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<document id="00E1A44C99287981065420D5F9B9CB1F" ID-DOI="10.11646/zootaxa.4810.1.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="3937036" ID-ZooBank="424415DC-807E-493A-8883-903D2ADE9427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594292134250" checkinUser="plazi" docAuthor="Zhu, Wenhui, Yao, Zhiyuan, Zheng, Guo &amp; Li, Shuqiang" docDate="2020" docId="039FEF343D0AFFC9FF3CF470947FFE74" docLanguage="en" docName="zootaxa.4810.1.12.pdf" docOrigin="Zootaxa 4810 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana maoer Zhu &amp; Yao &amp; Zheng &amp; Li 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="182" masterDocId="FFA6974C3D0EFFCEFFABF735972AFFDB" masterDocTitle="Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China" masterLastPageNumber="197" masterPageNumber="175" pageNumber="179" updateTime="1700277344912" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EC97CF41A536CA928038D1A5039D0F37">Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China</mods:title>
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<mods:namePart id="BFB9C60484F049270256659F5B888B63">Zhu, Wenhui</mods:namePart>
<mods:affiliation id="69B2343296B8EB167F2E6CDC0CAA8D38">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
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<mods:namePart id="3E6923FF433A9965267EE0EBFA9B6887">Yao, Zhiyuan</mods:namePart>
<mods:affiliation id="0309B3CB4C39D670A3B3723A0C73752C">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; yaozy @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 1631 - 0949</mods:affiliation>
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<mods:namePart id="A1935FD6BAC128E2D63069330D7A17C2">Zheng, Guo</mods:namePart>
<mods:affiliation id="9CDF1D09647ADADDC914635B4422535B">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhengguo @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 9735 - 9700</mods:affiliation>
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<mods:namePart id="687EF69F383CF0AA921F00C77232A7E9">Li, Shuqiang</mods:namePart>
<mods:affiliation id="9D3D4E0D11C9DEB8E3A5EC72A73EF900">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
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<mods:date id="0A9AB4C34AEB52C4D6FF5576FB6CE379">2020</mods:date>
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<emphasis id="B94282303D0AFFCAFF3CF4709518FC84" bold="true" box="[151,562,837,864]" pageId="4" pageNumber="179">
<taxonomicName id="4C3625A13D0AFFCAFF3CF47096E4FC84" ID-CoL="93VJF" authority="Yao &amp; Li" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[151,462,837,864]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="maoer" status="sp. nov.">
<emphasis id="B94282303D0AFFCAFF3CF470967FFC84" bold="true" box="[151,341,837,864]" italics="true" pageId="4" pageNumber="179">Belisana maoer</emphasis>
Yao &amp; Li
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<taxonomicNameLabel id="A2713F4B3D0AFFCAFE7EF4709518FC84" box="[469,562,837,863]" pageId="4" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B895E223D0AFFCAFF3CF45C969CFC58" blockId="4.[151,562,837,899]" box="[151,438,873,899]" pageId="4" pageNumber="179">
<figureCitation id="130D42A73D0AFFCAFF3CF45C97D1FC58" box="[151,251,873,899]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1875,1900]" captionStartId-1="6.[151,250,1875,1900]" captionTargetBox-0="[226,1361,181,1850]" captionTargetBox-1="[226,1361,181,1850]" captionTargetId-0="figure@5.[226,1361,181,1850]" captionTargetId-1="figure@6.[226,1361,181,1850]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." captionText-1="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.3937042" figureDoi-1="http://doi.org/10.5281/zenodo.3937044" httpUri-0="https://zenodo.org/record/3937042/files/figure.png" httpUri-1="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 34</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFEADF45C960AFC58" box="[262,288,873,899]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13</figureCitation>
CD, 14CD
</paragraph>
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<paragraph id="8B895E223D0AFFCAFF3CF484926FFBC8" blockId="4.[151,1437,944,2015]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF3CF484966CFC10" bold="true" box="[151,326,944,971]" pageId="4" pageNumber="179">Type material.</emphasis>
<materialsCitation id="3B5E547F3D0AFFCAFEE6F48497EBFBC8" collectingDate="2014-04-26" collectionCode="IZCAS" collectorName="Y. Lin &amp; H. Zhao &amp; Y. Li &amp; J. Wu &amp; F. Li" country="China" county="Xingwen County" elevation="345" latitude="28.294369" location="Maoer Cave" longLatPrecision="1" longitude="105.2212" municipality="Yibin" pageId="4" pageNumber="179" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
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<typeStatus id="548DE0803D0AFFCAFEE6F4849694FC10" box="[333,446,945,971]" pageId="4" pageNumber="179" type="holotype">Holotype</typeStatus>
:
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Male (
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Ar39850),
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(
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,
<geoCoordinate id="EE0238E53D0AFFCAFB95F48593DDFC10" box="[1086,1271,944,971]" degrees="105" direction="east" minutes="13" orientation="longitude" pageId="4" pageNumber="179" precision="1" seconds="16.32" value="105.2212">105°1316.32”E</geoCoordinate>
,
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elevation
<quantity id="4CCEF3C73D0AFFCAFAD9F485979AFC34" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="4" pageNumber="179" unit="m" value="345.0">345 m</quantity>
</elevation>
),
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,
<collectingCounty id="62E826AE3D0AFFCAFEC4F4E0951AFC34" box="[367,560,981,1007]" pageId="4" pageNumber="179">Xingwen County</collectingCounty>
,
<collectingMunicipality id="6BEDC4583D0AFFCAFD92F4E09550FC34" box="[569,634,981,1007]" pageId="4" pageNumber="179">Yibin</collectingMunicipality>
,
<emphasis id="B94282303D0AFFCAFD2EF4E195CFFC34" bold="true" box="[645,741,980,1007]" pageId="4" pageNumber="179">
<collectingRegion id="49F290C03D0AFFCAFD2EF4E195CFFC34" box="[645,741,980,1007]" country="China" name="Sichuan" pageId="4" pageNumber="179">Sichuan</collectingRegion>
</emphasis>
,
<emphasis id="B94282303D0AFFCAFD5FF4E19417FC34" bold="true" box="[756,829,980,1007]" pageId="4" pageNumber="179">
<collectingCountry id="F3211EB23D0AFFCAFD5FF4E19417FC34" box="[756,829,980,1007]" name="China" pageId="4" pageNumber="179">China</collectingCountry>
</emphasis>
,
<date id="FF8878E23D0AFFCAFCE0F4E094C3FC34" box="[843,1001,981,1007]" pageId="4" pageNumber="179" value="2014-04-26">
<collectingDate id="EFCC810A3D0AFFCAFCE0F4E094C3FC34" box="[843,1001,981,1007]" pageId="4" pageNumber="179" value="2014-04-26">26 April 2014</collectingDate>
</date>
,
<collectorName id="26C33BF43D0AFFCAFC59F4E09311FC34" box="[1010,1083,981,1007]" pageId="4" pageNumber="179">Y. Lin</collectorName>
,
<collectorName id="26C33BF43D0AFFCAFBEEF4E09389FC34" box="[1093,1187,981,1007]" pageId="4" pageNumber="179">H. Zhao</collectorName>
,
<collectorName id="26C33BF43D0AFFCAFB05F4E093CDFC34" box="[1198,1255,981,1007]" pageId="4" pageNumber="179">Y. Li</collectorName>
,
<collectorName id="26C33BF43D0AFFCAFB58F4E09218FC34" box="[1267,1330,981,1007]" pageId="4" pageNumber="179">J. Wu</collectorName>
and
<collectorName id="26C33BF43D0AFFCAFAC3F4E092B6FC34" box="[1384,1436,981,1007]" pageId="4" pageNumber="179">F. Li</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B5E547F3D0AFFCAFF6CF4CC95B7FBC8" box="[199,669,1016,1043]" collectionCode="IZCAS" country="China" pageId="4" pageNumber="179" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="B94282303D0AFFCAFF6CF4CC9662FBC8" bold="true" box="[199,328,1017,1043]" pageId="4" pageNumber="179">
<typeStatus id="548DE0803D0AFFCAFF6CF4CC9669FBC8" box="[199,323,1017,1043]" pageId="4" pageNumber="179" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D3095AB3D0AFFCAFEE4F4CC96B0FBC8" box="[335,410,1017,1043]" pageId="4" pageNumber="179" type="male">1 male</specimenCount>
(
<collectionCode id="ED27C6E73D0AFFCAFE02F4CC96D6FBC8" box="[425,508,1017,1043]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="179">IZCAS</collectionCode>
Ar39851) and
</materialsCitation>
<materialsCitation id="3B5E547F3D0AFFCAFD0EF4CC926BFBC8" box="[677,1345,1016,1043]" collectingDate="2014-04-26" collectionCode="IZCAS" collectorName="Y. Lin &amp; H. Zhao &amp; Y. Li &amp; J. Wu &amp; F. Li" country="China" county="Xingwen County" elevation="345" latitude="28.294369" location="Maoer Cave" longLatPrecision="1" longitude="105.2212" municipality="Yibin" pageId="4" pageNumber="179" specimenCount="2" specimenCount-female="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D3095AB3D0AFFCAFD0EF4CC943BFBC8" box="[677,785,1017,1043]" pageId="4" pageNumber="179" type="female">2 females</specimenCount>
(
<collectionCode id="ED27C6E73D0AFFCAFCB4F4CC9458FBC8" box="[799,882,1017,1043]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="179">IZCAS</collectionCode>
Ar3985239853), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0AFFCAFF6CF3289356FBEC" box="[199,1148,1053,1079]" pageId="4" pageNumber="179" type="etymology">
<paragraph id="8B895E223D0AFFCAFF6CF3289356FBEC" blockId="4.[151,1437,944,2015]" box="[199,1148,1053,1079]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF3289664FBEC" bold="true" box="[199,334,1053,1079]" pageId="4" pageNumber="179">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="548DE0803D0AFFCAFD1EF32895CFFBEC" box="[693,741,1053,1079]" pageId="4" pageNumber="179">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0AFFCAFF6CF3759651FA8C" pageId="4" pageNumber="179" type="diagnosis">
<paragraph id="8B895E223D0AFFCAFF6CF3759651FA8C" blockId="4.[151,1437,944,2015]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF3759668FB80" bold="true" box="[199,322,1088,1115]" pageId="4" pageNumber="179">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C3625A13D0AFFCAFDE0F37794FEFB80" authority="Zhang &amp; Peng, 2011" authorityName="Zhang &amp; Peng" authorityYear="2011" box="[587,980,1089,1115]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="galeiformis">
<emphasis id="B94282303D0AFFCAFDE0F37795C3FB80" box="[587,745,1089,1115]" italics="true" pageId="4" pageNumber="179">B. galeiformis</emphasis>
<bibRefCitation id="EFA723D33D0AFFCAFD44F37494FEFB80" author="Zhang, F. &amp; Peng, Y." box="[751,980,1089,1115]" pageId="4" pageNumber="179" pagination="51 - 68" refId="ref9108" refString="Zhang, F. &amp; Peng, Y. (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa, 2989 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1.2" type="journal article" year="2011">Zhang &amp; Peng, 2011</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFA723D33D0AFFCAFBA7F374923CFB80" author="Zhang, F. &amp; Peng, Y." box="[1036,1302,1088,1115]" pageId="4" pageNumber="179" pagination="51 - 68" refId="ref9108" refString="Zhang, F. &amp; Peng, Y. (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa, 2989 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1.2" type="journal article" year="2011">Zhang &amp; Peng 2011: 52</bibRefCitation>
, fig. 1) with similar male chelicerae (
<figureCitation id="130D42A73D0AFFCAFE05F3509523FBA4" box="[430,521,1125,1151]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Fig. 4D</figureCitation>
), bulbal apophyses (
<figureCitation id="130D42A73D0AFFCAFD58F3509460FBA4" box="[755,842,1125,1151]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Fig. 4C</figureCitation>
) and vulval pore plates (arrow pp in
<figureCitation id="130D42A73D0AFFCAFB58F3509264FBA4" box="[1267,1358,1125,1151]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 4B</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFAF7F35092A5FBA4" box="[1372,1423,1125,1151]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="4" pageNumber="179">14D</figureCitation>
), but males can be distinguished by presence of curved subdisto-prolateral sclerite on procursus (arrow
<quantity id="4CCEF3C73D0AFFCAFAA7F3BC921DFB78" box="[1292,1335,1161,1187]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="179" unit="in" value="1.0">1 in</quantity>
<figureCitation id="130D42A73D0AFFCAFA96F3BC92BDFB78" box="[1341,1431,1161,1187]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFF3CF39897E3FB1C" box="[151,201,1197,1223]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13C</figureCitation>
), by presence of long retrolatero-ventral apophysis on pedipalpal trochanter (arrow in
<figureCitation id="130D42A73D0AFFCAFB26F39893C8FB1C" box="[1165,1250,1197,1223]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Fig. 3B</figureCitation>
) and by absence of proximo-dorsal apophysis on pedipalpal femur (
<figureCitation id="130D42A73D0AFFCAFD49F3E49400FB30" box="[738,810,1233,1259]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3</figureCitation>
AB). Females can be distinguished by having frontal “epigynal” pockets anteriorly on sclerotized plate (arrow ep in
<figureCitation id="130D42A73D0AFFCAFCFFF3C094B3FAD4" box="[852,921,1269,1295]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 4</figureCitation>
AB, 14CD) and strongly protruding median ridge of anterior arch (arrow in
<figureCitation id="130D42A73D0AFFCAFE52F22C9578FAE8" box="[505,594,1305,1331]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 4B</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFDF5F22C95B8FAE8" box="[606,658,1305,1331]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="4" pageNumber="179">14D</figureCitation>
). Also distinguished by absence of median marks on carapace of both sexes (
<figureCitation id="130D42A73D0AFFCAFF48F2089645FA8C" box="[227,367,1341,1367]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 4E, 4G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0AFFC9FF6CF25593DAFE50" lastPageId="7" lastPageNumber="182" pageId="4" pageNumber="179" type="description">
<paragraph id="8B895E223D0AFFCAFF6CF25595D4F8A8" blockId="4.[151,1437,944,2015]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF25596B5FAA0" bold="true" box="[199,415,1376,1403]" pageId="4" pageNumber="179">Description. Male</emphasis>
(
<emphasis id="B94282303D0AFFCAFE19F2549533FAA0" bold="true" box="[434,537,1377,1403]" pageId="4" pageNumber="179">
<typeStatus id="548DE0803D0AFFCAFE19F2549533FAA0" box="[434,537,1377,1403]" pageId="4" pageNumber="179" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar39850): Total length 1.75 (1.88 with clypeus), carapace 0.73 long, 0.70 wide, opisthosoma 1.02 long, 0.86 wide. Leg I: 16.86 (4.23 + 0.26 + 4.36 + 6.73 + 1.28), leg II: 11.22 (3.21 + 0.25 + 2.76 + 4.17 + 0.83), leg III: 7.55 (2.24 + 0.19 + 1.92 + 2.69 + 0.51), leg IV: 9.70 (2.82 + 0.22 + 2.56 + 3.33 + 0.77); tibia I L/d: 70. Distance PME-PME 0.22, diameter PME 0.08, distance PME-ALE 0.09, AME absent. Sternum slightly wider than long (0.63/0.55). Habitus as in
<figureCitation id="130D42A73D0AFFCAFC91F2C494AAF9D0" box="[826,896,1521,1547]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 4</figureCitation>
EF. Carapace pale, with indistinct black lateral margins; sternum pale, without marks. Legs pale, without darker rings. Opisthosoma pale, with indistinct dark spots. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Chelicerae (
<figureCitation id="130D42A73D0AFFCAFB9FF10C93A0F988" box="[1076,1162,1593,1619]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Fig. 4D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of short, curved distal apophyses (distance between tips of distal apophyses: 0.31). Pedipalps as in
<figureCitation id="130D42A73D0AFFCAFEEEF1B496A3F940" box="[325,393,1665,1691]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3</figureCitation>
AB; coxa unmodified; trochanter with long retrolatero-ventral apophysis (arrow in
<figureCitation id="130D42A73D0AFFCAFA9CF1B492A4F940" box="[1335,1422,1665,1691]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Fig. 3B</figureCitation>
); femur without proximo-dorsal hump; procursus (
<figureCitation id="130D42A73D0AFFCAFD10F19095D5F964" box="[699,767,1701,1727]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3</figureCitation>
AD, 13CD) simple proximally but complex distally, with curved subdisto-prolateral sclerite (arrow
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<figureCitation id="130D42A73D0AFFCAFD03F1FC942EF938" box="[680,772,1737,1763]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFCB9F1FC9469F938" box="[786,835,1737,1763]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13C</figureCitation>
), long distal spine (arrow
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<figureCitation id="130D42A73D0AFFCAFB0EF1FC922BF938" box="[1189,1281,1737,1763]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFAA5F1FC926AF938" box="[1294,1344,1737,1763]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13C</figureCitation>
), nearly elliptic, distal membranous lamella (arrow
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<figureCitation id="130D42A73D0AFFCAFD07F1D8942CF8DC" box="[684,774,1773,1799]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFCB9F1D89468F8DC" box="[786,834,1773,1799]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13C</figureCitation>
) and narrow retrolateral membranous flap (arrow f in
<figureCitation id="130D42A73D0AFFCAFF3CF02497D8F8F0" box="[151,242,1809,1835]" captionStart="FIGURE 3" captionStartId="5.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@5.[226,1361,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Belisana maoer sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="4" pageNumber="179">Figs 3D</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFF55F024961BF8F0" box="[254,305,1809,1835]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13D</figureCitation>
), without subdisto-dorsal protruding part; bulb (
<figureCitation id="130D42A73D0AFFCAFCE1F02494B5F8F0" box="[842,927,1809,1835]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Fig. 4C</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia I at 10% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus I with 22 distinct pseudosegments.
</paragraph>
<paragraph id="8B895E223D0AFFC9FF6CF048945DFEC4" blockId="4.[151,1437,944,2015]" lastBlockId="7.[151,1437,153,431]" lastPageId="7" lastPageNumber="182" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF0489634F84C" bold="true" box="[199,286,1917,1943]" pageId="4" pageNumber="179">Female</emphasis>
(IZCAS Ar39852): Similar to male, habitus as in
<figureCitation id="130D42A73D0AFFCAFCCBF048948DF84C" box="[864,935,1917,1943]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="4" pageNumber="179">Figs 4</figureCitation>
GH. Total length 1.81 (1.90 with clypeus), carapace 0.75 long, 0.70 wide, opisthosoma 1.06 long, 0.81 wide. Tibia I: 3.01; tibia I L/d: 48. Distance PME-PME 0.24, diameter PME 0.08, distance PME-ALE 0.08, AME absent. Sternum slightly wider than long (0.55/0.47). Carapace with black margins. External female genitalia (
<figureCitation id="130D42A73D09FFC9FCB5F7AC9457FF68" box="[798,893,153,179]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="7" pageNumber="182">Figs 4A</figureCitation>
,
<figureCitation id="130D42A73D09FFC9FC21F7AC9490FF68" box="[906,954,153,179]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14C</figureCitation>
) simple and flat, light brown, with pair of frontal “epigynal” pockets anteriorly on sclerotized plate (arrow ep in
<figureCitation id="130D42A73D09FFC9FC00F78894DBFF0C" box="[939,1009,189,215]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="7" pageNumber="182">Figs 4</figureCitation>
AB, 14CD), 0.11 apart. Vulva (
<figureCitation id="130D42A73D09FFC9FAC1F7889790FF20" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="7" pageNumber="182">Figs 4B</figureCitation>
,
<figureCitation id="130D42A73D09FFC9FF6EF7D497DCFF20" box="[197,246,225,251]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14D</figureCitation>
) with W-shaped anterior arch (median ridge strongly protruding, arrow in
<figureCitation id="130D42A73D09FFC9FB86F7D493ACFF20" box="[1069,1158,225,251]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="7" pageNumber="182">Figs 4B</figureCitation>
,
<figureCitation id="130D42A73D09FFC9FB3AF7D493E8FF20" box="[1169,1218,225,251]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14D</figureCitation>
) and pair of narrow pore plates (arrow pp in
<figureCitation id="130D42A73D09FFC9FE01F6309529FEC4" box="[426,515,261,287]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1900]" captionTargetBox="[226,1361,181,1850]" captionTargetId="figure@6.[226,1361,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Belisana maoer sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="7" pageNumber="182">Figs 4B</figureCitation>
,
<figureCitation id="130D42A73D09FFC9FDBBF6309568FEC4" box="[528,578,261,287]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="7" pageNumber="182">14D</figureCitation>
), without serrated sclerites.
</paragraph>
<caption id="DF490EAA3D0BFFCBFF3CF0669638F827" ID-DOI="http://doi.org/10.5281/zenodo.3937042" ID-Zenodo-Dep="3937042" httpUri="https://zenodo.org/record/3937042/files/figure.png" pageId="5" pageNumber="180" startId="5.[151,250,1875,1900]" targetBox="[226,1361,181,1850]" targetPageId="5">
<paragraph id="8B895E223D0BFFCBFF3CF0669638F827" blockId="5.[151,1436,1875,2044]" pageId="5" pageNumber="180">
<emphasis id="B94282303D0BFFCBFF3CF0669639F8B7" bold="true" box="[151,275,1875,1900]" pageId="5" pageNumber="180">FIGURE 3.</emphasis>
<taxonomicName id="4C3625A13D0BFFCBFEB1F0619691F8B7" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[282,443,1876,1900]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="maoer" status="sp. nov.">
<emphasis id="B94282303D0BFFCBFEB1F0619691F8B7" box="[282,443,1876,1900]" italics="true" pageId="5" pageNumber="180">Belisana maoer</emphasis>
</taxonomicName>
<emphasis id="B94282303D0BFFCBFE69F0619539F8B7" bold="true" box="[450,531,1876,1900]" pageId="5" pageNumber="180">
<taxonomicNameLabel id="A2713F4B3D0BFFCBFE69F0619539F8B7" box="[450,531,1876,1900]" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at long retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-prolateral sclerite, arrow 2 points at long distal spine, arrow 3 points at nearly elliptic, distal membranous lamella; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD).
</paragraph>
</caption>
<caption id="DF490EAA3D08FFC8FF3CF06696B8F827" ID-DOI="http://doi.org/10.5281/zenodo.3937044" ID-Zenodo-Dep="3937044" httpUri="https://zenodo.org/record/3937044/files/figure.png" pageId="6" pageNumber="181" startId="6.[151,250,1875,1900]" targetBox="[226,1361,181,1850]" targetPageId="6">
<paragraph id="8B895E223D08FFC8FF3CF06696B8F827" blockId="6.[151,1437,1875,2044]" pageId="6" pageNumber="181">
<emphasis id="B94282303D08FFC8FF3CF066963FF8B7" bold="true" box="[151,277,1875,1900]" pageId="6" pageNumber="181">FIGURE 4.</emphasis>
<taxonomicName id="4C3625A13D08FFC8FEB5F06196EAF8B7" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[286,448,1876,1900]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="maoer" status="sp. nov.">
<emphasis id="B94282303D08FFC8FEB5F06196EAF8B7" box="[286,448,1876,1900]" italics="true" pageId="6" pageNumber="181">Belisana maoer</emphasis>
</taxonomicName>
<emphasis id="B94282303D08FFC8FE63F0619531F8B7" bold="true" box="[456,539,1876,1900]" pageId="6" pageNumber="181">
<taxonomicNameLabel id="A2713F4B3D08FFC8FE63F0619531F8B7" box="[456,539,1876,1900]" pageId="6" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH).
</paragraph>
</caption>
<paragraph id="8B895E223D09FFC9FF6CF61D9661FEBC" blockId="7.[151,1437,153,431]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF61D966BFE98" bold="true" box="[199,321,296,323]" pageId="7" pageNumber="182">Variation:</emphasis>
<materialsCitation id="3B5E547F3D09FFC9FEE7F61C966DFEBC" collectionCode="IZCAS" collectorName="Tibia I" pageId="7" pageNumber="182" specimenCount="2" typeStatus="paratype">
<collectorName id="26C33BF43D09FFC9FEE7F61C96B1FE98" box="[332,411,297,323]" pageId="7" pageNumber="182">Tibia I</collectorName>
in male
<typeStatus id="548DE0803D09FFC9FDA7F61C9546FE98" box="[524,620,297,323]" pageId="7" pageNumber="182" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27C6E73D09FFC9FD2BF61C95F9FE98" box="[640,723,297,323]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="7" pageNumber="182">IZCAS</collectionCode>
Ar39851): 3.76.
<collectorName id="26C33BF43D09FFC9FC0AF61C94DAFE98" box="[929,1008,297,323]" pageId="7" pageNumber="182">Tibia I</collectorName>
in another female
<typeStatus id="548DE0803D09FFC9FB7DF61C921CFE98" box="[1238,1334,297,323]" pageId="7" pageNumber="182" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27C6E73D09FFC9FAE1F61C92B7FE98" box="[1354,1437,297,323]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="7" pageNumber="182">IZCAS</collectionCode>
Ar39853): 2.05
</materialsCitation>
.
</paragraph>
<paragraph id="8B895E223D09FFC9FF6CF64493DAFE50" blockId="7.[151,1437,153,431]" box="[199,1264,368,395]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF64496AFFE50" bold="true" box="[199,389,368,395]" pageId="7" pageNumber="182">Natural history.</emphasis>
The species was found underside of rocks in the aphotic zone inside the cave.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D09FFC9FF6CF6A1947FFE74" box="[199,853,404,431]" pageId="7" pageNumber="182" type="distribution">
<paragraph id="8B895E223D09FFC9FF6CF6A1947FFE74" blockId="7.[151,1437,153,431]" box="[199,853,404,431]" pageId="7" pageNumber="182">
<emphasis id="B94282303D09FFC9FF6CF6A1964BFE74" bold="true" box="[199,353,404,431]" pageId="7" pageNumber="182">Distribution.</emphasis>
<collectingCountry id="F3211EB23D09FFC9FECCF6A09680FE74" box="[359,426,405,431]" name="China" pageId="7" pageNumber="182">China</collectingCountry>
(
<collectingRegion id="49F290C03D09FFC9FE12F6A09532FE74" box="[441,536,405,431]" country="China" name="Sichuan" pageId="7" pageNumber="182">Sichuan</collectingRegion>
,
<typeStatus id="548DE0803D09FFC9FD88F6A09579FE74" box="[547,595,405,431]" pageId="7" pageNumber="182">type</typeStatus>
locality only;
<figureCitation id="130D42A73D09FFC9FD5DF6A09462FE74" box="[758,840,404,431]" captionStart="FIGURE 15" captionStartId="20.[151,250,1181,1206]" captionTargetBox="[151,1436,181,1156]" captionTargetId="figure@20.[151,1436,181,1156]" captionTargetPageId="20" captionText="FIGURE 15. Distribution records of six new species of the genus Belisana from southern China. 1. B. guilin sp. nov.; 2. B. maoer sp. nov.; 3. B. naling sp. nov.; 4. B. xiyuan sp. nov.; 5. B. xuanguan sp. nov.; 6. B. yuexiu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3937066" httpUri="https://zenodo.org/record/3937066/files/figure.png" pageId="7" pageNumber="182">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>