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<document id="1F9537FC57066F31D8A1B2E9D0C3B677" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFFE4FFBDFD95FC94A839EC76" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia amoeboides Moonlight" docType="treatment" docVersion="7" lastPageNumber="69" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="66" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D5E9985220A0F1C23F756679A44BE169" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FBD3AE884844CF14980CAF4931C7F108">
<mods:title id="15477AE763ED8338C8B25B6E58A8CF57">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="A7A25619DCB02C1F9FD73401F891680A" type="personal">
<mods:role id="3D10A54403BEA18771A9EE06BFF02B82">
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<mods:namePart id="299988DBBA877605CDBD92DC5B7E3289">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="520D2BF08702D7C9C985AE669A93DB3C">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="34581F5BEA44B32D52C25D92A93A1879" type="email">moonligp@tcd.ie</mods:nameIdentifier>
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<mods:roleTerm id="B37FF3B0038B1BBDF2D678B60A0C11F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93CE82C959D7AB2E0E346839D140B10D">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="8CF6E4E81B1A9E69D48B5650F9AA7A1F">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="E05846CA26E66B9681569ED55192B80C" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="EAE03F25A8C1E620323601EBDD7F71F5" type="personal">
<mods:role id="59EE303467649CD9E03D504EF2B16112">
<mods:roleTerm id="BE8CFFDA1775AA92EDD8EDF11FBD78E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E8E5385A8AD233C5FFDC0A8266255AA">Purvis, David A.</mods:namePart>
<mods:affiliation id="436E3840BE8495333D974FE5E7C4C5EE">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="2EF10297C7C3CB553C57B1E34A9E6E07" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="CDE73D3D72A819E0D66761B3D12F349D" type="personal">
<mods:role id="199CB0A08D71055E20B24106CACFC165">
<mods:roleTerm id="18CBE8C6BD7D00113DA5D7B6CB8DD80C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F9252233D9554F9214AA5A585A3CC2B1">Delves, Jay</mods:namePart>
<mods:affiliation id="4AB431AF1C57E9AC0AC5206B93BE508C">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="2EC7CE25DCC73CA3D510F9D8D8942691" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="2C129EF6E784646121F842E777E94137" type="personal">
<mods:role id="0239947726AB4348C27FACE0B1A08FB5">
<mods:roleTerm id="47005B7CC2E6529E30E170ED5727824F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFC030228123C3D4CE690A00C9143E8B">Allen, Josh P.</mods:namePart>
<mods:affiliation id="E9E5EF35B8FFB917F646A1C90DC3ED73">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="2D9408E9E225F0EF0FB0140D43884133" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:role id="61784A0FEFF98857E7BDBD62BD96E0DA">
<mods:roleTerm id="2183D2B5B348E62AB8938ABBAE65D53B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BFA66310DA9488BCE11E005AC219495D">Reynel, Carlos</mods:namePart>
<mods:affiliation id="DA94B6C56630DF6B9C3D5C589674A764">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="CBF0573E8AE051ACB0A841F3908DA845" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="ACDA6323000761503067A671B2FB6BD8">text</mods:typeOfResource>
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<mods:title id="0B0DB04039B3EDE3F955A9BEAFD52CEE">European Journal of Taxonomy</mods:title>
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<mods:part id="8994B3506D9FAFB746FE2947CD1542BC">
<mods:date id="B1B9C125D69B951790EB1A0A7659D785">2023</mods:date>
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<mods:number id="90671D50BDB523183F71B9F03FF7536C">2023-07-18</mods:number>
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<mods:detail id="E33044D844854CC06077B980902761DF" type="volume">
<mods:number id="6FA3BF64BDC0897C8C21EECB8E2C25B7">881</mods:number>
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<mods:start id="F61FA5BF3DB1E5EE7C1B06E2086A30E7">1</mods:start>
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<mods:url id="CE1BCF869C3FABF783A2688314E8CC85">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
</mods:location>
<mods:classification id="82B05C5D8B1D5507D2D9DD24A8D9042C">journal article</mods:classification>
<mods:identifier id="7386696DA1DA5A90B13D358EE08FEE69" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="8E56DAB3FE8BCD1F5B905B37D3B43E9A" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="F7A81B6FD581821DA9597644EF5550AE" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFFE4FFBDFD95FC94A839EC76" LSID="urn:lsid:plazi:treatment:03B26B4BFFE4FFBDFD95FC94A839EC76" httpUri="http://treatment.plazi.org/id/03B26B4BFFE4FFBDFD95FC94A839EC76" lastPageId="68" lastPageNumber="69" pageId="65" pageNumber="66">
<subSubSection id="C30189D6FFE4FFB8FD95FC94AEC0E93C" box="[588,999,810,837]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="8BA4DA5DFFE4FFB8FD95FC94AEC0E93C" blockId="65.[588,999,810,873]" box="[588,999,810,837]" pageId="65" pageNumber="66">
9.
<taxonomicName id="4C1BA1DEFFE4FFB8FDB3FC94AEC0E93C" ID-CoL="7SDNZ" ID-ENA="3018801" authority="Moonlight" authorityName="Moonlight" box="[618,999,810,837]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE4FFB8FDB3FC94AE43E93C" bold="true" box="[618,868,810,837]" italics="true" pageId="65" pageNumber="66">Begonia amoeboides</emphasis>
Moonlight
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFB8FD17FCF1AE41E910" box="[718,870,846,873]" pageId="65" pageNumber="66" type="description">
<paragraph id="8BA4DA5DFFE4FFB8FD17FCF1AE41E910" blockId="65.[588,999,810,873]" box="[718,870,846,873]" pageId="65" pageNumber="66">
<figureCitation id="1320C6D8FFE4FFB8FD17FCF1AE1AE910" box="[718,829,846,873]" captionStart="Fig" captionStartId="66.[189,232,1787,1813]" captionTargetBox="[194,1391,514,1732]" captionTargetId="figure-13@66.[189,1398,504,1740]" captionTargetPageId="66" captionText="Fig. 24. Distribution ofBegonia sect.Cyathocnemis (Klotzsch)A.DC. in Peru and surrounding countries. A. B. amoeboides Moonlight (red) andB. imbrexiformis Moonlight sp. nov. (blue).B. B. bracteosaA.DC. (red) and B. longinqua Moonlight (blue). C. B. cyathophora Poepp. &amp; Endl. (red), B. serratistipula Moonlight sp. nov. (blue) and C. vargasii Moonlight sp. nov. (green). D. B. lamolina Moonlight (red), B. lophoptera Rolfe (blue) and B. obtecticaulis Irmsch. (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178339" httpUri="https://zenodo.org/record/8178339/files/figure.png" pageId="65" pageNumber="66">Figs 24A</figureCitation>
,
<figureCitation id="1320C6D8FFE4FFB8FC90FCF1AE41E910" box="[841,870,846,873]" captionStart="Fig" captionStartId="67.[189,232,1928,1954]" captionTargetBox="[270,1326,278,1859]" captionTargetId="figure-12@67.[246,1342,259,1875]" captionTargetPageId="67" captionText="Fig. 25. Begonia amoeboides Moonlight. A. Habit. B. Stamen, front view. C. Staminate flower, front view. D. Pistillate flower, side view. E. Pistils, side view. Illustration by Claire Banks from P.W. Moonlight &amp; A. Daza 150 (E)." figureDoi="http://doi.org/10.5281/zenodo.8178343" httpUri="https://zenodo.org/record/8178343/files/figure.png" pageId="65" pageNumber="66">25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFB8FF64FC2FAE37E9D3" box="[189,784,912,938]" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="8BA4DA5DFFE4FFB8FF64FC2FAE37E9D3" blockId="65.[189,1400,911,1153]" box="[189,784,912,938]" pageId="65" pageNumber="66">
<emphasis id="B96F064FFFE4FFB8FF64FC2FAC12E9D3" box="[189,309,912,938]" italics="true" pageId="65" pageNumber="66">Phytotaxa</emphasis>
381 (1): 119 (
<bibRefCitation id="EF8AA7ACFFE4FFB8FE09FC2FAE23E9D3" author="Moonlight &amp; Reynel" box="[464,772,912,938]" firstAuthor="Moonlight" pageId="65" pageNumber="66" pagination="116 - 126" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" type="journal article" year="2018">Moonlight &amp; Reynel 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFB8FCCDFC2EAC82EE6F" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8BA4DA5DFFE4FFB8FCCDFC2EAC82EE6F" blockId="65.[189,1400,911,1153]" pageId="65" pageNumber="66">
–
<emphasis id="B96F064FFFE4FFB8FCFDFC2FAE45E9D3" bold="true" box="[804,866,912,938]" pageId="65" pageNumber="66">Type</emphasis>
:
<materialsCitation id="3B73D000FFE4FFB8FCABFC2EAEF5E98B" collectingDate="2016-01-31" collectionCode="MOL" collectorName="P. W. Moonlight &amp; A. Daza" country="Peru" elevation="2088" latitude="-5.6894445" location="Dist. Yambrasbamba" longLatPrecision="21" longitude="-77.81972" municipality="Prov. Bongara" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Amazonas Region" typeStatus="holotype">
<collectingCountry id="F30C9ACDFFE4FFB8FCABFC2EAE9AE9D2" box="[882,957,913,939]" name="Peru" pageId="65" pageNumber="66">PERU</collectingCountry>
–
<emphasis id="B96F064FFFE4FFB8FC08FC2FA854E9D3" bold="true" box="[977,1395,911,938]" pageId="65" pageNumber="66">
<collectingRegion id="49DF14BFFFE4FFB8FC08FC2FA991E9D3" box="[977,1206,912,938]" country="Peru" name="Amazonas" pageId="65" pageNumber="66">Amazonas Region</collectingRegion>
:
<collectingMunicipality id="6BC04027FFE4FFB8FB19FC2FA854E9D3" box="[1216,1395,911,938]" pageId="65" pageNumber="66">Prov. Bongará</collectingMunicipality>
</emphasis>
•
<location id="8EC48C86FFE4FFB8FF24FC0BACD2E9B7" LSID="urn:lsid:plazi:treatment:03B26B4BFFE4FFBDFD95FC94A839EC76:8EC48C86FFE4FFB8FF24FC0BACD2E9B7" box="[253,501,948,974]" country="Peru" latitude="-5.6894445" longLatPrecision="21" longitude="-77.81972" municipality="Prov. Bongara" name="Dist. Yambrasbamba" pageId="65" pageNumber="66" stateProvince="Amazonas Region">Dist. Yambrasbamba</location>
,
<locationDeviation id="9A16C37FFFE4FFB8FDDAFC0BAE40E9B7" box="[515,871,948,975]" pageId="65" pageNumber="66">road from Amazonas to Rioja</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFFE4FFB8FCADFC0DAED0E9B7" box="[884,1015,946,974]" degrees="5" direction="south" minutes="41" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="22" value="-5.6894445">5°41′22″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFE4FFB8FBDCFC0DA986E9B6" box="[1029,1185,946,975]" degrees="77" direction="west" minutes="49" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="11" value="-77.81972">77°49′11″ W</geoCoordinate>
;
<elevation id="00363D6EFFE4FFB8FB77FC0BA86DE9B7" box="[1198,1354,948,975]" metricMagnitude="3" metricUnit="m" metricValue="2.088" pageId="65" pageNumber="66" unit="m" value="2088.0">
<quantity id="4CE377B8FFE4FFB8FB77FC0BA82EE9B6" box="[1198,1289,948,975]" metricMagnitude="3" metricUnit="m" metricValue="2.088" pageId="65" pageNumber="66" unit="m" value="2088.0">2088 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFFE4FFB8FA80FC0BAC44E988" pageId="65" pageNumber="66" value="2016-01-31">
<collectingDate id="EFE10575FFE4FFB8FA80FC0BAC44E988" pageId="65" pageNumber="66" value="2016-01-31">31 Jan. 2016</collectingDate>
</date>
;
<emphasis id="B96F064FFFE4FFB8FEADFC67AFDDE98B" box="[372,762,983,1010]" italics="true" pageId="65" pageNumber="66">
<collectorName id="26EEBF8BFFE4FFB8FEADFC67AF09E988" box="[372,558,983,1009]" pageId="65" pageNumber="66">P.W. Moonlight</collectorName>
&amp;
<collectorName id="26EEBF8BFFE4FFB8FD84FC67AFE6E988" box="[605,705,984,1010]" pageId="65" pageNumber="66">A. Daza</collectorName>
150
</emphasis>
; holotype:
<collectionCode id="ED0A4298FFE4FFB8FC50FC67AEE9E98B" box="[905,974,984,1010]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="65" pageNumber="66" type="Herbarium">MOL</collectionCode>
;
</materialsCitation>
<materialsCitation id="3B73D000FFE4FFB8FC07FC68AC82EE6F" collectingDate="2016-01-31" collectionCode="E, K, MO" collectorName="P. W. Moonlight &amp; A. Daza" country="Peru" elevation="2088" latitude="-5.6894445" location="Dist. Yambrasbamba" longLatPrecision="21" longitude="-77.81972" municipality="Prov. Bongara" pageId="65" pageNumber="66" specimenCode="E00934304, MO-3237376" specimenCount="1" stateProvince="Amazonas Region" typeStatus="isotype">
isotypes:
<collectionCode id="ED0A4298FFE4FFB8FB8AFC67A942E98B" box="[1107,1125,984,1010]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="65" pageNumber="66" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FBA3FC67A825E98B" box="[1146,1282,984,1010]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="65" pageNumber="66" type="Herbarium">E00934304</specimenCode>
],
<collectionCode id="ED0A4298FFE4FFB8FAC6FC67A810E98B" box="[1311,1335,984,1010]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="65" pageNumber="66" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED0A4298FFE4FFB8FA9EFC67A851E98B" box="[1351,1398,984,1010]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="65" pageNumber="66" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FF2DFC44ACB3EE6C" box="[244,404,1019,1045]" pageId="65" pageNumber="66">MO-3237376</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFB8FF64FB9FA850EEF8" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="8BA4DA5DFFE4FFB8FF64FB9FAC1FEE24" blockId="65.[189,1400,911,1153]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="AB0BFD73FFE4FFB8FF64FB9FAC1FEE24" pageId="65" pageNumber="66">
<taxonomicName id="4C1BA1DEFFE4FFB8FF64FB9FACD5EE40" ID-CoL="87YTG" authority="Rolfe" authorityName="Rolfe" box="[189,498,1055,1081]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="lophoptera">
<emphasis id="B96F064FFFE4FFB8FF64FB9FAC8EEE40" box="[189,425,1055,1081]" italics="true" pageId="65" pageNumber="66">Begonia lophoptera</emphasis>
Rolfe
</taxonomicName>
pro parte in R.Vásquez
<emphasis id="B96F064FFFE4FFB8FCC1FB9FAE73EE40" box="[792,852,1055,1081]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
,
<emphasis id="B96F064FFFE4FFB8FCBDFBA0AEF3EE40" box="[868,980,1055,1081]" italics="true" pageId="65" pageNumber="66">Arnaldoa</emphasis>
12 (1–2):
<treatmentCitation id="0ABAFC4CFFE4FFB8FB8DFBA0AC1FEE24" author="Vásquez et al." page="112-125" pageId="65" pageNumber="66" year="2005">
112–125 (
<bibRefCitation id="EF8AA7ACFFE4FFB8FB13FBA0AC0BEE24" author="Vasquez" etAl="et al." firstAuthor="Vasquez" pageId="65" pageNumber="66" pagination="112 - 125" refId="ref233185" refString="Vasquez R., Rojas R. G,. Monteagudo A. M., Meza K. V., van der Werff H., Ortiz-Gentry R. &amp; Catchpole D. 2005. Flora vascular de la selva central del Peru: una aproximacion de la copmposicion floristica de tres areas naturales protegidas. Arnaldoa 12 (1 - 2): 112 - 125." type="journal article" year="2005">
Vásquez
<emphasis id="B96F064FFFE4FFB8FAE2FB9FA85FEE40" box="[1339,1400,1055,1081]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
2005
</bibRefCitation>
).
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA4DA5DFFE4FFB8FF64FBD8A850EEF8" blockId="65.[189,1400,911,1153]" box="[189,1399,1126,1153]" pageId="65" pageNumber="66">
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<taxonomicName id="4C1BA1DEFFE4FFB8FF64FBD8ACA3EEF9" ID-CoL="7SHT4" authorityName="C. DC." box="[189,388,1126,1152]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="65" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiliifolia">
<emphasis id="B96F064FFFE4FFB8FF64FBD8ACA3EEF9" box="[189,388,1126,1152]" italics="true" pageId="65" pageNumber="66">Begonia tiliifolia</emphasis>
</taxonomicName>
auct. non C.DC.: R.Vásquez
<emphasis id="B96F064FFFE4FFB8FD0FFBD8AE36EEF9" box="[726,785,1126,1152]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
,
<emphasis id="B96F064FFFE4FFB8FCC6FBD9AEA8EEF9" box="[799,911,1126,1152]" italics="true" pageId="65" pageNumber="66">Arnaldoa</emphasis>
12 (1–2):
<treatmentCitation id="0ABAFC4CFFE4FFB8FBDCFBD9A850EEF8" author="Vásquez et al." box="[1029,1399,1126,1153]" page="112-125" pageId="65" pageNumber="66" year="2005">
112–125 (
<bibRefCitation id="EF8AA7ACFFE4FFB8FBA1FBD9A84CEEF8" author="Vasquez" box="[1144,1387,1126,1153]" etAl="et al." firstAuthor="Vasquez" pageId="65" pageNumber="66" pagination="112 - 125" refId="ref233185" refString="Vasquez R., Rojas R. G,. Monteagudo A. M., Meza K. V., van der Werff H., Ortiz-Gentry R. &amp; Catchpole D. 2005. Flora vascular de la selva central del Peru: una aproximacion de la copmposicion floristica de tres areas naturales protegidas. Arnaldoa 12 (1 - 2): 112 - 125." type="journal article" year="2005">
Vásquez
<emphasis id="B96F064FFFE4FFB8FB3AFBD8A802EEF9" box="[1251,1317,1126,1152]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
2005
</bibRefCitation>
).
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFB8FF64FB17AF7CEF6D" pageId="65" pageNumber="66" type="etymology">
<paragraph id="8BA4DA5DFFE4FFB8FF64FB17AC62EEBB" blockId="65.[189,325,1192,1218]" box="[189,325,1192,1218]" pageId="65" pageNumber="66">
<heading id="D0EC6D31FFE4FFB8FF64FB17AC62EEBB" bold="true" box="[189,325,1192,1218]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96F064FFFE4FFB8FF64FB17AC62EEBB" bold="true" box="[189,325,1192,1218]" pageId="65" pageNumber="66">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFE4FFB8FF64FB68AF7CEF6D" blockId="65.[189,1399,1239,1300]" pageId="65" pageNumber="66">The epithet refers to the irregularly multifid stigmas of the species, which resemble the irregularly branching pseudopods of amoebae.</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFB8FF64FA83A9BDED2D" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="8BA4DA5DFFE4FFB8FF64FA83AF09EF2F" blockId="65.[189,558,1340,1366]" box="[189,558,1340,1366]" pageId="65" pageNumber="66">
<heading id="D0EC6D31FFE4FFB8FF64FA83AF09EF2F" bold="true" box="[189,558,1340,1366]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96F064FFFE4FFB8FF64FA83AF09EF2F" bold="true" box="[189,558,1340,1366]" pageId="65" pageNumber="66">Selected specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFE4FFB8FF64FAD3A9BDED2D" blockId="65.[188,1399,1386,1877]" pageId="65" pageNumber="66">
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<collectingCountry id="F30C9ACDFFE4FFB8FF64FAD3AC2FEFFF" box="[189,264,1388,1414]" name="Peru" pageId="65" pageNumber="66">PERU</collectingCountry>
–
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<collectingRegion id="49DF14BFFFE4FFB8FEF5FAD4AF0DEFFC" box="[300,554,1386,1413]" country="Peru" name="San Martin" pageId="65" pageNumber="66">San Martín Region</collectingRegion>
:
<collectingMunicipality id="6BC04027FFE4FFB8FDE2FAD4AFEAEFFC" box="[571,717,1387,1413]" country="Peru" name="Prov. Rioja" pageId="65" pageNumber="66" stateProvince="San Martin">Prov. Rioja</collectingMunicipality>
</emphasis>
•
<collectingCounty id="62C5A2D1FFE4FFB8FD34FAD4AEFFEFFC" box="[749,984,1387,1413]" pageId="65" pageNumber="66">Dist. Pardo Miguel</collectingCounty>
,
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;
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;
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<collectorName id="26EEBF8BFFE4FFB8FC7BFA2FA90FEFD0" box="[930,1064,1423,1449]" pageId="65" pageNumber="66">D.N. Smith</collectorName>
6005
</emphasis>
;
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[
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],
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[
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.
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<materialsCitation id="3B73D000FFE4FFB8FE26FA0CAC23EC6D" collectingDate="2018-07-03" collectionCode="E, USM" collectorName="P. W. Moonlight" county="Dist. Yambrasbamba" elevation="2075" latitude="-5.687778" location="Road from Amazonas to Rioja" longLatPrecision="21" longitude="-77.78611" municipality="Prov. Bongara" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Amazonas Region">
–
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<collectingRegion id="49DF14BFFFE4FFB8FDC1FA0DAE24EFB5" box="[536,771,1458,1484]" country="United States of America" name="Amazonas Region" pageId="65" pageNumber="66">Amazonas Region</collectingRegion>
:
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•
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;
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<collectingDate id="EFE10575FFE4FFB8FC77FA69A91DEF89" box="[942,1082,1494,1520]" pageId="65" pageNumber="66" value="2018-07-03">3 Jul. 2018</collectingDate>
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;
<emphasis id="B96F064FFFE4FFB8FB93FA68A86DEF89" box="[1098,1354,1494,1520]" italics="true" pageId="65" pageNumber="66">
<collectorName id="26EEBF8BFFE4FFB8FB93FA68A824EF89" box="[1098,1283,1494,1520]" pageId="65" pageNumber="66">P.W. Moonlight</collectorName>
1258
</emphasis>
;
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,
<collectionCode id="ED0A4298FFE4FFB8FF64FA45AC27EC6D" box="[189,256,1530,1556]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="65" pageNumber="66">USM</collectionCode>
.
</materialsCitation>
<materialsCitation id="3B73D000FFE4FFB8FED7FA45A850EC22" collectingDate="2009-10-22" collectionCode="HOXA, E, MO, USM" collectorName="L. Valenzuela &amp; A. Monteagudo &amp; M. Cueva &amp; A. Pena &amp; J. Mateo &amp; R. Rivera" county="Dist. Huancabamba" elevation="2478" latitude="-10.314445" location="Sector Oso Playa" longLatPrecision="21" longitude="-75.57528" municipality="Prov. Oxapampa" pageId="65" pageNumber="66" specimenCode="E00934198, MO-2991353" specimenCount="1" stateProvince="Pasco Region">
–
<emphasis id="B96F064FFFE4FFB8FEFEFA45AF98EC6A" bold="true" box="[295,703,1529,1556]" pageId="65" pageNumber="66">
<collectingRegion id="49DF14BFFFE4FFB8FEFEFA45ACF1EC6A" box="[295,470,1529,1556]" country="United States of America" name="Pasco Region" pageId="65" pageNumber="66">Pasco Region</collectingRegion>
:
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</emphasis>
•
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,
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;
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<quantity id="4CE377B8FFE4FFB8FEBDF9A2AC99EC41" box="[356,446,1565,1592]" metricMagnitude="3" metricUnit="m" metricValue="2.478" pageId="65" pageNumber="66" unit="m" value="2478.0">2478 m</quantity>
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L.
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Valenzuela,
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,
<collectorName id="26EEBF8BFFE4FFB8FF66F9FDAC12EC22" box="[191,309,1601,1627]" pageId="65" pageNumber="66">M. Cueva</collectorName>
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<collectorName id="26EEBF8BFFE4FFB8FE9BF9FEAC83EC22" box="[322,420,1601,1627]" pageId="65" pageNumber="66">A. Peña</collectorName>
,
<collectorName id="26EEBF8BFFE4FFB8FE68F9FDAF3DEC22" box="[433,538,1602,1627]" pageId="65" pageNumber="66">J. Mateo</collectorName>
&amp;
<collectorName id="26EEBF8BFFE4FFB8FD9BF9FDAF96EC22" box="[578,689,1602,1627]" pageId="65" pageNumber="66">R. Rivera</collectorName>
13842;
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[
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•
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;
<emphasis id="B96F064FFFE4FFB8FBC5F937AF9FECBF" italics="true" pageId="65" pageNumber="66">
<collectorName id="26EEBF8BFFE4FFB8FBC5F937A9F2ECDB" box="[1052,1237,1672,1698]" pageId="65" pageNumber="66">A. Monteagudo</collectorName>
,
<collectorName id="26EEBF8BFFE4FFB8FB06F936A855ECDB" box="[1247,1394,1672,1698]" pageId="65" pageNumber="66">L. Cárdenas</collectorName>
,
<collectorName id="26EEBF8BFFE4FFB8FF65F913AC07ECBC" box="[188,288,1708,1734]" pageId="65" pageNumber="66">A. Peña</collectorName>
,
<collectorName id="26EEBF8BFFE4FFB8FEE9F913AC94ECBC" box="[304,435,1708,1733]" pageId="65" pageNumber="66">J.L. Mateo</collectorName>
&amp;
<collectorName id="26EEBF8BFFE4FFB8FE39F913AF46ECBC" box="[480,609,1708,1733]" pageId="65" pageNumber="66">R. Francis</collectorName>
12001
</emphasis>
;
<collectionCode id="ED0A4298FFE4FFB8FD12F913AE04ECBF" box="[715,803,1708,1734]" country="Peru" httpUri="http://grbio.org/cool/yriw-6hkr" name="Estación biológica del Jardin Botanico de Missouri" pageId="65" pageNumber="66" type="Herbarium">HOXA</collectionCode>
,
<collectionCode id="ED0A4298FFE4FFB8FCEBF913AE46ECBF" box="[818,865,1708,1734]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="65" pageNumber="66" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FCAFF913A932ECBF" box="[886,1045,1708,1734]" pageId="65" pageNumber="66">MO-2183947</specimenCode>
],
<collectionCode id="ED0A4298FFE4FFB8FBEBF913A970ECBF" box="[1074,1111,1708,1734]" pageId="65" pageNumber="66">US</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FBB5F914A821ECBF" box="[1132,1286,1707,1734]" pageId="65" pageNumber="66">US00951211</specimenCode>
],
<collectionCode id="ED0A4298FFE4FFB8FAFBF913A845ECBF" box="[1314,1378,1708,1734]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="65" pageNumber="66">USM</collectionCode>
</materialsCitation>
</collectingCounty>
</collectorName>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B73D000FFE4FFB8FF64F970AE0FED74" collectingDate="1984-07-13" collectionCode="MO, USM" collectorName="D. N. Smith" elevation="2700" latitude="-10.383333" location="Trail to summit of Cordillera Yanachaga via rio San Daniel" longLatPrecision="1298" longitude="-75.45" pageId="65" pageNumber="66" specimenCode="MO-1835942" specimenCount="1">
<location id="8EC48C86FFE4FFB8FF64F970AE59EC90" LSID="urn:lsid:plazi:treatment:03B26B4BFFE4FFBDFD95FC94A839EC76:8EC48C86FFE4FFB8FF64F970AE59EC90" box="[189,894,1743,1770]" latitude="-10.383333" longLatPrecision="1298" longitude="-75.45" name="Trail to summit of Cordillera Yanachaga via rio San Daniel" pageId="65" pageNumber="66">Trail to summit of Cordillera Yanachaga via río San Daniel</location>
;
<elevation id="00363D6EFFE4FFB8FC53F970A902EC90" box="[906,1061,1743,1770]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="65" pageNumber="66" unit="m" value="2700.0">
<quantity id="4CE377B8FFE4FFB8FC53F970AEC3EC93" box="[906,996,1743,1770]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="65" pageNumber="66" unit="m" value="2700.0">2700 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE2FBC9AFFE4FFB8FBEAF971A9BDEC90" box="[1075,1178,1742,1770]" degrees="10" direction="south" minutes="23" orientation="latitude" pageId="65" pageNumber="66" precision="925" value="-10.383333">10°23′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFE4FFB8FB71F971A83DEC93" box="[1192,1306,1742,1770]" degrees="75" direction="west" minutes="27" orientation="longitude" pageId="65" pageNumber="66" precision="925" value="-75.45">75°27′ W</geoCoordinate>
;
<date id="FFA5FC9DFFE4FFB8FAFEF970ADDCED74" pageId="65" pageNumber="66" value="1984-07-13">
<collectingDate id="EFE10575FFE4FFB8FAFEF970ADDCED74" pageId="65" pageNumber="66" value="1984-07-13">13 Jul. 1984</collectingDate>
</date>
;
<emphasis id="B96F064FFFE4FFB8FED3F94BACF1ED74" box="[266,470,1779,1805]" italics="true" pageId="65" pageNumber="66">
<collectorName id="26EEBF8BFFE4FFB8FED3F94BACB7ED74" box="[266,400,1779,1805]" pageId="65" pageNumber="66">D.N. Smith</collectorName>
7801
</emphasis>
;
<collectionCode id="ED0A4298FFE4FFB8FE30F94CAF3FED74" box="[489,536,1779,1805]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="65" pageNumber="66" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FDF4F94CAFEAED77" box="[557,717,1779,1806]" pageId="65" pageNumber="66">MO-1835942</specimenCode>
],
<collectionCode id="ED0A4298FFE4FFB8FD31F94CAE0FED74" box="[744,808,1779,1805]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="65" pageNumber="66">USM</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FFE4FFB8FCEBF94CA9B1ED2D" collectingDate="2009-02-26" collectionCode="HOXA, E, MO, USM" collectorName="R. Vasquez &amp; L. Valenzuela &amp; R. Rivera" elevation="1951" latitude="-10.456944" location="Parque Nacional Yanachaga-Chemillen" longLatPrecision="21" longitude="-75.45555" pageId="65" pageNumber="66" specimenCode="E00934197, MO-3009205" specimenCount="1">
•
<location id="8EC48C86FFE4FFB8FC9EF94CA83AED74" LSID="urn:lsid:plazi:treatment:03B26B4BFFE4FFBDFD95FC94A839EC76:8EC48C86FFE4FFB8FC9EF94CA83AED74" box="[839,1309,1779,1805]" latitude="-10.456944" longLatPrecision="21" longitude="-75.45555" name="Parque Nacional Yanachaga-Chemillen" pageId="65" pageNumber="66">Parque Nacional Yanachaga-Chemillén</location>
,
<location id="8EC48C86FFE4FFB8FAF5F94CAF08ED49" LSID="urn:lsid:plazi:treatment:03B26B4BFFE4FFBDFD95FC94A839EC76:8EC48C86FFE4FFB8FAF5F94CAF08ED49" latitude="-10.456944" longLatPrecision="21" longitude="-75.45555" name="Sector San Daniel-Quebrada Colmena" pageId="65" pageNumber="66">Sector San Daniel-Quebrada Colmena</location>
;
<elevation id="00363D6EFFE4FFB8FDE3F8A9AFF4ED49" box="[570,723,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.9509999999999998" pageId="65" pageNumber="66" unit="m" value="1951.0">
<quantity id="4CE377B8FFE4FFB8FDE3F8A9AFB4ED48" box="[570,659,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.9509999999999998" pageId="65" pageNumber="66" unit="m" value="1951.0">1951 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE2FBC9AFFE4FFB8FD39F8AAAE57ED49" box="[736,880,1813,1841]" degrees="10" direction="south" minutes="27" orientation="latitude" pageId="65" pageNumber="66" precision="15" seconds="25" value="-10.456944">10°27′25″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFE4FFB8FCA5F8AAA930ED48" box="[892,1047,1813,1841]" degrees="75" direction="west" minutes="27" orientation="longitude" pageId="65" pageNumber="66" precision="15" seconds="20" value="-75.45555">75°27′20″ W</geoCoordinate>
;
<date id="FFA5FC9DFFE4FFB8FBFAF8A9A9E7ED49" box="[1059,1216,1814,1840]" pageId="65" pageNumber="66" value="2009-02-26">
<collectingDate id="EFE10575FFE4FFB8FBFAF8A9A9E7ED49" box="[1059,1216,1814,1840]" pageId="65" pageNumber="66" value="2009-02-26">26 Feb. 2009</collectingDate>
</date>
;
<emphasis id="B96F064FFFE4FFB8FB15F8A8AF02ED2D" italics="true" pageId="65" pageNumber="66">
<collectorName id="26EEBF8BFFE4FFB8FB15F8A8A874ED48" box="[1228,1363,1815,1841]" pageId="65" pageNumber="66">R. Vásquez</collectorName>
,
<collectorName id="26EEBF8BFFE4FFB8FA86F8A8AC1AED2D" pageId="65" pageNumber="66">L. Valenzuela</collectorName>
&amp;
<collectorName id="26EEBF8BFFE4FFB8FEBAF884ACF4ED2D" box="[355,467,1851,1876]" pageId="65" pageNumber="66">R. Rivera</collectorName>
35403
</emphasis>
;
<collectionCode id="ED0A4298FFE4FFB8FDEDF885AFABED2D" box="[564,652,1850,1876]" collectionName="HOXA" country="Peru" httpUri="http://grbio.org/cool/yriw-6hkr" name="Estación biológica del Jardin Botanico de Missouri" pageId="65" pageNumber="66" type="Herbarium">HOXA</collectionCode>
,
<collectionCode id="ED0A4298FFE4FFB8FD41F885AF8DED2D" box="[664,682,1850,1876]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="65" pageNumber="66" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FD62F885AE65ED2D" box="[699,834,1850,1876]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="65" pageNumber="66" type="Herbarium">E00934197</specimenCode>
],
<collectionCode id="ED0A4298FFE4FFB8FC82F885AEADED2D" box="[859,906,1850,1876]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="65" pageNumber="66" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFE4FFB8FC42F885A91CED2C" box="[923,1083,1850,1877]" pageId="65" pageNumber="66">MO-3009205</specimenCode>
],
<collectionCode id="ED0A4298FFE4FFB8FB8AF885A9B1ED2D" box="[1107,1174,1850,1876]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="65" pageNumber="66">USM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE4FFBDFF64F8C4A83CE906" lastPageId="68" lastPageNumber="69" pageId="65" pageNumber="66" type="description">
<paragraph id="8BA4DA5DFFE4FFB8FF64F8C4AC68EDEC" blockId="65.[189,335,1915,1941]" box="[189,335,1915,1941]" pageId="65" pageNumber="66">
<heading id="D0EC6D31FFE4FFB8FF64F8C4AC68EDEC" bold="true" box="[189,335,1915,1941]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B96F064FFFE4FFB8FF64F8C4AC68EDEC" bold="true" box="[189,335,1915,1941]" pageId="65" pageNumber="66">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFE4FFBDFF64F814A83CE906" blockId="65.[188,1399,1963,2025]" lastBlockId="68.[187,1401,265,895]" lastPageId="68" lastPageNumber="69" pageId="65" pageNumber="66">
Caulescent herb, to
<quantity id="4CE377B8FFE4FFB8FE7DF814ACDDEDBC" box="[420,506,1963,1990]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="66" unit="cm" value="150.0">150 cm</quantity>
high.
<emphasis id="B96F064FFFE4FFB8FD9BF814AF5CEDBC" box="[578,635,1963,1989]" italics="true" pageId="65" pageNumber="66">Stem</emphasis>
erect, occasionally branching; internodes to
<quantity id="4CE377B8FFE4FFB8FB5FF814A9E9EDBC" box="[1158,1230,1963,1990]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="66" unit="cm" value="15.0">15 cm</quantity>
long, to
<quantity id="4CE377B8FFE4FFB8FAEDF814A850EDBC" box="[1332,1399,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="66" unit="mm" value="8.0">8 mm</quantity>
thick, succulent, dark green to dark red, sparsely to densely pilose.
<emphasis id="B96F064FFFE4FFB8FC18F871A938ED91" box="[961,1055,1998,2024]" italics="true" pageId="65" pageNumber="66">Stipules</emphasis>
deciduous, lanceolate, 10–40 ×
<quantity id="4CE377B8FFE7FFBBFF0DFEB6AC61EB5D" box="[212,326,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="66" pageNumber="67" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
, apex acute, translucent, pale green to dark brown, glabrous, margin entire, aciliate, the base surrounded by squamous hairs.
<emphasis id="B96F064FFFE7FFBBFDE3FE92AFABEB3F" box="[570,652,301,326]" italics="true" pageId="66" pageNumber="67">Leaves</emphasis>
&gt; 5, alternate, basifixed; petiole
<quantity id="4CE377B8FFE7FFBBFBCEFE93A98DEB3E" box="[1047,1194,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="13.5" metricValueMin="3.5" pageId="66" pageNumber="67" unit="cm" value="8.5" valueMax="13.5" valueMin="3.5">3.5–13.5 cm</quantity>
long, dark green to dark red, densely pilose; blade subsymmetric, reniform, to 11 ×
<quantity id="4CE377B8FFE7FFBBFC0DFEF0A907EB13" box="[980,1056,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="66" pageNumber="67" unit="cm" value="16.0">16 cm</quantity>
, sub-succulent, apex broadly rounded, base truncate to cordate, basal lobes not overlapping, sinus to
<quantity id="4CE377B8FFE7FFBBFBDBFECCA972EBF4" box="[1026,1109,371,397]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="mm" value="20.0">20 mm</quantity>
deep, margin irregularly double-dentate, with 2–8 cusps around the lamina, ciliate, upper surface green, sparsely pilose, lower surface pale green, sparsely pilose, densely pilose on the veins, veins palmate, 8–12 veined from the base.
<emphasis id="B96F064FFFE1FFBDFF62FE93AC44EB3F" box="[187,355,300,326]" italics="true" pageId="68" pageNumber="69">Inflorescences</emphasis>
1–3, bisexual, axillary, erect, cymose, with up to 8 branches, bearing up to 16 staminate flowers and 8 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FFE1FFBDFCB5FEEFAE91EB13" box="[876,950,336,362]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="69" unit="cm" value="10.0">10 cm</quantity>
long, pale green to pale red, sparsely to densely pilose, bracts deciduous, lanceolate to ovate, 6–13 ×
<quantity id="4CE377B8FFE1FFBDFC7FFECBA92DEBF7" box="[934,1034,372,398]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="68" pageNumber="69" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3–8 mm</quantity>
, translucent, white, glabrous to sparsely pilose, apex acute, margin entire, ciliate.
<emphasis id="B96F064FFFE1FFBDFCD5FE28AEF8EBC8" box="[780,991,407,433]" italics="true" pageId="68" pageNumber="69">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FFE1FFBDFBA1FE28A9ECEBC8" box="[1144,1227,407,434]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="68" pageNumber="69" unit="mm" value="15.0">15 mm</quantity>
long, sparsely pilose; tepals 2, spreading, ovate, 8–18 ×
<quantity id="4CE377B8FFE1FFBDFD67FE05AE12EBAC" box="[702,821,442,469]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="68" pageNumber="69" unit="mm" value="11.0" valueMax="15.0" valueMin="7.0">7–15 mm</quantity>
, apex acute, white or rarely pink, inner surface glabrous, outer surface pilose, margin entire to irregularly serrate, aciliate to ciliate; stamens 60–100, spreading, yellow, filaments
<quantity id="4CE377B8FFE1FFBDFDCDFDBEAFACE865" box="[532,651,513,540]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="68" pageNumber="69" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, free, anthers linear, ca 2 ×
<quantity id="4CE377B8FFE1FFBDFBC2FDBEA95DE865" box="[1051,1146,513,540]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="68" pageNumber="69" unit="mm" value="0.8">0.8 mm</quantity>
, dehiscing via lateral slits, connectives extending to
<quantity id="4CE377B8FFE1FFBDFDF2FD9AAFAEE846" box="[555,649,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="68" pageNumber="69" unit="mm" value="0.4">0.4 mm</quantity>
, symmetrically basifixed.
<emphasis id="B96F064FFFE1FFBDFC19FD9AA9AEE846" box="[960,1161,549,575]" italics="true" pageId="68" pageNumber="69">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FFE1FFBDFAFAFD9AA850E846" box="[1315,1399,549,576]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="68" pageNumber="69" unit="mm" value="15.0">15 mm</quantity>
long; bracteoles 3, positioned directly beneath the ovary, lanceolate to circular, 8–10 ×
<quantity id="4CE377B8FFE1FFBDFB17FDF7A815E81A" box="[1230,1330,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="68" pageNumber="69" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, apex acute, translucent, white, glabrous to short-pilose, margin entire, aciliate to ciliate; tepals 5, subequal, deciduous in fruit, projecting, lanceolate to ovate, 9–10 ×
<quantity id="4CE377B8FFE1FFBDFCBBFD30AEE1E8D3" box="[866,966,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="68" pageNumber="69" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, apex acute to obtuse, white or rarely pink, inner surface glabrous, outer surface sparsely pilose, margin entire, aciliate; ovary body ovoid, 4–11 ×
<quantity id="4CE377B8FFE1FFBDFEC5FD69ACA3E888" box="[284,388,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="68" pageNumber="69" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">2–6 mm</quantity>
, pale green, sparsely pilose, unequally 3-winged, wings triangular, the largest wing notched, 4–10 ×
<quantity id="4CE377B8FFE1FFBDFE51FD45ACC9E96D" box="[392,494,762,788]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="68" pageNumber="69" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 mm</quantity>
, smallest 4–9 ×
<quantity id="4CE377B8FFE1FFBDFD6AFD46AE61E96D" box="[691,838,761,788]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="68" pageNumber="69" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free,
<quantity id="4CE377B8FFE1FFBDFD3FFCA2AE6FE941" box="[742,840,797,824]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="68" pageNumber="69" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, many times divided, stigmatic papillae in a spirally-twisted band.
<emphasis id="B96F064FFFE1FFBDFE0DFCFEAFB3E923" box="[468,660,832,858]" italics="true" pageId="68" pageNumber="69">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FFE1FFBDFD61FCFFAE2DE922" box="[696,778,832,859]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="68" pageNumber="69" unit="mm" value="25.0">25 mm</quantity>
long.
<emphasis id="B96F064FFFE1FFBDFC8BFCFEAEE8E923" box="[850,975,832,858]" italics="true" pageId="68" pageNumber="69">Fruit body</emphasis>
ovoid, to 14 ×
<quantity id="4CE377B8FFE1FFBDFB59FCFEA9E1E922" box="[1152,1222,833,859]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="68" pageNumber="69" unit="mm" value="7.0">7 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 12 ×
<quantity id="4CE377B8FFE1FFBDFC58FCDBAEF0E906" box="[897,983,868,895]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="68" pageNumber="69" unit="mm" value="10.0">10 mm</quantity>
, the smallest to 12 ×
<quantity id="4CE377B8FFE1FFBDFB09FCDBA830E906" box="[1232,1303,868,895]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="68" pageNumber="69" unit="mm" value="4.0">4 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF648AD5FFE7FFBBFF64F944AC17ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178339" ID-Zenodo-Dep="8178339" httpUri="https://zenodo.org/record/8178339/files/figure.png" pageId="66" pageNumber="67" startId="66.[189,232,1787,1813]" targetBox="[194,1391,514,1732]" targetPageId="66">
<paragraph id="8BA4DA5DFFE7FFBBFF64F944AC17ED91" blockId="66.[185,1400,1787,2024]" pageId="66" pageNumber="67">
<emphasis id="B96F064FFFE7FFBBFF64F944AC3FED6C" bold="true" box="[189,280,1787,1813]" pageId="66" pageNumber="67">Fig. 24.</emphasis>
Distribution of
<taxonomicName id="4C1BA1DEFFE7FFBBFE13F943AEC0ED6C" authority="(Klotzsch) A. DC." authorityName="A. DC." baseAuthorityName="Klotzsch" box="[458,999,1787,1814]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="section" section="Cyathocnemis">
<emphasis id="B96F064FFFE7FFBBFE13F943AF0AED6C" box="[458,557,1788,1813]" italics="true" pageId="66" pageNumber="67">Begonia</emphasis>
sect.
<emphasis id="B96F064FFFE7FFBBFDB5F944AE33ED6C" box="[620,788,1787,1813]" italics="true" pageId="66" pageNumber="67">Cyathocnemis</emphasis>
(Klotzsch)A.DC.
</taxonomicName>
in Peru and surrounding countries.
<emphasis id="B96F064FFFE7FFBBFF62F8A1ADF7ED41" bold="true" box="[187,208,1822,1848]" pageId="66" pageNumber="67">A</emphasis>
.
<taxonomicName id="4C1BA1DEFFE7FFBBFF07F8A0AF2CED41" authority="Moonlight" authorityName="Moonlight" box="[222,523,1822,1848]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE7FFBBFF07F8A0ACADED41" box="[222,394,1822,1848]" italics="true" pageId="66" pageNumber="67">B. amoeboides</emphasis>
Moonlight
</taxonomicName>
(red) and
<taxonomicName id="4C1BA1DEFFE7FFBBFDA3F8A0AEE6ED41" authority="Moonlight" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[634,961,1822,1848]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="imbrexiformis" status="sp. nov.">
<emphasis id="B96F064FFFE7FFBBFDA3F8A0AE1AED41" box="[634,829,1822,1848]" italics="true" pageId="66" pageNumber="67">B. imbrexiformis</emphasis>
Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFE7FFBBFC1CF8A0A93AED40" box="[965,1053,1823,1849]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
(blue).
<emphasis id="B96F064FFFE7FFBBFBA8F8A1A9A2ED41" bold="true" box="[1137,1157,1822,1848]" pageId="66" pageNumber="67">B</emphasis>
.
<taxonomicName id="4C1BA1DEFFE7FFBBFB4AF8A0A850ED41" authority="A. DC." authorityName="A. DC." box="[1171,1399,1822,1848]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="bracteosa">
<emphasis id="B96F064FFFE7FFBBFB4AF8A0A800ED41" box="[1171,1319,1822,1848]" italics="true" pageId="66" pageNumber="67">B. bracteosa</emphasis>
A.DC.
</taxonomicName>
(red) and
<taxonomicName id="4C1BA1DEFFE7FFBBFEECF8FDAF70ED22" authority="Moonlight" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[309,599,1857,1883]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="longinqua">
<emphasis id="B96F064FFFE7FFBBFEECF8FDACF7ED22" box="[309,464,1857,1883]" italics="true" pageId="66" pageNumber="67">B. longinqua</emphasis>
Moonlight
</taxonomicName>
(blue).
<emphasis id="B96F064FFFE7FFBBFD60F8FEAFE9ED22" bold="true" box="[697,718,1857,1883]" pageId="66" pageNumber="67">C</emphasis>
.
<taxonomicName id="4C1BA1DEFFE7FFBBFD06F8FDA979ED22" authority="Poepp. &amp; Endl." authorityName="Poepp. &amp; Endl." box="[735,1118,1857,1884]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="cyathophora">
<emphasis id="B96F064FFFE7FFBBFD06F8FDAEBEED22" box="[735,921,1857,1883]" italics="true" pageId="66" pageNumber="67">B. cyathophora</emphasis>
Poepp. &amp; Endl.
</taxonomicName>
(red),
<taxonomicName id="4C1BA1DEFFE7FFBBFB6AF8FDAC1EED06" authority="Moonlight" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="serratistipula" status="sp. nov.">
<emphasis id="B96F064FFFE7FFBBFB6AF8FDA851ED22" box="[1203,1398,1857,1883]" italics="true" pageId="66" pageNumber="67">B. serratistipula</emphasis>
Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFE7FFBBFE98F8DAACBBED06" box="[321,412,1893,1919]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
(blue) and
<taxonomicName id="4C1BA1DEFFE7FFBBFDFCF8DAAE0BED06" authority="Moonlight" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[549,812,1893,1919]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="vargasii" status="sp. nov.">
<emphasis id="B96F064FFFE7FFBBFDFCF8DAAF8FED06" box="[549,680,1893,1919]" italics="true" pageId="66" pageNumber="67">C. vargasii</emphasis>
Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFE7FFBBFCEAF8DAAEA8ED06" box="[819,911,1893,1919]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
(green).
<emphasis id="B96F064FFFE7FFBBFC23F8DAA928ED06" bold="true" box="[1018,1039,1893,1919]" pageId="66" pageNumber="67">D</emphasis>
.
<taxonomicName id="4C1BA1DEFFE7FFBBFBC6F8D9A808ED06" authority="Moonlight" authorityName="Moonlight" box="[1055,1327,1893,1919]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="lamolina">
<emphasis id="B96F064FFFE7FFBBFBC6F8D9A98DED06" box="[1055,1194,1893,1919]" italics="true" pageId="66" pageNumber="67">B. lamolina</emphasis>
Moonlight
</taxonomicName>
(red),
<taxonomicName id="4C1BA1DEFFE7FFBBFF67F836AC89EDDB" authority="Rolfe" authorityName="Rolfe" box="[190,430,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="lophoptera">
<emphasis id="B96F064FFFE7FFBBFF67F836AC45EDDB" box="[190,354,1928,1954]" italics="true" pageId="66" pageNumber="67">B. lophoptera</emphasis>
Rolfe
</taxonomicName>
(blue) and
<taxonomicName id="4C1BA1DEFFE7FFBBFDE0F836AE71EDDB" authority="Irmsch." authorityName="Irmsch." box="[569,854,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B96F064FFFE7FFBBFDE0F836AFD4EDDB" box="[569,755,1928,1954]" italics="true" pageId="66" pageNumber="67">B. obtecticaulis</emphasis>
Irmsch.
</taxonomicName>
(green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation.
</paragraph>
</caption>
<caption id="DF648AD5FFE6FFBAFF64F837AF02ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178343" ID-Zenodo-Dep="8178343" httpUri="https://zenodo.org/record/8178343/files/figure.png" pageId="67" pageNumber="68" startId="67.[189,232,1928,1954]" targetBox="[270,1326,278,1859]" targetPageId="67">
<paragraph id="8BA4DA5DFFE6FFBAFF64F837AF02ED91" blockId="67.[189,1402,1928,2025]" pageId="67" pageNumber="68">
<emphasis id="B96F064FFFE6FFBAFF64F837AC06EDDB" bold="true" box="[189,289,1928,1954]" pageId="67" pageNumber="68">Fig. 25.</emphasis>
<taxonomicName id="4C1BA1DEFFE6FFBAFEF6F836AEF5EDDB" authority="Moonlight. A. Habit. B. Stamen" box="[303,978,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE6FFBAFEF6F836AF0AEDDB" box="[303,557,1928,1954]" italics="true" pageId="67" pageNumber="68">Begonia amoeboides</emphasis>
Moonlight.
<emphasis id="B96F064FFFE6FFBAFD14F837AFC5EDDB" bold="true" box="[717,738,1928,1954]" pageId="67" pageNumber="68">A</emphasis>
. Habit.
<emphasis id="B96F064FFFE6FFBAFC97F837AE45EDDB" bold="true" box="[846,866,1928,1954]" pageId="67" pageNumber="68">B</emphasis>
. Stamen
</taxonomicName>
, front view.
<emphasis id="B96F064FFFE6FFBAFBAEF837A9ABEDDB" bold="true" box="[1143,1164,1928,1954]" pageId="67" pageNumber="68">C</emphasis>
. Staminate flower, front view.
<emphasis id="B96F064FFFE6FFBAFE9EF814AC7CEDBC" bold="true" box="[327,347,1963,1989]" pageId="67" pageNumber="68">D</emphasis>
. Pistillate flower, side view.
<emphasis id="B96F064FFFE6FFBAFD6DF814AFEFEDBC" bold="true" box="[692,712,1963,1989]" pageId="67" pageNumber="68">E</emphasis>
. Pistils, side view. Illustration by Claire Banks from
<emphasis id="B96F064FFFE6FFBAFA9AF813ACD6ED91" italics="true" pageId="67" pageNumber="68">P.W. Moonlight &amp; A. Daza 150</emphasis>
(E).
</paragraph>
</caption>
<subSubSection id="C30189D6FFE1FFBDFF64FC19AE9AE996" pageId="68" pageNumber="69" type="conservation">
<paragraph id="8BA4DA5DFFE1FFBDFF64FC19AF4AE9B9" blockId="68.[189,621,933,960]" box="[189,621,933,960]" pageId="68" pageNumber="69">
<heading id="D0EC6D31FFE1FFBDFF64FC19AF4AE9B9" bold="true" box="[189,621,933,960]" fontSize="11" level="3" pageId="68" pageNumber="69" reason="3">
<emphasis id="B96F064FFFE1FFBDFF64FC19AF4AE9B9" bold="true" box="[189,621,933,960]" pageId="68" pageNumber="69">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFE1FFBDFF64FC6BAE9AE996" blockId="68.[189,957,980,1007]" box="[189,957,980,1007]" pageId="68" pageNumber="69">
Assessed by
<bibRefCitation id="EF8AA7ACFFE1FFBDFE8CFC6BAFB9E996" author="Moonlight &amp; Reynel" box="[341,670,980,1007]" firstAuthor="Moonlight" pageId="68" pageNumber="69" pagination="116 - 126" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" type="journal article" year="2018">Moonlight &amp; Reynel (2018)</bibRefCitation>
as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE1FFBDFF64FBA9AF36EF69" pageId="68" pageNumber="69" type="discussion">
<paragraph id="8BA4DA5DFFE1FFBDFF64FBA9AC97EE49" blockId="68.[189,432,1046,1072]" box="[189,432,1046,1072]" pageId="68" pageNumber="69">
<heading id="D0EC6D31FFE1FFBDFF64FBA9AC97EE49" bold="true" box="[189,432,1046,1072]" fontSize="11" level="3" pageId="68" pageNumber="69" reason="3">
<emphasis id="B96F064FFFE1FFBDFF64FBA9AC97EE49" bold="true" box="[189,432,1046,1072]" pageId="68" pageNumber="69">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFE1FFBDFF64FBFAAF36EF69" blockId="68.[188,1399,1093,1297]" pageId="68" pageNumber="69">
Within its range,
<taxonomicName id="4C1BA1DEFFE1FFBDFE57FBF9AF1AEE26" box="[398,573,1093,1119]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE1FFBDFE57FBF9AF1AEE26" box="[398,573,1093,1119]" italics="true" pageId="68" pageNumber="69">B. amoeboides</emphasis>
</taxonomicName>
can only be confused with
<taxonomicName id="4C1BA1DEFFE1FFBDFC53FBF9A90AEE26" box="[906,1069,1093,1119]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="lophoptera">
<emphasis id="B96F064FFFE1FFBDFC53FBF9A90AEE26" box="[906,1069,1093,1119]" italics="true" pageId="68" pageNumber="69">B. lophoptera</emphasis>
</taxonomicName>
. When in flower, these two species can easily be distinguished by the number of petals on the pistillate flower (five in
<taxonomicName id="4C1BA1DEFFE1FFBDFB10FBD6A851EEFB" box="[1225,1398,1128,1154]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE1FFBDFB10FBD6A851EEFB" box="[1225,1398,1128,1154]" italics="true" pageId="68" pageNumber="69">B. amoeboides</emphasis>
</taxonomicName>
vs two in
<taxonomicName id="4C1BA1DEFFE1FFBDFEEDFB32ACF0EEDF" box="[308,471,1164,1190]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="lophoptera">
<emphasis id="B96F064FFFE1FFBDFEEDFB32ACF0EEDF" box="[308,471,1164,1190]" italics="true" pageId="68" pageNumber="69">B. lophoptera</emphasis>
</taxonomicName>
) but they are difficult to distinguish when sterile.
<taxonomicName id="4C1BA1DEFFE1FFBDFBF0FB32A806EEDF" box="[1065,1313,1164,1190]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE1FFBDFBF0FB32A806EEDF" box="[1065,1313,1164,1190]" italics="true" pageId="68" pageNumber="69">Begonia amoeboides</emphasis>
</taxonomicName>
can be identified by its straight, reniform leaves, which lack a distinct apex (transverse, ovate, and with a distinct apex in
<taxonomicName id="4C1BA1DEFFE1FFBDFEAAFB6BAF32EE94" box="[371,533,1235,1261]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="lophoptera">
<emphasis id="B96F064FFFE1FFBDFEAAFB6BAF32EE94" box="[371,533,1235,1261]" italics="true" pageId="68" pageNumber="69">B. lophoptera</emphasis>
</taxonomicName>
) and its deciduous, lanceolate stipules (vs persistent and reniform to ovate or rarely broadly lanceolate).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFE1FFBDFF64FA87A839EC76" pageId="68" pageNumber="69" type="distribution">
<paragraph id="8BA4DA5DFFE1FFBDFF64FA87ACD2EF2B" blockId="68.[189,501,1336,1362]" box="[189,501,1336,1362]" pageId="68" pageNumber="69">
<heading id="D0EC6D31FFE1FFBDFF64FA87ACD2EF2B" bold="true" box="[189,501,1336,1362]" fontSize="11" level="3" pageId="68" pageNumber="69" reason="3">
<emphasis id="B96F064FFFE1FFBDFF64FA87ACD2EF2B" bold="true" box="[189,501,1336,1362]" pageId="68" pageNumber="69">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFE1FFBDFF64FAD8A839EC76" blockId="68.[188,1400,1383,1551]" pageId="68" pageNumber="69">
Endemic to
<collectingCountry id="F30C9ACDFFE1FFBDFE88FAD7ACA0EFFB" box="[337,391,1384,1410]" name="Peru" pageId="68" pageNumber="69">Peru</collectingCountry>
and known from Amazonas,
<collectingRegion id="49DF14BFFFE1FFBDFD28FAD8AE5CEFF8" box="[753,891,1383,1409]" country="Peru" name="San Martin" pageId="68" pageNumber="69">San Martín</collectingRegion>
, and
<collectingRegion id="49DF14BFFFE1FFBDFC66FAD7A924EFFB" box="[959,1027,1384,1410]" country="Peru" name="Pasco" pageId="68" pageNumber="69">Pasco</collectingRegion>
Regions (
<figureCitation id="1320C6D8FFE1FFBDFB5BFAD8A9D4EFF8" box="[1154,1267,1383,1409]" captionStart="Fig" captionStartId="66.[189,232,1787,1813]" captionTargetBox="[194,1391,514,1732]" captionTargetId="figure-13@66.[189,1398,504,1740]" captionTargetPageId="66" captionText="Fig. 24. Distribution ofBegonia sect.Cyathocnemis (Klotzsch)A.DC. in Peru and surrounding countries. A. B. amoeboides Moonlight (red) andB. imbrexiformis Moonlight sp. nov. (blue).B. B. bracteosaA.DC. (red) and B. longinqua Moonlight (blue). C. B. cyathophora Poepp. &amp; Endl. (red), B. serratistipula Moonlight sp. nov. (blue) and C. vargasii Moonlight sp. nov. (green). D. B. lamolina Moonlight (red), B. lophoptera Rolfe (blue) and B. obtecticaulis Irmsch. (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178339" httpUri="https://zenodo.org/record/8178339/files/figure.png" pageId="68" pageNumber="69">Fig. 24A</figureCitation>
). Found in upper and middle montane forest at elevations of
<quantity id="4CE377B8FFE1FFBDFCC3FA35AE98EFDC" box="[794,959,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" metricValueMax="2.93" metricValueMin="1.85" pageId="68" pageNumber="69" unit="m" value="2390.0" valueMax="2930.0" valueMin="1850.0">1850–2930 m</quantity>
a.s.l., where it is typically collected on wet banks. In
<collectingRegion id="49DF14BFFFE1FFBDFE51FA11AF43EFB1" box="[392,612,1454,1480]" country="Peru" name="Amazonas" pageId="68" pageNumber="69">Amazonas Region</collectingRegion>
,
<taxonomicName id="4C1BA1DEFFE1FFBDFDA9FA10AE83EFB1" authority="Moonlight" authorityName="Moonlight" box="[624,932,1454,1480]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="amoeboides">
<emphasis id="B96F064FFFE1FFBDFDA9FA10AE38EFB1" box="[624,799,1454,1480]" italics="true" pageId="68" pageNumber="69">B. amoeboides</emphasis>
Moonlight
</taxonomicName>
has been observed growing adjacent to
<taxonomicName id="4C1BA1DEFFE1FFBDFF67FA6DACEFEF92" authority="Moonlight" authorityName="Moonlight" box="[190,456,1489,1515]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="lamolina">
<emphasis id="B96F064FFFE1FFBDFF67FA6DAC60EF92" box="[190,327,1489,1515]" italics="true" pageId="68" pageNumber="69">B. lamolina</emphasis>
Moonlight
</taxonomicName>
and collections in
<collectingRegion id="49DF14BFFFE1FFBDFD7BFA6DAE65EF92" box="[674,834,1489,1516]" country="Peru" name="Pasco" pageId="68" pageNumber="69">Pasco Region</collectingRegion>
were made at the same locality as
<taxonomicName id="4C1BA1DEFFE1FFBDFB0AFA6DAC22EC76" authority="Rolfe." authorityName="Rolfe." class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="lophoptera">
<emphasis id="B96F064FFFE1FFBDFB0AFA6DA852EF92" box="[1235,1397,1489,1515]" italics="true" pageId="68" pageNumber="69">B. lophoptera</emphasis>
Rolfe.
</taxonomicName>
These three species are closely related, but no hybrids are known from either population.
</paragraph>
</subSubSection>
</treatment>
</document>
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