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<mods:title id="AECEB14A30F1FE3787535F02FFCB0B65">New taxa and taxonomic revisions to the Poraniidae (Valvatacea; Asteroidea) with Comments on Feeding Biology</mods:title>
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<mods:namePart id="91A3C3C17DA94B50D365911426B10AAA">Mah, Christopher L.</mods:namePart>
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<treatment id="039F453C95518724FF0BFEFEFC44F8E5" ID-DOI="http://doi.org/10.5281/zenodo.6133825" ID-GBIF-Taxon="119426365" ID-Zenodo-Dep="6133825" LSID="urn:lsid:plazi:treatment:039F453C95518724FF0BFEFEFC44F8E5" httpUri="http://treatment.plazi.org/id/039F453C95518724FF0BFEFEFC44F8E5" lastPageNumber="334" pageId="7" pageNumber="334">
<subSubSection id="C32CA7A195518724FF0BFEFEFD9CFEC5" box="[151,575,295,322]" pageId="7" pageNumber="334" type="nomenclature">
<paragraph id="8B89F42A95518724FF0BFEFEFD9CFEC5" blockId="7.[151,575,295,322]" box="[151,575,295,322]" pageId="7" pageNumber="334">
<heading id="D0C1434695518724FF0BFEFEFD9CFEC5" bold="true" box="[151,575,295,322]" fontSize="11" level="1" pageId="7" pageNumber="334" reason="1">
<taxonomicName id="4C368FA995518724FF0BFEFEFD9CFEC5" ID-CoL="69ZQ5" authority="Verrill 1878" authorityName="Verrill" authorityYear="1878" box="[151,575,295,322]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis">
<emphasis id="B942283895518724FF0BFEFEFD9CFEC5" bold="true" box="[151,575,295,322]" pageId="7" pageNumber="334">
<emphasis id="B942283895518724FF0BFEFEFE32FEC6" bold="true" box="[151,401,295,321]" italics="true" pageId="7" pageNumber="334">Chondraster grandis</emphasis>
(
<bibRefCitation id="EFA789DB95518724FE3CFEF1FD94FEC5" author="Verrill" box="[416,567,296,322]" pageId="7" pageNumber="334" refString="Verrill, A. E. (1878) Notice of recent additions to the marine fauna of the eastern coast of North America. 1 &amp; 2. American Journal of Science, 16, 207 - 215, 371 - 378. http: // dx. doi. org / 10.2475 / ajs. s 3 - 16.93.207" type="journal article" year="1878">Verrill 1878</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32CA7A195518724FF0BFEA9FBF2F90D" pageId="7" pageNumber="334" type="reference_group">
<paragraph id="8B89F42A95518724FF0BFEA9FDA0FE24" blockId="7.[151,1436,366,634]" pageId="7" pageNumber="334">
<treatmentCitationGroup id="AB26D30495518724FF0BFEA9FDA0FE24" infered="true" pageId="7" pageNumber="334">
<taxonomicName id="4C368FA995518724FF0BFEA9FDA5FE03" ID-CoL="4LVVF" authority="Verrill 1878: 371" authorityName="Verrill" authorityPageNumber="371" authorityYear="1878" box="[151,518,366,389]" class="Asteroidea" family="Poraniidae" genus="Porania" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis">
<emphasis id="B942283895518724FF0BFEA9FEE1FE03" box="[151,322,366,389]" italics="true" pageId="7" pageNumber="334">Porania grandis</emphasis>
<treatmentCitation id="0A97D23B95518724FED1FEB7FDA5FE03" author="Verrill" box="[333,518,366,388]" page="371" pageId="7" pageNumber="334" year="1878">
<bibRefCitation id="EFA789DB95518724FED1FEB7FE6DFE03" author="Verrill" box="[333,462,366,388]" pageId="7" pageNumber="334" refString="Verrill, A. E. (1878) Notice of recent additions to the marine fauna of the eastern coast of North America. 1 &amp; 2. American Journal of Science, 16, 207 - 215, 371 - 378. http: // dx. doi. org / 10.2475 / ajs. s 3 - 16.93.207" type="journal article" year="1878">Verrill 1878</bibRefCitation>
: 371
</treatmentCitation>
</taxonomicName>
; 1882: 218; 1884: 652; 1885: 542 (pt.), pl. 15, fig. 44, pl. 16, fig. 44a (non figs 45, 45a=
<taxonomicName id="4C368FA995518724FF6BFE54FE5FFE24" ID-CoL="7HV2B" box="[247,508,397,419]" class="Asteroidea" family="Poraniidae" genus="Porania" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="subSpecies" species="pulvillus" subSpecies="insignis">Porania pulvillus insignis</taxonomicName>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89F42A95518724FF0BFE75FC1EFE67" blockId="7.[151,1436,366,634]" pageId="7" pageNumber="334">
(possible synonym)
<taxonomicName id="4C368FA995518724FEFAFE75FCE2FE45" ID-CoL="7CYFW" authority="Danielssen &amp; Koren 1881: 186" authorityName="Danielssen &amp; Koren" authorityPageNumber="186" authorityYear="1881" box="[358,833,428,450]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="willei">
<emphasis id="B942283895518724FEFAFE75FE54FE45" box="[358,503,428,450]" italics="true" pageId="7" pageNumber="334">Tylaster willei</emphasis>
<bibRefCitation id="EFA789DB95518724FE63FE75FCAEFE45" author="Danielssen" box="[511,781,428,450]" pageId="7" pageNumber="334" refString="Danielssen, D. C. &amp; Koren, J. (1881) Fra den norske Nordhavsexpedition, Echinodermer. Nyt Magazin for Naturvidenskaberne, 26, 177 - 195." type="journal article" year="1881">Danielssen &amp; Koren 1881</bibRefCitation>
: 186
</taxonomicName>
; 1884: 64;
<bibRefCitation id="EFA789DB95518724FC2AFE75FBE1FE45" author="Ludwig" box="[950,1090,428,450]" pageId="7" pageNumber="334" refString="Ludwig, H. (1900) Arktische Seesterne. Fauna Arctica, 1 (3), 447 - 502." type="journal article" year="1900">Ludwig 1900</bibRefCitation>
: 459;
<bibRefCitation id="EFA789DB95518724FB1EFE75FA80FE45" author="Ostergren" box="[1154,1315,428,450]" pageId="7" pageNumber="334" refString="Ostergren, H. (1901) Uber die von der Schwedischen Zoologischen Polarexpedition 1900 eingeselten, Asteroidea, Echinoidea, Holothuroidea und Crinoidea. Zoologische Anzeiger, 24, 252 - 232." type="journal article" year="1901">Ostergren 1901</bibRefCitation>
: 252;
<bibRefCitation id="EFA789DB95518724FAFEFE75FF5EFE67" author="Grieg" pageId="7" pageNumber="334" refString="Grieg, J. A. (1907) Echinodermen von dem norwegischen Fischereidampfer Michael Sars in den Jahren 1900 - 1903 gesammelt. 3. Asteroidea. 1. Bergens Museums. Skrifter, 1906 (13), 1 - 87." type="journal article" year="1907">Grieg 1907</bibRefCitation>
: 32;
<bibRefCitation id="EFA789DB95518724FEB1FE13FE6DFE67" author="Djakonov" box="[301,462,458,480]" pageId="7" pageNumber="334" refString="Djakonov, A. M. (1933) Les Echinodermes des mers arctiques. Izdatel'stvo Akademii Nauk 8, 1 - 166." type="journal article" year="1933">Djakonov 1933</bibRefCitation>
: 44; 1950: 60 (1968: 51);
<bibRefCitation id="EFA789DB95518724FD49FE12FC25FE67" author="Clark" box="[725,902,458,480]" pageId="7" pageNumber="334" refString="Clark, A. M. (1993) An index of names of recent Asteroidea - Part 2, Valvatida. Echinoderm Studies, 4, 187 - 366." type="journal article" year="1993">A.M. Clark 1993</bibRefCitation>
: 238.
</paragraph>
<paragraph id="8B89F42A95518724FF0BFE30FD5BFE78" blockId="7.[151,1436,366,634]" box="[151,760,489,511]" pageId="7" pageNumber="334">
<treatmentCitationGroup id="AB26D30495518724FF0BFE30FD5BFE78" box="[151,760,489,511]" pageId="7" pageNumber="334">
?
<taxonomicName id="4C368FA995518724FF3EFE33FDEBFE78" ID-CoL="3Y56H" authority="Sladen 1889: 365" authorityName="Sladen" authorityPageNumber="365" authorityYear="1889" box="[162,584,489,511]" class="Asteroidea" family="Poraniidae" genus="Marginaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="fimbriatus">
<emphasis id="B942283895518724FF3EFE33FE2DFE78" box="[162,398,489,511]" italics="true" pageId="7" pageNumber="334">Marginaster fimbriatus</emphasis>
<treatmentCitation id="0A97D23B95518724FE08FE30FDEBFE78" author="Sladen" box="[404,584,489,511]" page="365" pageId="7" pageNumber="334" year="1889">
<bibRefCitation id="EFA789DB95518724FE08FE30FDB5FE78" author="Sladen" box="[404,534,489,511]" pageId="7" pageNumber="334" refString="Sladen, W. P. (1889) Asteroidea. Report of the Scientific Results of H. M. S. Challenger, 30, 1 - 893." type="journal article" year="1889">Sladen 1889</bibRefCitation>
: 365
</treatmentCitation>
</taxonomicName>
, pl. 58, figs. 46.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89F42A95518724FF0BFDD1FDA8FD99" blockId="7.[151,1436,366,634]" box="[151,523,520,542]" pageId="7" pageNumber="334">
<treatmentCitationGroup id="AB26D30495518724FF0BFDD1FDA8FD99" box="[151,523,520,542]" pageId="7" pageNumber="334">
?
<taxonomicName id="4C368FA995518724FF3EFDD1FDA8FD99" ID-CoL="TRQC" authority="Bell 1893: 81" authorityName="Bell" authorityPageNumber="81" authorityYear="1893" box="[162,523,520,542]" class="Asteroidea" family="Poraniidae" genus="Cheilaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="fimbriatus">
<emphasis id="B942283895518724FF3EFDD1FED9FD99" box="[162,378,520,542]" italics="true" pageId="7" pageNumber="334">Cheilaster fimbriatus</emphasis>
<treatmentCitation id="0A97D23B95518724FE1CFDD1FDA8FD99" author="Bell" box="[384,523,520,542]" page="81" pageId="7" pageNumber="334" year="1893">
<bibRefCitation id="EFA789DB95518724FE1CFDD1FE4BFD99" author="Bell" box="[384,488,520,542]" pageId="7" pageNumber="334" refString="Bell, F. J. (1893) Catalogue of the British echinoderms in the British Museum (Natural History). London, xvii + 202 pp., 16 pls." type="book" year="1893">Bell 1893</bibRefCitation>
: 81
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89F42A95518724FF0BFDF1FD2BFDBB" blockId="7.[151,1436,366,634]" box="[151,648,550,573]" pageId="7" pageNumber="334">
<treatmentCitationGroup id="AB26D30495518724FF0BFDF1FD2BFDBB" box="[151,648,550,573]" pageId="7" pageNumber="334">
<taxonomicName id="4C368FA995518724FF0BFDF1FD2BFDBB" ID-CoL="4LVVF" authority="Verrill 1895: 137" authorityName="Verrill" authorityPageNumber="137" authorityYear="1895" box="[151,648,550,573]" class="Asteroidea" family="Poraniidae" genus="Porania" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis" subGenus="Chondraster">
<emphasis id="B942283895518724FF0BFDF1FE77FDBB" box="[151,468,550,573]" italics="true" pageId="7" pageNumber="334">Porania (Chondraster) grandis</emphasis>
<treatmentCitation id="0A97D23B95518724FE47FDFFFD2BFDBB" author="Verrill" box="[475,648,550,572]" page="137" pageId="7" pageNumber="334" year="1895">
<bibRefCitation id="EFA789DB95518724FE47FDFFFDFBFDBB" author="Verrill" box="[475,600,550,572]" pageId="7" pageNumber="334" refString="Verrill, A. E. (1895) Distribution of the echinoderms of north - eastern America. American Journal of Science, 49, 127 - 141." type="journal article" year="1895">Verrill 1895</bibRefCitation>
: 137
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89F42A95518724FF0BFD9CFBC6FDDC" blockId="7.[151,1436,366,634]" box="[151,1125,581,603]" pageId="7" pageNumber="334">
<treatmentCitationGroup id="AB26D30495518724FF0BFD9CFBC6FDDC" box="[151,1125,581,603]" pageId="7" pageNumber="334">
<taxonomicName id="4C368FA995518724FF0BFD9CFE99FDDC" box="[151,314,581,603]" class="Asteroidea" family="Oreasteridae" genus="Culcita" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="borealis">
<emphasis id="B942283895518724FF0BFD9CFE99FDDC" box="[151,314,581,603]" italics="true" pageId="7" pageNumber="334">Culcita borealis</emphasis>
</taxonomicName>
(pt.)
<treatmentCitation id="0A97D23B95518724FEEDFD9FFDB0FDDC" author="Farran" box="[369,531,581,603]" page="15" pageId="7" pageNumber="334" year="1913">
<bibRefCitation id="EFA789DB95518724FEEDFD9FFE53FDDC" author="Farran" box="[369,496,581,603]" pageId="7" pageNumber="334" refString="Farran, G. P. (1913) The deep water Asteroidea, Ophiuroidea, and Echinoidea of the west coast of Ireland. Scientific Investigations of the Fish Board of Ireland, 1912 (6), 1 - 66." type="journal article" year="1913">Farran 1913</bibRefCitation>
: 15
</treatmentCitation>
(st. SR 483) [Non-C.
<taxonomicName id="4C368FA995518724FD6FFD9CFBFCFDDC" authority="Sussbach &amp; Breckner 1911" authorityName="Sussbach &amp; Breckner" authorityYear="1911" box="[755,1119,581,603]" class="Asteroidea" family="Oreasteridae" genus="Culcita" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="borealis">
borealis
<bibRefCitation id="EFA789DB95518724FCD5FD9CFBFCFDDC" author="Sussbach" box="[841,1119,581,603]" pageId="7" pageNumber="334" refString="Sussbach, S. &amp; Breckner, A. (1911) Die Seeigel, Seesterne und Schlangensterne der Nord - und Ostee. Wissenschaftliche Meereesuntersuchungen Kiel, 12, 167 - 300." type="journal article" year="1911">Süssbach &amp; Breckner 1911</bibRefCitation>
</taxonomicName>
]
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89F42A95518724FF0BFDBDFA24FDFD" blockId="7.[151,1436,366,634]" box="[151,1415,612,634]" pageId="7" pageNumber="334">
<treatmentCitationGroup id="AB26D30495518724FF0BFDBDFA24FDFD" box="[151,1415,612,634]" pageId="7" pageNumber="334">
<taxonomicName id="4C368FA995518724FF0BFDBDFDE5FDFD" ID-CoL="69ZQ5" authority="A.M. Clark 1984: 27" authorityName="A.M. Clark" authorityPageNumber="27" authorityYear="1984" box="[151,582,612,634]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis">
<emphasis id="B942283895518724FF0BFDBDFEC9FDFD" box="[151,362,612,634]" italics="true" pageId="7" pageNumber="334">Chondraster grandis</emphasis>
<treatmentCitation id="0A97D23B95518724FEECFDBDFDE5FDFD" author="Clark" box="[368,582,612,634]" page="27" pageId="7" pageNumber="334" year="1984">
<bibRefCitation id="EFA789DB95518724FEECFDBDFD82FDFD" author="Clark" box="[368,545,612,634]" pageId="7" pageNumber="334" refString="Clark, A. M. (1984) Notes on Atlantic and other Asteroidea. 4. Families Poraniidae and Asteropseidae. Bulletin of the British Museum of Natural History (Zoology), 47 (1), 19 - 51." type="journal article" year="1984">A.M. Clark 1984</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
, figs. 4A, B, 5A, 6A, B, 7d;
<treatmentCitation id="0A97D23B95518724FCFBFDBCFBF8FDFD" author="Harvey" box="[871,1115,612,634]" page="163" pageId="7" pageNumber="334" year="1988">
<bibRefCitation id="EFA789DB95518724FCFBFDBCFB8BFDFD" author="Harvey" box="[871,1064,612,634]" pageId="7" pageNumber="334" refString="Harvey, R., Gage, J. D., Billet, D. S. M., Clark, A. M. &amp; Paterson, G. L. J. (1988) Echinoderms of the Rockall Through and adjacent areas. 3. Additional records. Bulletin of the British Museum of Natural History (Zoology), 54 (4), 53 - 198." type="journal article" year="1988">
Harvey
<emphasis id="B942283895518724FC25FDBCFC4FFDFD" box="[953,1004,612,634]" italics="true" pageId="7" pageNumber="334">et al.</emphasis>
1988
</bibRefCitation>
: 163
</treatmentCitation>
;
<treatmentCitation id="0A97D23B95518724FBF9FDBDFA20FDFD" author="Clark" box="[1125,1411,612,634]" page="203" pageId="7" pageNumber="334" year="1992">
<bibRefCitation id="EFA789DB95518724FBF9FDBDFAF3FDFD" author="Clark" box="[1125,1360,612,634]" pageId="7" pageNumber="334" refString="Clark, A. M. &amp; Downey, M. E. (1992) Starfishes of the Atlantic. Chapman and Hall, London, 820 pp." type="book" year="1992">Clark &amp; Downey 1992</bibRefCitation>
: 203
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89F42A95518724FF0BFD7FFC73FD65" blockId="7.[151,1437,678,1891]" pageId="7" pageNumber="334">
<emphasis id="B942283895518724FF0BFD7FFE8AFD38" bold="true" box="[151,297,678,703]" pageId="7" pageNumber="334">Occurrence.</emphasis>
Off Cape Cod (c. 42º) S to SE of Cape May(c. 38º), Lydonia Canyon (south side of Georges Bank), Baltimore Canyon, SE of Cape Mayin the NW Atlantic.
<emphasis id="B942283895518724FC8CFD13FC6AFD64" bold="true" box="[784,969,714,739]" pageId="7" pageNumber="334">Bear Seamount</emphasis>
.
</paragraph>
<paragraph id="8B89F42A95518724FF5BFD36FCCEFCAD" blockId="7.[151,1437,678,1891]" pageId="7" pageNumber="334">
Lousy Bank, SW of the Faeroe Islands, Rockall Trough, Porcupine Seabight, and the Wyville Thomson Ridge south to the southern Bay of Biscay (c. 44ºN, 4ºW).
<quantity id="4CCE59CF95518724FD7DFCCAFCCEFCAD" box="[737,877,787,811]" metricMagnitude="3" metricUnit="m" metricValue="1.395" metricValueMax="2.4899999999999998" metricValueMin="0.3" pageId="7" pageNumber="334" unit="m" value="1395.0" valueMax="2490.0" valueMin="300.0">3002490 m</quantity>
</paragraph>
<paragraph id="8B89F42A95518724FF5BFCEFFE82FC58" blockId="7.[151,1437,678,1891]" pageId="7" pageNumber="334">
<emphasis id="B942283895518724FF5BFCEFFEF7FCC9" box="[199,340,822,847]" italics="true" pageId="7" pageNumber="334">Other Notes</emphasis>
: This species has been observed
<emphasis id="B942283895518724FD78FCEEFC8DFCC9" box="[740,814,823,846]" italics="true" pageId="7" pageNumber="334">in situ</emphasis>
in several Northeast Atlantic localities by Remotely Operated Vehicles deployed by the
<taxonomicName id="4C368FA995518724FDB7FC83FD5BFCF4" authority="Explorer" authorityName="Explorer" box="[555,760,858,883]" class="Insecta" family="Pentatomidae" genus="Okeanos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B942283895518724FDB7FC83FD5BFCF4" box="[555,760,858,883]" italics="true" pageId="7" pageNumber="334">Okeanos Explorer</emphasis>
</taxonomicName>
(pers. observations). Video observations of this species are recorded from Heezen, Hydrographer, and Nygren canyons where it has been observed on soft-sediment covered substrates in conjunction with octocorals and encrusting colonial invertebrates. It displays a purple/pink-white color in life.
</paragraph>
<paragraph id="8B89F42A95518724FF5BFC33FE62FB14" blockId="7.[151,1437,678,1891]" pageId="7" pageNumber="334">
<emphasis id="B942283895518724FF5BFC33FD40FB84" bold="true" box="[199,739,1002,1027]" pageId="7" pageNumber="334">
Synonymy of
<taxonomicName id="4C368FA995518724FEF3FC33FE4AFB84" ID-CoL="69ZQ5" box="[367,489,1002,1027]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis">
<emphasis id="B942283895518724FEF3FC33FE4AFB84" bold="true" box="[367,489,1002,1027]" italics="true" pageId="7" pageNumber="334">C. grandis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C368FA995518724FDADFC33FD76FB84" ID-CoL="7CYFW" box="[561,725,1002,1027]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="willei">
<emphasis id="B942283895518724FDADFC33FD76FB84" bold="true" box="[561,725,1002,1027]" italics="true" pageId="7" pageNumber="334">Tylaster willei</emphasis>
</taxonomicName>
?
</emphasis>
<taxonomicName id="4C368FA995518724FD70FC33FB77FB84" ID-CoL="7CYFW" authority="Danielssen and Koren 1881" authorityName="Danielssen and Koren" authorityYear="1881" box="[748,1236,1002,1027]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="willei">
<emphasis id="B942283895518724FD70FC33FC2EFB84" box="[748,909,1002,1027]" italics="true" pageId="7" pageNumber="334">Tylaster willei</emphasis>
<bibRefCitation id="EFA789DB95518724FC0BFC33FB77FB84" author="Danielssen" box="[919,1236,1002,1027]" pageId="7" pageNumber="334" refString="Danielssen, D. C. &amp; Koren, J. (1881) Fra den norske Nordhavsexpedition, Echinodermer. Nyt Magazin for Naturvidenskaberne, 26, 177 - 195." type="journal article" year="1881">Danielssen and Koren 1881</bibRefCitation>
</taxonomicName>
has been seldom recorded since its original description, and in spite of its occurrence in the Norwegian Sea and adjacent regions it is conspicuously absent from Clark and Downeys (1992) comprehensive monograph of Atlantic asteroids. Subsequent accounts [e.g.,
<bibRefCitation id="EFA789DB95518724FE43FB8FFD36FBE8" author="Djakonov" box="[479,661,1110,1135]" pageId="7" pageNumber="334" refString="Djakonov, A. M. (1950) Morskie Zvezdy Morei SSSR. Tableaux analytiquesdelafaune deU. R. S. S., 34, 1 - 203. [Translated as Dyakonov. A. M. 1968. Sea stars (Asteroids) of the USSR Seas. Keys to the Fauna of the USSR, 34. Zoological Institute of the Academy of Sciences of the USSR, ed. by A. A. Strelkov. Israel Program for scientific translations Ltd. Jerusalem, 183 pp.]" type="book chapter" year="1950">Djakonov 1950</bibRefCitation>
(1968)] have done little to elaborate on the poorly understood description of this species.
</paragraph>
<paragraph id="8B89F42A95518724FF5BFB46FBF2F90D" blockId="7.[151,1437,678,1891]" pageId="7" pageNumber="334">
A review of the original description and comparison with further specimens of
<taxonomicName id="4C368FA995518724FBABFB47FA83FB30" ID-CoL="69ZQ5" box="[1079,1312,1182,1207]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis">
<emphasis id="B942283895518724FBABFB47FA83FB30" box="[1079,1312,1182,1207]" italics="true" pageId="7" pageNumber="334">Chondraster grandis</emphasis>
</taxonomicName>
suggests it is consistent with the original description and figures as illustrated by
<bibRefCitation id="EFA789DB95518724FC27FB1BFAAAFB5C" author="Danielssen" box="[955,1289,1218,1243]" pageId="7" pageNumber="334" refString="Danielssen, D. C. &amp; Koren, J. (1881) Fra den norske Nordhavsexpedition, Echinodermer. Nyt Magazin for Naturvidenskaberne, 26, 177 - 195." type="journal article" year="1881">Danielssen and Koren (1881)</bibRefCitation>
with several characters shared between
<taxonomicName id="4C368FA995518724FE4CFB3EFDE9FB78" ID-CoL="69ZQ5" box="[464,586,1254,1279]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="grandis">
<emphasis id="B942283895518724FE4CFB3EFDE9FB78" box="[464,586,1254,1279]" italics="true" pageId="7" pageNumber="334">C. grandis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C368FA995518724FD15FB3EFD44FB78" ID-CoL="7CYFW" box="[649,743,1254,1279]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="willei">
<emphasis id="B942283895518724FD15FB3EFD44FB78" box="[649,743,1254,1279]" italics="true" pageId="7" pageNumber="334">T. willei</emphasis>
</taxonomicName>
. This includes the poorly decalcified abactinal surface, the distinct rows of actinal intermediate plates and tissue gaps between actinal plate rows, the fine spination present on the curved actinal series and a similar number of furrow spines present in each.
<bibRefCitation id="EFA789DB95518724FBD4FAF7FA3FFAC0" author="Danielssen" box="[1096,1436,1326,1351]" pageId="7" pageNumber="334" refString="Danielssen, D. C. &amp; Koren, J. (1881) Fra den norske Nordhavsexpedition, Echinodermer. Nyt Magazin for Naturvidenskaberne, 26, 177 - 195." type="journal article" year="1881">Danielssen and Koren (1881)</bibRefCitation>
describe 3 adambulacral spines for a specimen with R=4.0 cm. A comparably sized specimen (USNM
<accessionNumber id="946569C995518724FAA0FA8BFA36FAEC" box="[1340,1429,1362,1387]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E30694" pageId="7" pageNumber="334" type="EnaNcbi">E30694</accessionNumber>
) shows variable furrow spine number (2 or 3 furrow spines). It is also possible that Danielssen and Koren may have mistaken the subambulacral spines, of which there are three, for the furrow spines. Character differences between
<taxonomicName id="4C368FA995518724FF0BFA67FE86FA50" ID-CoL="7NSRH" box="[151,293,1470,1495]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="genus">
<emphasis id="B942283895518724FF0BFA67FE86FA50" box="[151,293,1470,1495]" italics="true" pageId="7" pageNumber="334">Chondraster</emphasis>
</taxonomicName>
and
<taxonomicName id="4C368FA995518724FEC6FA67FE11FA50" ID-CoL="7Q5L9" box="[346,434,1470,1495]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="genus">
<emphasis id="B942283895518724FEC6FA67FE11FA50" box="[346,434,1470,1495]" italics="true" pageId="7" pageNumber="334">Tylaster</emphasis>
</taxonomicName>
summarized by
<bibRefCitation id="EFA789DB95518724FDF7FA66FC9FFA50" author="Clark" box="[619,828,1470,1495]" pageId="7" pageNumber="334" refString="Clark, A. M. (1984) Notes on Atlantic and other Asteroidea. 4. Families Poraniidae and Asteropseidae. Bulletin of the British Museum of Natural History (Zoology), 47 (1), 19 - 51." type="journal article" year="1984">A.M. Clark (1984)</bibRefCitation>
such as the fine spinules on the “dorsal body wall” are present on the surface of
<taxonomicName id="4C368FA995518724FE33FA3BFD98FA7C" ID-CoL="7NSRH" box="[431,571,1506,1531]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="genus">
<emphasis id="B942283895518724FE33FA3BFD98FA7C" box="[431,571,1506,1531]" italics="true" pageId="7" pageNumber="334">Chondraster</emphasis>
</taxonomicName>
, but below the epidermis. It is possible that
<taxonomicName id="4C368FA995518724FBBBFA3DFB23FA7C" ID-CoL="7CYFW" box="[1063,1152,1506,1531]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="willei">
<emphasis id="B942283895518724FBBBFA3DFB23FA7C" box="[1063,1152,1506,1531]" italics="true" pageId="7" pageNumber="334">T. willei</emphasis>
</taxonomicName>
may show damage to the body wall. Fine spinelets on the actinal surface are also present in both taxa. Further specimens, including the
<typeStatus id="548D4A8895518724FAF0F9DEFA3FF998" box="[1388,1436,1543,1567]" pageId="7" pageNumber="334">type</typeStatus>
will need to be studied in order to make a conclusive synonymy. In that event,
<taxonomicName id="4C368FA995518724FB99F9F3FA3FF9C4" ID-CoL="7Q5L9" authority="Danielssen and Koren 1881" authorityName="Danielssen and Koren" authorityYear="1881" box="[1029,1436,1578,1603]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="genus">
<emphasis id="B942283895518724FB99F9F3FBFEF9C4" box="[1029,1117,1578,1603]" italics="true" pageId="7" pageNumber="334">Tylaster</emphasis>
<bibRefCitation id="EFA789DB95518724FBFAF9F3FA3FF9C4" author="Danielssen" box="[1126,1436,1578,1603]" pageId="7" pageNumber="334" refString="Danielssen, D. C. &amp; Koren, J. (1881) Fra den norske Nordhavsexpedition, Echinodermer. Nyt Magazin for Naturvidenskaberne, 26, 177 - 195." type="journal article" year="1881">Danielssen and Koren 1881</bibRefCitation>
</taxonomicName>
would be the senior name for
<taxonomicName id="4C368FA995518724FD8FF997FCE3F9E0" ID-CoL="7NSRH" authority="Verrill 1895" authorityName="Verrill" authorityYear="1895" box="[531,832,1614,1639]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="genus">
<emphasis id="B942283895518724FD8FF997FD03F9E0" box="[531,672,1614,1639]" italics="true" pageId="7" pageNumber="334">Chondraster</emphasis>
<bibRefCitation id="EFA789DB95518724FD33F997FCE3F9E0" author="Verrill" box="[687,832,1614,1639]" pageId="7" pageNumber="334" refString="Verrill, A. E. (1895) Distribution of the echinoderms of north - eastern America. American Journal of Science, 49, 127 - 141." type="journal article" year="1895">Verrill 1895</bibRefCitation>
</taxonomicName>
. For now, we retain
<taxonomicName id="4C368FA995518724FBD7F997FB7BF9E0" ID-CoL="7NSRH" box="[1099,1240,1614,1639]" class="Asteroidea" family="Poraniidae" genus="Chondraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="genus">
<emphasis id="B942283895518724FBD7F997FB7BF9E0" box="[1099,1240,1614,1639]" italics="true" pageId="7" pageNumber="334">Chondraster</emphasis>
</taxonomicName>
pending further examination of
<typeStatus id="548D4A8895518724FEDAF9AAFED5F90C" box="[326,374,1651,1675]" pageId="7" pageNumber="334">type</typeStatus>
specimens and possible suppression of the seldom used
<taxonomicName id="4C368FA995518724FC6CF9ADFBE9F90C" ID-CoL="7CYFW" box="[1008,1098,1650,1675]" class="Asteroidea" family="Poraniidae" genus="Tylaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="334" phylum="Echinodermata" rank="species" species="willei">
<emphasis id="B942283895518724FC6CF9ADFBE9F90C" box="[1008,1098,1650,1675]" italics="true" pageId="7" pageNumber="334">T. willei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CA7A195518724FF5BF94FFC44F8E5" pageId="7" pageNumber="334" type="materials_examined">
<paragraph id="8B89F42A95518724FF5BF94FFC44F8E5" blockId="7.[151,1437,678,1891]" pageId="7" pageNumber="334">
<emphasis id="B942283895518724FF5BF94FFE13F928" bold="true" box="[199,432,1686,1711]" pageId="7" pageNumber="334">Material examined.</emphasis>
<collectionCode id="ED276CEF95518724FE24F94EFDAAF928" LSID="urn:lsid:biocol.org:col:34871" box="[440,521,1687,1711]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="334">USNM</collectionCode>
<accessionNumber id="946569C995518724FD8DF94FFDCBF928" box="[529,616,1686,1711]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E30694" pageId="7" pageNumber="334" type="EnaNcbi">E30694</accessionNumber>
Lydonia Canyon, North Atlantic
<geoCoordinate id="EE0292ED95518724FC7FF94EFBCFF928" box="[995,1132,1687,1711]" direction="north" orientation="latitude" pageId="7" pageNumber="334" precision="15" value="40.36361">40°2149”N</geoCoordinate>
<geoCoordinate id="EE0292ED95518724FBE8F94FFAA0F928" box="[1140,1283,1686,1711]" direction="west" orientation="longitude" pageId="7" pageNumber="334" precision="15" value="-67.662224">67°3944”W</geoCoordinate>
,
<quantity id="4CCE59CF95518724FA93F94FFAFAF929" box="[1295,1369,1686,1711]" metricMagnitude="2" metricUnit="m" metricValue="6.92" pageId="7" pageNumber="334" unit="m" value="692.0">692 m</quantity>
, Coll. BLM submersible
<emphasis id="B942283895518724FEE6F963FE16F954" box="[378,437,1722,1747]" italics="true" pageId="7" pageNumber="334">Alvin</emphasis>
. (1 dry spec. R=5.6, r=3.2);
<collectionCode id="ED276CEF95518724FCB5F962FCD9F954" LSID="urn:lsid:biocol.org:col:34871" box="[809,890,1723,1747]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="334">USNM</collectionCode>
<accessionNumber id="946569C995518724FC15F963FC43F954" box="[905,992,1722,1747]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E50308" pageId="7" pageNumber="334" type="EnaNcbi">E50308</accessionNumber>
NE of Virginia Beach,
<geoCoordinate id="EE0292ED95518724FA8FF963FA3FF954" box="[1299,1436,1722,1747]" direction="north" orientation="latitude" pageId="7" pageNumber="334" precision="15" value="36.995">36°5942”N</geoCoordinate>
74°3330W to
<geoCoordinate id="EE0292ED95518724FEA3F907FE73F970" box="[319,464,1758,1783]" direction="north" orientation="latitude" pageId="7" pageNumber="334" precision="15" value="36.963333">36°57 48”N</geoCoordinate>
<geoCoordinate id="EE0292ED95518724FE44F906FDC4F970" box="[472,615,1759,1783]" direction="west" orientation="longitude" pageId="7" pageNumber="334" precision="15" value="-74.57166">74°3418”W</geoCoordinate>
,
<quantity id="4CCE59CF95518724FDE8F907FD55F971" box="[628,758,1758,1783]" metricMagnitude="2" metricUnit="m" metricValue="7.565" metricValueMax="8.64" metricValueMin="6.49" pageId="7" pageNumber="334" unit="m" value="756.5" valueMax="864.0" valueMin="649.0">649864 m</quantity>
, coll. R/V Gilliss (2 dry specs. R=5.1, r=3.3; R=5.8, r=4.8);
<collectionCode id="ED276CEF95518724FF0BF8DAFF4BF89C" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,1795,1819]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="334">USNM</collectionCode>
<accessionNumber id="946569C995518724FF72F8DBFEE6F89C" box="[238,325,1794,1819]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E50540" pageId="7" pageNumber="334" type="EnaNcbi">E50540</accessionNumber>
Baltimore Canyon,
<geoCoordinate id="EE0292ED95518724FDBBF8DAFD01F89C" box="[551,674,1795,1819]" direction="north" orientation="latitude" pageId="7" pageNumber="334" precision="15" value="38.155556">38°920”N</geoCoordinate>
<geoCoordinate id="EE0292ED95518724FD34F8DBFC9BF89C" box="[680,824,1794,1819]" direction="west" orientation="longitude" pageId="7" pageNumber="334" precision="15" value="-73.84139">73°5029”W</geoCoordinate>
,
<quantity id="4CCE59CF95518724FCDFF8DBFC28F89D" box="[835,907,1794,1819]" metricMagnitude="2" metricUnit="m" metricValue="6.21" pageId="7" pageNumber="334" unit="m" value="621.0">621 m</quantity>
, Coll. DSR/V Alvin (1 dry spec. R=7.9, r=4.2);
<collectionCode id="ED276CEF95518724FF0BF8FEFF4BF8B8" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,1831,1855]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="334">USNM</collectionCode>
1016057 Bear Seamount, 39.94139, -67.38908,
<quantity id="4CCE59CF95518724FC83F8FEFC1CF8B9" box="[799,959,1831,1855]" metricMagnitude="3" metricUnit="m" metricValue="1.51" metricValueMax="1.888" metricValueMin="1.132" pageId="7" pageNumber="334" unit="m" value="1510.0" valueMax="1888.0" valueMin="1132.0">11321888 m</quantity>
, coll. R/V Delaware
<collectionCode id="ED276CEF95518724FB26F8FEFB6CF8B9" box="[1210,1231,1831,1854]" pageId="7" pageNumber="334">II</collectionCode>
;
<collectionCode id="ED276CEF95518724FB43F8FEFA93F8B8" LSID="urn:lsid:biocol.org:col:34871" box="[1247,1328,1831,1855]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="334">USNM</collectionCode>
1016073 Bear Seamount 39.93442, -67.39419,
<quantity id="4CCE59CF95518724FDDDF893FD78F8E5" box="[577,731,1866,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.326" metricValueMax="1.541" metricValueMin="1.111" pageId="7" pageNumber="334" unit="m" value="1326.0" valueMax="1541.0" valueMin="1111.0">11111541 m</quantity>
, Coll. R/V Delaware
<collectionCode id="ED276CEF95518724FC51F892FC40F8E5" box="[973,995,1867,1890]" pageId="7" pageNumber="334">II</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>