250 lines
37 KiB
XML
250 lines
37 KiB
XML
<document id="0E03711AA92004062A1DFD45B0244E30" ID-DOI="10.5281/zenodo.13319975" ID-ISSN="0253-116X" ID-Zenodo-Dep="13319975" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722341373250" checkinUser="felipe" docAuthor="Assing, V." docDate="2006" docId="03AD6616005EE108FEF8D9B465FEEF9D" docLanguage="en" docName="LinzerbiolBeitr.2006.38.2.1143-1156.pdf" docOrigin="Linzer biologische Beiträge 38 (2)" docStyle="DocumentStyle:6260912E6386EE56293601EB6FBA9D25.4:LinzerbiolBeitr.2006-.journal_article.type2" docStyleId="6260912E6386EE56293601EB6FBA9D25" docStyleName="LinzerbiolBeitr.2006-.journal_article.type2" docStyleVersion="4" docTitle="Atheta (Microdota) foliacea Assing, 2006, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1156" masterDocId="FF941E6E005DE10DFFACDA1E6146EB01" masterDocTitle="Six new species of micropterous Atheta and Apimela from northern Yunnan, China (Coleoptera: Staphylinidae, Aleocharinae)" masterLastPageNumber="1156" masterPageNumber="1143" pageNumber="1143" updateTime="1723635334303" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="813FC11F13192667D0AC043A16D193A2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BFD37A01BAB953F6484EA87B4872605B">
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<mods:title id="086D23CFAC1DDFD937A061352E6087AD">Six new species of micropterous Atheta and Apimela from northern Yunnan, China (Coleoptera: Staphylinidae, Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B6930711AF93C9A7B3EA210B44CDE7BC" type="personal">
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<mods:role id="13EA9EA14F23C58801E4E540825D3CA7">
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<mods:roleTerm id="2F7EEC48362491A97CEF7E36224F1311">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5F568660D7C2B4AAF4D53ABAE2C6CDCE">Assing, V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8205ED1B3C97D2EE173C1BD62FF7A8BB">text</mods:typeOfResource>
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<mods:relatedItem id="6F023636F2CEE2E527487B10444520FC" type="host">
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<mods:titleInfo id="45E066335F72AABE5B2999507C9B311E">
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<mods:title id="75D3DE671AD61C16B48A60D9FF8E5DF3">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="67FA1F6D953A54920E25C10D5D05A713">
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<mods:date id="B074BFD7FC21447E65EA6EEDFB830C4C">2006</mods:date>
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<mods:detail id="8B2CDF213BFC3ED42077817CFF4C7F8F" type="pubDate">
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<mods:number id="32DC8D4D051300531FE2558AD6FFC906">2006-12-29</mods:number>
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</mods:detail>
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<mods:detail id="51CD97A6EDDA0D810D06608DC05E1967" type="volume">
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<mods:number id="652BEB832814A1BA1FCE1E74675FC4BE">38</mods:number>
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</mods:detail>
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<mods:detail id="6D1A4824C17ECBB3084012BC38BFE633" type="issue">
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<mods:number id="ACD348ED13DBBE175736E91D20B67B89">2</mods:number>
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</mods:detail>
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<mods:extent id="8F6BE99FDB8ECD09EB0B1E98357ADD60" unit="page">
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<mods:start id="FF228652673B0BD171AE3FCF3CBC5772">1143</mods:start>
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<mods:end id="9E4501517605FFF506D3B5D212490A5B">1156</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="5A142920A55D6972524A4EE1B56F2E9C">journal article</mods:classification>
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<mods:identifier id="C133B13FC28FB1F34FA42CCC29EC46EA" type="DOI">10.5281/zenodo.13319975</mods:identifier>
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<mods:identifier id="60821085EF6E2F2915A986E46864CE07" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="2A8360154CD1069F895EB9B251BB72A0" type="Zenodo-Dep">13319975</mods:identifier>
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</mods:mods>
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<treatment id="03AD6616005EE108FEF8D9B465FEEF9D" LSID="urn:lsid:plazi:treatment:03AD6616005EE108FEF8D9B465FEEF9D" httpUri="http://treatment.plazi.org/id/03AD6616005EE108FEF8D9B465FEEF9D" lastPageId="5" pageId="3">
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<subSubSection id="C31E848B005EE10EFEF8D9B4620AE8C1" box="[340,844,938,960]" pageId="3" type="nomenclature">
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<paragraph id="8BBBD700005EE10EFEF8D9B4620AE8C1" blockId="3.[340,844,938,960]" box="[340,844,938,960]" pageId="3">
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<heading id="D0F3606C005EE10EFEF8D9B4620AE8C1" box="[340,844,938,960]" fontSize="8" level="2" pageId="3" reason="2">
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<emphasis id="B9700B12005EE10EFEF8D9B4638EE8C1" bold="true" box="[340,712,938,960]" pageId="3">
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<taxonomicName id="4C04AC83005EE10EFEF8D9B46330E8C1" ID-CoL="JD3V" authorityName="Assing" authorityYear="2006" box="[340,630,938,960]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="foliacea" status="sp. nov." subGenus="Microdota">
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<emphasis id="B9700B12005EE10EFEF8D9B460DFE8C1" bold="true" box="[340,409,938,960]" italics="true" pageId="3">Atheta</emphasis>
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(
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<emphasis id="B9700B12005EE10EFE04D9B46352E8C1" bold="true" box="[424,532,938,960]" italics="true" pageId="3">Microdota</emphasis>
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)
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<emphasis id="B9700B12005EE10EFD8FD9B46330E8C1" bold="true" box="[547,630,938,960]" italics="true" pageId="3">foliacea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A243B669005EE10EFDD0D9B4638EE8C1" box="[636,712,938,960]" pageId="3" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="133FCB85005EE10EFD7AD9B46202E8C1" box="[726,836,938,960]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Figs 14-26</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E848B005EE108FEF8D9CE65FEEF9D" lastPageId="5" pageId="3" type="materials_examined">
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<paragraph id="8BBBD700005EE10EFEF8D9CE658EEF28" blockId="3.[340,1248,975,1065]" pageId="3">
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<materialsCitation id="3B6CDD5D005EE10EFEF8D9CE6554EF13" collectingDate="2005-06-05" collectorName="M. Schulke" country="China" county="Zhongdian Co." elevation="3895" latitude="27.618334" location="Xue Shan" longLatPrecision="123" longitude="99.64167" municipality="Diqing" pageId="3" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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<typeStatus id="54BF69A2005EE10EFEF8D9CE60EFE8E5" box="[340,425,976,996]" pageId="3" type="holotype">Holotype</typeStatus>
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Ƌ:
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<collectingCountry id="F3139790005EE10EFE7FD9CE634BE8E5" box="[467,525,976,996]" name="China" pageId="3">China</collectingCountry>
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: N-Yunnan [C2005-05],
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<collectingMunicipality id="6BDF4D7A005EE10EFD55D9CE627EE8E5" box="[761,824,976,996]" pageId="3">Diqing</collectingMunicipality>
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<collectingRegion id="49C019E2005EE10EFCEDD9CE6232E8E5" box="[833,884,976,996]" country="China" name="Xizang" pageId="3">Tibet</collectingRegion>
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. Aut. Pref.,
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<collectingCounty id="62DAAF8C005EE10EFC46D9CE653EE8E5" box="[1002,1144,976,996]" pageId="3">Zhongdian Co.</collectingCounty>
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,
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<location id="8EDB81DB005EE10EFB2FD9CE6599E8E5" LSID="urn:lsid:plazi:treatment:03AD6616005EE108FEF8D9B465FEEF9D:8EDB81DB005EE10EFB2FD9CE6599E8E5" box="[1155,1247,976,996]" country="China" county="Zhongdian Co." latitude="27.618334" longLatPrecision="123" longitude="99.64167" municipality="Diqing" name="Xue Shan" pageId="3" stateProvince="Xizang">Xue Shan</location>
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near lake
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<locationDeviation id="9A09CE22005EE10EFE01D9F96323E8FA" box="[429,613,999,1019]" location="Zhongdian" pageId="3">
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<quantity id="4CFC7AE5005EE10EFE01D9F960A1E8FA" box="[429,487,999,1019]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="3" unit="km" value="23.0">23 km</quantity>
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S Zhongdian
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</locationDeviation>
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,
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<quantity id="4CFC7AE5005EE10EFDC2D9F963F2E8FA" box="[622,692,999,1019]" metricMagnitude="3" metricUnit="m" metricValue="3.895" pageId="3" unit="m" value="3895.0">
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<elevation id="00293033005EE10EFDC2D9F963F2E8FA" box="[622,692,999,1019]" metricMagnitude="3" metricUnit="m" metricValue="3.895" pageId="3" unit="m" value="3895.0">3895 m</elevation>
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</quantity>
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/
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<geoCoordinate id="EE30B1C7005EE10EFD6AD9F96262E8FA" box="[710,804,999,1019]" degrees="27" direction="north" minutes="37.1" orientation="latitude" pageId="3" precision="92" value="27.618334">27°37.1'N</geoCoordinate>
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,
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<geoCoordinate id="EE30B1C7005EE10EFC82D9F962CFE8FA" box="[814,905,999,1019]" degrees="99" direction="east" minutes="38.5" orientation="longitude" pageId="3" precision="92" value="99.64167">99°38.5'E</geoCoordinate>
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, devast. mixed forest, meadows, lake border, leaf litter, dead wood, sifted,
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<date id="FFBAF1C0005EE10EFD05D9E0624EEF13" box="[681,776,1022,1042]" pageId="3" value="2005-06-05">
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<collectingDate id="EFFE0828005EE10EFD05D9E0624EEF13" box="[681,776,1022,1042]" pageId="3" value="2005-06-05">5.VI.2005</collectingDate>
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</date>
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, leg.
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<collectorName id="26F1B2D6005EE10EFC95D9E062E5EF13" box="[825,931,1022,1042]" pageId="3">M. Schülke</collectorName>
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[C2005-05]
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</materialsCitation>
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/
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<typeStatus id="54BF69A2005EE10EFB8AD9E065C3EF13" box="[1062,1157,1022,1042]" pageId="3" type="holotype">Holotypus</typeStatus>
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Ƌ
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<taxonomicName id="4C04AC83005EE10EFB08D9E060DBEF28" authorityName="Assing" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="foliacea" status="sp. nov.">
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<emphasis id="B9700B12005EE10EFB08D9E060DBEF28" italics="true" pageId="3">Atheta foliacea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A243B669005EE10EFE0FDE0B6092EF28" box="[419,468,1045,1065]" pageId="3" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<collectionCode id="ED154FC5005EE10EFDADDE0B6352EF28" box="[513,532,1045,1065]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
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. Assing 2006 (cAss).
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<typeStatus id="54BF69A2005EE10EFD77DE0B6271EF28" box="[731,823,1045,1065]" pageId="3" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D021C89005EE10EFC93DE0B623EEF28" box="[831,888,1045,1065]" count="5" pageId="3" type="generic">5 exs.</specimenCount>
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: same data as
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<typeStatus id="54BF69A2005EE10EFC55DE0B650FEF28" box="[1017,1097,1045,1065]" pageId="3" type="holotype">holotype</typeStatus>
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(cSch, cAss).
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</paragraph>
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<paragraph id="8BBBD700005EE10EFEF8DE2663A2EFA4" blockId="3.[340,1248,1079,1189]" pageId="3">
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D e s c r i p t i o n:
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<quantity id="4CFC7AE5005EE10EFD9DDE2963F6EF4C" box="[561,688,1079,1101]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="3" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4-2.9 mm</quantity>
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. Habitus as in
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<figureCitation id="133FCB85005EE10EFCF3DE2962F6EF4C" box="[863,944,1079,1101]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 14</figureCitation>
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. Coloration: head brown to blackish brown; pronotum and elytra brown to dark brown; abdomen blackish brown, with the apex slightly paler; legs light brown; antennae brown, with the basal 3-4 antennomeres yellowish to yellowish brown.
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</paragraph>
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<paragraph id="8BBBD700005EE10EFEF8DEAA6596EE7B" blockId="3.[340,1248,1204,1402]" pageId="3">
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Head, with sexual dimorphism, slightly wider than long, of suborbicular shape (
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<figureCitation id="133FCB85005EE10EFB2ADEAA6595EFCB" box="[1158,1235,1204,1226]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 15</figureCitation>
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); eyes very small, composed of only approximately 10 ommatidia (
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<figureCitation id="133FCB85005EE10EFC43DECF657CEFE6" box="[1007,1082,1233,1255]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 16</figureCitation>
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), not protruding from lateral outline of head, less than 1/4 the length of postocular region in dorsal view; integument with distinct microreticulation and almost matt; puncturation sparse, fine, and very shallow, barely noticeable in the microsculpture. Antenna distinctly incrassate apically (
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<figureCitation id="133FCB85005EE10EFE18DF586347EE5D" box="[436,513,1350,1372]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 17</figureCitation>
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); antennomere III distinctly shorter than II; IV weakly transverse; V-X of increasing width and increasingly transverse; X approximately twice as wide as long.
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</paragraph>
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<paragraph id="8BBBD700005EE10EFEF8DF9463A1EEF6" blockId="3.[340,1248,1417,1527]" pageId="3">
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Pronotum, with sexual dimorphism, 1.15-1.20 times as wide as long and approximately 1.15 times as wide as head (
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<figureCitation id="133FCB85005EE10EFD27DFB8639DEEBD" box="[651,731,1446,1468]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 15</figureCitation>
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); maximal width in anterior half; pubescence of midline directed cephalad in anterior half and caudad in posterior half; puncturation and microsculpture similar to those of head.
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</paragraph>
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<paragraph id="8BBBD700005EE10EFEF8DC186509ED56" blockId="3.[340,1248,1542,1623]" pageId="3">
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Elytra with sexual dimorphism, 1.05-1.10 times as wide and at suture less than 0.6 times as long as pronotum (
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<figureCitation id="133FCB85005EE10EFD92DC3D63CBED38" box="[574,653,1571,1593]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 15</figureCitation>
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); microreticulation pronounced, puncturation denser and slightly more distinct than that of head and pronotum. Hind wings reduced.
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</paragraph>
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<paragraph id="8BBBD700005EE10EFEF8DC7860DEEDD5" blockId="3.[340,1248,1638,1748]" pageId="3">
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Abdomen slightly (1.10-1.15 x) wider than elytra, maximal width at segments
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<collectionCode id="ED154FC5005EE10EFBD8DC7865C0ED7D" box="[1140,1158,1638,1660]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
|
|
/
|
|
<collectionCode id="ED154FC5005EE10EFB22DC7865EEED7D" box="[1166,1192,1638,1660]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3">VI</collectionCode>
|
|
(
|
|
<figureCitation id="133FCB85005EE10EFB1ADC786036ED98" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 14</figureCitation>
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|
); puncturation fine, much sparser on posterior than on anterior tergites; microreticulation distinct; posterior margin of tergite VII with very narrow rudiment of a palisade fringe.
|
|
</paragraph>
|
|
<paragraph id="8BBBD700005EE10EFEF8DCFC65A6EC17" blockId="3.[340,1248,1762,1814]" pageId="3">
|
|
Ƌ: head and pronotum with extensive median impressions (
|
|
<figureCitation id="133FCB85005EE10EFC19DCFD6547EDF8" box="[949,1025,1763,1785]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 15</figureCitation>
|
|
); elytra on either side of suture somewhat elevated (
|
|
<figureCitation id="133FCB85005EE10EFD26DD1E6391EC17" box="[650,727,1792,1814]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="3">Fig. 15</figureCitation>
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); posterior margin of tergite VIII broadly concave
|
|
</paragraph>
|
|
<paragraph id="8BBBD7000059E109FCAEDB7B6274EA7B" blockId="4.[770,818,357,378]" box="[770,818,357,378]" pageId="4">1147</paragraph>
|
|
<paragraph id="8BBBD7000059E109FEF8DBB160C8E91C" blockId="4.[340,1248,431,541]" pageId="4">
|
|
in the middle (
|
|
<figureCitation id="133FCB850059E109FE41DBB1637CEAC4" box="[493,570,431,453]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="4">Fig. 18</figureCitation>
|
|
); sternite VIII much longer than tergite VIII, its posterior margin strongly convex (
|
|
<figureCitation id="133FCB850059E109FDABDBD36312EAE2" box="[519,596,461,483]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="4">Fig. 19</figureCitation>
|
|
); median lobe of aedeagus with thin, blade-like ventral process (
|
|
<figureCitation id="133FCB850059E109FEF7DBF46097E901" box="[347,465,490,512]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="4">Figs 22-23</figureCitation>
|
|
); apical lobe of paramere shaped as in
|
|
<figureCitation id="133FCB850059E109FCD8DBF46284E901" box="[884,962,490,512]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="4">Fig. 24</figureCitation>
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, with three relatively short setae.
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|
</paragraph>
|
|
<caption id="DF7B87880059E109FEF8DC6D655AEDCA" ID-DOI="http://doi.org/10.5281/zenodo.13319979" ID-Zenodo-Dep="13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="4" startId="4.[340,380,1651,1671]" targetBox="[344,1253,579,1626]" targetPageId="4" targetType="figure">
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|
<paragraph id="8BBBD7000059E109FEF8DC6D655AEDCA" blockId="4.[340,1248,1650,1739]" pageId="4">
|
|
<emphasis id="B9700B120059E109FEF8DC6D60FAED86" bold="true" box="[340,444,1651,1671]" pageId="4">Figs 14-26</emphasis>
|
|
:
|
|
<taxonomicName id="4C04AC830059E109FE60DC6D631CED86" authorityName="Assing" authorityYear="2006" box="[460,602,1651,1671]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="foliacea">
|
|
<emphasis id="B9700B120059E109FE60DC6D631CED86" box="[460,602,1651,1671]" italics="true" pageId="4">Atheta foliacea</emphasis>
|
|
</taxonomicName>
|
|
: (
|
|
<emphasis id="B9700B120059E109FDDEDC6D63CFED86" bold="true" box="[626,649,1651,1671]" pageId="4">14</emphasis>
|
|
) Ƌ habitus; (
|
|
<emphasis id="B9700B120059E109FCBDDC6D626EED86" bold="true" box="[785,808,1651,1671]" pageId="4">15</emphasis>
|
|
) Ƌ forebody; (
|
|
<emphasis id="B9700B120059E109FC6DDC6D629EED86" bold="true" box="[961,984,1651,1671]" pageId="4">16</emphasis>
|
|
) head in lateral view; (
|
|
<emphasis id="B9700B120059E109FB6DDC6D659EED86" bold="true" box="[1217,1240,1651,1671]" pageId="4">17</emphasis>
|
|
) antenna; (
|
|
<emphasis id="B9700B120059E109FE03DC946080ED9F" bold="true" box="[431,454,1674,1694]" pageId="4">18</emphasis>
|
|
) Ƌ tergite VIII; (
|
|
<emphasis id="B9700B120059E109FDCBDC946338ED9F" bold="true" box="[615,638,1674,1694]" pageId="4">19</emphasis>
|
|
) Ƌ sternite VIII; (
|
|
<emphasis id="B9700B120059E109FC85DC946206ED9F" bold="true" box="[809,832,1674,1694]" pageId="4">20</emphasis>
|
|
) ♀ tergite VIII; (
|
|
<emphasis id="B9700B120059E109FC4DDC9462BEED9F" bold="true" box="[993,1016,1674,1694]" pageId="4">21</emphasis>
|
|
) ♀ sternite VIII; (
|
|
<emphasis id="B9700B120059E109FB0FDC94659EED9F" bold="true" box="[1187,1240,1674,1694]" pageId="4">22-23</emphasis>
|
|
) median lobe of aedeagus in lateral and in ventral view; (
|
|
<emphasis id="B9700B120059E109FCF1DCBF6232EDB4" bold="true" box="[861,884,1697,1717]" pageId="4">24</emphasis>
|
|
) apical lobe of paramere; (
|
|
<emphasis id="B9700B120059E109FBC3DCBF65E2EDB4" bold="true" box="[1135,1188,1697,1717]" pageId="4">25-26</emphasis>
|
|
) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BBBD7000058E108FCAEDB7B6274EA7B" blockId="5.[770,818,357,378]" box="[770,818,357,378]" pageId="5">1148</paragraph>
|
|
<paragraph id="8BBBD7000058E108FEF8DBAD650EE91C" blockId="5.[340,1248,431,541]" pageId="5">
|
|
<specimenCount id="9D021C890058E108FEF8DBAD602FEACB" box="[340,361,435,458]" count="1" pageId="5" type="female">♀</specimenCount>
|
|
: head and pronotum without or with very indistinct impressions; elytra not distinctly elevated on either side of suture; tergite VIII similar to that of Ƌ (
|
|
<figureCitation id="133FCB850058E108FBACDBD36508EAE2" box="[1024,1102,461,483]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="5">Fig. 20</figureCitation>
|
|
); sternite VIII longer than tergite VIII, its posterior margin in the middle distinctly concave and with distinctly modified marginal setae (
|
|
<figureCitation id="133FCB850058E108FD17D8196240E91C" box="[699,774,519,541]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="5">Fig. 21</figureCitation>
|
|
); spermatheca as in
|
|
<figureCitation id="133FCB850058E108FC78D819650EE91C" box="[980,1096,519,541]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="5">Figs 25-26.</figureCitation>
|
|
</paragraph>
|
|
<paragraph id="8BBBD7000058E108FEF8D833630DE961" blockId="5.[340,1247,557,608]" pageId="5">E t y m o l o g y: The name (Lat., adj.: shaped like a leaf) refers to the thin ventral process of the aedeagus.</paragraph>
|
|
<paragraph id="8BBBD7000058E108FEF8D87163B0EF3D" blockId="5.[340,1248,623,1084]" pageId="5">
|
|
C o m p a r a t i v e n o t e s: Based on the modifications of the male head and pronotum, as well as on the morphology of the aedeagus,
|
|
<taxonomicName id="4C04AC830058E108FC64D893657CE9A3" authorityName="Assing" authorityYear="2006" box="[968,1082,652,674]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="foliacea">
|
|
<emphasis id="B9700B120058E108FC64D893657CE9A3" box="[968,1082,652,674]" italics="true" pageId="5">A. foliacea</emphasis>
|
|
</taxonomicName>
|
|
is very closely related to
|
|
<taxonomicName id="4C04AC830058E108FE6FD8B46352E9C1" box="[451,532,682,704]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="elisa">
|
|
<emphasis id="B9700B120058E108FE6FD8B46352E9C1" box="[451,532,682,704]" italics="true" pageId="5">A. elisa</emphasis>
|
|
</taxonomicName>
|
|
from the Daba Shan in
|
|
<collectingRegion id="49C019E20058E108FC8ED8B46231E9C1" box="[802,887,682,704]" country="China" name="Shaanxi" pageId="5">Shaanxi</collectingRegion>
|
|
. From this species,
|
|
<taxonomicName id="4C04AC830058E108FBFDD8B46582E9C1" authorityName="Assing" authorityYear="2006" box="[1105,1220,682,704]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="foliacea">
|
|
<emphasis id="B9700B120058E108FBFDD8B46582E9C1" box="[1105,1220,682,704]" italics="true" pageId="5">A. foliacea</emphasis>
|
|
</taxonomicName>
|
|
is distinguished especially by the darker coloration of the forebody, the less pronounced impressions on the male head and pronotum, the modified male elytra, the different shapes of tergite and sternite VIII in both sexes, the more evenly curved (lateral view), broader and apically less acute (ventral view) ventral process of the aedeagus, as well as by the slightly differently shaped spermatheca. For illustrations of the sexual characters of
|
|
<taxonomicName id="4C04AC830058E108FEDAD944608EE86E" box="[374,456,857,879]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="elisa">
|
|
<emphasis id="B9700B120058E108FEDAD944608EE86E" box="[374,456,857,879]" italics="true" pageId="5">A. elisa</emphasis>
|
|
</taxonomicName>
|
|
see
|
|
<bibRefCitation id="EF95AAF10058E108FDAED94763E7E86E" author="ASSING V" box="[514,673,857,879]" pageId="5" pagination="953 - 969" refId="ref6277" refString="ASSING V. (2002): On some micropterous species of Athetini from Nepal and China (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biol. Beitr. 34: 953 - 969." type="journal article" year="2002">ASSING (2002)</bibRefCitation>
|
|
. From the syntopic
|
|
<taxonomicName id="4C04AC830058E108FC2FD94462A1E86E" authorityName="Assing" authorityYear="2006" box="[899,999,858,879]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="xueica">
|
|
<emphasis id="B9700B120058E108FC2FD94462A1E86E" box="[899,999,858,879]" italics="true" pageId="5">A. xueica</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C04AC830058E108FC56D9446529E86E" authorityName="Assing" authorityYear="2006" box="[1018,1135,857,879]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="foliacea">
|
|
<emphasis id="B9700B120058E108FC56D9446529E86E" box="[1018,1135,857,879]" italics="true" pageId="5">A. foliacea</emphasis>
|
|
</taxonomicName>
|
|
is readily separated by much paler coloration of the forebody and the appendages, by the more transverse antennomeres IV-X, the much smaller eyes, the subcircular head shape, the sexual dimorphism of head, pronotum, and elytra, the smaller and less transverse pronotum, the distinctly shorter elytra, the narrow rudiment of a palisade fringe at the posterior margin of tergite VII, the different shapes of tergite and sternite VIII in both sexes, as well as by the completely different morphology of the primary sexual characters; for comparison see
|
|
<figureCitation id="133FCB850058E108FD23DE3863B0EF3D" box="[655,758,1062,1084]" captionStart-0="Figs 1-13" captionStart-1="Figs 14-26" captionStartId-0="2.[340,380,1652,1672]" captionStartId-1="4.[340,380,1651,1671]" captionTargetBox-0="[340,1250,608,1621]" captionTargetBox-1="[344,1253,579,1626]" captionTargetId-0="figure-63@2.[340,1253,674,1153]" captionTargetId-1="figure-48@4.[340,1253,674,1153]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="Figs 1-13: Atheta xueica: (1) habitus; (2) forebody; (3) head in lateral view; (4) antenna; (5) Ƌ tergite VIII; (6) Ƌ sternite VIII; (7) posterior margin of ♀ tergite VIII; (8) ♀ sternite VIII; (9-10) median lobe of aedeagus in lateral and in ventral view; (11) apex of median lobe of aedeagus in ventral view; (12) apical lobe of paramere; (13) spermatheca. Scale bars: 1: 1.0 mm; 2-3: 0.5 mm; 4-8: 0.2 mm; 9-13: 0.1 mm." captionText-1="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13319977" figureDoi-1="http://doi.org/10.5281/zenodo.13319979" httpUri-0="https://zenodo.org/record/13319977/files/figure.png" httpUri-1="https://zenodo.org/record/13319979/files/figure.png" pageId="5">Figs 1-14.</figureCitation>
|
|
</paragraph>
|
|
<paragraph id="8BBBD7000058E108FEF8DE5265FEEF9D" blockId="5.[340,1247,1100,1180]" pageId="5">
|
|
D i s t r i b u t i o n a n d b i o n o m i c s: Thetypelocality is situated in the Xue Shan in northern
|
|
<collectingRegion id="49C019E20058E108FDA4DE74631DEF7E" box="[520,603,1130,1151]" country="China" name="Yunnan" pageId="5">Yunnan</collectingRegion>
|
|
; for an illustration see ASSING (2006:
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<figureCitation id="133FCB850058E108FC4BDE776572EF7E" box="[999,1076,1129,1151]" captionStart="Figs 14-26" captionStartId="4.[340,380,1651,1671]" captionTargetBox="[344,1253,579,1626]" captionTargetId="figure-48@4.[340,1253,674,1153]" captionTargetPageId="4" captionText="Figs 14-26: Atheta foliacea: (14) Ƌ habitus; (15) Ƌ forebody; (16) head in lateral view; (17) antenna; (18) Ƌ tergite VIII; (19) Ƌ sternite VIII; (20) ♀ tergite VIII; (21) ♀ sternite VIII; (22-23) median lobe of aedeagus in lateral and in ventral view; (24) apical lobe of paramere; (25-26) spermatheca. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-21: 0.2 mm; 22-26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13319979" httpUri="https://zenodo.org/record/13319979/files/figure.png" pageId="5">Fig. 20</figureCitation>
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). The specimens were sifted from the forest floor in a mixed forest at an altitude of approximately
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<quantity id="4CFC7AE50058E108FBC9DE9865F2EF9D" box="[1125,1204,1158,1180]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="5" unit="m" value="3900.0">3900 m</quantity>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |