292 lines
42 KiB
XML
292 lines
42 KiB
XML
<document id="D742FC2464B4E8DE22D3B9A88264901F" ID-CLB-Dataset="32767" ID-DOI="10.11646/zootaxa.4286.1.1" ID-GBIF-Dataset="a614f1d9-bda9-4df0-a486-aef5af7813fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="828206" ID-ZooBank="6A22EB50-0146-465A-8E2A-523E2F8FE470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108241184" checkinUser="plazi" docAuthor="Galvis, William" docDate="2017" docId="0386A7195725907EFF21FE83FA8FFAE4" docLanguage="en" docName="zootaxa.4286.1.1.pdf" docOrigin="Zootaxa 4286 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus quinini Galvis, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="15" masterDocId="FFBFDF6157289070FFB6FFCFFF81FFE8" masterDocTitle="Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation" masterLastPageNumber="40" masterPageNumber="1" pageNumber="14" updateTime="1698465056562" updateUser="ExternalLinkService">
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<mods:title id="666E8155FB4B9FAC683EAF674602EEE0">Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation</mods:title>
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<mods:name id="B843AF6A5EE15C40DC245A3F31F62679" type="personal">
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<mods:namePart id="7B387FE0725ADD763EA2D2D22ECD2559">Galvis, William</mods:namePart>
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<mods:title id="BE22A00A8465CDEC6384334D7EDF030C">Zootaxa</mods:title>
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<mods:date id="66C3CF491C0175AC5D177E68D79649AD">2017</mods:date>
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<treatment id="0386A7195725907EFF21FE83FA8FFAE4" ID-DOI="http://doi.org/10.5281/zenodo.6045647" ID-GBIF-Taxon="132548512" ID-Zenodo-Dep="6045647" LSID="urn:lsid:plazi:treatment:0386A7195725907EFF21FE83FA8FFAE4" httpUri="http://treatment.plazi.org/id/0386A7195725907EFF21FE83FA8FFAE4" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<subSubSection id="C33545845725907DFF21FE83FDF2FE6F" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8B90160F5725907DFF21FE83FE70FE8E" blockId="13.[151,627,332,391]" box="[151,497,332,358]" pageId="13" pageNumber="14">
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<heading id="D0D8A1635725907DFF21FE83FE70FE8E" bold="true" box="[151,497,332,358]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="B95BCA1D5725907DFF21FE83FE70FE8E" bold="true" box="[151,497,332,358]" pageId="13" pageNumber="14">
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<taxonomicName id="4C2F6D8C5725907DFF21FE83FE0DFE8E" ID-CoL="D2GN" box="[151,396,332,358]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">
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<emphasis id="B95BCA1D5725907DFF21FE83FE0DFE8E" bold="true" box="[151,396,332,358]" italics="true" pageId="13" pageNumber="14">Amphidraus quinini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26877665725907DFE25FE83FE70FE8E" box="[403,497,332,358]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8B90160F5725907DFF21FEA1FDF2FE6F" blockId="13.[151,627,332,391]" box="[151,627,366,391]" pageId="13" pageNumber="14">
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<figureCitation id="13140A8A5725907DFF21FEA1FE86FE6F" box="[151,263,366,391]" captionStart="FIGURES 8" captionStartId="13.[151,264,1877,1899]" captionTargetBox="[195,1401,678,1837]" captionTargetId="figure@13.[176,1420,654,1862]" captionTargetPageId="13" captionText="FIGURES 8 a – d. A. quinini sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EDS = embolic disc spur, EF = embolic filament, fd = fertilization duct, HaD = distal haematodocha, PED = process on embolic disc, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828222/files/figure.png" pageId="13" pageNumber="14">Figs 8a–d</figureCitation>
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,
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<figureCitation id="13140A8A5725907DFEA5FEA1FEDBFE6F" box="[275,346,366,391]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="13" pageNumber="14">17d–e</figureCitation>
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,
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<figureCitation id="13140A8A5725907DFED3FEA1FE29FE6F" box="[357,424,366,391]" captionStart="FIGURES 21" captionStartId="32.[151,264,1917,1939]" captionTargetBox="[187,1405,208,1895]" captionTargetId="figure@32.[151,1436,193,1896]" captionTargetPageId="32" captionText="FIGURES 21 a – k. Amphidraus spp, left male palp. A. guatavita sp. nov., a) ventral view; b) same, retrolateral view; c) same, dorsal view of the tibia. A. pae sp. nov., d) ventral view; e) same, retrolateral view. A. quimbaya sp. nov., f) ventral view; g) same, retrolateral view. A. quinini sp. nov., h) ventral view, partially expanded; i) same, retrolateral view. A. sie sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828248/files/figure.png" pageId="13" pageNumber="14">21h–i</figureCitation>
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,
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<figureCitation id="13140A8A5725907DFE05FEA0FD94FE6F" box="[435,533,366,391]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a – l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="13" pageNumber="14">24e, 24k</figureCitation>
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,
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<figureCitation id="13140A8A5725907DFD97FEA1FDCDFE6F" box="[545,588,366,391]" captionStart="FIGURES 26" captionStartId="37.[151,264,1181,1203]" captionTargetBox="[151,1436,193,1159]" captionTargetId="figure@37.[151,1436,193,1159]" captionTargetPageId="37" captionText="FIGURES 26 a – d. Different altitudinal ecosystems where some of the species of Amphidraus here described were found in Colombia. a) Páramo ecosystem, at Las Lagunas Encantadas Natural Reserve, Duitama, Colombia, habitat of A. tundama sp. nov. (~ 3800 m); b) Andean wet forest at Cueva de los Guácharos National Natural Park, Huila, Colombia, habitat of A. guaitipan sp. nov. (~ 2000 m); c) Andean Oak forest (Quercus humboldtii) at Quinini Natural Reserve, Cundinamarca, Colombia, habitat of A. quinini sp. nov. (~ 1700 m); d) Secondary lowland semi wet forest on the outskirts of the city of Villavicencio, Meta, Colombia, habitat of A. sikuani sp. nov. (~ 500 m)." httpUri="https://zenodo.org/record/828258/files/figure.png" pageId="13" pageNumber="14">26c</figureCitation>
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,
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<figureCitation id="13140A8A5725907DFDE1FEA0FDF2FE6F" box="[599,627,367,391]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="13" pageNumber="14">27</figureCitation>
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</paragraph>
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<materialsCitation id="3B471C525725907DFF21FE79FF5AFDFF" ID-GBIF-Occurrence="1572324833" collectingDate="2015-10-17" collectingDateMax="2015-10-24" collectingDateMin="2015-10-17" collectionCode="ICN" country="Colombia" elevation="1740" latitude="4.336805" location="Reserva Natural Cerro" longLatPrecision="1" longitude="-74.49611" municipality="Tibacuy" pageId="13" pageNumber="14" specimenCount="2" stateProvince="Cundinamarca" typeStatus="holotype">
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<typeStatus id="5494A8AD5725907DFF21FE79FF5EFE27" box="[151,223,438,463]" pageId="13" pageNumber="14">Types</typeStatus>
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.
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<typeStatus id="5494A8AD5725907DFF45FE79FEE5FE27" box="[243,356,438,463]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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male from
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<taxonomicName id="4C2F6D8C5725907DFCA9FE79FCF6FE27" box="[799,887,438,463]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">Quininí</taxonomicName>
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,
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<collectingMunicipality id="6BF48C755725907DFC3DFE79FC66FE27" box="[907,999,438,463]" pageId="13" pageNumber="14">Tibacuy</collectingMunicipality>
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,
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<collectingRegion id="49EBD8ED5725907DFC4CFE79FB21FE27" box="[1018,1184,438,463]" country="Colombia" name="Cundinamarca" pageId="13" pageNumber="14">Cundinamarca</collectingRegion>
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,
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<collectingCountry id="F338569F5725907DFB05FE79FAA5FE27" box="[1203,1316,438,463]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
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,
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,
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<geoCoordinate id="EE1B70C85725907DFE85FE15FE4FFE1B" box="[307,462,474,499]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-74.49611">74.496111°W</geoCoordinate>
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,
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<date id="FF9130CF5725907DFE69FE15FD03FE1B" box="[479,642,474,499]" pageId="13" pageNumber="14" value="2015-10-17" valueMax="2015-10-24" valueMin="2015-10-17">17–24.X.2015</date>
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</collectingDate>
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, Grupo Artropofauna Universidad Nacional de
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<collectingCountry id="F338569F5725907DFB0DFE15FAABFE1B" box="[1211,1322,474,499]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
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(
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<collectionCode id="ED3E8ECA5725907DFA88FE14FAEEFE1B" box="[1342,1391,475,499]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="13" pageNumber="14">ICN</collectionCode>
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–Ar 8266)
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</materialsCitation>
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.
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<materialsCitation id="3B471C525725907DFF50FE31FF61FDB7" ID-GBIF-Occurrence="1572324838" collectingDate="2015-10-17" collectingDateMax="2015-10-24" collectingDateMin="2015-10-17" collectionCode="ICN" country="Colombia" elevation="1887" latitude="4.324833" location="Two" longLatPrecision="1" longitude="-74.4933" municipality="Tibacuy" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Cundinamarca" typeStatus="paratype">
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<emphasis id="B95BCA1D5725907DFF50FE31FEE6FDFF" bold="true" box="[230,359,510,535]" pageId="13" pageNumber="14">
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<typeStatus id="5494A8AD5725907DFF50FE31FEE3FDFF" box="[230,354,510,535]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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one female with same data as holotype (
|
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<collectionCode id="ED3E8ECA5725907DFC9DFE30FCDDFDFF" box="[811,860,511,535]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="13" pageNumber="14">ICN</collectionCode>
|
||
–Ar 8267).
|
||
<location id="8EF040D45725907DFC68FE30FB91FDFE" LSID="urn:lsid:plazi:treatment:0386A7195725907EFF21FE83FA8FFAE4:8EF040D45725907DFC68FE30FB91FDFE" box="[990,1040,511,534]" country="Colombia" latitude="4.324833" longLatPrecision="1" longitude="-74.4933" name="Two" pageId="13" pageNumber="14">Two</location>
|
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males from the same locality,
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,
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,
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<geoCoordinate id="EE1B70C85725907DFEE2FDEDFE70FDD3" box="[340,497,546,571]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-74.4933">74.493305°W</geoCoordinate>
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,
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<collectingDate id="EFD5C9275725907DFE48FDEDFD20FDD3" box="[510,673,546,571]" pageId="13" pageNumber="14" value="2015-10-17" valueMax="2015-10-24" valueMin="2015-10-17">
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<date id="FF9130CF5725907DFE48FDEDFD20FDD3" box="[510,673,546,571]" pageId="13" pageNumber="14" value="2015-10-17" valueMax="2015-10-24" valueMin="2015-10-17">17–24.X.2015</date>
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</collectingDate>
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, Grupo Artropofauna Universidad Nacional de
|
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<collectingCountry id="F338569F5725907DFB76FDEDFAAFFDD3" box="[1216,1326,546,571]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
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(
|
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<collectionCode id="ED3E8ECA5725907DFA88FDECFAEEFDD3" box="[1342,1391,547,571]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="13" pageNumber="14">ICN</collectionCode>
|
||
–Ar 8270).
|
||
</materialsCitation>
|
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</paragraph>
|
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</subSubSection>
|
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<paragraph id="8B90160F5725907DFF21F89AFEECF819" blockId="13.[151,1436,1877,2033]" pageId="13" pageNumber="14">
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<emphasis id="B95BCA1D5725907DFF21F89AFECBF883" bold="true" box="[151,330,1877,1899]" pageId="13" pageNumber="14">FIGURES 8a–d.</emphasis>
|
||
<taxonomicName id="4C2F6D8C5725907DFEE4F899FE39F883" box="[338,440,1878,1899]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">
|
||
<emphasis id="B95BCA1D5725907DFEE4F899FE39F883" box="[338,440,1878,1899]" italics="true" pageId="13" pageNumber="14">A. quinini</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26877665725907DFE76F89AFD93F883" box="[448,530,1877,1899]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="B95BCA1D5725907DFE76F89AFD93F883" bold="true" box="[448,530,1877,1899]" pageId="13" pageNumber="14">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B95BCA1D5725907DFD97F89AFDB4F883" bold="true" box="[545,565,1877,1899]" pageId="13" pageNumber="14">a)</emphasis>
|
||
left male palp, ventral view;
|
||
<emphasis id="B95BCA1D5725907DFCDDF89AFC00F883" bold="true" box="[875,897,1877,1899]" pageId="13" pageNumber="14">b)</emphasis>
|
||
same, retrolateral view;
|
||
<emphasis id="B95BCA1D5725907DFB32F89AFB16F883" bold="true" box="[1156,1175,1877,1899]" pageId="13" pageNumber="14">c)</emphasis>
|
||
epigyne, cleared, ventral view;
|
||
<emphasis id="B95BCA1D5725907DFF6AF8B9FF72F864" bold="true" box="[220,243,1910,1932]" pageId="13" pageNumber="14">d)</emphasis>
|
||
same, dorsal view. Abbreviations: cd=copulatory duct, co=copulatory opening, e=embolus, ED=embolic disc, EDS=embolic disc spur, EF=embolic filament, fd=fertilization duct, HaD=distal haematodocha, PED=process on embolic disc, po=epigynal pocket, RTA=retrolateral tibial apophysis of the male palp, SD=sperm duct, sp=spermatheca, TL=tegular lobe. Scale bars: 0.10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33545845726907EFF71FF58FF46FEF4" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="8B90160F5726907EFF71FF58FF46FEF4" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FF58FECFFF58" bold="true" box="[199,334,151,176]" pageId="14" pageNumber="15">Etymology.</emphasis>
|
||
The epithet is a noun in apposition that honors the Natural Reserve where the species was found, the Natural Reserve Cerro
|
||
<taxonomicName id="4C2F6D8C5726907EFE7CFF73FDA3FF3D" box="[458,546,188,213]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="38" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">Quininí</taxonomicName>
|
||
, ancient territory of the Panches or Tolimas indigenous people of the Cariban language family, known by the Spanish invaders as fearsome warriors and cannibals, whose lives revolved around war.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33545845726907EFF71FEE8FA18FE38" pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph id="8B90160F5726907EFF71FEE8FA18FE38" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FEE8FEC0FEA8" bold="true" box="[199,321,295,320]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C2F6D8C5726907EFE05FEE7FDA0FEA8" box="[435,545,296,320]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">
|
||
<emphasis id="B95BCA1D5726907EFE05FEE7FDA0FEA8" box="[435,545,296,320]" italics="true" pageId="14" pageNumber="15">A. quinini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26877665726907EFD9FFEE8FD05FEA8" box="[553,644,295,320]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B95BCA1D5726907EFD9FFEE8FD05FEA8" bold="true" box="[553,644,295,320]" pageId="14" pageNumber="15">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be easily distinguished from those of the remaining species of the genus by the presence of a bilobed RTA, with the ventral lobe long and finger-like, and the dorsal lobe serrated and wide (
|
||
<figureCitation id="13140A8A5726907EFF6AFEA0FECCFE60" box="[220,333,367,392]" captionStart="FIGURES 8" captionStartId="13.[151,264,1877,1899]" captionTargetBox="[195,1401,678,1837]" captionTargetId="figure@13.[176,1420,654,1862]" captionTargetPageId="13" captionText="FIGURES 8 a – d. A. quinini sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EDS = embolic disc spur, EF = embolic filament, fd = fertilization duct, HaD = distal haematodocha, PED = process on embolic disc, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828222/files/figure.png" pageId="14" pageNumber="15">Figs 8a–b</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A5726907EFEEEFEA0FE1DFE60" box="[344,412,367,392]" captionStart="FIGURES 21" captionStartId="32.[151,264,1917,1939]" captionTargetBox="[187,1405,208,1895]" captionTargetId="figure@32.[151,1436,193,1896]" captionTargetPageId="32" captionText="FIGURES 21 a – k. Amphidraus spp, left male palp. A. guatavita sp. nov., a) ventral view; b) same, retrolateral view; c) same, dorsal view of the tibia. A. pae sp. nov., d) ventral view; e) same, retrolateral view. A. quimbaya sp. nov., f) ventral view; g) same, retrolateral view. A. quinini sp. nov., h) ventral view, partially expanded; i) same, retrolateral view. A. sie sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828248/files/figure.png" pageId="14" pageNumber="15">21h–i</figureCitation>
|
||
). Females of
|
||
<taxonomicName id="4C2F6D8C5726907EFD86FEBFFD1FFE60" box="[560,670,368,392]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quinini" status="sp. nov.">
|
||
<emphasis id="B95BCA1D5726907EFD86FEBFFD1FFE60" box="[560,670,368,392]" italics="true" pageId="14" pageNumber="15">A. quinini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26877665726907EFD12FEA0FD7FFE60" box="[676,766,367,392]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B95BCA1D5726907EFD12FEA0FD7FFE60" bold="true" box="[676,766,367,392]" pageId="14" pageNumber="15">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are similar to those of
|
||
<taxonomicName id="4C2F6D8C5726907EFC4BFEBFFB09FE60" box="[1021,1160,368,392]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="guaitipan" status="sp. nov.">
|
||
<emphasis id="B95BCA1D5726907EFC4BFEBFFB09FE60" box="[1021,1160,368,392]" italics="true" pageId="14" pageNumber="15">A. guaitipan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26877665726907EFB3BFEA0FB66FE60" box="[1165,1255,367,392]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B95BCA1D5726907EFB3BFEA0FB66FE60" bold="true" box="[1165,1255,367,392]" pageId="14" pageNumber="15">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by their general disposition of internal structures of the epigyne, but can be distinguished from those by their longer copulatory ducts, with the posterior epigynal border concave, and a semi-quadrangular epigynal pocket (
|
||
<figureCitation id="13140A8A5726907EFB1DFE78FA9DFE38" box="[1195,1308,439,464]" captionStart="FIGURES 8" captionStartId="13.[151,264,1877,1899]" captionTargetBox="[195,1401,678,1837]" captionTargetId="figure@13.[176,1420,654,1862]" captionTargetPageId="13" captionText="FIGURES 8 a – d. A. quinini sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EDS = embolic disc spur, EF = embolic filament, fd = fertilization duct, HaD = distal haematodocha, PED = process on embolic disc, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828222/files/figure.png" pageId="14" pageNumber="15">Figs 8c–d</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A5726907EFA9EFE77FA0CFE38" box="[1320,1421,439,464]" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a – l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="14" pageNumber="15">24e, 24k</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33545845726907EFF71FE13FEB6FB48" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8B90160F5726907EFF71FE13FAE0FCFD" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FE13FDA8FE1D" bold="true" box="[199,553,476,501]" pageId="14" pageNumber="15">
|
||
Description. Male (
|
||
<typeStatus id="5494A8AD5726907EFE06FE13FD9DFE1D" box="[432,540,476,501]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Total length: 2.39. Carapace lustrous black, 1.24 long, 0.91 wide, 0.72 high (
|
||
<figureCitation id="13140A8A5726907EFF29FE30FE84FDF0" box="[159,261,511,536]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="14" pageNumber="15">Fig. 17d</figureCitation>
|
||
). OC black lustrous, 0.63 long. Anterior eye row 0.96 wide and posterior 0.85 wide. Sternum black lustrous, 0.49 long, 0.38 wide. Labium black lustrous, 0.08 long, 0.13 wide. Chelicerae black with four retromarginal and two promarginal teeth. Palp brown with distal femur and patellae with long white hairs, with a bilobed RTA, with the ventral lobe long and finger-like, and the dorsal lobe serrated and wide, rounded tegular lobe (TL) and long embolic filament (EF) (
|
||
<figureCitation id="13140A8A5726907EFDD4FD40FD58FD40" box="[610,729,655,680]" captionStart="FIGURES 8" captionStartId="13.[151,264,1877,1899]" captionTargetBox="[195,1401,678,1837]" captionTargetId="figure@13.[176,1420,654,1862]" captionTargetPageId="13" captionText="FIGURES 8 a – d. A. quinini sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EDS = embolic disc spur, EF = embolic filament, fd = fertilization duct, HaD = distal haematodocha, PED = process on embolic disc, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828222/files/figure.png" pageId="14" pageNumber="15">Figs 8a–b</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A5726907EFD5CFD40FCAFFD40" box="[746,814,655,680]" captionStart="FIGURES 21" captionStartId="32.[151,264,1917,1939]" captionTargetBox="[187,1405,208,1895]" captionTargetId="figure@32.[151,1436,193,1896]" captionTargetPageId="32" captionText="FIGURES 21 a – k. Amphidraus spp, left male palp. A. guatavita sp. nov., a) ventral view; b) same, retrolateral view; c) same, dorsal view of the tibia. A. pae sp. nov., d) ventral view; e) same, retrolateral view. A. quimbaya sp. nov., f) ventral view; g) same, retrolateral view. A. quinini sp. nov., h) ventral view, partially expanded; i) same, retrolateral view. A. sie sp. nov., j) ventral view; k) same, retrolateral view." httpUri="https://zenodo.org/record/828248/files/figure.png" pageId="14" pageNumber="15">21h–i</figureCitation>
|
||
). Legs 4312, all black with I–II with metatarsi–tarsi yellowish, and III–IV yellow. Leg macrosetae: femur, I–II d 1 di; III d 2 di; IV
|
||
<date id="FF9130CF5726907EFBBFFD7AFBADFD24" box="[1033,1068,692,716]" pageId="14" pageNumber="15">v 1</date>
|
||
di; patella, III–IV r 1 me; tibia, I
|
||
<date id="FF9130CF5726907EFF21FD17FF55FD18" box="[151,212,728,752]" pageId="14" pageNumber="15">v 2-2</date>
|
||
-2; II
|
||
<date id="FF9130CF5726907EFEA6FD17FECBFD18" box="[272,330,728,752]" pageId="14" pageNumber="15">v 2-1</date>
|
||
-2, p 1 di; III–IV v 0-1-0, p 0-1-1, r 0-1-1; metatarsus, I
|
||
<date id="FF9130CF5726907EFC0CFD17FC76FD18" box="[954,1015,728,752]" pageId="14" pageNumber="15">v 2-2</date>
|
||
; II
|
||
<date id="FF9130CF5726907EFBAAFD17FBD9FD18" box="[1052,1112,728,752]" pageId="14" pageNumber="15">v 2-2</date>
|
||
, p 1 di; III
|
||
<date id="FF9130CF5726907EFB63FD17FB74FD18" box="[1237,1269,728,752]" pageId="14" pageNumber="15">v 1</date>
|
||
-0-1, p 1-0-2, r 2 di; IV
|
||
<date id="FF9130CF5726907EFF43FD32FE97FCFC" box="[245,278,764,788]" pageId="14" pageNumber="15">v 1</date>
|
||
-0-1, p 1-0-1, r 1-0-2. Abdomen brown with cream-colored posterior marks and stripes (
|
||
<figureCitation id="13140A8A5726907EFB45FD33FAD5FCFD" box="[1267,1364,764,789]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="14" pageNumber="15">Fig. 17d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B90160F5726907EFF71FCD0FB90FCD0" blockId="14.[151,1437,151,1293]" box="[199,1041,799,824]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FCD0FEB9FCD0" bold="true" box="[199,312,799,824]" pageId="14" pageNumber="15">Variation</emphasis>
|
||
(n=
|
||
<specimenCount id="9D29DD865726907EFED1FCEFFE40FCD0" box="[359,449,799,824]" pageId="14" pageNumber="15" type="male">3 males</specimenCount>
|
||
). Total length 2.01–2.39. Carapace length 1.11–1.27.
|
||
</paragraph>
|
||
<paragraph id="8B90160F5726907EFF71FC8BFEB6FB48" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FC8BFDDFFCB5" bold="true" box="[199,606,836,861]" pageId="14" pageNumber="15">
|
||
Female (
|
||
<typeStatus id="5494A8AD5726907EFE98FC8BFE1CFCB5" box="[302,413,836,861]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
|
||
, ICN–Ar 8267).
|
||
</emphasis>
|
||
Total length 2.86. Carapace dark brown with a dorsal lighter mark, 1.31 long, 0.95 wide, 0.63 high (
|
||
<figureCitation id="13140A8A5726907EFE63FCA8FDB7FC68" box="[469,566,871,896]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="14" pageNumber="15">Fig. 17e</figureCitation>
|
||
). OC black, 0.63 long. Anterior eye row 0.96 wide and posterior 0.87 wide. Sternum dark brown, 0.54 long, 0.46 wide. Labium dark brown, 0.11 long, 0.24 wide. Chelicerae dark brown, with five retromarginal and two promarginal teeth. Legs 4312, all yellow with lateral dark brown sides. Leg macrosetae: femur, I–II, IV d 1 di; III d 2 di; patella, III–IV r 1 me; tibiae, I
|
||
<date id="FF9130CF5726907EFCECFC1AFC16FC04" box="[858,919,980,1004]" pageId="14" pageNumber="15">v 2-2</date>
|
||
-2; II
|
||
<date id="FF9130CF5726907EFC62FC1AFB90FC04" box="[980,1041,980,1004]" pageId="14" pageNumber="15">v 1-2</date>
|
||
-1, p 1 di; III
|
||
<date id="FF9130CF5726907EFB12FC1AFB46FC04" box="[1188,1223,980,1004]" pageId="14" pageNumber="15">v 1</date>
|
||
me, p 1-1-0, r 1-1- 0; IV
|
||
<date id="FF9130CF5726907EFF61FC37FF7BFBF8" box="[215,250,1016,1040]" pageId="14" pageNumber="15">v 1</date>
|
||
pr, p 1-0-1, r 1-0-1; metatarsus, I
|
||
<date id="FF9130CF5726907EFDC5FC37FD31FBF8" box="[627,688,1016,1040]" pageId="14" pageNumber="15">v 2-2</date>
|
||
; II
|
||
<date id="FF9130CF5726907EFD63FC37FC90FBF8" box="[725,785,1016,1040]" pageId="14" pageNumber="15">v 2-2</date>
|
||
, p 1 di; III
|
||
<date id="FF9130CF5726907EFC3AFC37FC2CFBF8" box="[908,941,1016,1040]" pageId="14" pageNumber="15">v 1</date>
|
||
-0-1, p 1-0-2, r 1-0-2; IV
|
||
<date id="FF9130CF5726907EFB7FFC37FB6DFBF8" box="[1225,1260,1016,1040]" pageId="14" pageNumber="15">v 1</date>
|
||
pr, p 1-0-2, r 1- 0-2. Abdomen dark brown with dorsal and lateral light brown marks and spots (
|
||
<figureCitation id="13140A8A5726907EFBA7FBD3FBF1FBDC" box="[1041,1136,1052,1077]" captionStart="FIGURES 17" captionStartId="28.[151,264,1660,1682]" captionTargetBox="[340,1244,413,1635]" captionTargetId="figure@28.[340,1247,411,1639]" captionTargetPageId="28" captionText="FIGURES 17 a – i. A. guatavita sp. nov., a) male holotype (ANDES – IN 3155). A. pae sp. nov., b) male holotype (ICN – Ar 6576). A. quimbaya sp. nov., c) male holotype (ICN – Ar 7746). A. quinini sp. nov., d) male holotype (ICN – Ar 8266); e) female paratype (ICN – Ar 8267). A. sie sp. nov., f) male holotype (ICN – Ar 8254); g) female paratype (ICN – Ar 8255). A. sikuani sp. nov., h) male holotype (ICN – Ar 6558); i) female paratype (ICN – Ar 7185). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828240/files/figure.png" pageId="14" pageNumber="15">Fig. 17e</figureCitation>
|
||
). Epigyne (
|
||
<figureCitation id="13140A8A5726907EFB46FBD3FAE0FBDD" box="[1264,1377,1052,1077]" captionStart="FIGURES 8" captionStartId="13.[151,264,1877,1899]" captionTargetBox="[195,1401,678,1837]" captionTargetId="figure@13.[176,1420,654,1862]" captionTargetPageId="13" captionText="FIGURES 8 a – d. A. quinini sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EDS = embolic disc spur, EF = embolic filament, fd = fertilization duct, HaD = distal haematodocha, PED = process on embolic disc, po = epigynal pocket, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, sp = spermatheca, TL = tegular lobe. Scale bars: 0.10 mm." httpUri="https://zenodo.org/record/828222/files/figure.png" pageId="14" pageNumber="15">Figs 8c–d</figureCitation>
|
||
,
|
||
<figureCitation id="13140A8A5726907EFADBFBD3FF43FBB0" captionStart="FIGURES 24" captionStartId="35.[151,264,1828,1850]" captionTargetBox="[155,1418,206,1799]" captionTargetId="figure@35.[151,1436,193,1807]" captionTargetPageId="35" captionText="FIGURES 24 a – l. Amphidraus spp, epigyne. A. bochica sp. nov., a) ventral view; g) disposition of the internal structures. A. chie sp. nov., b) ventral view; h) disposition of the internal structures. A. guaitipan sp. nov., c) ventral view; i) disposition of the internal structures. A. mae sp. nov., d) ventral view; j) disposition of the internal structures. A. quinini sp. nov., e) ventral view; k) disposition of the internal structures. A. sie sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828254/files/figure.png" pageId="14" pageNumber="15">24e, 24k</figureCitation>
|
||
) with anterior and medial copulatory openings, long copulatory ducts with the anterior portion concave in disposition, semi-concave posterior border and semi-quadrangular epigynal pocket, and oval anterior spermathecae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33545845726907EFF71FB63FE5CFB00" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="8B90160F5726907EFF71FB63FE5CFB00" blockId="14.[151,1437,151,1293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FB63FECEFB2D" bold="true" box="[199,335,1196,1221]" pageId="14" pageNumber="15">Comments.</emphasis>
|
||
The
|
||
<typeStatus id="5494A8AD5726907EFE21FB62FE46FB2D" box="[407,455,1197,1221]" pageId="14" pageNumber="15">type</typeStatus>
|
||
material was collected in pitfall and Winkler traps in an Andean Oak (
|
||
<taxonomicName id="4C2F6D8C5726907EFA88FB62FE91FB00" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="humboldtii">
|
||
<emphasis id="B95BCA1D5726907EFA88FB62FE91FB00" italics="true" pageId="14" pageNumber="15">Quercus humboldtii</emphasis>
|
||
</taxonomicName>
|
||
) forest (
|
||
<figureCitation id="13140A8A5726907EFED9FB00FE50FB00" box="[367,465,1231,1256]" captionStart="FIGURES 26" captionStartId="37.[151,264,1181,1203]" captionTargetBox="[151,1436,193,1159]" captionTargetId="figure@37.[151,1436,193,1159]" captionTargetPageId="37" captionText="FIGURES 26 a – d. Different altitudinal ecosystems where some of the species of Amphidraus here described were found in Colombia. a) Páramo ecosystem, at Las Lagunas Encantadas Natural Reserve, Duitama, Colombia, habitat of A. tundama sp. nov. (~ 3800 m); b) Andean wet forest at Cueva de los Guácharos National Natural Park, Huila, Colombia, habitat of A. guaitipan sp. nov. (~ 2000 m); c) Andean Oak forest (Quercus humboldtii) at Quinini Natural Reserve, Cundinamarca, Colombia, habitat of A. quinini sp. nov. (~ 1700 m); d) Secondary lowland semi wet forest on the outskirts of the city of Villavicencio, Meta, Colombia, habitat of A. sikuani sp. nov. (~ 500 m)." httpUri="https://zenodo.org/record/828258/files/figure.png" pageId="14" pageNumber="15">Fig. 26c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33545845726907EFF71FB3BFA8FFAE4" box="[199,1294,1268,1293]" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8B90160F5726907EFF71FB3BFA8FFAE4" blockId="14.[151,1437,151,1293]" box="[199,1294,1268,1293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B95BCA1D5726907EFF71FB3BFEE1FAE5" bold="true" box="[199,352,1268,1293]" pageId="14" pageNumber="15">Distribution.</emphasis>
|
||
<collectingCountry id="F338569F5726907EFED1FB3BFE54FAE5" box="[359,469,1268,1293]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49EBD8ED5726907EFE52FB3BFD0AFAE5" box="[484,651,1268,1293]" country="Colombia" name="Cundinamarca" pageId="14" pageNumber="15">Cundinamarca</collectingRegion>
|
||
) (
|
||
<figureCitation id="13140A8A5726907EFD14FB3BFD77FAE4" box="[674,758,1268,1293]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="14" pageNumber="15">Fig. 27</figureCitation>
|
||
). Known altitudinal distribution:
|
||
<quantity id="4CD7BBEA5726907EFBDDFB3BFA8BFAE4" box="[1131,1290,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="1.8135" metricValueMax="1.8869999999999998" metricValueMin="1.7399999999999998" pageId="14" pageNumber="15" unit="m" value="1813.5" valueMax="1887.0" valueMin="1740.0">1740–1887 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |