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<document id="2FA84674DB2119CC40D7E7A3DF870AAB" ID-CLB-Dataset="32767" ID-DOI="10.11646/zootaxa.4286.1.1" ID-GBIF-Dataset="a614f1d9-bda9-4df0-a486-aef5af7813fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="828206" ID-ZooBank="6A22EB50-0146-465A-8E2A-523E2F8FE470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108241184" checkinUser="plazi" docAuthor="Galvis, William" docDate="2017" docId="0386A719572C9075FF21FF58FB3DFAD8" docLanguage="en" docName="zootaxa.4286.1.1.pdf" docOrigin="Zootaxa 4286 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus chie Galvis, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FFBFDF6157289070FFB6FFCFFF81FFE8" masterDocTitle="Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation" masterLastPageNumber="40" masterPageNumber="1" pageNumber="5" updateTime="1698465056562" updateUser="ExternalLinkService">
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<mods:title id="AD74B1B7419210AD57AF0B3D79C5F44C">Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation</mods:title>
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<mods:namePart id="4D078992913C8320BA383C38D1CFEAFF">Galvis, William</mods:namePart>
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<mods:date id="A100088AE2F641F4D823A35C9305BE8D">2017</mods:date>
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<treatment id="0386A719572C9075FF21FF58FB3DFAD8" ID-DOI="http://doi.org/10.5281/zenodo.6045627" ID-GBIF-Taxon="132548509" ID-Zenodo-Dep="6045627" LSID="urn:lsid:plazi:treatment:0386A719572C9075FF21FF58FB3DFAD8" httpUri="http://treatment.plazi.org/id/0386A719572C9075FF21FF58FB3DFAD8" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C3354584572C9074FF21FF58FDC2FF3B" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B90160F572C9074FF21FF58FE4AFF5A" blockId="4.[151,579,151,211]" box="[151,459,151,178]" pageId="4" pageNumber="5">
<heading id="D0D8A163572C9074FF21FF58FE4AFF5A" bold="true" box="[151,459,151,178]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B95BCA1D572C9074FF21FF58FE4AFF5A" bold="true" box="[151,459,151,178]" pageId="4" pageNumber="5">
<taxonomicName id="4C2F6D8C572C9074FF21FF58FEE7FF59" ID-CoL="D2G4" box="[151,358,151,177]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">
<emphasis id="B95BCA1D572C9074FF21FF58FEE7FF59" bold="true" box="[151,358,151,177]" italics="true" pageId="4" pageNumber="5">Amphidraus chie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766572C9074FED8FF57FE4AFF5A" box="[366,459,152,178]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B90160F572C9074FF21FF75FDC2FF3B" blockId="4.[151,579,151,211]" box="[151,579,186,211]" pageId="4" pageNumber="5">
<figureCitation id="13140A8A572C9074FF21FF75FE89FF3B" box="[151,264,186,211]" captionStart="FIGURES 2" captionStartId="4.[151,264,1909,1931]" captionTargetBox="[193,1411,455,1885]" captionTargetId="figure@4.[173,1417,443,1894]" captionTargetPageId="4" captionText="FIGURES 2 a d. A. chie sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: bwe = basal widening of the embolus, cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm (a b), 0.10 mm (c d)." httpUri="https://zenodo.org/record/828210/files/figure.png" pageId="4" pageNumber="5">Figs 2ad</figureCitation>
,
<figureCitation id="13140A8A572C9074FEA5FF75FEDBFF3B" box="[275,346,186,211]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a i. A. bochica sp. nov., a) male holotype (ICN Ar 8276); b) female paratype (ICN Ar 8269). A. chie sp. nov., c) male holotype (ICN Ar 8273); d) female paratype (ICN Ar 8274). A. colombianus sp. nov., e) male holotype (ICN Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN Ar 8257); g) female paratype (ANDES IN 3871). A. mae sp. nov., h) male holotype (ICN Ar 8259); i) female paratype (ICN Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="4" pageNumber="5">16cd</figureCitation>
,
<figureCitation id="13140A8A572C9074FED3FF75FE2DFF3B" box="[357,428,186,211]" captionStart="FIGURES 20" captionStartId="31.[151,264,1906,1928]" captionTargetBox="[248,1341,212,1891]" captionTargetId="figure@31.[228,1365,193,1894]" captionTargetPageId="31" captionText="FIGURES 20 a k. Amphidraus spp, left male palp. A. bochica sp. nov., a) ventral view; b) same, retrolateral view. A. chie sp. nov., c) ventral view; d) same, retrolateral view. A. colombianus sp. nov., e) ventral view; f) same, retroventral view; g) same, retrolateral view. A. guaitipan sp. nov., h) ventral view (arrow shows the retrolateral macrosetae at the base of the RTA); i) same, retrolateral view. A. mae sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828246/files/figure.png" pageId="4" pageNumber="5">20cd</figureCitation>
,
<figureCitation id="13140A8A572C9074FE01FF75FD9DFF3B" box="[439,540,186,211]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="4" pageNumber="5">24b, 24h</figureCitation>
,
<figureCitation id="13140A8A572C9074FD91FF74FDC2FF3B" box="[551,579,187,211]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="4" pageNumber="5">27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3354584572C9074FF21FECDFDE8FE6F" pageId="4" pageNumber="5" type="description">
<paragraph id="8B90160F572C9074FF21FECDFDE8FE6F" blockId="4.[151,1436,258,391]" pageId="4" pageNumber="5">
<materialsCitation id="3B471C52572C9074FF21FECDFCA0FED7" ID-GBIF-Occurrence="1572324816" collectingDate="2016-12-06" collectionCode="ICN" collectorName="Reserva Natural La Almenara &amp; R. Anderson" country="Colombia" elevation="1126" latitude="4.87907" location="Reserva Natural La Almenara" longLatPrecision="1" longitude="-73.25333" municipality="Santa Maria" pageId="4" pageNumber="5" specimenCount="2" stateProvince="Boyaca" typeStatus="holotype">
<emphasis id="B95BCA1D572C9074FF21FECDFEE0FEF3" bold="true" box="[151,353,258,283]" pageId="4" pageNumber="5">
<typeStatus id="5494A8AD572C9074FF21FECDFF5EFEF3" box="[151,223,258,283]" pageId="4" pageNumber="5">Types</typeStatus>
.
<typeStatus id="5494A8AD572C9074FF5DFECDFEDDFEF3" box="[235,348,258,283]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
:
</emphasis>
male from
<collectorName id="26DA73D9572C9074FE52FECCFCB7FEF3" box="[484,822,258,283]" pageId="4" pageNumber="5">
<location id="8EF040D4572C9074FE52FECCFCB7FEF3" LSID="urn:lsid:plazi:treatment:0386A719572C9075FF21FF58FB3DFAD8:8EF040D4572C9074FE52FECCFCB7FEF3" box="[484,822,258,283]" country="Colombia" latitude="4.87907" longLatPrecision="1" longitude="-73.25333" municipality="Santa Maria" name="Reserva Natural La Almenara" pageId="4" pageNumber="5" stateProvince="Boyaca">Reserva Natural La Almenara</location>
</collectorName>
,
<collectingMunicipality id="6BF48C75572C9074FCF4FECCFC4FFEF3" box="[834,974,259,283]" pageId="4" pageNumber="5">Santa María</collectingMunicipality>
,
<collectingRegion id="49EBD8ED572C9074FC6FFECCFBB1FEF3" box="[985,1072,259,283]" country="Colombia" name="Boyaca" pageId="4" pageNumber="5">Boyacá</collectingRegion>
,
<collectingCountry id="F338569F572C9074FB8DFECDFB2DFEF3" box="[1083,1196,258,283]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
,
<elevation id="0002F13C572C9074FB0EFECDFA8FFEF2" box="[1208,1294,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.126" pageId="4" pageNumber="5" unit="m" value="1126.0">
<quantity id="4CD7BBEA572C9074FB0EFECDFA8FFEF2" box="[1208,1294,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.126" pageId="4" pageNumber="5" unit="m" value="1126.0">1126 m</quantity>
</elevation>
,
<geoCoordinate id="EE1B70C8572C9074FAACFECCFA19FEF3" box="[1306,1432,259,283]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="4.87907">4.87907°N</geoCoordinate>
,
<geoCoordinate id="EE1B70C8572C9074FF21FEE9FEA7FED7" box="[151,294,294,319]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="-73.25333">73.25333°W</geoCoordinate>
,
<collectingDate id="EFD5C927572C9074FE8EFEE9FE34FED7" box="[312,437,294,319]" pageId="4" pageNumber="5" value="2016-12-06">
<date id="FF9130CF572C9074FE8EFEE9FE34FED7" box="[312,437,294,319]" pageId="4" pageNumber="5" value="2016-12-06">6.XII.2016</date>
</collectingDate>
,
<collectorName id="26DA73D9572C9074FE71FEE8FDDDFED7" box="[455,604,294,319]" pageId="4" pageNumber="5">R. Anderson</collectorName>
(
<collectionCode id="ED3E8ECA572C9074FDC7FEE8FD23FED7" box="[625,674,295,319]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="5">ICN</collectionCode>
Ar 8273)
</materialsCitation>
.
<materialsCitation id="3B471C52572C9074FC84FEE9FDE8FE6F" ID-GBIF-Occurrence="1572324824" collectingDate="2016-12-07" collectionCode="ICN" collectorName="R. Anderson" country="Colombia" elevation="1152" latitude="4.88749" location="One" longLatPrecision="1" longitude="-73.24989" municipality="Santa Maria" pageId="4" pageNumber="5" specimenCount="3" stateProvince="Boyaca" typeStatus="paratype">
<emphasis id="B95BCA1D572C9074FC84FEE9FC35FED7" bold="true" box="[818,948,294,319]" pageId="4" pageNumber="5">
<typeStatus id="5494A8AD572C9074FC84FEE9FC2EFED7" box="[818,943,294,319]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
:
</emphasis>
one female with same data as holotype (
<collectionCode id="ED3E8ECA572C9074FF28FE84FF4EFE8B" box="[158,207,331,355]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="5">ICN</collectionCode>
Ar 8274).
<location id="8EF040D4572C9074FED7FE84FE11FE8B" LSID="urn:lsid:plazi:treatment:0386A719572C9075FF21FF58FB3DFAD8:8EF040D4572C9074FED7FE84FE11FE8B" box="[353,400,331,355]" latitude="4.88749" longLatPrecision="1" longitude="-73.24989" name="One" pageId="4" pageNumber="5">One</location>
male
<typeStatus id="5494A8AD572C9074FE55FE84FDC2FE8B" box="[483,579,331,355]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
with the same locality as
<typeStatus id="5494A8AD572C9074FC24FE85FC76FE8B" box="[914,1015,330,355]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
,
<elevation id="0002F13C572C9074FBBFFE85FBE7FE8A" box="[1033,1126,330,355]" metricMagnitude="3" metricUnit="m" metricValue="1.152" pageId="4" pageNumber="5" unit="m" value="1152.0">
<quantity id="4CD7BBEA572C9074FBBFFE85FBE7FE8A" box="[1033,1126,330,355]" metricMagnitude="3" metricUnit="m" metricValue="1.152" pageId="4" pageNumber="5" unit="m" value="1152.0">1152 m</quantity>
</elevation>
,
<geoCoordinate id="EE1B70C8572C9074FBCCFE84FB76FE8B" box="[1146,1271,331,355]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="4.88749">4.88749°N</geoCoordinate>
,
<geoCoordinate id="EE1B70C8572C9074FABFFE84FA19FE8B" box="[1289,1432,331,355]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="-73.24989">73.24989°W</geoCoordinate>
,
<collectingDate id="EFD5C927572C9074FF21FEA1FE95FE6F" box="[151,276,366,391]" pageId="4" pageNumber="5" value="2016-12-07">
<date id="FF9130CF572C9074FF21FEA1FE95FE6F" box="[151,276,366,391]" pageId="4" pageNumber="5" value="2016-12-07">7.XII.2016</date>
</collectingDate>
,
<collectorName id="26DA73D9572C9074FE96FEA0FE2CFE6F" box="[288,429,366,391]" pageId="4" pageNumber="5">R. Anderson</collectorName>
(
<collectionCode id="ED3E8ECA572C9074FE0AFEA0FE6CFE6F" box="[444,493,367,391]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="5">ICN</collectionCode>
Ar 8275).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF504687572C9074FF21F8BAFBDAF807" httpUri="https://zenodo.org/record/828210/files/figure.png" pageId="4" pageNumber="5" targetBox="[193,1411,455,1885]" targetPageId="4">
<paragraph id="8B90160F572C9074FF21F8BAFBDAF807" blockId="4.[151,1436,1909,2031]" pageId="4" pageNumber="5">
<emphasis id="B95BCA1D572C9074FF21F8BAFEC9F863" bold="true" box="[151,328,1909,1931]" pageId="4" pageNumber="5">FIGURES 2ad.</emphasis>
<taxonomicName id="4C2F6D8C572C9074FEF8F8BAFE14F862" box="[334,405,1909,1930]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">
<emphasis id="B95BCA1D572C9074FEF8F8BAFE14F862" box="[334,405,1909,1930]" italics="true" pageId="4" pageNumber="5">A. chie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766572C9074FE2DF8BAFE6AF863" box="[411,491,1909,1931]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B95BCA1D572C9074FE2DF8BAFE6AF863" bold="true" box="[411,491,1909,1931]" pageId="4" pageNumber="5">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="B95BCA1D572C9074FE4FF8BAFD8CF863" bold="true" box="[505,525,1909,1931]" pageId="4" pageNumber="5">a)</emphasis>
left male palp, ventral view;
<emphasis id="B95BCA1D572C9074FC8EF8BAFCCEF863" bold="true" box="[824,847,1909,1931]" pageId="4" pageNumber="5">b)</emphasis>
same, retrolateral view;
<emphasis id="B95BCA1D572C9074FBFCF8BAFBDFF863" bold="true" box="[1098,1118,1909,1931]" pageId="4" pageNumber="5">c)</emphasis>
epigyne, cleared, ventral view;
<emphasis id="B95BCA1D572C9074FF21F859FF2FF844" bold="true" box="[151,174,1942,1964]" pageId="4" pageNumber="5">d)</emphasis>
same, dorsal view. Abbreviations: bwe=basal widening of the embolus, cd=copulatory duct, co=copulatory opening, e=embolus, ED=embolic disc, EF=embolic filament, fd=fertilization duct, sp=spermathecal, RTA=retrolateral tibial apophysis of the male palp, SD=sperm duct, TL=tegular lobe. Scale bars: 0.20 mm
<emphasis id="B95BCA1D572C9074FCCFF816FC30F807" bold="true" box="[889,945,2009,2031]" pageId="4" pageNumber="5">(ab)</emphasis>
, 0.10 mm
<emphasis id="B95BCA1D572C9074FBA8F816FBD4F807" bold="true" box="[1054,1109,2009,2031]" pageId="4" pageNumber="5">(cd)</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C3354584572D9075FF71FF58FCE8FF10" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8B90160F572D9075FF71FF58FCE8FF10" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FF58FECFFF58" bold="true" box="[199,334,151,176]" pageId="5" pageNumber="6">Etymology.</emphasis>
The epithet is a noun in apposition that honors
<taxonomicName id="4C2F6D8C572D9075FCD7FF58FC18FF58" box="[865,921,151,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">Chíe</taxonomicName>
, the goddess of moon (also known as Chía) in the Muisca mythology, indigenous people that inhabited the region where the species was found, in the central Andean highlands of the present-day Colombia's Eastern Range.
</paragraph>
</subSubSection>
<subSubSection id="C3354584572D9075FF71FECBFC19FE45" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B90160F572D9075FF71FECBFC19FE45" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FECBFEC0FEF5" bold="true" box="[199,321,260,285]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C2F6D8C572D9075FE0EFECBFD88FEF5" box="[440,521,260,285]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">
<emphasis id="B95BCA1D572D9075FE0EFECBFD88FEF5" box="[440,521,260,285]" italics="true" pageId="5" pageNumber="6">A. chie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766572D9075FDA4FECBFDEFFEF5" box="[530,622,260,285]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B95BCA1D572D9075FDA4FECBFDEFFEF5" bold="true" box="[530,622,260,285]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easily distinguished from those of the remaining species of the genus by their smaller body size and darker coloration, the double-pointed RTA (
<figureCitation id="13140A8A572D9075FB90FEE8FB18FEA8" box="[1062,1177,295,320]" captionStart="FIGURES 2" captionStartId="4.[151,264,1909,1931]" captionTargetBox="[193,1411,455,1885]" captionTargetId="figure@4.[173,1417,443,1894]" captionTargetPageId="4" captionText="FIGURES 2 a d. A. chie sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: bwe = basal widening of the embolus, cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm (a b), 0.10 mm (c d)." httpUri="https://zenodo.org/record/828210/files/figure.png" pageId="5" pageNumber="6">Figs 2ab</figureCitation>
,
<figureCitation id="13140A8A572D9075FB12FEE8FB6FFEA8" box="[1188,1262,295,320]" captionStart="FIGURES 20" captionStartId="31.[151,264,1906,1928]" captionTargetBox="[248,1341,212,1891]" captionTargetId="figure@31.[228,1365,193,1894]" captionTargetPageId="31" captionText="FIGURES 20 a k. Amphidraus spp, left male palp. A. bochica sp. nov., a) ventral view; b) same, retrolateral view. A. chie sp. nov., c) ventral view; d) same, retrolateral view. A. colombianus sp. nov., e) ventral view; f) same, retroventral view; g) same, retrolateral view. A. guaitipan sp. nov., h) ventral view (arrow shows the retrolateral macrosetae at the base of the RTA); i) same, retrolateral view. A. mae sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828246/files/figure.png" pageId="5" pageNumber="6">20cd</figureCitation>
). Females of
<taxonomicName id="4C2F6D8C572D9075FA32FEE7FF47FE8D" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">
<emphasis id="B95BCA1D572D9075FA32FEE7FF47FE8D" italics="true" pageId="5" pageNumber="6">A. chie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766572D9075FF7BFE83FEA9FE8D" box="[205,296,332,357]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B95BCA1D572D9075FF7BFE83FEA9FE8D" bold="true" box="[205,296,332,357]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
are similar to those of
<taxonomicName id="4C2F6D8C572D9075FD86FE83FD2BFE8D" box="[560,682,332,357]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bochica" status="sp. nov.">
<emphasis id="B95BCA1D572D9075FD86FE83FD2BFE8D" box="[560,682,332,357]" italics="true" pageId="5" pageNumber="6">A. bochica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766572D9075FD07FE83FC8DFE8D" box="[689,780,332,357]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B95BCA1D572D9075FD07FE83FC8DFE8D" bold="true" box="[689,780,332,357]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2F6D8C572D9075FCF2FE83FC34FE8D" box="[836,949,332,357]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sikuani" status="sp. nov.">
<emphasis id="B95BCA1D572D9075FCF2FE83FC34FE8D" box="[836,949,332,357]" italics="true" pageId="5" pageNumber="6">A. sikuani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766572D9075FC0BFE83FB96FE8D" box="[957,1047,332,357]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B95BCA1D572D9075FC0BFE83FB96FE8D" bold="true" box="[957,1047,332,357]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
by the short copulatory ducts, but can be distinguished from those by the fact that the copulatory ducts connect proximally to the spermathecae, and posterior concave border of the epigynal plate (
<figureCitation id="13140A8A572D9075FD1EFE5BFC9BFE45" box="[680,794,404,429]" captionStart="FIGURES 2" captionStartId="4.[151,264,1909,1931]" captionTargetBox="[193,1411,455,1885]" captionTargetId="figure@4.[173,1417,443,1894]" captionTargetPageId="4" captionText="FIGURES 2 a d. A. chie sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: bwe = basal widening of the embolus, cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm (a b), 0.10 mm (c d)." httpUri="https://zenodo.org/record/828210/files/figure.png" pageId="5" pageNumber="6">Figs 2cd</figureCitation>
,
<figureCitation id="13140A8A572D9075FC90FE5BFC0DFE45" box="[806,908,404,429]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="5" pageNumber="6">24b, 24h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3354584572D9075FF71FE78FA8DFBB0" pageId="5" pageNumber="6" type="description">
<paragraph id="8B90160F572D9075FF71FE78FC0DFD18" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FE78FDA5FE38" bold="true" box="[199,548,439,464]" pageId="5" pageNumber="6">
Description. Male (
<typeStatus id="5494A8AD572D9075FE1AFE78FD99FE38" box="[428,536,439,464]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length: 1.81. Carapace black, 1.06 long, 0.78 wide, 0.57 high (
<figureCitation id="13140A8A572D9075FA98FE78FA11FE38" box="[1326,1424,439,464]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a i. A. bochica sp. nov., a) male holotype (ICN Ar 8276); b) female paratype (ICN Ar 8269). A. chie sp. nov., c) male holotype (ICN Ar 8273); d) female paratype (ICN Ar 8274). A. colombianus sp. nov., e) male holotype (ICN Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN Ar 8257); g) female paratype (ANDES IN 3871). A. mae sp. nov., h) male holotype (ICN Ar 8259); i) female paratype (ICN Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="5" pageNumber="6">Fig. 16c</figureCitation>
). OC black, 0.54 long. Anterior eye row 0.85 wide and posterior 0.77 wide. Sternum black, 0.46 long, 0.34 wide. Labium black, 0.09 long, 0.17 wide. Chelicerae black with four retromarginal and two promarginal teeth. Palp black and extremely small, with a long and bifurcated-tipped RTA, and long and rounded tegular lobe (TL), rounded embolic disc (ED), and a short embolus with basal widening (bwe) (
<figureCitation id="13140A8A572D9075FBA0FD88FB0AFD88" box="[1046,1163,583,608]" captionStart="FIGURES 4" captionStartId="7.[151,264,1904,1926]" captionTargetBox="[275,1321,710,1880]" captionTargetId="figure@7.[268,1321,690,1891]" captionTargetPageId="7" captionText="FIGURES 4 a d. A. guaitipan sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828214/files/figure.png" pageId="5" pageNumber="6">Figs 4ab</figureCitation>
,
<figureCitation id="13140A8A572D9075FB2FFD88FB63FD88" box="[1177,1250,583,608]" captionStart="FIGURES 20" captionStartId="31.[151,264,1906,1928]" captionTargetBox="[248,1341,212,1891]" captionTargetId="figure@31.[228,1365,193,1894]" captionTargetPageId="31" captionText="FIGURES 20 a k. Amphidraus spp, left male palp. A. bochica sp. nov., a) ventral view; b) same, retrolateral view. A. chie sp. nov., c) ventral view; d) same, retrolateral view. A. colombianus sp. nov., e) ventral view; f) same, retroventral view; g) same, retrolateral view. A. guaitipan sp. nov., h) ventral view (arrow shows the retrolateral macrosetae at the base of the RTA); i) same, retrolateral view. A. mae sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828246/files/figure.png" pageId="5" pageNumber="6">20cd</figureCitation>
). Legs 4312, all black, with the III with metatarsustarsus yellowish, and IIIIV with femurtarsus with yellow marks. Leg macrosetae: femur, IIV d 1 di; tibia, I
<date id="FF9130CF572D9075FDFDFD5FFD06FD40" box="[587,647,656,680]" pageId="5" pageNumber="6">v 2-2</date>
-2; II
<date id="FF9130CF572D9075FD72FD5FFD7AFD40" box="[708,763,656,680]" pageId="5" pageNumber="6">v 1-1</date>
-2; IIIIV
<date id="FF9130CF572D9075FCC4FD5FFC14FD40" box="[882,917,656,680]" pageId="5" pageNumber="6">v 1</date>
pr, p 0-1-1, r 0-1-1; metatarsus, I
<date id="FF9130CF572D9075FAB8FD5FFACDFD40" box="[1294,1356,656,680]" pageId="5" pageNumber="6">v 2-2</date>
; II
<date id="FF9130CF572D9075FAC6FD5FFF26FD24" pageId="5" pageNumber="6">v 2- 2</date>
, p 1 di; III
<date id="FF9130CF572D9075FE91FD7AFECBFD24" box="[295,330,692,716]" pageId="5" pageNumber="6">v 1</date>
di, p 1-0-1, r 2 di; IV
<date id="FF9130CF572D9075FDF1FD7AFDE8FD24" box="[583,617,692,716]" pageId="5" pageNumber="6">v 1</date>
-0-1, p 1-0-2, r 2 di. Abdomen black with transversal thin cream-colored stripes, and two pairs of lateral and posterior white spots (
<figureCitation id="13140A8A572D9075FCA9FD18FC01FD18" box="[799,896,727,752]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a i. A. bochica sp. nov., a) male holotype (ICN Ar 8276); b) female paratype (ICN Ar 8269). A. chie sp. nov., c) male holotype (ICN Ar 8273); d) female paratype (ICN Ar 8274). A. colombianus sp. nov., e) male holotype (ICN Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN Ar 8257); g) female paratype (ANDES IN 3871). A. mae sp. nov., h) male holotype (ICN Ar 8259); i) female paratype (ICN Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="5" pageNumber="6">Fig. 16c</figureCitation>
).
</paragraph>
<paragraph id="8B90160F572D9075FF71FD33FB93FCFD" blockId="5.[151,1437,151,1328]" box="[199,1042,764,789]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FD33FEB9FCFD" bold="true" box="[199,312,764,789]" pageId="5" pageNumber="6">Variation</emphasis>
(n=
<specimenCount id="9D29DD86572D9075FED3FD33FE40FCFD" box="[357,449,764,789]" pageId="5" pageNumber="6" type="male">2 males</specimenCount>
). Total length 1.811.84. Carapace length 1.051.06.
</paragraph>
<paragraph id="8B90160F572D9075FF71FCD0FA8DFBB0" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FCD0FDE3FCD0" bold="true" box="[199,610,799,824]" pageId="5" pageNumber="6">
Female (
<typeStatus id="5494A8AD572D9075FE86FCD0FE1EFCD0" box="[304,415,799,824]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
, ICNAr 8274).
</emphasis>
Total length 2.42. Carapace dark brown with posterior side with black wide marks, 1.12 long, 0.76 wide, 0.57 high (
<figureCitation id="13140A8A572D9075FD18FC8BFC92FCB5" box="[686,787,836,861]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a i. A. bochica sp. nov., a) male holotype (ICN Ar 8276); b) female paratype (ICN Ar 8269). A. chie sp. nov., c) male holotype (ICN Ar 8273); d) female paratype (ICN Ar 8274). A. colombianus sp. nov., e) male holotype (ICN Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN Ar 8257); g) female paratype (ANDES IN 3871). A. mae sp. nov., h) male holotype (ICN Ar 8259); i) female paratype (ICN Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="5" pageNumber="6">Fig. 16d</figureCitation>
). OC black, 0.57 long. Anterior eye row 0.82 wide and posterior 0.77 wide. Sternum yellow, 0.49 long, 0.37 wide. Labium yellow, 0.14 long, 0.18 wide. Chelicerae yellow, with four retromarginal and two promarginal teeth. Legs 4312, all yellow. Leg macrosetae: femur, III, IV d 1 di, III d 2 di; tibiae, I
<date id="FF9130CF572D9075FE64FC7FFD93FC20" box="[466,530,944,968]" pageId="5" pageNumber="6">v 2-2</date>
-2; II
<date id="FF9130CF572D9075FDE0FC7FFD14FC20" box="[598,661,944,968]" pageId="5" pageNumber="6">v 2-2</date>
-2, p 1 di; III
<date id="FF9130CF572D9075FC8AFC7FFCE3FC20" box="[828,866,944,968]" pageId="5" pageNumber="6">v 1</date>
pr, p 0-1-1, r 0-1-1; IV
<date id="FF9130CF572D9075FB33FC7FFB2BFC20" box="[1157,1194,944,968]" pageId="5" pageNumber="6">v 1</date>
-0-1, p 1 di, r 1-0-1; metatarsus, I
<date id="FF9130CF572D9075FE99FC1AFEEFFC04" box="[303,366,980,1004]" pageId="5" pageNumber="6">v 2-2</date>
; II
<date id="FF9130CF572D9075FE23FC1AFE55FC04" box="[405,468,980,1004]" pageId="5" pageNumber="6">v 2-2</date>
, p 1 di; III
<date id="FF9130CF572D9075FDE0FC1AFDFAFC04" box="[598,635,980,1004]" pageId="5" pageNumber="6">v 1</date>
pr, p 1-0-2, r 1-0-1; IV
<date id="FF9130CF572D9075FC3BFC1AFC31FC04" box="[909,944,980,1004]" pageId="5" pageNumber="6">v 1</date>
-0-1, p 1-0-2, r 1-0-1. Abdomen black with transversal thin cream-colored stripes, and two pairs of lateral and posterior white spots, and two anterior and dorsal gray stripes (
<figureCitation id="13140A8A572D9075FECDFBD3FE61FBDD" box="[379,480,1052,1077]" captionStart="FIGURES 16" captionStartId="27.[151,264,1766,1788]" captionTargetBox="[344,1247,526,1743]" captionTargetId="figure@27.[340,1247,517,1745]" captionTargetPageId="27" captionText="FIGURES 16 a i. A. bochica sp. nov., a) male holotype (ICN Ar 8276); b) female paratype (ICN Ar 8269). A. chie sp. nov., c) male holotype (ICN Ar 8273); d) female paratype (ICN Ar 8274). A. colombianus sp. nov., e) male holotype (ICN Ar 7292). A. guaitipan sp. nov., f) male holotype (ICN Ar 8257); g) female paratype (ANDES IN 3871). A. mae sp. nov., h) male holotype (ICN Ar 8259); i) female paratype (ICN Ar 8260). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828238/files/figure.png" pageId="5" pageNumber="6">Fig. 16d</figureCitation>
). Epigyne (
<figureCitation id="13140A8A572D9075FDD3FBD3FD58FBDD" box="[613,729,1052,1077]" captionStart="FIGURES 2" captionStartId="4.[151,264,1909,1931]" captionTargetBox="[193,1411,455,1885]" captionTargetId="figure@4.[173,1417,443,1894]" captionTargetPageId="4" captionText="FIGURES 2 a d. A. chie sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: bwe = basal widening of the embolus, cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm (a b), 0.10 mm (c d)." httpUri="https://zenodo.org/record/828210/files/figure.png" pageId="5" pageNumber="6">Figs 2cd</figureCitation>
,
<figureCitation id="13140A8A572D9075FD51FBD3FCCFFBDD" box="[743,846,1052,1077]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="5" pageNumber="6">24b, 24h</figureCitation>
) with anterior and wide copulatory openings, short copulatory ducts, rounded posterior spermathecae, and concave posterior border of the epigynal plate.
</paragraph>
</subSubSection>
<subSubSection id="C3354584572D9075FF71FBABFE59FAE5" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8B90160F572D9075FF71FBABFE59FAE5" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FBABFECEFB95" bold="true" box="[199,335,1124,1149]" pageId="5" pageNumber="6">Comments.</emphasis>
The
<typeStatus id="5494A8AD572D9075FE27FBAAFE40FB95" box="[401,449,1125,1149]" pageId="5" pageNumber="6">type</typeStatus>
material was collected in Winkler traps, in a secondary Andean forest, at the same locality where the
<typeStatus id="5494A8AD572D9075FED1FB46FE16FB49" box="[359,407,1161,1185]" pageId="5" pageNumber="6">type</typeStatus>
specimens of
<emphasis id="B95BCA1D572D9075FD8EFB47FDFEFB48" box="[568,639,1160,1184]" italics="true" pageId="5" pageNumber="6">A. sua</emphasis>
<emphasis id="B95BCA1D572D9075FD30FB48FD5EFB48" bold="true" box="[646,735,1159,1184]" pageId="5" pageNumber="6">sp. nov.</emphasis>
were collected, so here they are considered to be sympatric. It is interesting to point out that this species was collected in the same sample with numerous specimens of an undescribed species of the genus
<taxonomicName id="4C2F6D8C572D9075FDBCFB00FC2CFB00" authority="O. Pickard-Cambridge, 1897" authorityName="O. Pickard-Cambridge" authorityYear="1897" box="[522,941,1231,1256]" class="Arachnida" family="Zodariidae" genus="Tenedos" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B95BCA1D572D9075FDBCFB00FDE4FB00" box="[522,613,1231,1256]" italics="true" pageId="5" pageNumber="6">Tenedos</emphasis>
O. Pickard-Cambridge, 1897
</taxonomicName>
(
<taxonomicName id="4C2F6D8C572D9075FC0AFB00FBBAFB00" box="[956,1083,1231,1256]" class="Arachnida" family="Zodariidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Zodariidae</taxonomicName>
: Stonerinae), which presented a similar pattern of coloration.
</paragraph>
</subSubSection>
<subSubSection id="C3354584572D9075FF71FAD8FB3DFAD8" box="[199,1212,1303,1328]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8B90160F572D9075FF71FAD8FB3DFAD8" blockId="5.[151,1437,151,1328]" box="[199,1212,1303,1328]" pageId="5" pageNumber="6">
<emphasis id="B95BCA1D572D9075FF71FAD8FEE1FAD8" bold="true" box="[199,352,1303,1328]" pageId="5" pageNumber="6">Distribution.</emphasis>
<collectingCountry id="F338569F572D9075FED1FAD8FE54FAD8" box="[359,469,1303,1328]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
(
<collectingRegion id="49EBD8ED572D9075FE52FAD7FDBAFAD8" box="[484,571,1304,1328]" country="Colombia" name="Boyaca" pageId="5" pageNumber="6">Boyacá</collectingRegion>
) (
<figureCitation id="13140A8A572D9075FDE4FAD8FD26FAD8" box="[594,679,1303,1328]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="5" pageNumber="6">Fig. 27</figureCitation>
). Known altitudinal distribution:
<quantity id="4CD7BBEA572D9075FBAAFAD8FB39FAD8" box="[1052,1208,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.139" metricValueMax="1.152" metricValueMin="1.126" pageId="5" pageNumber="6" unit="m" value="1139.0" valueMax="1152.0" valueMin="1126.0">11261152 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>