154 lines
17 KiB
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154 lines
17 KiB
XML
<document id="7E3457A7EEDE28D9F1CEFF7CBF58CAC3" ID-CLB-Dataset="20378" ID-DOI="10.11646/zootaxa.5104.4.4" ID-GBIF-Dataset="c3a77909-4443-494a-a3f2-cf57eed196c2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332181" ID-ZooBank="44C79B8A-12BC-4066-9864-E42DCBA0EF6B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646595398227" checkinUser="plazi" docAuthor="Poorani, J." docDate="2022" docId="039FCF4AFFB4FF842AB26E8DFBAFF9ED" docLanguage="en" docName="zootaxa.5104.4.4.pdf" docOrigin="Zootaxa 5104 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Horniolus Weise 1901" docType="treatment" docVersion="7" lastPageNumber="546" masterDocId="FFA6B732FFB5FF852A256D22FFF6FFC4" masterDocTitle="A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species" masterLastPageNumber="556" masterPageNumber="545" pageNumber="546" updateTime="1699316983446" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="53A4400B0F37B642F7ECC1DEC5BBAC91">
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<mods:title id="CAEB28CC06A40B4DF0404452CC3B0993">A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species</mods:title>
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<mods:name id="6D798B32BFF12ACE36D835BB2BBBBEB5" type="personal">
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<mods:namePart id="98DCE59D964EEC7F8490EA7DE17AE00A">Poorani, J.</mods:namePart>
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<mods:title id="29E84CEF7CD69F3AA6D0C9B4A1417B55">Zootaxa</mods:title>
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<mods:part id="A833A617E737878A14C2A2887BAC562E">
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<mods:date id="F3E79B90B0081B548535095DC47EEC38">2022</mods:date>
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<mods:number id="21D1EDA411C20590D2AC3B8442300997">2022-03-01</mods:number>
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<mods:number id="2B1012C1D4445DB77CC778ACFE02156C">5104</mods:number>
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<mods:start id="19C146BC8AD13D4C6F3FB16B777FCA2E">545</mods:start>
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<mods:identifier id="8231C2010028DA9D6E5E14A5746FF80A" type="DOI">10.11646/zootaxa.5104.4.4</mods:identifier>
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<mods:identifier id="8E9C6700576EC448D7A1A7AFFBBEE48D" type="GBIF-Dataset">c3a77909-4443-494a-a3f2-cf57eed196c2</mods:identifier>
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<treatment id="039FCF4AFFB4FF842AB26E8DFBAFF9ED" ID-DOI="http://doi.org/10.5281/zenodo.6333320" ID-GBIF-Taxon="193878385" ID-Zenodo-Dep="6333320" LSID="urn:lsid:plazi:treatment:039FCF4AFFB4FF842AB26E8DFBAFF9ED" httpUri="http://treatment.plazi.org/id/039FCF4AFFB4FF842AB26E8DFBAFF9ED" lastPageNumber="546" pageId="1" pageNumber="546">
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<subSubSection id="C32C2DD7FFB4FF842AB26E8DFE4CFC0D" box="[151,442,943,970]" pageId="1" pageNumber="546" type="nomenclature">
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<paragraph id="8B897E5CFFB4FF842AB26E8DFE4CFC0D" blockId="1.[151,442,943,970]" box="[151,442,943,970]" pageId="1" pageNumber="546">
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<heading id="D0C1C930FFB4FF842AB26E8DFE4CFC0D" bold="true" box="[151,442,943,970]" fontSize="11" level="1" pageId="1" pageNumber="546" reason="1">
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<emphasis id="B942A24EFFB4FF842AB26E8DFE4CFC0D" bold="true" box="[151,442,943,970]" pageId="1" pageNumber="546">
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Genus
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<taxonomicName id="4C3605DFFFB4FF842ACB6E8DFE4CFC0D" authority="Weise" authorityName="Weise" authorityYear="1901" box="[238,442,943,970]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="genus">
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<emphasis id="B942A24EFFB4FF842ACB6E8DFE9CFC0E" bold="true" box="[238,362,943,970]" italics="true" pageId="1" pageNumber="546">Horniolus</emphasis>
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Weise
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C2DD7FFB4FF842AB26ED6FBEFFBC8" box="[151,1049,1012,1036]" pageId="1" pageNumber="546" type="reference_group">
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<paragraph id="8B897E5CFFB4FF842AB26ED6FBEFFBC8" blockId="1.[151,1049,1012,1036]" box="[151,1049,1012,1036]" pageId="1" pageNumber="546">
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<treatmentCitationGroup id="AB265972FFB4FF842AB26ED6FBEFFBC8" box="[151,1049,1012,1036]" pageId="1" pageNumber="546">
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<treatmentCitation id="0A97584DFFB4FF842AB26ED6FE41FBC8" author="Weise, J." box="[151,439,1012,1036]" page="442" pageId="1" pageNumber="546" year="1901">
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<taxonomicName id="4C3605DFFFB4FF842AB26ED6FE41FBC8" authority="Weise, 1901: 442" authorityName="Weise" authorityPageNumber="442" authorityYear="1901" box="[151,439,1012,1036]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="genus">
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<emphasis id="B942A24EFFB4FF842AB26ED6FF08FBC8" box="[151,254,1012,1036]" italics="true" pageId="1" pageNumber="546">Horniolus</emphasis>
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<bibRefCitation id="EFA703ADFFB4FF842B216ED6FE41FBC8" author="Weise, J." box="[260,439,1012,1036]" pageId="1" pageNumber="546" pagination="417 - 448" refId="ref5851" refString="Weise, J. (1901) Coccinelliden aus Ceylon gesammelt von Dr. Horn. Deutsche Entomologische Zeitschrift, 44, 417 - 448." type="journal article" year="1901">Weise, 1901: 442</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="548DC0FEFFB4FF842BE56ED6FE04FBC8" box="[448,498,1012,1036]" pageId="1" pageNumber="546">Type</typeStatus>
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species:
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<taxonomicName id="4C3605DFFFB4FF8428756ED6FC75FBC8" authority="Weise, 1901" authorityName="Weise, R. G. Booth" authorityYear="1901" box="[592,899,1012,1036]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="species" species="dispar">
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<emphasis id="B942A24EFFB4FF8428756ED6FD08FBC8" box="[592,766,1012,1036]" italics="true" pageId="1" pageNumber="546">Horniolus dispar</emphasis>
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<bibRefCitation id="EFA703ADFFB4FF8429216ED6FC75FBC8" author="Weise, J." box="[772,899,1012,1036]" pageId="1" pageNumber="546" pagination="417 - 448" refId="ref5851" refString="Weise, J. (1901) Coccinelliden aus Ceylon gesammelt von Dr. Horn. Deutsche Entomologische Zeitschrift, 44, 417 - 448." type="journal article" year="1901">Weise, 1901</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C2DD7FFB4FF842AE2691BFC52FA5C" pageId="1" pageNumber="546" type="diagnosis">
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<paragraph id="8B897E5CFFB4FF842AE2691BFCECFACC" blockId="1.[151,1437,1081,1577]" pageId="1" pageNumber="546">
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<emphasis id="B942A24EFFB4FF842AE2691BFECCFB97" bold="true" box="[199,314,1081,1107]" pageId="1" pageNumber="546">Diagnosis</emphasis>
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. Form elongate, oblong to broadly oval, dorsum moderately to strongly convex, densely pubescent, elytral pubescence often comprising a mixture of dark and paler hairs. Head with expanded clypeus, eyes with a distinct but narrow ocular canthus.Antennae with 11 antennomeres. Scutellar shield large, triangular and prominent. Prosternal carinae inverted Y-shaped with a short stem. Abdominal postcoxal line complete, deeply semicircular, or apically obliterated and apparently incomplete (
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<bibRefCitation id="EFA703ADFFB4FF8428E869E8FC7BFB21" author="Chen, X. & Xie, X. & Ren, S. & Wang, X." box="[717,909,1226,1253]" pageId="1" pageNumber="546" pagination="105 - 123" refId="ref4677" refString="Chen, X., Xie, X., Ren, S. & Wang, X. (2016) A taxonomic review of the genus Horniolus Weise from China, with description of a new species (Coleoptera, Coccinellidae). ZooKeys, 623, 105 - 123. https: // doi. org / 10.3897 / zookeys. 623.10191" type="journal article" year="2016">
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Chen
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<emphasis id="B942A24EFFB4FF84293469E9FCBFFB20" box="[785,841,1226,1252]" italics="true" pageId="1" pageNumber="546">et al.</emphasis>
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2016
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</bibRefCitation>
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). Tarsal formula 4-4-4. Female genitalia with a complex, coiled spermatheca, coxites elongate triangular.
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</paragraph>
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<paragraph id="8B897E5CFFB4FF842AE26830FC52FA5C" blockId="1.[151,1437,1081,1577]" pageId="1" pageNumber="546">
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<emphasis id="B942A24EFFB4FF842AE26830FEF0FAE8" bold="true" box="[199,262,1298,1324]" pageId="1" pageNumber="546">Note.</emphasis>
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Many species of this genus have a similar external appearance with the ground colour black to dark pitchy brown with yellowish-orange elytral maculae, often in the form of four orange yellow elytral spots of various sizes / shapes. This genus is generally poorly represented in institutional collections and due to its general resemblance to other
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<taxonomicName id="4C3605DFFFB4FF842AFD685CFEB0FA5C" box="[216,326,1406,1432]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="tribe" tribe="Scymnini">Scymnini</taxonomicName>
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, it is likely to be difficult to spot in mixed collections.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C2DD7FFB4FF842AE26883FBAFF9ED" pageId="1" pageNumber="546" type="biology_ecology">
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<paragraph id="8B897E5CFFB4FF842AE26883FBAFF9ED" blockId="1.[151,1437,1081,1577]" pageId="1" pageNumber="546">
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<emphasis id="B942A24EFFB4FF842AE26883FE8DFA78" bold="true" box="[199,379,1441,1468]" pageId="1" pageNumber="546">Biology/ Hosts.</emphasis>
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Members of
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<taxonomicName id="4C3605DFFFB4FF8428376880FD72FA78" authorityName="Weise" authorityYear="1901" box="[530,644,1442,1468]" class="Insecta" family="Coccinellidae" genus="Horniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="genus">
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<emphasis id="B942A24EFFB4FF8428376880FD72FA78" box="[530,644,1442,1468]" italics="true" pageId="1" pageNumber="546">Horniolus</emphasis>
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</taxonomicName>
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mostly prey on mealybugs, such as
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<taxonomicName id="4C3605DFFFB4FF842E3B6880FA6BFA79" authority="(Cockerell)" baseAuthorityName="Cockerell" baseAuthorityYear="1905" box="[1054,1437,1442,1469]" class="Insecta" family="Pseudococcidae" genus="Planococcus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="species" species="lilacinus">
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<emphasis id="B942A24EFFB4FF842E3B6880FAE3FA78" box="[1054,1301,1442,1468]" italics="true" pageId="1" pageNumber="546">Planococcus lilacinus</emphasis>
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(Cockerell)
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</taxonomicName>
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and
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<taxonomicName id="4C3605DFFFB4FF842AEF68E5FDA7FA25" authority="(Cockerell)" baseAuthorityName="Cockerell" baseAuthorityYear="1893" box="[202,593,1478,1505]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="species" species="brevipes">
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<emphasis id="B942A24EFFB4FF842AEF68E5FE30FA24" box="[202,454,1478,1504]" italics="true" pageId="1" pageNumber="546">Dysmicoccus brevipes</emphasis>
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(Cockerell)
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</taxonomicName>
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(
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<taxonomicName id="4C3605DFFFB4FF84284768E4FD28FA24" box="[610,734,1478,1504]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
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:
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<taxonomicName id="4C3605DFFFB4FF8428C968E4FC52FA25" box="[748,932,1478,1505]" class="Insecta" family="Pseudococcidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="family">Pseudococcidae</taxonomicName>
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), infesting coffee, tea and pineapple. These beetles also feed on spiralling whitefly,
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<taxonomicName id="4C3605DFFFB4FF84284268C8FC4BF9C0" authority="Russell" authorityName="Russell" authorityYear="1965" box="[615,957,1514,1540]" class="Insecta" family="Aleyrodidae" genus="Aleurodicus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="species" species="dispersus">
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<emphasis id="B942A24EFFB4FF84284268C8FC97F9C0" box="[615,865,1514,1540]" italics="true" pageId="1" pageNumber="546">Aleurodicus dispersus</emphasis>
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Russell
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</taxonomicName>
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(
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<taxonomicName id="4C3605DFFFB4FF8429EA68C8FBBDF9C0" box="[975,1099,1514,1540]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
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:
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<taxonomicName id="4C3605DFFFB4FF842E7268C8FB15F9C1" authorityName="Westwood" authorityYear="1840" box="[1111,1251,1514,1541]" class="Insecta" family="Aleyrodidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="family">Aleyrodidae</taxonomicName>
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) (
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<bibRefCitation id="EFA703ADFFB4FF842ED868C8FF24F9EC" author="Irulandi, S. & Kumar, P. K. V. & Seetharama, H. G. & Sreedharan, K." pageId="1" pageNumber="546" pagination="18 - 24" refId="ref4843" refString="Irulandi, S., Kumar, P. K. V., Seetharama, H. G. & Sreedharan, K. (2001) Biology of Horniolus vietnamicus, a newly recorded coccinellid predator of the coffee mealybug, Planococcus lilacinus Cockerell. Journal of Coffee Research, 29 (1 / 2), 18 - 24." type="journal article" year="2001">
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Irulandi
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<emphasis id="B942A24EFFB4FF842F4468C9FA63F9C0" box="[1377,1429,1514,1540]" italics="true" pageId="1" pageNumber="546">et al</emphasis>
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. 2001
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</bibRefCitation>
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;
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<bibRefCitation id="EFA703ADFFB4FF842AFB6B2CFE41F9EC" author="Ramani, S. & Poorani, J. & Bhumannavar, B. S." box="[222,439,1550,1576]" pageId="1" pageNumber="546" pagination="55 - 62" refId="ref5378" refString="Ramani, S., Poorani, J. & Bhumannavar, B. S. (2002) Spiralling whitefly, Aleurodicus dispersus Russell, in India. Biocontrol News and Information, 23 (2), 55 - 62." type="journal article" year="2002">
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Ramani
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<emphasis id="B942A24EFFB4FF842B196B2DFE9BF9EC" box="[316,365,1550,1576]" italics="true" pageId="1" pageNumber="546">et al</emphasis>
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. 2002
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</bibRefCitation>
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;
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<bibRefCitation id="EFA703ADFFB4FF842BE76B2CFD95F9EC" author="He, Y. B. & Xu, Z. F. & Zhan, R. L. & Liu, Y. H. & Sun, G. M. & Zhao, Y. L." box="[450,611,1550,1576]" pageId="1" pageNumber="546" pagination="473 - 477" refId="ref4761" refString="He, Y. B., Xu, Z. F., Zhan, R. L., Liu, Y. H., Sun, G. M. & Zhao, Y. L. (2013) Survey of beneficial organisms and predation of Horniolus hisamatsui Miyatake on Dysmicoccus brevipes (Cockerell). Journal of Environmental Entomology, 35 (4), 473 - 477. [in Chinese with English summary]" type="journal article" year="2013">
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He
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<emphasis id="B942A24EFFB4FF842BCC6B2DFDEDF9EC" box="[489,539,1550,1576]" italics="true" pageId="1" pageNumber="546">et al</emphasis>
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. 2013
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</bibRefCitation>
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;
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<bibRefCitation id="EFA703ADFFB4FF84284A6B2CFCFEF9ED" author="Poorani, J." box="[623,776,1550,1577]" pageId="1" pageNumber="546" pagination="5296" refId="ref5332" refString="Poorani, J. (2015) Two new species of Scymnini (Coleoptera: Coccinellidae) from Karnataka, India. Biodiversity Data Journal, 3, e 5296. https: // doi. org / 10.3897 / BDJ. 3. e 5296" type="journal article" year="2015">Poorani 2015</bibRefCitation>
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;
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<bibRefCitation id="EFA703ADFFB4FF8429316B2CFC27F9EC" author="Chen, X. & Xie, X. & Ren, S. & Wang, X." box="[788,977,1550,1576]" pageId="1" pageNumber="546" pagination="105 - 123" refId="ref4677" refString="Chen, X., Xie, X., Ren, S. & Wang, X. (2016) A taxonomic review of the genus Horniolus Weise from China, with description of a new species (Coleoptera, Coccinellidae). ZooKeys, 623, 105 - 123. https: // doi. org / 10.3897 / zookeys. 623.10191" type="journal article" year="2016">
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Chen
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<emphasis id="B942A24EFFB4FF8429736B2DFC71F9EC" box="[854,903,1550,1576]" italics="true" pageId="1" pageNumber="546">et al</emphasis>
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. 2016
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</bibRefCitation>
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; label data).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |