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<document id="99EE305021E7886890E1077349025617" ID-DOI="10.11646/zootaxa.5584.4.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="14853712" ID-ZooBank="5E57B4DE-C868-4D1E-A6D7-37F612A91E71" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1739312307337" checkinUser="plazi" docAuthor="Feng, Zi-Yan, Chen, Bin &amp; Li, Ting-Jing" docDate="2025" docId="03ABD609A02EFFDEFF5B9709B384F9FC" docLanguage="en" docName="zootaxa.5584.4.10.pdf" docOrigin="Zootaxa 5584 (4)" docSource="https://doi.org/10.11646/zootaxa.5584.4.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyanotiphia breviclypeola Feng, Chen &amp; Li 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="595" masterDocId="FF92AE71A02FFFDDFFCC9042B017FFF5" masterDocTitle="A taxonomic review of the newly recorded genus Cyanotiphia Cameron, 1907 (Hymenoptera, Tiphiidae, Tiphiinae) from China, with a key to all known species" masterLastPageNumber="598" masterPageNumber="592" pageNumber="593" updateTime="1739365256552" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="19922E172FB43BD6F558230936F585F1">A taxonomic review of the newly recorded genus Cyanotiphia Cameron, 1907 (Hymenoptera, Tiphiidae, Tiphiinae) from China, with a key to all known species</mods:title>
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<mods:namePart id="D5D36EF703EB1F77F85C683052A0EF7B">Feng, Zi-Yan</mods:namePart>
<mods:nameIdentifier id="2563C33407F99D65F2697DA49C5E064F" type="ORCID">0009-0006-3418-1999</mods:nameIdentifier>
<mods:affiliation id="F8264843417047C70727A91D5E26FA98">Chongqing Key Laboratory of Vector Control and Utilization; Institute of Entomology and Molecular Biology, Chongqing Normal University, Chongqing 401331, China</mods:affiliation>
<mods:nameIdentifier id="96B0053011FCF5264F7B818B0BEDCB82" type="email">2023210513054@stu.cqnu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="50C30A2B9E6CD1C9B5D61813311C5ED2">Chen, Bin</mods:namePart>
<mods:nameIdentifier id="791EDA0453C7554790526482294CE6AD" type="ORCID">0000-0002-5227-7736</mods:nameIdentifier>
<mods:affiliation id="A94FCF0BC2AD2EC32216DC7BC4BC4059">Chongqing Key Laboratory of Vector Control and Utilization; Institute of Entomology and Molecular Biology, Chongqing Normal University, Chongqing 401331, China</mods:affiliation>
<mods:nameIdentifier id="3A24C78CEA5676CACB167B124C583DC0" type="email">c_bin@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="8119A0BB22B5BB0BA40D28B6767EB46B">Li, Ting-Jing</mods:namePart>
<mods:nameIdentifier id="C0EC8C6051AAFF89A0F12BC4E069C9DF" type="ORCID">0000-0001-7175-2697</mods:nameIdentifier>
<mods:affiliation id="ED6D536F2ADB3687295F81421EAD130E">Chongqing Key Laboratory of Vector Control and Utilization; Institute of Entomology and Molecular Biology, Chongqing Normal University, Chongqing 401331, China</mods:affiliation>
<mods:nameIdentifier id="1101B628D4FB6B44546B5EA8D96EFE79" type="email">ltjing1979@cqnu.edu.cn</mods:nameIdentifier>
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<mods:title id="5646128AB60407B4EE4ACE4DC96E9E48">Zootaxa</mods:title>
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<mods:date id="739371CE22423AE062F1C7144D260A5E">2025</mods:date>
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<mods:number id="82C64D0EED102529619BFDAC16CDB5AA">2025-02-11</mods:number>
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<treatment id="03ABD609A02EFFDEFF5B9709B384F9FC" ID-DOI="http://doi.org/10.5281/zenodo.14853720" ID-Zenodo-Dep="14853720" LSID="urn:lsid:plazi:treatment:03ABD609A02EFFDEFF5B9709B384F9FC" httpUri="http://treatment.plazi.org/id/03ABD609A02EFFDEFF5B9709B384F9FC" lastPageId="3" lastPageNumber="595" pageId="1" pageNumber="593">
<subSubSection id="C3183494A02EFFDCFF5B9709B30FF893" box="[151,792,1867,1894]" pageId="1" pageNumber="593" type="nomenclature">
<paragraph id="8BBD671FA02EFFDCFF5B9709B30FF893" blockId="1.[151,792,1867,1930]" box="[151,792,1867,1894]" pageId="1" pageNumber="593">
<heading id="D0F5D073A02EFFDCFF5B9709B30FF893" bold="true" box="[151,792,1867,1894]" fontSize="11" level="1" pageId="1" pageNumber="593" reason="1">
<emphasis id="B976BB0DA02EFFDCFF5B9709B30FF893" bold="true" box="[151,792,1867,1894]" pageId="1" pageNumber="593">
<taxonomicName id="4C021C9CA02EFFDCFF5B9709B2BAF893" authority="Feng, Chen &amp; Li" authorityName="Feng, Chen &amp; Li" authorityYear="2025" box="[151,685,1867,1894]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="593" phylum="Arthropoda" rank="species" species="breviclypeola" status="sp. nov.">
<emphasis id="B976BB0DA02EFFDCFF5B9709B1C4F893" bold="true" box="[151,467,1867,1894]" italics="true" pageId="1" pageNumber="593">Cyanotiphia breviclypeola</emphasis>
Feng, Chen &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="A2450676A02EFFDCFD77970EB30FF893" box="[699,792,1868,1894]" pageId="1" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3183494A02EFFDCFF5B9732B11CF87F" box="[151,267,1904,1930]" pageId="1" pageNumber="593" type="description">
<paragraph id="8BBD671FA02EFFDCFF5B9732B11CF87F" blockId="1.[151,792,1867,1930]" box="[151,267,1904,1930]" pageId="1" pageNumber="593">
(
<figureCitation id="13397B9AA02EFFDCFF539732B115F87F" box="[159,258,1904,1930]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="1" pageNumber="593">Figs 1–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3183494A02EFFDCFF5B97FAB303F803" pageId="1" pageNumber="593" type="materials_examined">
<paragraph id="8BBD671FA02EFFDCFF5B97FAB303F803" blockId="1.[151,1436,1976,2039]" pageId="1" pageNumber="593">
<emphasis id="B976BB0DA02EFFDCFF5B97FAB12CF827" bold="true" box="[151,315,1976,2002]" pageId="1" pageNumber="593">Type material</emphasis>
.
<materialsCitation id="3B6A6D42A02EFFDCFE8B97FAB303F803" collectingDate="2019-08-18" collectionCode="CNU" collectorName="Xun-Dong Li" country="China" county="Wuxi County" elevation="731" latitude="31.52389" location="Hongchiba Town" longLatPrecision="20" longitude="108.95111" municipality="Chongqing City" pageId="1" pageNumber="593" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="holotype">
<typeStatus id="54B9D9BDA02EFFDCFE8B97FAB1A5F827" box="[327,434,1976,2002]" pageId="1" pageNumber="593" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D04AC96A02EFFDCFE7797FBB1C5F826" box="[443,466,1977,2003]" count="1" pageId="1" pageNumber="593" type="male">♂</specimenCount>
,
<collectingCountry id="F315278FA02EFFDCFE1797FAB236F827" box="[475,545,1976,2002]" name="China" pageId="1" pageNumber="593">China</collectingCountry>
,
<collectingMunicipality id="6BD9FD65A02EFFDCFDE597FAB2CCF827" box="[553,731,1976,2002]" pageId="1" pageNumber="593">Chongqing City</collectingMunicipality>
,
<collectingCounty id="62DC1F93A02EFFDCFD2897FAB36EF827" box="[740,889,1976,2002]" pageId="1" pageNumber="593">Wuxi County</collectingCounty>
,
<location id="8EDD31C4A02EFFDCFC4E97FAB454F826" LSID="urn:lsid:plazi:treatment:03ABD609A02EFFDEFF5B9709B384F9FC:8EDD31C4A02EFFDCFC4E97FAB454F826" box="[898,1091,1976,2003]" country="China" county="Wuxi County" latitude="31.52389" longLatPrecision="20" longitude="108.95111" municipality="Chongqing City" name="Hongchiba Town" pageId="1" pageNumber="593" stateProvince="Chongqing">Hongchiba Town</location>
,
<location id="8EDD31C4A02EFFDCFB8797FAB513F827" LSID="urn:lsid:plazi:treatment:03ABD609A02EFFDEFF5B9709B384F9FC:8EDD31C4A02EFFDCFB8797FAB513F827" box="[1099,1284,1976,2002]" country="China" county="Wuxi County" latitude="31.52389" longLatPrecision="20" longitude="108.95111" municipality="Chongqing City" name="Chashan Village" pageId="1" pageNumber="593" stateProvince="Chongqing">Chashan Village</location>
,
<geoCoordinate id="EE3601D8A02EFFDCFAC097FAB58FF827" box="[1292,1432,1976,2002]" degrees="31" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="593" precision="15" seconds="26" value="31.52389">31°31’26”N</geoCoordinate>
,
<geoCoordinate id="EE3601D8A02EFFDCFF5B979EB137F802" box="[151,288,2012,2039]" degrees="108" direction="east" minutes="57" orientation="longitude" pageId="1" pageNumber="593" precision="15" seconds="4" value="108.95111">108°57’4”E</geoCoordinate>
,
<quantity id="4CFACAFAA02EFFDCFEE7979EB163F802" box="[299,372,2012,2039]" metricMagnitude="2" metricUnit="m" metricValue="7.31" pageId="1" pageNumber="593" unit="m" value="731.0">
<elevation id="002F802CA02EFFDCFEE7979EB163F802" box="[299,372,2012,2039]" metricMagnitude="2" metricUnit="m" metricValue="7.31" pageId="1" pageNumber="593" unit="m" value="731.0">731 m</elevation>
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,
<date id="FFBC41DFA02EFFDCFE4C979EB202F803" box="[384,533,2012,2038]" pageId="1" pageNumber="593" value="2019-08-18">
<collectingDate id="EFF8B837A02EFFDCFE4C979EB202F803" box="[384,533,2012,2038]" pageId="1" pageNumber="593" value="2019-08-18">18.VIII.2019</collectingDate>
</date>
,
<collectorName id="26F702C9A02EFFDCFDEC979FB2AFF803" box="[544,696,2012,2039]" pageId="1" pageNumber="593">Xun-Dong Li</collectorName>
(
<collectionCode id="ED13FFDAA02EFFDCFD0B979EB31FF803" box="[711,776,2012,2038]" pageId="1" pageNumber="593">CNU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF7D3797A02DFFDFFF5B9672B28FF965" ID-DOI="http://doi.org/10.5281/zenodo.14853716" ID-Zenodo-Dep="14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594" startId="2.[151,264,1584,1608]" targetBox="[151,1436,184,1560]" targetPageId="2" targetType="figure">
<paragraph id="8BBD671FA02DFFDFFF5B9672B28FF965" blockId="2.[151,1437,1584,1680]" pageId="2" pageNumber="594">
<emphasis id="B976BB0DA02DFFDFFF5B9672B123F9BD" bold="true" box="[151,308,1584,1608]" pageId="2" pageNumber="594">FIGURES 1–8</emphasis>
.
<taxonomicName id="4C021C9CA02DFFDFFE8C9672B25BF9BD" authorityName="Feng &amp; Chen &amp; Li" authorityYear="2025" box="[320,588,1584,1608]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="594" phylum="Arthropoda" rank="species" species="breviclypeola" status="sp. nov.">
<emphasis id="B976BB0DA02DFFDFFE8C9672B25BF9BD" box="[320,588,1584,1608]" italics="true" pageId="2" pageNumber="594">Cyanotiphia breviclypeola</emphasis>
</taxonomicName>
<emphasis id="B976BB0DA02DFFDFFD9D9672B2B6F9BD" bold="true" box="[593,673,1584,1608]" pageId="2" pageNumber="594">
<taxonomicNameLabel id="A2450676A02DFFDFFD9D9672B2B6F9BD" box="[593,673,1584,1608]" pageId="2" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (♂).
<emphasis id="B976BB0DA02DFFDFFCF09672B358F9BD" bold="true" box="[828,847,1584,1608]" pageId="2" pageNumber="594">1.</emphasis>
Habitus in dorsal view.
<emphasis id="B976BB0DA02DFFDFFB8E9672B442F9BD" bold="true" box="[1090,1109,1584,1608]" pageId="2" pageNumber="594">2.</emphasis>
Head in frontal view.
<emphasis id="B976BB0DA02DFFDFFAF99672B550F9BD" bold="true" box="[1333,1351,1584,1608]" pageId="2" pageNumber="594">3.</emphasis>
Head in dorsal view.
<emphasis id="B976BB0DA02DFFDFFED49616B13CF999" bold="true" box="[280,299,1620,1644]" pageId="2" pageNumber="594">4.</emphasis>
Dorsal face of propodeum.
<emphasis id="B976BB0DA02DFFDFFD859616B24CF999" bold="true" box="[585,603,1620,1644]" pageId="2" pageNumber="594">5.</emphasis>
S1 in ventral view.
<emphasis id="B976BB0DA02DFFDFFCEB9616B32DF999" bold="true" box="[807,826,1620,1644]" pageId="2" pageNumber="594">6.</emphasis>
Lateral face of propodeum.
<emphasis id="B976BB0DA02DFFDFFB909616B478F999" bold="true" box="[1116,1135,1620,1644]" pageId="2" pageNumber="594">7.</emphasis>
Metasoma in ventral view.
<emphasis id="B976BB0DA02DFFDFFA459616B58BF999" bold="true" box="[1417,1436,1620,1644]" pageId="2" pageNumber="594">8.</emphasis>
T7 in dorsal view (the arrow shows lateral carina).
</paragraph>
</caption>
<subSubSection id="C3183494A02DFFDEFF0B96A0B413FE90" lastPageId="3" lastPageNumber="595" pageId="2" pageNumber="594" type="diagnosis">
<paragraph id="8BBD671FA02DFFDFFF0B96A0B590F80D" blockId="2.[151,1436,1762,2041]" pageId="2" pageNumber="594">
<emphasis id="B976BB0DA02DFFDFFF0B96A0B12DF909" bold="true" box="[199,314,1762,1788]" pageId="2" pageNumber="594">Diagnosis</emphasis>
. The male of this species is similar to that of
<taxonomicName id="4C021C9CA02DFFDFFCFC96A0B505F909" authority="(Tsuneki, 1986)" authorityName="Feng &amp; Chen &amp; Li" authorityYear="2025" baseAuthorityName="Tsuneki" baseAuthorityYear="1986" box="[816,1298,1762,1788]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="594" phylum="Arthropoda" rank="species" species="fenchihuensis" status="comb. nov.">
<emphasis id="B976BB0DA02DFFDFFCFC96A0B44EF909" box="[816,1113,1762,1788]" italics="true" pageId="2" pageNumber="594">Cyanotiphia fenchihuensis</emphasis>
(
<bibRefCitation id="EF931AEEA02DFFDFFBAB96A0B51DF909" author="Tsuneki, K." box="[1127,1290,1762,1788]" pageId="2" pageNumber="594" pagination="1 - 8" refId="ref4671" refString="Tsuneki, K. (1986) A contribution to the knowledge of the Taiwanese Tiphiidae (Hymenoptera, Tiphiidae). Special Publications of the Japan Hymenopterists Association, 33, 1-8." type="journal article" year="1986">Tsuneki, 1986</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B976BB0DA02DFFDFFAD196A0B58BF909" bold="true" box="[1309,1436,1762,1788]" pageId="2" pageNumber="594">
<taxonomicNameLabel id="A2450676A02DFFDFFAD196A0B58BF909" box="[1309,1436,1762,1788]" pageId="2" pageNumber="594" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
in body color, the angle on transverse carina of anterior T1, and the distribution of punctures on body, but distinctly differs in the following characters: anterior margin of clypeus almost straight (
<figureCitation id="13397B9AA02DFFDFFBC29768B444F8B1" box="[1038,1107,1834,1860]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Fig. 2</figureCitation>
); lower frons without medial longitudinal carina (
<figureCitation id="13397B9AA02DFFDFFEBA970CB1CFF89D" box="[374,472,1870,1896]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Figs 2, 3</figureCitation>
); dorsal pronotum posteriorly with impunctate margin (
<figureCitation id="13397B9AA02DFFDFFB8D970CB496F89D" box="[1089,1153,1870,1896]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Fig. 1</figureCitation>
); mesopleuron anteriorly with vertical carina not extending to ventral side; lateral longitudinal carina of disc of propodeum oblique outward, with short oblique and straight wrinkles between longitudinal carina and propodeal areola, propodeal areola short and wide (
<figureCitation id="13397B9AA02DFFDFFEC697F8B159F821" box="[266,334,1978,2004]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Fig. 4</figureCitation>
); and punctures on T1 denser, almost reticulate, and laterally even denser (
<figureCitation id="13397B9AA02DFFDFFB4997F8B4D2F821" box="[1157,1221,1978,2004]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Fig. 1</figureCitation>
); S1 with few deep punctures along lateral carina, and without wrinkles (
<figureCitation id="13397B9AA02DFFDFFD27979CB327F80C" box="[747,816,2014,2041]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Fig. 5</figureCitation>
); pygidium of T7 present, with lateral carina (
<figureCitation id="13397B9AA02DFFDFFAF8979CB56EF80D" box="[1332,1401,2014,2040]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="2" pageNumber="594">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BBD671FA02CFFDEFF0B90D5B413FE90" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="595">
It also can be distinguished from the
<typeStatus id="54B9D9BDA02CFFDEFDB390D5B2B8FF44" box="[639,687,151,177]" pageId="3" pageNumber="595">type</typeStatus>
species
<taxonomicName id="4C021C9CA02CFFDEFCD990D5B4C6FF44" authority="Cameron, 1907" authorityName="Cameron" authorityYear="1907" box="[789,1233,151,177]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="ruficauda">
<emphasis id="B976BB0DA02CFFDEFCD990D5B400FF44" box="[789,1047,151,177]" italics="true" pageId="3" pageNumber="595">Cyanotiphia ruficauda</emphasis>
<bibRefCitation id="EF931AEEA02CFFDEFBED90D5B4C6FF44" author="Cameron, P." box="[1057,1233,151,177]" pageId="3" pageNumber="595" pagination="287 - 288" refId="ref4392" refString="Cameron, P. (1907) On the bornean Tiphiidae, including a new genus. The Entomologist, 40, 287-288." type="journal article" year="1907">Cameron, 1907</bibRefCitation>
</taxonomicName>
by the following characters: body black, without metallic blue (
<figureCitation id="13397B9AA02CFFDEFD6C90F9B2F5FF20" box="[672,738,187,213]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); punctures shallower on head and mesosoma, but deeper and denser on metasoma; anterior margin of clypeus almost straight (
<figureCitation id="13397B9AA02CFFDEFCA4909DB3BBFF0C" box="[872,940,223,249]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 2</figureCitation>
); propodeal areola shorter,
<collectionCode id="ED13FFDAA02CFFDEFB17909DB53BFF0C" box="[1243,1324,223,249]" pageId="3" pageNumber="595">APWL</collectionCode>
= 2.2: 1.4: 1.4 (
<figureCitation id="13397B9AA02CFFDEFF059141B119FEE8" box="[201,270,259,285]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 4</figureCitation>
); lateral longitudinal carina of disc of propodeum oblique outward (
<figureCitation id="13397B9AA02CFFDEFBC99141B45DFEE8" box="[1029,1098,259,285]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 4</figureCitation>
); transverse carina on anterior
<specimenCode id="DBA4CF64A02CFFDEFF5B9165B0A1FEB4" box="[151,182,295,321]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="595">T1</specimenCode>
angled distinctly backward in the middle (
<figureCitation id="13397B9AA02CFFDEFD679165B2E7FEB4" box="[683,752,295,321]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); the glossy cambered area on posterior
<specimenCode id="DBA4CF64A02CFFDEFB029165B4FAFEB4" box="[1230,1261,295,321]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="595">T1</specimenCode>
with a distinct anterior edge (
<figureCitation id="13397B9AA02CFFDEFEF59109B16CFE90" box="[313,379,331,357]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); and pygidium of
<specimenCode id="DBA4CF64A02CFFDEFD9F9109B265FE90" box="[595,626,331,357]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="595">T7</specimenCode>
present, with lateral carina (
<figureCitation id="13397B9AA02CFFDEFC7E9109B3E1FE90" box="[946,1014,331,357]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3183494A02CFFDEFF0B912CB189FA68" pageId="3" pageNumber="595" type="description">
<paragraph id="8BBD671FA02CFFDEFF0B912CB47FFE7C" blockId="3.[151,1437,151,1545]" box="[199,1128,366,393]" pageId="3" pageNumber="595">
<emphasis id="B976BB0DA02CFFDEFF0B912CB146FE7D" bold="true" box="[199,337,366,392]" pageId="3" pageNumber="595">Description</emphasis>
. Male,
<typeStatus id="54B9D9BDA02CFFDEFE6B912DB21EFE7C" box="[423,521,367,393]" pageId="3" pageNumber="595" type="holotype">holotype</typeStatus>
(
<figureCitation id="13397B9AA02CFFDEFDD4912DB24DFE7C" box="[536,602,367,393]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
). Body length
<quantity id="4CFACAFAA02CFFDEFCCF912CB34CFE7C" box="[771,859,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="595" unit="mm" value="9.5">9.5 mm</quantity>
, forewing length
<quantity id="4CFACAFAA02CFFDEFBD3912DB474FE7C" box="[1055,1123,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="595" unit="mm" value="7.0">7 mm</quantity>
.
</paragraph>
<paragraph id="8BBD671FA02CFFDEFF0B91D1B285FE00" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="595">
Color. Body almost entirely black; spur of fore leg brown (
<figureCitation id="13397B9AA02CFFDEFCBB91D1B3C8FE58" box="[887,991,403,429]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Figs 1–3</figureCitation>
); wings semitransparent, and marginal cell of forewing dark (
<figureCitation id="13397B9AA02CFFDEFE5991F5B1CFFE24" box="[405,472,439,465]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); setae of head and mesosoma brown mixed with white dorsally and white ventrally; metasomal setae brown mixed white (
<figureCitation id="13397B9AA02CFFDEFD8C9199B293FE00" box="[576,644,475,501]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BBD671FA02CFFDEFF0B91BDB4C4FD5C" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="595">
Head. Mandible with preapical denticle, and without distinct medial transverse groove; clypeus short, its length subequal to WAS, slightly raised in the middle, anterior margin almost straight and with punctures and dense setae (
<figureCitation id="13397B9AA02CFFDEFF539205B0F2FD94" box="[159,229,583,609]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 2</figureCitation>
); W: OW: L: IOD=8.5: 3.9: 3.5: 4.6; OOD: POD: Od= 10: 9: 9; AOD: WAS: IAD: CL= 5: 7: 2: 8.5; lower frons with dense and reticulate punctures, and without medial longitudinal narrow carina (
<figureCitation id="13397B9AA02CFFDEFB5C9229B4C2FD70" box="[1168,1237,619,645]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 2</figureCitation>
); upper frons and vertex with moderate punctures, interspaces between punctures more or less polished (
<figureCitation id="13397B9AA02CFFDEFBAC92CDB4D2FD5C" box="[1120,1221,655,681]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Figs 2, 3</figureCitation>
).
</paragraph>
<paragraph id="8BBD671FA02CFFDEFF0B92F1B2FEFBAC" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="595">
Mesosoma. Pronotal antero-dorsal carina distinct, medially sinuate backward triangularly; dorsal and lateral pronotum with dense punctures and with an impunctate posterior margin; mesoscutum, mesopleuron, scutellum and metanotum with dense punctures similar to reticulation; mesopleuron with anterior vertical carina extending from superior margin to lower margin of lateral side; propodeal areola highly raised and trapezoidal, APWL= 2.2: 1.4: 1.4, medial longitudinal carina reaching posterior 1/3 of areola (
<figureCitation id="13397B9AA02CFFDEFC6A9301B3FAFCA8" box="[934,1005,835,861]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 4</figureCitation>
); lateral longitudinal carina of disc of propodeum oblique outward, with short oblique and straight wrinkles between longitudinal carina and propodeal areola, propodeal areola short and wide, almost as long as wide (
<figureCitation id="13397B9AA02CFFDEFC4093C9B3C3FC50" box="[908,980,907,933]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 4</figureCitation>
); lateral surface of propodeum dorsally with distinct, long and oblique wrinkles, ventrally impunctate (
<figureCitation id="13397B9AA02CFFDEFC9193EDB3B5FC3C" box="[861,930,943,969]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 6</figureCitation>
); posterior surface of propodeum with radial wrinkles; tegula large, reaching posterior 1/3 of mesoscutellum, opaque with shallow and dense punctures on anterior margin and interior half, and impunctate on external half (
<figureCitation id="13397B9AA02CFFDEFCF893B5B36FFBE4" box="[820,888,1015,1041]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); marginal cell of fore wing completely closed, the first submarginal cell incompletely separated by incomplete vein
<emphasis id="B976BB0DA02CFFDEFC539459B3DCFBC0" box="[927,971,1051,1077]" italics="true" pageId="3" pageNumber="595">2RS</emphasis>
(
<figureCitation id="13397B9AA02CFFDEFC179459B40AFBC0" box="[987,1053,1051,1077]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); fore tibia with one serrated spur apically; mid and hind tibiae with two spurs apically.
</paragraph>
<paragraph id="8BBD671FA02CFFDEFF0B9421B566FA8C" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="595">
Metasoma. T1 separated by distinct transverse carina followed by short longitudinal wrinkles and angled backward in the middle (
<figureCitation id="13397B9AA02CFFDEFE6B94C5B1FCFB54" box="[423,491,1159,1185]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Figs 1</figureCitation>
); punctures on T1 dense, almost reticulate, and even denser on lateral side (
<figureCitation id="13397B9AA02CFFDEFAEB94C5B571FB54" box="[1319,1382,1159,1185]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); T1 with impunctate margin posteriorly (
<figureCitation id="13397B9AA02CFFDEFDFE94E9B264FB30" box="[562,627,1195,1221]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); S1 with few deep punctures along lateral carina, and without wrinkles (
<figureCitation id="13397B9AA02CFFDEFF53948DB0F0FB1C" box="[159,231,1230,1257]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 5</figureCitation>
); T2 anteriorly with row of short longitudinal wrinkles; posterior margin of T2 with narrow impunctate cambered area; other part of T2 distinctly punctate, punctures in the middle much sparser than laterally (
<figureCitation id="13397B9AA02CFFDEFA8B94B1B59DFAF8" box="[1351,1418,1267,1293]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 1</figureCitation>
); T3–T6 with dense punctures, anteriorly smaller and sparser; S2–S5 with sparse punctures and denser on posterior margin (
<figureCitation id="13397B9AA02CFFDEFF389579B120FAA0" box="[244,311,1339,1365]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 7</figureCitation>
); S5 without lateral denticle; S6 with moderate punctures denser on posterior margin; pygidium of T7 present, with lateral carina; T7 densely puncto-reticulate and medially with longitudinal narrow carina (
<figureCitation id="13397B9AA02CFFDEFAD3951DB573FA8C" box="[1311,1380,1375,1401]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1584,1608]" captionTargetBox="[151,1436,184,1560]" captionTargetId="figure-18@2.[151,1436,181,1560]" captionTargetPageId="2" captionText="FIGURES 1–8. Cyanotiphia breviclypeola sp. nov., holotype (♂). 1. Habitus in dorsal view. 2. Head in frontal view. 3. Head in dorsal view. 4. Dorsal face of propodeum. 5. S1 in ventral view. 6. Lateral face of propodeum. 7. Metasoma in ventral view. 8. T7 in dorsal view (the arrow shows lateral carina)." figureDoi="http://doi.org/10.5281/zenodo.14853716" httpUri="https://zenodo.org/record/14853716/files/figure.png" pageId="3" pageNumber="595">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BBD671FA02CFFDEFF0B95C1B189FA68" blockId="3.[151,1437,151,1545]" box="[199,414,1411,1437]" pageId="3" pageNumber="595">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3183494A02CFFDEFF0B95E4B25FFA34" box="[199,584,1446,1473]" pageId="3" pageNumber="595" type="distribution">
<paragraph id="8BBD671FA02CFFDEFF0B95E4B25FFA34" blockId="3.[151,1437,151,1545]" box="[199,584,1446,1473]" pageId="3" pageNumber="595">
<emphasis id="B976BB0DA02CFFDEFF0B95E4B14EFA35" bold="true" box="[199,345,1446,1472]" pageId="3" pageNumber="595">Distribution</emphasis>
.
<collectingCountry id="F315278FA02CFFDEFEAB95E5B1BDFA34" box="[359,426,1447,1473]" name="China" pageId="3" pageNumber="595">China</collectingCountry>
(
<collectingRegion id="49C6A9FDA02CFFDEFE7595E5B22BFA34" box="[441,572,1447,1473]" country="China" name="Chongqing" pageId="3" pageNumber="595">Chongqing</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3183494A02CFFDEFF0B9589B384F9FC" pageId="3" pageNumber="595" type="etymology">
<paragraph id="8BBD671FA02CFFDEFF0B9589B384F9FC" blockId="3.[151,1437,151,1545]" pageId="3" pageNumber="595">
<emphasis id="B976BB0DA02CFFDEFF0B9589B15FFA10" bold="true" box="[199,328,1483,1509]" pageId="3" pageNumber="595">Etymology</emphasis>
. The specific name of this species
<taxonomicName id="4C021C9CA02CFFDEFD189589B37CFA10" authorityName="Feng &amp; Chen &amp; Li" authorityYear="2025" box="[724,875,1483,1509]" class="Insecta" family="Vespidae" genus="Cyanotiphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="595" phylum="Arthropoda" rank="species" species="breviclypeola">
<emphasis id="B976BB0DA02CFFDEFD189589B37CFA10" box="[724,875,1483,1509]" italics="true" pageId="3" pageNumber="595">breviclypeola</emphasis>
</taxonomicName>
is derived from two Latin words:
<emphasis id="B976BB0DA02CFFDEFB239589B525FA10" box="[1263,1330,1483,1509]" italics="true" pageId="3" pageNumber="595">brevis</emphasis>
(= short) +
<emphasis id="B976BB0DA02CFFDEFF6295ADB10FF9FC" box="[174,280,1519,1545]" italics="true" pageId="3" pageNumber="595">clypeolus</emphasis>
(= clypeus), referring to the characteristic short clypeus.
</paragraph>
</subSubSection>
</treatment>
</document>
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