9D623782D025FFDBF7C8FE53FE3BFDD4" box="[412,507,509,533]" pageId="8" pageNumber="11" type="female">1 female</specimenCount>
(11.3 ×
<quantity id="4C9C51EED025FFDBF40DFE53FD7BFDD4" box="[601,699,509,533]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="8" pageNumber="11" unit="mm" value="10.7">10.7 mm</quantity>
) (
<specimenCode id="DBC25470D025FFDBF485FE53FF3EFDF4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="11">ZRC 2014.0268</specimenCode>
), same data as holotype. Others
</materialsCitation>
:
<materialsCitation id="3B0CF656D025FFDBF432FDB3FE62FDB4" collectingDate="2011-04-12" collectionCode="ZRC" country="Indonesia" location="Cilacap town" pageId="8" pageNumber="11" specimenCode="ZRC 2014.0272" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Timur" typeStatus="paratype">
<specimenCount id="9D623782D025FFDBF432FDB3FD70FDF4" box="[614,688,541,565]" pageId="8" pageNumber="11" type="male">1 male</specimenCount>
(12.9 ×
<quantity id="4C9C51EED025FFDBF6DCFD93FF26FD94" box="[136,230,573,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="8" pageNumber="11" unit="mm" value="11.8">11.8 mm</quantity>
) (
<specimenCode id="DBC25470D025FFDBF6ADFD93FE67FD94" box="[249,423,573,597]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="11">ZRC 2014.0272</specimenCode>
), from aquarium dealer in
<location id="8EBBAAD0D025FFDBF4E2FD93FF00FDB5" LSID="urn:lsid:plazi:treatment:03CD4D1DD025FFDBF747FF13FC7AFD14:8EBBAAD0D025FFDBF4E2FD93FF00FDB5" country="Indonesia" name="Cilacap town" pageId="8" pageNumber="11" stateProvince="Jawa Timur">Cilacap town</location>
,
<collectingRegion id="49A032E9D025FFDBF69FFDF3FF3DFDB5" box="[203,253,605,628]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="11">Java</collectingRegion>
,
<date id="FFDADACBD025FFDBF75CFDF3FE62FDB4" box="[264,418,605,629]" pageId="8" pageNumber="11" value="2011-04-12">
<collectingDate id="EF9E2323D025FFDBF75CFDF3FE62FDB4" box="[264,418,605,629]" pageId="8" pageNumber="11" value="2011-04-12">12 April 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B0CF656D025FFDBF7F8FDF3FD58FD74" collectingDate="2010-06-09" collectionCode="ZRC" collectorName="O. Mengedoht" country="Indonesia" location="Java" pageId="8" pageNumber="11" specimenCode="ZRC 2014.0267" specimenCount="2" specimenCount-male="2" stateProvince="Jawa Timur" typeStatus="paratype">
<specimenCount id="9D623782D025FFDBF7F8FDF3FE3EFDB4" box="[428,510,605,629]" pageId="8" pageNumber="11" type="male">2 males</specimenCount>
(larger 12.6 ×
<quantity id="4C9C51EED025FFDBF4CBFDF3FD3FFDB4" box="[671,767,605,629]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="8" pageNumber="11" unit="mm" value="11.6">11.6 mm</quantity>
) (
<specimenCode id="DBC25470D025FFDBF6DBFDD3FE89FD54" box="[143,329,637,661]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="11">ZRC 2014.0267</specimenCode>
), from aquarium trade, ostensibly from Cilacap,
<collectingRegion id="49A032E9D025FFDBF6BDFD33FEDBFD75" box="[233,283,669,692]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="11">Java</collectingRegion>
, from
<collectorName id="269199DDD025FFDBF730FD33FDC3FD74" box="[356,515,669,693]" pageId="8" pageNumber="11">O. Mengedoht</collectorName>
,
<date id="FFDADACBD025FFDBF45BFD33FD58FD74" box="[527,664,669,693]" pageId="8" pageNumber="11" value="2010-06-09">
<collectingDate id="EF9E2323D025FFDBF45BFD33FD58FD74" box="[527,664,669,693]" pageId="8" pageNumber="11" value="2010-06-09">9 June 2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37EAF80D025FFDBF6DCFD73FE46F974" pageId="8" pageNumber="11" type="description">
<paragraph id="8BDBFC0BD025FFDBF6DCFD73FD4BFAF4" blockId="8.[136,775,733,1333]" pageId="8" pageNumber="11">
<emphasis id="B9102019D025FFDBF6DCFD73FED4FD34" bold="true" box="[136,276,733,757]" pageId="8" pageNumber="11">Description.</emphasis>
Carapace squarish, slightly wider than long, lateral margins subparallel (
<figureCitation id="135FE08ED025FFDBF7E6FD53FDE7FCD4" box="[434,551,765,789]" captionStart="Fig" captionStartId="4.[136,171,1121,1143]" captionTargetBox="[136,1474,189,1106]" captionTargetId="figure-8@4.[136,1474,189,1108]" captionTargetPageId="4" captionText="Fig. 3. A, B, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; C, D, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, C, overall view; B, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504260" httpUri="https://zenodo.org/record/4504260/files/figure.png" pageId="8" pageNumber="11">Fig. 3C, D</figureCitation>
); dorsal surface with well-defined regions, anterior regions very densely covered with small rounded granules (
<figureCitation id="135FE08ED025FFDBF780FC93FD90FC94" box="[468,592,829,853]" captionStart="Fig" captionStartId="4.[136,171,1121,1143]" captionTargetBox="[136,1474,189,1106]" captionTargetId="figure-8@4.[136,1474,189,1108]" captionTargetPageId="4" captionText="Fig. 3. A, B, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; C, D, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, C, overall view; B, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504260" httpUri="https://zenodo.org/record/4504260/files/figure.png" pageId="8" pageNumber="11">Fig. 3C, D</figureCitation>
); front deflexed, frontal lobes broad with convex margins; postfrontal cristae prominent (
<figureCitation id="135FE08ED025FFDBF707FCD3FE0AFC54" box="[339,458,893,917]" captionStart="Fig" captionStartId="4.[136,171,1121,1143]" captionTargetBox="[136,1474,189,1106]" captionTargetId="figure-8@4.[136,1474,189,1108]" captionTargetPageId="4" captionText="Fig. 3. A, B, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; C, D, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, C, overall view; B, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504260" httpUri="https://zenodo.org/record/4504260/files/figure.png" pageId="8" pageNumber="11">Fig. 3C, D</figureCitation>
); external orbital tooth large, triangular, curving obliquely outwards, tip protruding beyond level of lateral carapace margin (
<figureCitation id="135FE08ED025FFDBF7BBFC13FD86FC14" box="[495,582,957,981]" captionStart="Fig" captionStartId="4.[136,171,1121,1143]" captionTargetBox="[136,1474,189,1106]" captionTargetId="figure-8@4.[136,1474,189,1108]" captionTargetPageId="4" captionText="Fig. 3. A, B, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; C, D, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, C, overall view; B, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504260" httpUri="https://zenodo.org/record/4504260/files/figure.png" pageId="8" pageNumber="11">Fig. 3D</figureCitation>
). Exopod of third maxilliped relatively slender, with long flagellum extending beyond merus width (
<figureCitation id="135FE08ED025FFDBF72AFC53FE19FBD4" box="[382,473,1021,1045]" captionStart="Fig" captionStartId="4.[136,171,2034,2056]" captionTargetBox="[305,1305,1212,2019]" captionTargetId="figure-56@4.[297,1312,1203,2022]" captionTargetPageId="4" captionText="Fig. 4. AG, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; HM, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, H, left third maxilliped; B, I, male abdominal somites 36 and telson; CG, JM, left G1 (drawn from different angles). All structures denuded. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4504262" httpUri="https://zenodo.org/record/4504262/files/figure.png" pageId="8" pageNumber="11">Fig. 4H</figureCitation>
). Outer surface of palm of adult male chelae granulated; dorsal margin of dactylus with 79 tubercles (distal one may be very low), each with pectinated tip (
<figureCitation id="135FE08ED025FFDBF766FBF3FE4AFBB4" box="[306,394,1117,1141]" captionStart="Fig" captionStartId="5.[113,148,1865,1887]" captionTargetBox="[113,1451,302,1850]" captionTargetId="figure-121@5.[113,1451,301,1851]" captionTargetPageId="5" captionText="Fig. 5. A, B, Geosesarma noduliferum (De Man, 1892), lectotype male (11.4 × 10.9 mm) (RMNH 1246a), Bogor, Java; C, G. bicolor Ng &amp; Davie, 1995, holotype male (11.0 × 10.5 mm) (ZRC 1995.279), Ujung Kulon, Java; D, F, G. dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; E, G, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java; H, I, G. confertum (Ortmann, 1894), male (10.7 × 9.7 mm) (ZRC 1970.3.7.11), Tjibodas, Java. A, H, anterior male thoracic sternum and abdomen; BE, I, outer surface of right chela; F, G, left last ambulatory leg" figureDoi="http://doi.org/10.5281/zenodo.4504264" httpUri="https://zenodo.org/record/4504264/files/figure.png" pageId="8" pageNumber="11">Fig. 5E</figureCitation>
). Ambulatory legs with relatively broad meri, with sharp subdistal spine on dorsal margin, surfaces rugose (
<figureCitation id="135FE08ED025FFDBF711FB33FE67FB74" box="[325,423,1181,1205]" captionStart="Fig" captionStartId="4.[136,171,1121,1143]" captionTargetBox="[136,1474,189,1106]" captionTargetId="figure-8@4.[136,1474,189,1108]" captionTargetPageId="4" captionText="Fig. 3. A, B, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; C, D, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, C, overall view; B, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504260" httpUri="https://zenodo.org/record/4504260/files/figure.png" pageId="8" pageNumber="11">Figs. 3C</figureCitation>
,
<figureCitation id="135FE08ED025FFDBF7E3FB33FE1BFB74" box="[439,475,1181,1205]" captionStart="Fig" captionStartId="5.[113,148,1865,1887]" captionTargetBox="[113,1451,302,1850]" captionTargetId="figure-121@5.[113,1451,301,1851]" captionTargetPageId="5" captionText="Fig. 5. A, B, Geosesarma noduliferum (De Man, 1892), lectotype male (11.4 × 10.9 mm) (RMNH 1246a), Bogor, Java; C, G. bicolor Ng &amp; Davie, 1995, holotype male (11.0 × 10.5 mm) (ZRC 1995.279), Ujung Kulon, Java; D, F, G. dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; E, G, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java; H, I, G. confertum (Ortmann, 1894), male (10.7 × 9.7 mm) (ZRC 1970.3.7.11), Tjibodas, Java. A, H, anterior male thoracic sternum and abdomen; BE, I, outer surface of right chela; F, G, left last ambulatory leg" figureDoi="http://doi.org/10.5281/zenodo.4504264" httpUri="https://zenodo.org/record/4504264/files/figure.png" pageId="8" pageNumber="11">5G</figureCitation>
). Male abdomen relatively broad; telson semicircular; somite 6 with strongly convex lateral margins (
<figureCitation id="135FE08ED025FFDBF763FB73FE42FB34" box="[311,386,1245,1269]" captionStart="Fig" captionStartId="4.[136,171,2034,2056]" captionTargetBox="[305,1305,1212,2019]" captionTargetId="figure-56@4.[297,1312,1203,2022]" captionTargetPageId="4" captionText="Fig. 4. AG, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; HM, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, H, left third maxilliped; B, I, male abdominal somites 36 and telson; CG, JM, left G1 (drawn from different angles). All structures denuded. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4504262" httpUri="https://zenodo.org/record/4504262/files/figure.png" pageId="8" pageNumber="11">Fig. 4I</figureCitation>
). G1 slender, proximal part straight, distal part pectinated, bent, elongate, spatuliform; tip entire, subtruncate when viewed dorsally (
<figureCitation id="135FE08ED025FFDBF45EFAB3FD8CFAF4" box="[522,588,1309,1333]" captionStart="Fig" captionStartId="4.[136,171,2034,2056]" captionTargetBox="[305,1305,1212,2019]" captionTargetId="figure-56@4.[297,1312,1203,2022]" captionTargetPageId="4" captionText="Fig. 4. AG, Geosesarma dennerle, new species, holotype male (14.4 × 14.0 mm) (MZB 4061), Cilacap, Java; HM, G. hagen, new species, holotype male (13.9 × 13.3 mm) (MZB 4062), Cilacap, Java. A, H, left third maxilliped; B, I, male abdominal somites 36 and telson; CG, JM, left G1 (drawn from different angles). All structures denuded. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4504262" httpUri="https://zenodo.org/record/4504262/files/figure.png" pageId="8" pageNumber="11">Fig. 4</figureCitation>
JM).
</paragraph>
<paragraph id="8BDBFC0BD025FFDBF6DCFAF3FE46F974" blockId="8.[136,775,1373,1717]" pageId="8" pageNumber="11">
<emphasis id="B9102019D025FFDBF6DCFAF3FF1FFAB4" bold="true" box="[136,223,1373,1397]" pageId="8" pageNumber="11">Colour.</emphasis>
Adult coloration in life: dark brown on anterior half or third of carapace and ambulatory legs; posterior half or two thirds of carapace orange or yellow; thoracic sternum greyish-white to grey with numerous grey specks; abdomen dark grey with small white spots; chelae bright orange; eyes bright yellow (
<figureCitation id="135FE08ED025FFDBF774FA53FEA6F9D4" box="[288,358,1533,1557]" captionStart="Fig" captionStartId="6.[136,172,2006,2028]" captionTargetBox="[136,1474,625,1991]" captionTargetId="figure-265@6.[136,1474,625,1991]" captionTargetPageId="6" captionText="Fig. 6. Colours in life. AC, Geosesarma dennerle, new species, Cilacap, Java; DF, G. hagen, new species, Cilacap, Java. A, B, specimens from type series; C, male (13.1 × 11.7 mm) (ZRC 2014.0271). A, B: 7°2559”S, 108°5550”E, Cilacap, Java; F, 7°2750”S, 108°5016”E, Cilacap, Java; CE, from aquarium trade, from Cilacap, Java (Photographs: Chris Lukhaup [A, B, F]; Tan Heok Hui [C]; Oliver Mengedoht (specimens not preserved) [D, E])." figureDoi="http://doi.org/10.5281/zenodo.4504266" httpUri="https://zenodo.org/record/4504266/files/figure.png" pageId="8" pageNumber="11">Figs. 6</figureCitation>
DF, 7E, F). The extent of bright orange on the dorsal surface of the carapace varies considerably, with some specimens being almost completely orange (
<figureCitation id="135FE08ED025FFDBF4F8F993FF5CF9B4" captionStart="Fig" captionStartId="6.[136,172,2006,2028]" captionTargetBox="[136,1474,625,1991]" captionTargetId="figure-265@6.[136,1474,625,1991]" captionTargetPageId="6" captionText="Fig. 6. Colours in life. AC, Geosesarma dennerle, new species, Cilacap, Java; DF, G. hagen, new species, Cilacap, Java. A, B, specimens from type series; C, male (13.1 × 11.7 mm) (ZRC 2014.0271). A, B: 7°2559”S, 108°5550”E, Cilacap, Java; F, 7°2750”S, 108°5016”E, Cilacap, Java; CE, from aquarium trade, from Cilacap, Java (Photographs: Chris Lukhaup [A, B, F]; Tan Heok Hui [C]; Oliver Mengedoht (specimens not preserved) [D, E])." figureDoi="http://doi.org/10.5281/zenodo.4504266" httpUri="https://zenodo.org/record/4504266/files/figure.png" pageId="8" pageNumber="11">Fig. 6D, E</figureCitation>
). The chelae in adults are bright or reddish-orange (
<figureCitation id="135FE08ED025FFDBF485F9F3FF68F954" captionStart="Fig" captionStartId="6.[136,172,2006,2028]" captionTargetBox="[136,1474,625,1991]" captionTargetId="figure-265@6.[136,1474,625,1991]" captionTargetPageId="6" captionText="Fig. 6. Colours in life. AC, Geosesarma dennerle, new species, Cilacap, Java; DF, G. hagen, new species, Cilacap, Java. A, B, specimens from type series; C, male (13.1 × 11.7 mm) (ZRC 2014.0271). A, B: 7°2559”S, 108°5550”E, Cilacap, Java; F, 7°2750”S, 108°5016”E, Cilacap, Java; CE, from aquarium trade, from Cilacap, Java (Photographs: Chris Lukhaup [A, B, F]; Tan Heok Hui [C]; Oliver Mengedoht (specimens not preserved) [D, E])." figureDoi="http://doi.org/10.5281/zenodo.4504266" httpUri="https://zenodo.org/record/4504266/files/figure.png" pageId="8" pageNumber="11">Figs. 6E</figureCitation>
,
<figureCitation id="135FE08ED025FFDBF6E0F9D3FF32F954" box="[180,242,1661,1685]" captionStart="Fig" captionStartId="7.[113,148,2009,2031]" captionTargetBox="[113,1451,733,1993]" captionTargetId="figure-298@7.[113,1451,733,1994]" captionTargetPageId="7" captionText="Fig. 7. A, type locality of Geosesarma dennerle, new species; B, burrows by the side of a stream bank dug by of G. dennerle, new species; C, a young female G. dennerle, new species, on the stream bank; D, type locality of G. hagen, new species; E, adult G. hagen, new species, on bank of stream; F, an adult male of G. hagen, new species, on moss covered stream bank. AC: 7°2559”S, 108°5550”E, Cilacap, Java; DF, 7°2750”S, 108°5016”E, Cilacap, Java. Specimens not preserved (Photographs: Chris Lukhaup)." figureDoi="http://doi.org/10.5281/zenodo.4504268" httpUri="https://zenodo.org/record/4504268/files/figure.png" pageId="8" pageNumber="11">7E, F</figureCitation>
), but those from smaller specimens are generally more reddish (
<figureCitation id="135FE08ED025FFDBF773F933FEBAF974" box="[295,378,1693,1717]" captionStart="Fig" captionStartId="6.[136,172,2006,2028]" captionTargetBox="[136,1474,625,1991]" captionTargetId="figure-265@6.[136,1474,625,1991]" captionTargetPageId="6" captionText="Fig. 6. Colours in life. AC, Geosesarma dennerle, new species, Cilacap, Java; DF, G. hagen, new species, Cilacap, Java. A, B, specimens from type series; C, male (13.1 × 11.7 mm) (ZRC 2014.0271). A, B: 7°2559”S, 108°5550”E, Cilacap, Java; F, 7°2750”S, 108°5016”E, Cilacap, Java; CE, from aquarium trade, from Cilacap, Java (Photographs: Chris Lukhaup [A, B, F]; Tan Heok Hui [C]; Oliver Mengedoht (specimens not preserved) [D, E])." figureDoi="http://doi.org/10.5281/zenodo.4504266" httpUri="https://zenodo.org/record/4504266/files/figure.png" pageId="8" pageNumber="11">Fig. 6F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37EAF80D025FFDBF6DCF973FE21F8B4" pageId="8" pageNumber="11" type="etymology">
<paragraph id="8BDBFC0BD025FFDBF6DCF973FE21F8B4" blockId="8.[136,775,1757,1909]" pageId="8" pageNumber="11">
<emphasis id="B9102019D025FFDBF6DCF973FECAF934" bold="true" box="[136,266,1757,1781]" pageId="8" pageNumber="11">Etymology.</emphasis>
<taxonomicName id="4C648788D025FFDBF746F973FE21F934" authorityName="Ng &amp; Schubart &amp; Lukhaup" authorityYear="2015" box="[274,481,1757,1781]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="11" phylum="Arthropoda" rank="species" species="hagen">
<emphasis id="B9102019D025FFDBF746F973FE21F934" box="[274,481,1757,1781]" italics="true" pageId="8" pageNumber="11">Geosesarma hagen</emphasis>
</taxonomicName>
is named after the Rolf C. Hagen Group of Companies, a major pet supplies company in
<collectingCountry id="F373BC9BD025FFDBF6DCF8B3FF2FF8F4" box="[136,239,1821,1845]" name="Germany" pageId="8" pageNumber="11">Germany</collectingCountry>
. They kindly supported the third authors projects in Java and subsequent work by the second author. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37EAF80D025FFDBF6DCF833FB5BFE74" pageId="8" pageNumber="11" type="biology_ecology">
<paragraph id="8BDBFC0BD025FFDBF6DCF833FB5BFE74" blockId="8.[136,776,1949,2101]" lastBlockId="8.[835,1474,189,437]" pageId="8" pageNumber="11">
<emphasis id="B9102019D025FFDBF6DCF833FF2CF874" bold="true" box="[136,236,1949,1973]" pageId="8" pageNumber="11">Ecology.</emphasis>
The crabs were found on a banana and rubber plantation on the side of a small hill, around
<quantity id="4C9C51EED025FFDBF4D6F813FD09F815" box="[642,713,1981,2005]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="11" unit="m" value="300.0">300 m</quantity>
from the road (
<figureCitation id="135FE08ED025FFDBF6A6F873FE84F834" box="[242,324,2013,2037]" captionStart="Fig" captionStartId="7.[113,148,2009,2031]" captionTargetBox="[113,1451,733,1993]" captionTargetId="figure-298@7.[113,1451,733,1994]" captionTargetPageId="7" captionText="Fig. 7. A, type locality of Geosesarma dennerle, new species; B, burrows by the side of a stream bank dug by of G. dennerle, new species; C, a young female G. dennerle, new species, on the stream bank; D, type locality of G. hagen, new species; E, adult G. hagen, new species, on bank of stream; F, an adult male of G. hagen, new species, on moss covered stream bank. AC: 7°2559”S, 108°5550”E, Cilacap, Java; DF, 7°2750”S, 108°5016”E, Cilacap, Java. Specimens not preserved (Photographs: Chris Lukhaup)." figureDoi="http://doi.org/10.5281/zenodo.4504268" httpUri="https://zenodo.org/record/4504268/files/figure.png" pageId="8" pageNumber="11">Fig 7D</figureCitation>
). There are many trickles of water and a somewhat larger rivulet in the small area where they occur. Adult crabs are terrestrial, hiding under the dense bottom vegetation, usually under and between rocks, but they also sometimes excavate burrows at the edge of the water bodies (
<figureCitation id="135FE08ED025FFDBF5CEFF53FC30FED4" box="[922,1008,253,277]" captionStart="Fig" captionStartId="7.[113,148,2009,2031]" captionTargetBox="[113,1451,733,1993]" captionTargetId="figure-298@7.[113,1451,733,1994]" captionTargetPageId="7" captionText="Fig. 7. A, type locality of Geosesarma dennerle, new species; B, burrows by the side of a stream bank dug by of G. dennerle, new species; C, a young female G. dennerle, new species, on the stream bank; D, type locality of G. hagen, new species; E, adult G. hagen, new species, on bank of stream; F, an adult male of G. hagen, new species, on moss covered stream bank. AC: 7°2559”S, 108°5550”E, Cilacap, Java; DF, 7°2750”S, 108°5016”E, Cilacap, Java. Specimens not preserved (Photographs: Chris Lukhaup)." figureDoi="http://doi.org/10.5281/zenodo.4504268" httpUri="https://zenodo.org/record/4504268/files/figure.png" pageId="8" pageNumber="11">Fig. 7E</figureCitation>
). They seem to be carnivorous, feeding at least partially on insects inhabiting the ground. While adult crabs are seldom found in the water and mostly live on the moist banks (
<figureCitation id="135FE08ED025FFDBF58EFEF3FBEEFEB4" box="[986,1070,349,373]" captionStart="Fig" captionStartId="7.[113,148,2009,2031]" captionTargetBox="[113,1451,733,1993]" captionTargetId="figure-298@7.[113,1451,733,1994]" captionTargetPageId="7" captionText="Fig. 7. A, type locality of Geosesarma dennerle, new species; B, burrows by the side of a stream bank dug by of G. dennerle, new species; C, a young female G. dennerle, new species, on the stream bank; D, type locality of G. hagen, new species; E, adult G. hagen, new species, on bank of stream; F, an adult male of G. hagen, new species, on moss covered stream bank. AC: 7°2559”S, 108°5550”E, Cilacap, Java; DF, 7°2750”S, 108°5016”E, Cilacap, Java. Specimens not preserved (Photographs: Chris Lukhaup)." figureDoi="http://doi.org/10.5281/zenodo.4504268" httpUri="https://zenodo.org/record/4504268/files/figure.png" pageId="8" pageNumber="11">Fig. 7F</figureCitation>
), juveniles are often in the water or very close to the waterline, and are much less frequently seen on the drier forest ground.
</paragraph>
</subSubSection>
<subSubSection id="C37EAF80D025FFDBF517FE73FC7AFD14" pageId="8" pageNumber="11" type="discussion">
<paragraph id="8BDBFC0BD025FFDBF517FE73FBEAFDB4" blockId="8.[835,1475,477,629]" pageId="8" pageNumber="11">
<emphasis id="B9102019D025FFDBF517FE73FC6CFE34" bold="true" box="[835,940,477,501]" pageId="8" pageNumber="11">Remarks</emphasis>
. Material obtained from the aquarium trade by
<bibRefCitation id="EFF581FAD025FFDBF517FE53FB6AFDD4" author="Rademacher M &amp; Mengedoht O" box="[835,1194,509,533]" pageId="8" pageNumber="11" refId="ref7463" refString="Rademacher M &amp; Mengedoht O (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, Germany, 95 pp." type="book" year="2011">Rademacher &amp; Mengedoht (2011)</bibRefCitation>
identified as “
<taxonomicName id="4C648788D025FFDBF36AFE50FA02FDD4" authority="Rot" authorityName="De Man" authorityYear="1892" box="[1342,1474,510,533]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9102019D025FFDBF36AFE50FA02FDD4" box="[1342,1474,510,533]" italics="true" pageId="8" pageNumber="11">Geosesarma</emphasis>
</taxonomicName>
“Rot” Vampirkrabbe” (p. 44) are
<taxonomicName id="4C648788D025FFDBF28FFDB0FA7AFDF4" authority="Ng &amp; Schubart &amp; Lukhaup, 2015" authorityName="Ng &amp; Schubart &amp; Lukhaup" authorityYear="2015" box="[1243,1466,541,565]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="11" phylum="Arthropoda" rank="species" species="hagen" status="sp. nov.">
<emphasis id="B9102019D025FFDBF28FFDB0FA7AFDF4" box="[1243,1466,541,565]" italics="true" pageId="8" pageNumber="11">Geosesarma hagen</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2239D62D025FFDBF517FD90FC0AFD94" box="[835,970,573,597]" pageId="8" pageNumber="11" rank="species">new species</taxonomicNameLabel>
, on the basis of their colour photographs and specimens examined.
</paragraph>
<paragraph id="8BDBFC0BD025FFDBF517FD30FC7AFD14" blockId="8.[835,1473,669,725]" pageId="8" pageNumber="11">For comparisons with closest congeners, see general discussion.</paragraph>
</subSubSection>
</treatment>
</document>