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<document id="8E757E02D134A66AC1293D102ED36391" ID-DOI="10.11646/phytotaxa.261.3.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13671398" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725399484773" checkinUser="felipe" docAuthor="Abrahamczyk, Stefan, Janssens, Steven, Xixima, Lázaro, Ditsch, Barbara &amp; Fischer, Eberhard" docDate="2016" docId="03867B16997DBF6127CBB6FCFF3FFE22" docLanguage="en" docName="phytotaxa.261.3.3.pdf" docOrigin="Phytotaxa 261 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.261.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Impatiens pinganoensis Abrah., S. B. Janssens, Xixima, Ditsch &amp; Eb. Fisch. 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="249" masterDocId="FFBF036E997EBF682743B178FFE6FF97" masterDocTitle="Impatiens pinganoensis (Balsaminaceae), a new species from Angola" masterLastPageNumber="250" masterPageNumber="240" pageNumber="243" updateTime="1725421055326" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FD7DD64A6AD6478EA560942F34229B4C">Impatiens pinganoensis (Balsaminaceae), a new species from Angola</mods:title>
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<mods:affiliation id="781AED8187F8C434B9BBB8588E23AE09">Nees-Institut for Plant Biodiversity, University of Bonn, Meckenheimer Allee 170, 53115 Bonn, Germany;</mods:affiliation>
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<mods:namePart id="569297EE35EE457B5749321195746006">Xixima, Lázaro</mods:namePart>
<mods:affiliation id="A50F92638F1C0E35B66FEFD51AF9AC56">Universidade Kimpa Vita, Escola Superior Politecnica do Uíge, Rua Henrique Freitas N 1, Bairro Popular, Caixa Postal 77, Uíge, Angola</mods:affiliation>
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<emphasis id="B95B1612997DBF6B27CBB6FCFE47F809" bold="true" box="[136,417,1924,1950]" italics="true" pageId="3" pageNumber="243">Impatiens pinganoensis</emphasis>
Abrah., S.B.Janssens, Xixima, Ditsch &amp; Eb.Fisch.
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,
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(
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,
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)
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Impatienti filicornu
<emphasis id="B95B1612997DBF6B2613B6A8FAB0F79E" italics="true" pageId="3" pageNumber="243">affinis sed petiolo decurrente, regulas suberosas crescentibus, petalis superioribus lateralibus petalos inferiores superantibus, floribus albis centro violaceo et petalibus lateralibus superioribus maculis purpureis instructis valde differt.</emphasis>
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:—
<collectingCountry id="F3388A90997ABF6C2797B1D8FEDDFF22" box="[212,315,160,181]" name="Angola" pageId="4" pageNumber="244">ANGOLA</collectingCountry>
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,
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, near
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,
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, (
<figureCitation id="1314D685997ABF6C2325B1D8FB7AFF22" box="[1126,1180,160,181]" captionStart="FIGURE 3" captionStartId="3.[136,229,1586,1607]" captionTargetBox="[153,1433,193,1561]" captionTargetId="figure-18@3.[151,1436,190,1562]" captionTargetPageId="3" captionText="FIGURE 3. Distribution map of Impatiens pinganoensis." figureDoi="http://doi.org/10.5281/zenodo.13671408" httpUri="https://zenodo.org/record/13671408/files/figure.png" pageId="4" pageNumber="244">Fig 3</figureCitation>
),
<collectorName id="26DAAFD6997ABF6C23F1B1D8FA92FF22" box="[1202,1396,160,181]" pageId="4" pageNumber="244">Barbara Ditsch, BD</collectorName>
546a (
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! (
<figureCitation id="1314D685997ABF6C2625B1BCFE47FF4E" box="[358,417,196,217]" captionStart="FIGURE 4" captionStartId="5.[136,229,2000,2021]" captionTargetBox="[151,1436,857,1974]" captionTargetId="figure-366@5.[151,1436,857,1974]" captionTargetPageId="5" captionText="FIGURE 4. Holotype specimen of Impatiens pinganoensis, phot. B. Ditsch." figureDoi="http://doi.org/10.5281/zenodo.13671410" httpUri="https://zenodo.org/record/13671410/files/figure.png" pageId="4" pageNumber="244">Fig. 4</figureCitation>
);
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!.
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<paragraph id="8B90CA00997ABF6C27CBB074FAF5FEB6" blockId="4.[136,1299,268,289]" box="[136,1299,268,289]" pageId="4" pageNumber="244">
<emphasis id="B95B1612997ABF6C27CBB074FF16FEB6" bold="true" box="[136,240,268,289]" pageId="4" pageNumber="244">TABLE 1.</emphasis>
Characters separating
<taxonomicName id="4C2FB183997ABF6C2688B074FD49FEB6" box="[459,687,268,289]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B1612997ABF6C2688B074FD49FEB6" box="[459,687,268,289]" italics="true" pageId="4" pageNumber="244">Impatiens pinganoensis</emphasis>
</taxonomicName>
from the other species of the core
<taxonomicName id="4C2FB183997ABF6C24B8B074FB55FEB6" box="[1019,1203,268,289]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B1612997ABF6C24B8B074FB55FEB6" box="[1019,1203,268,289]" italics="true" pageId="4" pageNumber="244">Impatiens filicornu</emphasis>
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complex.
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<paragraph id="8B90CA00997ABF6C260EB04EFAACF787" pageId="4" pageNumber="244">
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<emphasis id="B95B1612997ABF6C260EB04EFE23FEDC" bold="true" box="[333,453,310,331]" pageId="4" pageNumber="244">Petiole base</emphasis>
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<emphasis id="B95B1612997ABF6C26B6B04EFD65FEDC" bold="true" box="[501,643,310,331]" pageId="4" pageNumber="244">Lateral petals</emphasis>
</th>
<th id="76CEA13E997A409724C1B04EFBF6FEDC" box="[898,1040,310,331]" gridcol="3" gridrow="0" pageId="4" pageNumber="244">
<emphasis id="B95B1612997ABF6C24C1B04EFC19FEDC" bold="true" box="[898,1023,310,331]" pageId="4" pageNumber="244">Dorsal petal</emphasis>
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<emphasis id="B95B1612997ABF6C2376B04EFB81FEDC" bold="true" box="[1077,1127,310,331]" pageId="4" pageNumber="244">Spur</emphasis>
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<emphasis id="B95B1612997ABF6C2251B04EFA72FEDC" bold="true" box="[1298,1428,310,331]" pageId="4" pageNumber="244">Flower color</emphasis>
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<th id="76CEA13E997A409727D4B01EFEC9FEEC" box="[151,303,358,379]" gridcol="0" gridrow="1" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B01EFECCFEEC" authority="Decurrent" authorityName="Decurrent" box="[151,298,358,379]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B1612997ABF6C27D4B01EFECCFEEC" box="[151,298,358,379]" italics="true" pageId="4" pageNumber="244">I. pinganoensis</emphasis>
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<td id="76CEA13E997A4097260EB01EFE30FEEC" box="[333,470,358,379]" gridcol="1" gridrow="1" pageId="4" pageNumber="244">Decurrent,</td>
<td id="76CEA13E997A409726B6B01EFC87FEEC" box="[501,865,358,379]" gridcol="2" gridrow="1" pageId="4" pageNumber="244">Upper petal straight to slightly curved</td>
<td id="76CEA13E997A409724C1B01EFBF6FEEC" box="[898,1040,358,379]" gridcol="3" gridrow="1" pageId="4" pageNumber="244">Ovate-</td>
<td id="76CEA13E997A40972376B01EFB0FFEEC" box="[1077,1257,358,379]" gridcol="4" gridrow="1" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B01EFA7DFEEC" box="[1298,1435,358,379]" gridcol="5" gridrow="1" pageId="4" pageNumber="244">White with</td>
</tr>
<tr id="351FC842997A409727D4B0F2FA7DFE08" box="[151,1435,394,415]" gridrow="2" pageId="4" pageNumber="244" rowspan-0="1">
<td id="76CEA13E997A4097260EB0F2FE30FE08" box="[333,470,394,415]" gridcol="1" gridrow="2" pageId="4" pageNumber="244">developing</td>
<td id="76CEA13E997A409726B6B0F2FC87FE08" box="[501,865,394,415]" gridcol="2" gridrow="2" pageId="4" pageNumber="244">upwards, broadly obtuse at apex;</td>
<td id="76CEA13E997A409724C1B0F2FBF6FE08" box="[898,1040,394,415]" gridcol="3" gridrow="2" pageId="4" pageNumber="244">lanceolate,</td>
<td id="76CEA13E997A40972376B0F2FB0FFE08" box="[1077,1257,394,415]" gridcol="4" gridrow="2" pageId="4" pageNumber="244">navicular, abruptly</td>
<td id="76CEA13E997A40972251B0F2FA7DFE08" box="[1298,1435,394,415]" gridcol="5" gridrow="2" pageId="4" pageNumber="244">mauve centre</td>
</tr>
<tr id="351FC842997A409727D4B0D6FA7DFE54" box="[151,1435,430,451]" gridrow="3" pageId="4" pageNumber="244" rowspan-0="1">
<td id="76CEA13E997A4097260EB0D6FE30FE54" box="[333,470,430,451]" gridcol="1" gridrow="3" pageId="4" pageNumber="244">into cork</td>
<td id="76CEA13E997A409726B6B0D6FC87FE54" box="[501,865,430,451]" gridcol="2" gridrow="3" pageId="4" pageNumber="244">lower petal with rounded indistinct</td>
<td id="76CEA13E997A409724C1B0D6FBF6FE54" box="[898,1040,430,451]" gridcol="3" gridrow="3" pageId="4" pageNumber="244">helmet-like,</td>
<td id="76CEA13E997A40972376B0D6FB0FFE54" box="[1077,1257,430,451]" gridcol="4" gridrow="3" pageId="4" pageNumber="244">constricted into</td>
<td id="76CEA13E997A40972251B0D6FA7DFE54" box="[1298,1435,430,451]" gridcol="5" gridrow="3" pageId="4" pageNumber="244">and wing-like</td>
</tr>
<tr id="351FC842997A409727D4B0AAFA7DFE70" box="[151,1435,466,487]" gridrow="4" pageId="4" pageNumber="244" rowspan-0="1">
<td id="76CEA13E997A4097260EB0AAFE30FE70" box="[333,470,466,487]" gridcol="1" gridrow="4" pageId="4" pageNumber="244">ridges at older</td>
<td id="76CEA13E997A409726B6B0AAFC87FE70" box="[501,865,466,487]" gridcol="2" gridrow="4" pageId="4" pageNumber="244">upper lobe and obtuse lower lobe;</td>
<td id="76CEA13E997A409724C1B0AAFBF6FE70" box="[898,1040,466,487]" gridcol="3" gridrow="4" pageId="4" pageNumber="244">folded</td>
<td id="76CEA13E997A40972376B0AAFB0FFE70" box="[1077,1257,466,487]" gridcol="4" gridrow="4" pageId="4" pageNumber="244">broad, filiform,</td>
<td id="76CEA13E997A40972251B0AAFA7DFE70" box="[1298,1435,466,487]" gridcol="5" gridrow="4" pageId="4" pageNumber="244">pattern on the</td>
</tr>
<tr id="351FC842997A409727D4B08EFA7DFD9C" box="[151,1435,502,523]" gridrow="5" pageId="4" pageNumber="244" rowspan-0="1" rowspan-3="1">
<td id="76CEA13E997A4097260EB08EFE30FD9C" box="[333,470,502,523]" gridcol="1" gridrow="5" pageId="4" pageNumber="244">stems</td>
<td id="76CEA13E997A409726B6B08EFC87FD9C" box="[501,865,502,523]" gridcol="2" gridrow="5" pageId="4" pageNumber="244">upper petals laterally overtopping the</td>
<td id="76CEA13E997A40972376B08EFB0FFD9C" box="[1077,1257,502,523]" gridcol="4" gridrow="5" pageId="4" pageNumber="244">curved spur, apex</td>
<td id="76CEA13E997A40972251B08EFA7DFD9C" box="[1298,1435,502,523]" gridcol="5" gridrow="5" pageId="4" pageNumber="244">upper lateral</td>
</tr>
<tr id="351FC842997A409727D4B362FA7DFDB8" box="[151,1435,538,559]" gridrow="6" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B362FC87FDB8" box="[501,865,538,559]" gridcol="2" gridrow="6" pageId="4" pageNumber="244">lower petals</td>
<td id="76CEA13E997A40972376B362FB0FFDB8" box="[1077,1257,538,559]" gridcol="4" gridrow="6" pageId="4" pageNumber="244">obtuse</td>
<td id="76CEA13E997A40972251B362FA7DFDB8" box="[1298,1435,538,559]" gridcol="5" gridrow="6" pageId="4" pageNumber="244">petals</td>
</tr>
<tr id="351FC842997A409727D4B330FA7DFDCA" box="[151,1435,584,605]" gridrow="7" pageId="4" pageNumber="244">
<th id="76CEA13E997A409727D4B330FEC9FDCA" box="[151,303,584,605]" gridcol="0" gridrow="7" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B330FF18FDCA" authority="Not" authorityName="Not" box="[151,254,584,605]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B1612997ABF6C27D4B330FF18FDCA" box="[151,254,584,605]" italics="true" pageId="4" pageNumber="244">I. filicornu</emphasis>
</taxonomicName>
</th>
<td id="76CEA13E997A4097260EB330FE30FDCA" box="[333,470,584,605]" gridcol="1" gridrow="7" pageId="4" pageNumber="244">Not decurrent</td>
<td id="76CEA13E997A409726B6B330FC87FDCA" box="[501,865,584,605]" gridcol="2" gridrow="7" pageId="4" pageNumber="244">Upper petal curved upwards, widest</td>
<td id="76CEA13E997A409724C1B330FBF6FDCA" box="[898,1040,584,605]" gridcol="3" gridrow="7" pageId="4" pageNumber="244">Ovate-</td>
<td id="76CEA13E997A40972376B330FB0FFDCA" box="[1077,1257,584,605]" gridcol="4" gridrow="7" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B330FA7DFDCA" box="[1298,1435,584,605]" gridcol="5" gridrow="7" pageId="4" pageNumber="244">Pink or purple</td>
</tr>
<tr id="351FC842997A409727D4B314FA7DFD16" box="[151,1435,620,641]" gridrow="8" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B314FC87FD16" box="[501,865,620,641]" gridcol="2" gridrow="8" pageId="4" pageNumber="244">at rounded apex; lower petal with</td>
<td id="76CEA13E997A409724C1B314FBF6FD16" box="[898,1040,620,641]" gridcol="3" gridrow="8" pageId="4" pageNumber="244">lanceolate,</td>
<td id="76CEA13E997A40972376B314FB0FFD16" box="[1077,1257,620,641]" gridcol="4" gridrow="8" pageId="4" pageNumber="244">navicular, abruptly</td>
</tr>
<tr id="351FC842997A409727D4B3E8FA7DFD32" box="[151,1435,656,677]" gridrow="9" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B3E8FC87FD32" box="[501,865,656,677]" gridcol="2" gridrow="9" pageId="4" pageNumber="244">large rounded upper lobe and short,</td>
<td id="76CEA13E997A409724C1B3E8FBF6FD32" box="[898,1040,656,677]" gridcol="3" gridrow="9" pageId="4" pageNumber="244">more or less</td>
<td id="76CEA13E997A40972376B3E8FB0FFD32" box="[1077,1257,656,677]" gridcol="4" gridrow="9" pageId="4" pageNumber="244">constricted into a</td>
</tr>
<tr id="351FC842997A409727D4B3CCFA7DFD5E" box="[151,1435,692,713]" gridrow="10" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B3CCFC87FD5E" box="[501,865,692,713]" gridcol="2" gridrow="10" pageId="4" pageNumber="244">slightly curved lower lobe; upper</td>
<td id="76CEA13E997A409724C1B3CCFBF6FD5E" box="[898,1040,692,713]" gridcol="3" gridrow="10" pageId="4" pageNumber="244">flattened</td>
<td id="76CEA13E997A40972376B3CCFB0FFD5E" box="[1077,1257,692,713]" gridcol="4" gridrow="10" pageId="4" pageNumber="244">narrow, filiform,</td>
</tr>
<tr id="351FC842997A409727D4B3A0FA7DFD7A" box="[151,1435,728,749]" gridrow="11" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B3A0FC87FD7A" box="[501,865,728,749]" gridcol="2" gridrow="11" pageId="4" pageNumber="244">petal not laterally overtopping the</td>
<td id="76CEA13E997A40972376B3A0FB0FFD7A" box="[1077,1257,728,749]" gridcol="4" gridrow="11" pageId="4" pageNumber="244">curved spur, apex</td>
</tr>
<tr id="351FC842997A409727D4B384FA7DFC86" box="[151,1435,764,785]" gridrow="12" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B384FC87FC86" box="[501,865,764,785]" gridcol="2" gridrow="12" pageId="4" pageNumber="244">lobe of the lower petal, both apices</td>
<td id="76CEA13E997A40972376B384FB0FFC86" box="[1077,1257,764,785]" gridcol="4" gridrow="12" pageId="4" pageNumber="244">acuminate</td>
</tr>
<tr id="351FC842997A409727D4B258FA7DFCA2" box="[151,1435,800,821]" gridrow="13" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B258FC87FCA2" box="[501,865,800,821]" gridcol="2" gridrow="13" pageId="4" pageNumber="244">almost reaching the same level</td>
</tr>
<tr id="351FC842997A409727D4B237FA7DFCF3" box="[151,1435,847,868]" gridrow="14" pageId="4" pageNumber="244">
<th id="76CEA13E997A409727D4B237FEC9FCF3" box="[151,303,847,868]" gridcol="0" gridrow="14" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B237FEF4FCF3" authority="Not" authorityName="Not" box="[151,274,847,868]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="gongolana">
<emphasis id="B95B1612997ABF6C27D4B237FEF4FCF3" box="[151,274,847,868]" italics="true" pageId="4" pageNumber="244">I. gongolana</emphasis>
</taxonomicName>
</th>
<td id="76CEA13E997A4097260EB237FE30FCF3" box="[333,470,847,868]" gridcol="1" gridrow="14" pageId="4" pageNumber="244">Not decurrent</td>
<td id="76CEA13E997A409726B6B237FC87FCF3" box="[501,865,847,868]" gridcol="2" gridrow="14" pageId="4" pageNumber="244">Upper petal short, curved upwards,</td>
<td id="76CEA13E997A409724C1B237FBF6FCF3" box="[898,1040,847,868]" gridcol="3" gridrow="14" pageId="4" pageNumber="244">Ovate-</td>
<td id="76CEA13E997A40972376B237FB0FFCF3" box="[1077,1257,847,868]" gridcol="4" gridrow="14" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B237FA7DFCF3" box="[1298,1435,847,868]" gridcol="5" gridrow="14" pageId="4" pageNumber="244">Rose to</td>
</tr>
<tr id="351FC842997A409727D4B20BFA7DFC1F" box="[151,1435,883,904]" gridrow="15" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B20BFC87FC1F" box="[501,865,883,904]" gridcol="2" gridrow="15" pageId="4" pageNumber="244">acuminate; lower petal with large,</td>
<td id="76CEA13E997A409724C1B20BFBF6FC1F" box="[898,1040,883,904]" gridcol="3" gridrow="15" pageId="4" pageNumber="244">rectangular,</td>
<td id="76CEA13E997A40972376B20BFB0FFC1F" box="[1077,1257,883,904]" gridcol="4" gridrow="15" pageId="4" pageNumber="244">navicular, tapering</td>
<td id="76CEA13E997A40972251B20BFA7DFC1F" box="[1298,1435,883,904]" gridcol="5" gridrow="15" pageId="4" pageNumber="244">purple-pink</td>
</tr>
<tr id="351FC842997A409727D4B2EFFA7DFC3B" box="[151,1435,919,940]" gridrow="16" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B2EFFC87FC3B" box="[501,865,919,940]" gridcol="2" gridrow="16" pageId="4" pageNumber="244">rounded upper lobe and short</td>
<td id="76CEA13E997A409724C1B2EFFBF6FC3B" box="[898,1040,919,940]" gridcol="3" gridrow="16" pageId="4" pageNumber="244">more or less</td>
<td id="76CEA13E997A40972376B2EFFB0FFC3B" box="[1077,1257,919,940]" gridcol="4" gridrow="16" pageId="4" pageNumber="244">
into a narrow,
<tr id="351FC842997A409727D4B2C3FA7DFC47" box="[151,1435,955,976]" gridrow="17" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B2C3FC87FC47" box="[501,865,955,976]" gridcol="2" gridrow="17" pageId="4" pageNumber="244">uacuminate lower lobe; upper lobe of</td>
<td id="76CEA13E997A409724C1B2C3FBF6FC47" box="[898,1040,955,976]" gridcol="3" gridrow="17" pageId="4" pageNumber="244">flattened,</td>
<td id="76CEA13E997A40972376B2C3FB0FFC47" box="[1077,1257,955,976]" gridcol="4" gridrow="17" pageId="4" pageNumber="244">shaped spur, apex</td>
</tr>
</td>
</tr>
<tr id="351FC842997A409727D4B2A7FA7DFC63" box="[151,1435,991,1012]" gridrow="18" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B2A7FC87FC63" box="[501,865,991,1012]" gridcol="2" gridrow="18" pageId="4" pageNumber="244">lower petal laterally overtopping the</td>
<td id="76CEA13E997A409724C1B2A7FBF6FC63" box="[898,1040,991,1012]" gridcol="3" gridrow="18" pageId="4" pageNumber="244">emarginated at</td>
<td id="76CEA13E997A40972376B2A7FB0FFC63" box="[1077,1257,991,1012]" gridcol="4" gridrow="18" pageId="4" pageNumber="244">obtuse</td>
</tr>
<tr id="351FC842997A409727D4B57BFA7DFB8F" box="[151,1435,1027,1048]" gridrow="19" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B57BFC87FB8F" box="[501,865,1027,1048]" gridcol="2" gridrow="19" pageId="4" pageNumber="244">much shorter upper petal</td>
<td id="76CEA13E997A409724C1B57BFBF6FB8F" box="[898,1040,1027,1048]" gridcol="3" gridrow="19" pageId="4" pageNumber="244">apex</td>
</tr>
<tr id="351FC842997A409727D4B54AFA7DFBD0" box="[151,1435,1074,1095]" gridrow="20" pageId="4" pageNumber="244">
<th id="76CEA13E997A409727D4B54AFEC9FBD0" box="[151,303,1074,1095]" gridcol="0" gridrow="20" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B54AFEC9FBD0" authority="Not" authorityName="Not" box="[151,303,1074,1095]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="kamerunensis">
<emphasis id="B95B1612997ABF6C27D4B54AFEC9FBD0" box="[151,303,1074,1095]" italics="true" pageId="4" pageNumber="244">I. kamerunensis</emphasis>
</taxonomicName>
</th>
<td id="76CEA13E997A4097260EB54AFE30FBD0" box="[333,470,1074,1095]" gridcol="1" gridrow="20" pageId="4" pageNumber="244">Not decurrent</td>
<td id="76CEA13E997A409726B6B54AFC87FBD0" box="[501,865,1074,1095]" gridcol="2" gridrow="20" pageId="4" pageNumber="244">Upper petal curved upwards, apex</td>
<td id="76CEA13E997A409724C1B54AFBF6FBD0" box="[898,1040,1074,1095]" gridcol="3" gridrow="20" pageId="4" pageNumber="244">Rounded-</td>
<td id="76CEA13E997A40972376B54AFB0FFBD0" box="[1077,1257,1074,1095]" gridcol="4" gridrow="20" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B54AFA7DFBD0" box="[1298,1435,1074,1095]" gridcol="5" gridrow="20" pageId="4" pageNumber="244">Pink with</td>
</tr>
<tr id="351FC842997A409727D4B52EFA7DFBFC" box="[151,1435,1110,1131]" gridrow="21" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B52EFC87FBFC" box="[501,865,1110,1131]" gridcol="2" gridrow="21" pageId="4" pageNumber="244">obtuse-emarginate, lower petal with</td>
<td id="76CEA13E997A409724C1B52EFBF6FBFC" box="[898,1040,1110,1131]" gridcol="3" gridrow="21" pageId="4" pageNumber="244">orbicular,</td>
<td id="76CEA13E997A40972376B52EFB0FFBFC" box="[1077,1257,1110,1131]" gridcol="4" gridrow="21" pageId="4" pageNumber="244">navicular, abruptly</td>
<td id="76CEA13E997A40972251B52EFA7DFBFC" box="[1298,1435,1110,1131]" gridcol="5" gridrow="21" pageId="4" pageNumber="244">white and</td>
</tr>
<tr id="351FC842997A409727D4B502FA7DFB18" box="[151,1435,1146,1167]" gridrow="22" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B502FC87FB18" box="[501,865,1146,1167]" gridcol="2" gridrow="22" pageId="4" pageNumber="244">distinct rounded upper lobe and</td>
<td id="76CEA13E997A409724C1B502FBF6FB18" box="[898,1040,1146,1167]" gridcol="3" gridrow="22" pageId="4" pageNumber="244">more or less</td>
<td id="76CEA13E997A40972376B502FB0FFB18" box="[1077,1257,1146,1167]" gridcol="4" gridrow="22" pageId="4" pageNumber="244">constricted into a</td>
<td id="76CEA13E997A40972251B502FA7DFB18" box="[1298,1435,1146,1167]" gridcol="5" gridrow="22" pageId="4" pageNumber="244">yellow</td>
</tr>
<tr id="351FC842997A409727D4B5E6FA7DFB24" box="[151,1435,1182,1203]" gridrow="23" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B5E6FC87FB24" box="[501,865,1182,1203]" gridcol="2" gridrow="23" pageId="4" pageNumber="244">straight, obtuse lower lobe; upper</td>
<td id="76CEA13E997A409724C1B5E6FBF6FB24" box="[898,1040,1182,1203]" gridcol="3" gridrow="23" pageId="4" pageNumber="244">flattened</td>
<td id="76CEA13E997A40972376B5E6FB0FFB24" box="[1077,1257,1182,1203]" gridcol="4" gridrow="23" pageId="4" pageNumber="244">narrow, filiform,</td>
<td id="76CEA13E997A40972251B5E6FA7DFB24" box="[1298,1435,1182,1203]" gridcol="5" gridrow="23" pageId="4" pageNumber="244">markings on</td>
</tr>
<tr id="351FC842997A409727D4B5BAFA7DFB40" box="[151,1435,1218,1239]" gridrow="24" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B5BAFC87FB40" box="[501,865,1218,1239]" gridcol="2" gridrow="24" pageId="4" pageNumber="244">petals laterally slightly overtopping</td>
<td id="76CEA13E997A40972376B5BAFB0FFB40" box="[1077,1257,1218,1239]" gridcol="4" gridrow="24" pageId="4" pageNumber="244">curved spur, apex</td>
<td id="76CEA13E997A40972251B5BAFA7DFB40" box="[1298,1435,1218,1239]" gridcol="5" gridrow="24" pageId="4" pageNumber="244">the upper and</td>
</tr>
<tr id="351FC842997A409727D4B59EFA7DFB6C" box="[151,1435,1254,1275]" gridrow="25" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B59EFC87FB6C" box="[501,865,1254,1275]" gridcol="2" gridrow="25" pageId="4" pageNumber="244">the lower petals</td>
<td id="76CEA13E997A40972376B59EFB0FFB6C" box="[1077,1257,1254,1275]" gridcol="4" gridrow="25" pageId="4" pageNumber="244">acuminate</td>
<td id="76CEA13E997A40972251B59EFA7DFB6C" box="[1298,1435,1254,1275]" gridcol="5" gridrow="25" pageId="4" pageNumber="244">lower lateral</td>
</tr>
<tr id="351FC842997A409727D4B472FA7DFA88" box="[151,1435,1290,1311]" gridrow="26" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="76CEA13E997A40972251B472FA7DFA88" box="[1298,1435,1290,1311]" gridcol="5" gridrow="26" pageId="4" pageNumber="244">petals</td>
</tr>
<tr id="351FC842997A409727D4B441FA7DFAD9" box="[151,1435,1337,1358]" gridrow="27" pageId="4" pageNumber="244">
<th id="76CEA13E997A409727D4B441FEC9FAD9" box="[151,303,1337,1358]" gridcol="0" gridrow="27" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B441FEF2FAD9" authority="Not" authorityName="Not" box="[151,276,1337,1358]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="nigeriensis">
<emphasis id="B95B1612997ABF6C27D4B441FEF2FAD9" box="[151,276,1337,1358]" italics="true" pageId="4" pageNumber="244">I. nigeriensis</emphasis>
</taxonomicName>
</th>
<td id="76CEA13E997A4097260EB441FE30FAD9" box="[333,470,1337,1358]" gridcol="1" gridrow="27" pageId="4" pageNumber="244">Not decurrent</td>
<td id="76CEA13E997A409726B6B441FC87FAD9" box="[501,865,1337,1358]" gridcol="2" gridrow="27" pageId="4" pageNumber="244">Upper petal more or less straight,</td>
<td id="76CEA13E997A409724C1B441FBF6FAD9" box="[898,1040,1337,1358]" gridcol="3" gridrow="27" pageId="4" pageNumber="244">Ovate, helmet-</td>
<td id="76CEA13E997A40972376B441FB0FFAD9" box="[1077,1257,1337,1358]" gridcol="4" gridrow="27" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B441FA7DFAD9" box="[1298,1435,1337,1358]" gridcol="5" gridrow="27" pageId="4" pageNumber="244">Whitish to</td>
</tr>
<tr id="351FC842997A409727D4B425FA7DFAE5" box="[151,1435,1373,1394]" gridrow="28" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B425FC87FAE5" box="[501,865,1373,1394]" gridcol="2" gridrow="28" pageId="4" pageNumber="244">acuminate at apex; lower petal only</td>
<td id="76CEA13E997A409724C1B425FBF6FAE5" box="[898,1040,1373,1394]" gridcol="3" gridrow="28" pageId="4" pageNumber="244">like, folded</td>
<td id="76CEA13E997A40972376B425FB0FFAE5" box="[1077,1257,1373,1394]" gridcol="4" gridrow="28" pageId="4" pageNumber="244">navicular, abruptly</td>
<td id="76CEA13E997A40972251B425FA7DFAE5" box="[1298,1435,1373,1394]" gridcol="5" gridrow="28" pageId="4" pageNumber="244">pale mauve</td>
</tr>
<tr id="351FC842997A409727D4B4F9FA7DFA01" box="[151,1435,1409,1430]" gridrow="29" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B4F9FC87FA01" box="[501,865,1409,1430]" gridcol="2" gridrow="29" pageId="4" pageNumber="244">with one rounded lobe, lower lobe</td>
<td id="76CEA13E997A40972376B4F9FB0FFA01" box="[1077,1257,1409,1430]" gridcol="4" gridrow="29" pageId="4" pageNumber="244">constricted into a</td>
</tr>
<tr id="351FC842997A409727D4B4DDFA7DFA2D" box="[151,1435,1445,1466]" gridrow="30" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B4DDFC87FA2D" box="[501,865,1445,1466]" gridcol="2" gridrow="30" pageId="4" pageNumber="244">lacking; upper petal not laterally</td>
<td id="76CEA13E997A40972376B4DDFB0FFA2D" box="[1077,1257,1445,1466]" gridcol="4" gridrow="30" pageId="4" pageNumber="244">narrow, filiform,</td>
</tr>
<tr id="351FC842997A409727D4B4B1FA7DFA49" box="[151,1435,1481,1502]" gridrow="31" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B4B1FC87FA49" box="[501,865,1481,1502]" gridcol="2" gridrow="31" pageId="4" pageNumber="244">overtopping the lobe of the lower</td>
<td id="76CEA13E997A40972376B4B1FB0FFA49" box="[1077,1257,1481,1502]" gridcol="4" gridrow="31" pageId="4" pageNumber="244">curved spur, apex</td>
</tr>
<tr id="351FC842997A409727D4B495FA7DF995" box="[151,1435,1517,1538]" gridrow="32" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B495FC87F995" box="[501,865,1517,1538]" gridcol="2" gridrow="32" pageId="4" pageNumber="244">petal, both apices almost reaching the</td>
<td id="76CEA13E997A40972376B495FB0FF995" box="[1077,1257,1517,1538]" gridcol="4" gridrow="32" pageId="4" pageNumber="244">acuminate</td>
</tr>
<tr id="351FC842997A409727D4B769FA7DF9B1" box="[151,1435,1553,1574]" gridrow="33" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="76CEA13E997A409726B6B769FC87F9B1" box="[501,865,1553,1574]" gridcol="2" gridrow="33" pageId="4" pageNumber="244">same level</td>
</tr>
<tr id="351FC842997A409727D4B738FA7DF9C2" box="[151,1435,1600,1621]" gridrow="34" pageId="4" pageNumber="244">
<th id="76CEA13E997A409727D4B738FEC9F9C2" box="[151,303,1600,1621]" gridcol="0" gridrow="34" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B738FEECF9C2" authority=", Not" authorityName="Not" box="[151,266,1600,1621]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="nzabiana">
<emphasis id="B95B1612997ABF6C27D4B738FEECF9C2" box="[151,266,1600,1621]" italics="true" pageId="4" pageNumber="244">I. nzabiana,</emphasis>
</taxonomicName>
</th>
<td id="76CEA13E997A4097260EB738FE30F9C2" box="[333,470,1600,1621]" gridcol="1" gridrow="34" pageId="4" pageNumber="244">Not decurrent</td>
<td id="76CEA13E997A409726B6B738FC87F9C2" box="[501,865,1600,1621]" gridcol="2" gridrow="34" pageId="4" pageNumber="244">Upper petal wing-like, curved</td>
<td id="76CEA13E997A409724C1B738FBF6F9C2" box="[898,1040,1600,1621]" gridcol="3" gridrow="34" pageId="4" pageNumber="244">Orbicular-</td>
<td id="76CEA13E997A40972376B738FB0FF9C2" box="[1077,1257,1600,1621]" gridcol="4" gridrow="34" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B738FA7DF9C2" box="[1298,1435,1600,1621]" gridcol="5" gridrow="34" pageId="4" pageNumber="244">Pink with</td>
</tr>
<tr id="351FC842997A409727D4B71CFA7DF9EE" box="[151,1435,1636,1657]" gridrow="35" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B71CFC87F9EE" box="[501,865,1636,1657]" gridcol="2" gridrow="35" pageId="4" pageNumber="244">upwards, acuminate at apex; lower</td>
<td id="76CEA13E997A409724C1B71CFBF6F9EE" box="[898,1040,1636,1657]" gridcol="3" gridrow="35" pageId="4" pageNumber="244">cordate,</td>
<td id="76CEA13E997A40972376B71CFB0FF9EE" box="[1077,1257,1636,1657]" gridcol="4" gridrow="35" pageId="4" pageNumber="244">navicular, abruptly</td>
<td id="76CEA13E997A40972251B71CFA7DF9EE" box="[1298,1435,1636,1657]" gridcol="5" gridrow="35" pageId="4" pageNumber="244">white</td>
</tr>
<tr id="351FC842997A409727D4B7F0FA7DF90A" box="[151,1435,1672,1693]" gridrow="36" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B7F0FC87F90A" box="[501,865,1672,1693]" gridcol="2" gridrow="36" pageId="4" pageNumber="244">petal with short rounded upper lobe</td>
<td id="76CEA13E997A409724C1B7F0FBF6F90A" box="[898,1040,1672,1693]" gridcol="3" gridrow="36" pageId="4" pageNumber="244">more or less</td>
<td id="76CEA13E997A40972376B7F0FB0FF90A" box="[1077,1257,1672,1693]" gridcol="4" gridrow="36" pageId="4" pageNumber="244">constricted into a</td>
<td id="76CEA13E997A40972251B7F0FA7DF90A" box="[1298,1435,1672,1693]" gridcol="5" gridrow="36" pageId="4" pageNumber="244">markings on</td>
</tr>
<tr id="351FC842997A409727D4B7D4FA7DF956" box="[151,1435,1708,1729]" gridrow="37" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B7D4FC87F956" box="[501,865,1708,1729]" gridcol="2" gridrow="37" pageId="4" pageNumber="244">and straight acuminate lower lobe;</td>
<td id="76CEA13E997A409724C1B7D4FBF6F956" box="[898,1040,1708,1729]" gridcol="3" gridrow="37" pageId="4" pageNumber="244">flattened</td>
<td id="76CEA13E997A40972376B7D4FB0FF956" box="[1077,1257,1708,1729]" gridcol="4" gridrow="37" pageId="4" pageNumber="244">narrow, filiform,</td>
<td id="76CEA13E997A40972251B7D4FA7DF956" box="[1298,1435,1708,1729]" gridcol="5" gridrow="37" pageId="4" pageNumber="244">the upper and</td>
</tr>
<tr id="351FC842997A409727D4B7A8FA7DF972" box="[151,1435,1744,1765]" gridrow="38" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B7A8FC87F972" box="[501,865,1744,1765]" gridcol="2" gridrow="38" pageId="4" pageNumber="244">upper petals laterally overtopping the</td>
<td id="76CEA13E997A40972376B7A8FB0FF972" box="[1077,1257,1744,1765]" gridcol="4" gridrow="38" pageId="4" pageNumber="244">curved spur, apex</td>
<td id="76CEA13E997A40972251B7A8FA7DF972" box="[1298,1435,1744,1765]" gridcol="5" gridrow="38" pageId="4" pageNumber="244">lower lateral</td>
</tr>
<tr id="351FC842997A409727D4B78CFA7DF89E" box="[151,1435,1780,1801]" gridrow="39" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B78CFC87F89E" box="[501,865,1780,1801]" gridcol="2" gridrow="39" pageId="4" pageNumber="244">lower petals</td>
<td id="76CEA13E997A40972376B78CFB0FF89E" box="[1077,1257,1780,1801]" gridcol="4" gridrow="39" pageId="4" pageNumber="244">acuminate</td>
<td id="76CEA13E997A40972251B78CFA7DF89E" box="[1298,1435,1780,1801]" gridcol="5" gridrow="39" pageId="4" pageNumber="244">petals</td>
</tr>
<tr id="351FC842997A409727D4B65BFA7DF8AF" box="[151,1435,1827,1848]" gridrow="40" pageId="4" pageNumber="244">
<th id="76CEA13E997A409727D4B65BFEC9F8AF" box="[151,303,1827,1848]" gridcol="0" gridrow="40" pageId="4" pageNumber="244">
<taxonomicName id="4C2FB183997ABF6C27D4B65BFF15F8AF" authority="Not" authorityName="Not" box="[151,243,1827,1848]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="244" phylum="Tracheophyta" rank="species" species="oumina">
<emphasis id="B95B1612997ABF6C27D4B65BFF15F8AF" box="[151,243,1827,1848]" italics="true" pageId="4" pageNumber="244">I. oumina</emphasis>
</taxonomicName>
</th>
<td id="76CEA13E997A4097260EB65BFE30F8AF" box="[333,470,1827,1848]" gridcol="1" gridrow="40" pageId="4" pageNumber="244">Not decurrent</td>
<td id="76CEA13E997A409726B6B65BFC87F8AF" box="[501,865,1827,1848]" gridcol="2" gridrow="40" pageId="4" pageNumber="244">Upper petal narrow lanceolate,</td>
<td id="76CEA13E997A409724C1B65BFBF6F8AF" box="[898,1040,1827,1848]" gridcol="3" gridrow="40" pageId="4" pageNumber="244">Broadly ovate,</td>
<td id="76CEA13E997A40972376B65BFB0FF8AF" box="[1077,1257,1827,1848]" gridcol="4" gridrow="40" pageId="4" pageNumber="244">Lateral sepal</td>
<td id="76CEA13E997A40972251B65BFA7DF8AF" box="[1298,1435,1827,1848]" gridcol="5" gridrow="40" pageId="4" pageNumber="244">Pink with</td>
</tr>
<tr id="351FC842997A409727D4B63FFA7DF8CB" box="[151,1435,1863,1884]" gridrow="41" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B63FFC87F8CB" box="[501,865,1863,1884]" gridcol="2" gridrow="41" pageId="4" pageNumber="244">distinctly curved upwards, acuminate</td>
<td id="76CEA13E997A409724C1B63FFBF6F8CB" box="[898,1040,1863,1884]" gridcol="3" gridrow="41" pageId="4" pageNumber="244">helmet-like,</td>
<td id="76CEA13E997A40972376B63FFB0FF8CB" box="[1077,1257,1863,1884]" gridcol="4" gridrow="41" pageId="4" pageNumber="244">navicular, abruptly</td>
<td id="76CEA13E997A40972251B63FFA7DF8CB" box="[1298,1435,1863,1884]" gridcol="5" gridrow="41" pageId="4" pageNumber="244">white and</td>
</tr>
<tr id="351FC842997A409727D4B613FA7DF817" box="[151,1435,1899,1920]" gridrow="42" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1">
<td id="76CEA13E997A409726B6B613FC87F817" box="[501,865,1899,1920]" gridcol="2" gridrow="42" pageId="4" pageNumber="244">at apex; lower petal lanceolate, with</td>
<td id="76CEA13E997A409724C1B613FBF6F817" box="[898,1040,1899,1920]" gridcol="3" gridrow="42" pageId="4" pageNumber="244">slightly folded</td>
<td id="76CEA13E997A40972376B613FB0FF817" box="[1077,1257,1899,1920]" gridcol="4" gridrow="42" pageId="4" pageNumber="244">constricted into a</td>
<td id="76CEA13E997A40972251B613FA7DF817" box="[1298,1435,1899,1920]" gridcol="5" gridrow="42" pageId="4" pageNumber="244">yellow</td>
</tr>
<tr id="351FC842997A409727D4B6F7FA7DF833" box="[151,1435,1935,1956]" gridrow="43" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B6F7FC87F833" box="[501,865,1935,1956]" gridcol="2" gridrow="43" pageId="4" pageNumber="244">short rounded upper lobe near apex</td>
<td id="76CEA13E997A40972376B6F7FB0FF833" box="[1077,1257,1935,1956]" gridcol="4" gridrow="43" pageId="4" pageNumber="244">narrow, filiform,</td>
<td id="76CEA13E997A40972251B6F7FA7DF833" box="[1298,1435,1935,1956]" gridcol="5" gridrow="43" pageId="4" pageNumber="244">markings on</td>
</tr>
<tr id="351FC842997A409727D4B6CBFA7DF85F" box="[151,1435,1971,1992]" gridrow="44" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B6CBFC87F85F" box="[501,865,1971,1992]" gridcol="2" gridrow="44" pageId="4" pageNumber="244">and short straight obtuse lower lobe;</td>
<td id="76CEA13E997A40972376B6CBFB0FF85F" box="[1077,1257,1971,1992]" gridcol="4" gridrow="44" pageId="4" pageNumber="244">curved spur, apex</td>
<td id="76CEA13E997A40972251B6CBFA7DF85F" box="[1298,1435,1971,1992]" gridcol="5" gridrow="44" pageId="4" pageNumber="244">the upper and</td>
</tr>
<tr id="351FC842997A409727D4B6AFFA7DF87B" box="[151,1435,2007,2028]" gridrow="45" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76CEA13E997A409726B6B6AFFC87F87B" box="[501,865,2007,2028]" gridcol="2" gridrow="45" pageId="4" pageNumber="244">upper petals laterally overtopping the</td>
<td id="76CEA13E997A40972376B6AFFB0FF87B" box="[1077,1257,2007,2028]" gridcol="4" gridrow="45" pageId="4" pageNumber="244">acuminate</td>
<td id="76CEA13E997A40972251B6AFFA7DF87B" box="[1298,1435,2007,2028]" gridcol="5" gridrow="45" pageId="4" pageNumber="244">lower lateral</td>
</tr>
<tr id="351FC842997A409727D4B683FA7DF787" box="[151,1435,2043,2064]" gridrow="46" pageId="4" pageNumber="244" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td id="76CEA13E997A409726B6B683FC87F787" box="[501,865,2043,2064]" gridcol="2" gridrow="46" pageId="4" pageNumber="244">lower petals</td>
<td id="76CEA13E997A40972251B683FA7DF787" box="[1298,1435,2043,2064]" gridcol="5" gridrow="46" pageId="4" pageNumber="244">petals</td>
</tr>
</table>
</paragraph>
<paragraph id="8B90CA00997BBF6D27CBB1D8FAA4FCD6" blockId="5.[136,1452,160,833]" pageId="5" pageNumber="245">
Decumbent to slightly ascendant, perennial herb, up to 20(25) cm high. Stems strongly branching, rooting at and in between the lower nodes. Plant entirely glabrous. Leaves spirally arranged, deep green above, pale green below; the main veins, petioles and inflorescences reddish; the petiole base decurrent, developing into cork ridges at older stems; the petiole
<quantity id="4CD767E5997BBF6D2644B074FE64FEB2" box="[263,386,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="5" pageNumber="245" unit="mm" value="25.0" valueMax="40.0" valueMin="10.0">1040 mm</quantity>
long, continuously growing. Lamina (60)
<quantity id="4CD767E5997BBF6D2429B074FC12FEB2" box="[874,1012,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" metricValueMax="10.2" metricValueMin="8.0" pageId="5" pageNumber="245" unit="mm" value="91.0" valueMax="102.0" valueMin="80.0">80102 mm</quantity>
x (30)
<quantity id="4CD767E5997BBF6D230EB074FB2DFEB2" box="[1101,1227,268,293]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.2" metricValueMin="3.8" pageId="5" pageNumber="245" unit="mm" value="45.0" valueMax="52.0" valueMin="38.0">3852 mm</quantity>
, leathery, elliptic to lanceolate, the apex acute, the basis shortly attenuate; lateral veins 45(6) pairs; margin crenate; extrafloral nectaries
<quantity id="4CD767E5997BBF6D27CBB02CFF03FEFA" box="[136,229,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="245" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
, linear, at the lamina margin between each pair of teeth and at the tip of the apex, half erect, prominent especially at young leaves. Inflorescence a (4)8 flowered raceme. Flowers white with mauve centre and wing-like pattern on the upper lateral petals and occasionally a pair of small pink dots on the upper part of the lower lateral petals. Peduncle
<quantity id="4CD767E5997BBF6D27CBB0B8FEE9FE4E" box="[136,271,448,473]" metricMagnitude="-2" metricUnit="m" metricValue="8.35" metricValueMax="13.0" metricValueMin="3.7" pageId="5" pageNumber="245" unit="mm" value="83.5" valueMax="130.0" valueMin="37.0">37130 mm</quantity>
long, upright. Bracts
<quantity id="4CD767E5997BBF6D2541B0B8FDB9FE4E" box="[514,607,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="245" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
x
<quantity id="4CD767E5997BBF6D2538B0B8FD3CFE4E" box="[635,730,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="245" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
, ovate-lanceolate. Pedicels
<quantity id="4CD767E5997BBF6D234DB0B8FB9EFE4E" box="[1038,1144,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="5" pageNumber="245" unit="mm" value="8.5" valueMax="11.0" valueMin="6.0">611 mm</quantity>
long, slender, continuously growing until fruit ripeness to 1215(20) mm. Lateral sepals 2,
<quantity id="4CD767E5997BBF6D241EB09CFC5CFE6A" box="[861,954,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="245" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
x
<quantity id="4CD767E5997BBF6D2495B09CFBACFE6A" box="[982,1098,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="245" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
, ovate-lanceolate, acute. Lower sepal
<quantity id="4CD767E5997BBF6D278BB370FEC3FDB6" box="[200,293,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="245" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long, 46(7) mm deep, obliquely naviculate, white, abruptly peaking into a light greenish,
<quantity id="4CD767E5997BBF6D2264B370FA8EFDB6" box="[1319,1384,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="245" unit="mm" value="1.0">1 mm</quantity>
down facing tip and abruptly constricted into an (8)
<quantity id="4CD767E5997BBF6D25D3B354FCEAFDD2" box="[656,780,556,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="5" pageNumber="245" unit="mm" value="15.0" valueMax="17.0" valueMin="13.0">1317 mm</quantity>
, broad, curved, filiform, white, pinkish overflown spur with obtuse apex. Dorsal petal
<quantity id="4CD767E5997BBF6D26EAB328FDE0FDFE" box="[425,518,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="245" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
x (3)
<quantity id="4CD767E5997BBF6D250CB328FD49FDFE" box="[591,687,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="245" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, ovate-lanceolate, acute, helmet-like, folded, dorsally with a narrow, light greenish crest. Lateral united petals (10)
<quantity id="4CD767E5997BBF6D25D0B30CFCEBFD1A" box="[659,781,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="5" pageNumber="245" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
long, with petals different in size and shape; upper petal 9 10(11) mm x 34(5) mm, falcate to narrowly-oblong; lower petal 910(12) mm x
<quantity id="4CD767E5997BBF6D2303B3E0FB78FD26" box="[1088,1182,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="245" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
, suborbicular to oblong, distally drawn out along the inner margin into a
<quantity id="4CD767E5997BBF6D25F2B3C4FCE9FD42" box="[689,783,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="245" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
x
<quantity id="4CD767E5997BBF6D246DB3C4FC6AFD42" box="[814,908,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="245" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
ligulate appendage. Appendages overlapping to spreading. Ovary glabrous. Fruit
<quantity id="4CD767E5997BBF6D26BEB398FD91FD6E" box="[509,631,736,761]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="5" pageNumber="245" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 mm</quantity>
x
<quantity id="4CD767E5997BBF6D25D7B398FD15FD6E" box="[660,755,736,761]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="245" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, fusiform, glabrous, producing 521 ovate, smooth seeds of
<quantity id="4CD767E5997BBF6D22DDB398FF52FC8A" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="245" unit="mm" value="2.0">2 mm</quantity>
x
<quantity id="4CD767E5997BBF6D278CB27CFEE9FC8A" box="[207,271,772,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="245" unit="mm" value="1.0">1 mm</quantity>
having a tuff of numerous, dead, spiral hairs at the hilum and few additional ones at the narrow side of the seeds. Hairs tear along the spirally thickened cell walls developing into relatively tear-resistant, spiral fibres.
</paragraph>
<caption id="DF509A88997BBF6D27CBB6A8FC83F872" ID-DOI="http://doi.org/10.5281/zenodo.13671410" ID-Zenodo-Dep="13671410" box="[136,869,2000,2021]" httpUri="https://zenodo.org/record/13671410/files/figure.png" pageId="5" pageNumber="245" startId="5.[136,229,2000,2021]" targetBox="[151,1436,857,1974]" targetPageId="5" targetType="figure">
<paragraph id="8B90CA00997BBF6D27CBB6A8FC83F872" blockId="5.[136,869,2000,2021]" box="[136,869,2000,2021]" pageId="5" pageNumber="245">
<emphasis id="B95B1612997BBF6D27CBB6A8FF18F872" bold="true" box="[136,254,2000,2021]" pageId="5" pageNumber="245">FIGURE 4.</emphasis>
Holotype specimen of
<taxonomicName id="4C2FB183997BBF6D269EB6A8FD27F872" box="[477,705,2000,2021]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="245" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B1612997BBF6D269EB6A8FD27F872" box="[477,705,2000,2021]" italics="true" pageId="5" pageNumber="245">Impatiens pinganoensis</emphasis>
</taxonomicName>
, phot. B. Ditsch.
</paragraph>
</caption>
<caption id="DF509A889978BF6E27CBB6D2FCCCF828" ID-DOI="http://doi.org/10.5281/zenodo.13671414" ID-Zenodo-Dep="13671414" box="[136,810,1962,1983]" httpUri="https://zenodo.org/record/13671414/files/figure.png" pageId="6" pageNumber="246" startId="6.[136,229,1962,1983]" targetBox="[189,1398,191,1937]" targetPageId="6" targetType="figure">
<paragraph id="8B90CA009978BF6E27CBB6D2FCCCF828" blockId="6.[136,810,1962,1983]" box="[136,810,1962,1983]" pageId="6" pageNumber="246">
<emphasis id="B95B16129978BF6E27CBB6D2FF1BF828" bold="true" box="[136,253,1962,1983]" pageId="6" pageNumber="246">FIGURE 5.</emphasis>
AC, Habitat of
<taxonomicName id="4C2FB1839978BF6E26E1B6D2FD60F828" box="[418,646,1962,1983]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="246" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B16129978BF6E26E1B6D2FD60F828" box="[418,646,1962,1983]" italics="true" pageId="6" pageNumber="246">Impatiens pinganoensis</emphasis>
</taxonomicName>
, phot. B. Ditsch.
</paragraph>
</caption>
<caption id="DF509A889979BF6F27CBB4C3FE63FA63" ID-DOI="http://doi.org/10.5281/zenodo.13671418" ID-Zenodo-Dep="13671418" httpUri="https://zenodo.org/record/13671418/files/figure.png" pageId="7" pageNumber="247" startId="7.[136,229,1467,1488]" targetBox="[151,1436,190,1442]" targetPageId="7" targetType="figure">
<paragraph id="8B90CA009979BF6F27CBB4C3FE63FA63" blockId="7.[136,1452,1467,1524]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27CBB4C3FF18FA47" bold="true" box="[136,254,1467,1488]" pageId="7" pageNumber="247">FIGURE 6.</emphasis>
AD, Transections of the cork ridges at the older stems showing numerous rhaphides, phot. E. Fischer. A: Scale bar 1 mm, BC: 0.1 mm, D: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B90CA009979BF6F27FEB730FD4FF912" blockId="7.[136,1452,1608,2065]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27FEB730FEC5F9F6" bold="true" box="[189,291,1608,1633]" pageId="7" pageNumber="247">Habitat:</emphasis>
—On clastic sedimentary rocks of quartz or other silicate minerals (Direcção dos serviços de geologia e minas 1980) at the spray zone of a creek,
<specimenCount id="9D2901899979BF6F2518B714FD4FF912" box="[603,681,1644,1669]" count="930" pageId="7" pageNumber="247" type="generic">
<quantity id="4CD767E59979BF6F2518B714FD43F912" box="[603,677,1644,1669]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="7" pageNumber="247" unit="m" value="930.0">930 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C335998B9979BF6F27FEB7E8FDD2F8AE" pageId="7" pageNumber="247" type="distribution">
<paragraph id="8B90CA009979BF6F27FEB7E8FC4CF95A" blockId="7.[136,1452,1608,2065]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27FEB7E8FEBFF93E" bold="true" box="[189,345,1680,1705]" pageId="7" pageNumber="247">Distribution:</emphasis>
<collectingCountry id="F3388A909979BF6F2635B7E8FE2CF93E" box="[374,458,1680,1705]" name="Angola" pageId="7" pageNumber="247">Angola</collectingCountry>
,
<collectingRegion id="49EB04E29979BF6F2695B7E8FDE9F93E" box="[470,527,1680,1705]" country="Angola" name="Uige" pageId="7" pageNumber="247">Uíge</collectingRegion>
, Serra do Pingano, Bem Vindo, only known from the
<typeStatus id="549474A29979BF6F232FB7E8FB7AF93E" box="[1132,1180,1680,1705]" pageId="7" pageNumber="247">type</typeStatus>
collection and the cliffs of a waterfall nearby (
<geoCoordinate id="EE1BACC79979BF6F26C2B7CCFDF0F95A" box="[385,534,1716,1741]" degrees="7" direction="south" minutes="40" orientation="latitude" pageId="7" pageNumber="247" precision="1" seconds="22.7" value="-7.672972">S 7°4022.7”</geoCoordinate>
,
<geoCoordinate id="EE1BACC79979BF6F2562B7CCFD23F95A" box="[545,709,1716,1741]" degrees="14" direction="east" minutes="56" orientation="longitude" pageId="7" pageNumber="247" precision="1" seconds="17.7" value="14.93825">E 14°5617.7”</geoCoordinate>
,
<quantity id="4CD767E59979BF6F2592B7CCFCFDF95A" box="[721,795,1716,1741]" metricMagnitude="2" metricUnit="m" metricValue="6.15" pageId="7" pageNumber="247" unit="m" value="615.0">615 m</quantity>
;
<figureCitation id="1314D6859979BF6F2464B7CCFC8EF95A" box="[807,872,1716,1741]" captionStart="FIGURE 3" captionStartId="3.[136,229,1586,1607]" captionTargetBox="[153,1433,193,1561]" captionTargetId="figure-18@3.[151,1436,190,1562]" captionTargetPageId="3" captionText="FIGURE 3. Distribution map of Impatiens pinganoensis." figureDoi="http://doi.org/10.5281/zenodo.13671408" httpUri="https://zenodo.org/record/13671408/files/figure.png" pageId="7" pageNumber="247">Fig. 3</figureCitation>
&amp;
<figureCitation id="1314D6859979BF6F24CFB7CCFC7AF95A" box="[908,924,1716,1741]" captionStart="FIGURE 5" captionStartId="6.[136,229,1962,1983]" captionTargetBox="[189,1398,191,1937]" captionTargetId="figure-14@6.[189,1399,190,1937]" captionTargetPageId="6" captionText="FIGURE 5. AC, Habitat of Impatiens pinganoensis, phot. B. Ditsch." figureDoi="http://doi.org/10.5281/zenodo.13671414" httpUri="https://zenodo.org/record/13671414/files/figure.png" pageId="7" pageNumber="247">5</figureCitation>
).
</paragraph>
<paragraph id="8B90CA009979BF6F27FEB7A0FDD2F8AE" blockId="7.[136,1452,1608,2065]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27FEB7A0FEA5F966" bold="true" box="[189,323,1752,1777]" pageId="7" pageNumber="247">Phenology:</emphasis>
—Flowers and fruits in its habitat during the rainy season (October to April). Data for the dry season are not available. In greenhouses in the Botanical Gardens of Bonn and Dresden, it flowers and fruits around the year, especially between April and October.
</paragraph>
</subSubSection>
<subSubSection id="C335998B9979BF6127FEB63CFF3FFE22" lastPageId="9" lastPageNumber="249" pageId="7" pageNumber="247" type="etymology">
<paragraph id="8B90CA009979BF6F27FEB63CFB75F8CA" blockId="7.[136,1452,1608,2065]" box="[189,1171,1860,1885]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27FEB63CFEAEF8CA" bold="true" box="[189,328,1860,1885]" pageId="7" pageNumber="247">Etymology:</emphasis>
—Named after the Serra do Pingano, a mountain range in northern
<collectingCountry id="F3388A909979BF6F237AB63CFB69F8CA" box="[1081,1167,1860,1885]" name="Angola" pageId="7" pageNumber="247">Angola</collectingCountry>
.
</paragraph>
<paragraph id="8B90CA009979BF6F27FEB610FC96F85E" blockId="7.[136,1452,1608,2065]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27FEB610FE28F816" bold="true" box="[189,462,1896,1921]" pageId="7" pageNumber="247">Conservation status:—</emphasis>
We suggest to categorize the new species as critically endangered (CR) since it has a very limited distribution range and occurs only in the cliffs of two waterfalls with a very special microclimate. This habitat is endangered by deforestation and a planned construction project.
</paragraph>
<paragraph id="8B90CA009979BF6F27FEB6ACFC6EF786" blockId="7.[136,1452,1608,2065]" pageId="7" pageNumber="247">
<emphasis id="B95B16129979BF6F27FEB6ACFD16F87A" bold="true" box="[189,752,2004,2029]" pageId="7" pageNumber="247">
Additional specimens examined (
<typeStatus id="549474A29979BF6F250AB6ACFD23F87A" box="[585,709,2004,2029]" pageId="7" pageNumber="247" type="paratype">paratypes</typeStatus>
):—
</emphasis>
<collectingCountry id="F3388A909979BF6F25B3B6ACFCA3F87A" box="[752,837,2004,2029]" name="Angola" pageId="7" pageNumber="247">Angola</collectingCountry>
,
<collectingRegion id="49EB04E29979BF6F2412B6ACFC6CF87A" box="[849,906,2004,2029]" country="Angola" name="Uige" pageId="7" pageNumber="247">Uíge</collectingRegion>
, Serra do Pingano, Bem Vindo,
<geoCoordinate id="EE1BACC79979BF6F23BBB6ACFA68F87A" box="[1272,1422,2004,2029]" degrees="7" direction="south" minutes="40" orientation="latitude" pageId="7" pageNumber="247" precision="1" seconds="22.7" value="-7.672972">S 7°4022.7”</geoCoordinate>
,
<geoCoordinate id="EE1BACC79979BF6F22D9B6ACFEF2F786" degrees="14" direction="east" minutes="56" orientation="longitude" pageId="7" pageNumber="247" precision="1" seconds="17.7" value="14.93825">E 14°5617.7”</geoCoordinate>
,
<quantity id="4CD767E59979BF6F2663B680FE81F786" box="[288,359,2040,2065]" metricMagnitude="2" metricUnit="m" metricValue="6.15" pageId="7" pageNumber="247" unit="m" value="615.0">615 m</quantity>
Anne Göhre 25/2015 af, P!, B!, BR!, BONN!).
</paragraph>
<caption id="DF509A889976BF6027CBB552FC9BFBA8" ID-DOI="http://doi.org/10.5281/zenodo.13671422" ID-Zenodo-Dep="13671422" box="[136,893,1066,1087]" httpUri="https://zenodo.org/record/13671422/files/figure.png" pageId="8" pageNumber="248" startId="8.[136,229,1066,1087]" targetBox="[151,1436,190,1041]" targetPageId="8" targetType="figure">
<paragraph id="8B90CA009976BF6027CBB552FC9BFBA8" blockId="8.[136,893,1066,1087]" box="[136,893,1066,1087]" pageId="8" pageNumber="248">
<emphasis id="B95B16129976BF6027CBB552FF18FBA8" bold="true" box="[136,254,1066,1087]" pageId="8" pageNumber="248">FIGURE 7.</emphasis>
Pollinator (
<taxonomicName id="4C2FB1839976BF602633B552FE28FBA8" box="[368,462,1066,1087]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="248" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95B16129976BF602633B552FE4AFBA8" box="[368,428,1066,1087]" italics="true" pageId="8" pageNumber="248">Empis</emphasis>
sp.
</taxonomicName>
) of
<taxonomicName id="4C2FB1839976BF6026B6B552FD3FFBA8" box="[501,729,1066,1087]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B16129976BF6026B6B552FD3FFBA8" box="[501,729,1066,1087]" italics="true" pageId="8" pageNumber="248">Impatiens pinganoensis</emphasis>
</taxonomicName>
, phot. B. Ditsch.
</paragraph>
</caption>
<caption id="DF509A889976BF6027CBB726FAB9F9E4" ID-DOI="http://doi.org/10.5281/zenodo.13671426" ID-Zenodo-Dep="13671426" box="[136,1375,1630,1651]" httpUri="https://zenodo.org/record/13671426/files/figure.png" pageId="8" pageNumber="248" startId="8.[136,229,1630,1651]" targetBox="[466,1121,1113,1605]" targetPageId="8" targetType="figure">
<paragraph id="8B90CA009976BF6027CBB726FAB9F9E4" blockId="8.[136,1375,1630,1651]" box="[136,1375,1630,1651]" pageId="8" pageNumber="248">
<emphasis id="B95B16129976BF6027CBB726FF1BF9E4" bold="true" box="[136,253,1630,1651]" pageId="8" pageNumber="248">FIGURE 8.</emphasis>
Seed hairs with spirally thickened cell walls developing into relatively tear-resistant, spiral fibres, phot. M. Günther.
</paragraph>
</caption>
<paragraph id="8B90CA009976BF6027FEB7DBFD5AF803" blockId="8.[136,1452,1699,2084]" pageId="8" pageNumber="248">
The new species is part of the
<taxonomicName id="4C2FB1839976BF60256AB7DBFCE7F92B" box="[553,769,1699,1724]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B16129976BF60256AB7DBFCE7F92B" box="[553,769,1699,1724]" italics="true" pageId="8" pageNumber="248">Impatiens filicornu</emphasis>
</taxonomicName>
complex and closely related to
<taxonomicName id="4C2FB1839976BF602339B7DBFB13F92B" box="[1146,1269,1699,1724]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B16129976BF602339B7DBFB13F92B" box="[1146,1269,1699,1724]" italics="true" pageId="8" pageNumber="248">I. filicornu</emphasis>
</taxonomicName>
occurring from
<collectingCountry id="F3388A909976BF6027CBB7BFFF1BF977" box="[136,253,1735,1760]" name="Cameroon" pageId="8" pageNumber="248">Cameroon</collectingCountry>
to the
<collectingCountry id="F3388A909976BF602615B7BFFD22F977" box="[342,708,1735,1760]" name="Democratic Republic of the Congo" pageId="8" pageNumber="248">Democratic Republic of Congo</collectingCountry>
. The internal, phylogenetic relationships of this group of seven species have not been investigated, yet.
<taxonomicName id="4C2FB1839976BF602512B793FBBDF893" authority="sensu Grey-Wilson (1980)" authorityName="sensu Grey-Wilson" authorityYear="1980" box="[593,1115,1771,1796]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B16129976BF602512B793FCC1F893" box="[593,807,1771,1796]" italics="true" pageId="8" pageNumber="248">Impatiens filicornu</emphasis>
sensu
<bibRefCitation id="EFBEB7F19976BF602434B793FBBDF893" author="Grey-Wilson, C." box="[887,1115,1771,1796]" pageId="8" pageNumber="248" refId="ref3327" refString="Grey-Wilson, C. (1980) Impatiens of Africa. A. A. Balkema, Rotterdam." type="book" year="1980">Grey-Wilson (1980)</bibRefCitation>
</taxonomicName>
comprises apparently several undescribed taxa, which are currently studied by the authors. However,
<taxonomicName id="4C2FB1839976BF602486B677FB37F8BF" box="[965,1233,1807,1832]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B16129976BF602486B677FB37F8BF" box="[965,1233,1807,1832]" italics="true" pageId="8" pageNumber="248">Impatiens pinganoensis</emphasis>
</taxonomicName>
, the southernmost, local endemic representative differs from all other species of the
<taxonomicName id="4C2FB1839976BF602432B64BFC0CF8DB" authorityName="sensu Grey-Wilson" authorityYear="1980" box="[881,1002,1843,1868]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B16129976BF602432B64BFC0CF8DB" box="[881,1002,1843,1868]" italics="true" pageId="8" pageNumber="248">I. filicornu</emphasis>
</taxonomicName>
complex by several characters (
<tableCitation id="C6ADFFBB9976BF602218B64BFA43F8DB" box="[1371,1445,1843,1868]" captionStart="TABLE 1" captionStartId="4.[136,216,268,289]" captionText="TABLE 1. Characters separating Impatiens pinganoensis from the other species of the core Impatiens filicornu complex." pageId="8" pageNumber="248">Tab. 1</tableCitation>
;
<figureCitation id="1314D6859976BF6027CBB62FFF2CF8E7" box="[136,202,1879,1904]" captionStart="FIGURE 6" captionStartId="7.[136,229,1467,1488]" captionTargetBox="[151,1436,190,1442]" captionTargetId="figure-18@7.[151,1436,190,1442]" captionTargetPageId="7" captionText="FIGURE 6. AD, Transections of the cork ridges at the older stems showing numerous rhaphides, phot. E. Fischer. A: Scale bar 1 mm, BC: 0.1 mm, D: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13671418" httpUri="https://zenodo.org/record/13671418/files/figure.png" pageId="8" pageNumber="248">Fig. 6</figureCitation>
) and has probably been isolated from the populations of
<taxonomicName id="4C2FB1839976BF602404B62FFC59F8E7" authorityName="sensu Grey-Wilson" authorityYear="1980" box="[839,959,1879,1904]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="filicornu">
<emphasis id="B95B16129976BF602404B62FFC59F8E7" box="[839,959,1879,1904]" italics="true" pageId="8" pageNumber="248">I. filicornu</emphasis>
</taxonomicName>
, the closest relative further north during the drying up of tropical Africa since the last ice ages.
</paragraph>
<paragraph id="8B90CA009976BF6027FEB6E7FDFDF7B3" blockId="8.[136,1452,1699,2084]" pageId="8" pageNumber="248">
Cork ridges, first considered to be a unique character of
<taxonomicName id="4C2FB1839976BF60247BB6E7FBA4F82F" box="[824,1090,1951,1976]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B16129976BF60247BB6E7FBA4F82F" box="[824,1090,1951,1976]" italics="true" pageId="8" pageNumber="248">Impatiens pinganoensis</emphasis>
</taxonomicName>
, have already been observed by
<bibRefCitation id="EFBEB7F19976BF6027CBB6BBFEC4F84B" author="Weber, H." box="[136,290,1987,2012]" pageId="8" pageNumber="248" pagination="1 - 30" refId="ref4018" refString="Weber, H. (1949) I. Uber die Verbreitung von Spaltoffnungen an den Sprossachsen krautiger Pflanzen. In: Troll, W. &amp; Weber, H. (Eds.) Morphologische und anatomische Studien an hoheren Pflanzen Sitzungsberichte der Heidelberger Akademie der Wissenschaften Math. - Naturwiss. Klasse 6: 1 - 30." type="journal article" year="1949">Weber (1949)</bibRefCitation>
in
<taxonomicName id="4C2FB1839976BF602605B6BBFDE1F84B" authority="Hook. f." authorityName="Hook. f." box="[326,519,1987,2012]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="sultani">
<emphasis id="B95B16129976BF602605B6BBFE4CF84B" box="[326,426,1987,2012]" italics="true" pageId="8" pageNumber="248">I. sultani</emphasis>
Hook.f.
</taxonomicName>
(1882: pl. 6643) (=
<taxonomicName id="4C2FB1839976BF6025A8B6BBFB81F84B" authority="Hook. f. (1868: 302)" authorityName="Hook. f." authorityPageNumber="302" authorityYear="1868" box="[747,1127,1987,2012]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="248" phylum="Tracheophyta" rank="species" species="walleriana">
<emphasis id="B95B16129976BF6025A8B6BBFC9AF84B" box="[747,892,1987,2012]" italics="true" pageId="8" pageNumber="248">I. walleriana</emphasis>
Hook.f. (1868: 302)
</taxonomicName>
.
<bibRefCitation id="EFBEB7F19976BF602332B6BBFAEDF84B" author="Weber, H." box="[1137,1291,1987,2012]" pageId="8" pageNumber="248" pagination="1 - 30" refId="ref4018" refString="Weber, H. (1949) I. Uber die Verbreitung von Spaltoffnungen an den Sprossachsen krautiger Pflanzen. In: Troll, W. &amp; Weber, H. (Eds.) Morphologische und anatomische Studien an hoheren Pflanzen Sitzungsberichte der Heidelberger Akademie der Wissenschaften Math. - Naturwiss. Klasse 6: 1 - 30." type="journal article" year="1949">Weber (1949)</bibRefCitation>
states that the small, weak and irregular cork ridges are formed by the epidermis by transforming the epidermis cells to phellogene which protrude towards the outside.
</paragraph>
<paragraph id="8B90CA009977BF6127FEB1D8FBA8FEB2" blockId="9.[136,1452,160,437]" pageId="9" pageNumber="249">
During field observations in the Serra do Pingano a long-proboscid fly (
<taxonomicName id="4C2FB1839977BF61248FB1D8FBDFFF2E" box="[972,1081,160,185]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95B16129977BF61248FB1D8FBF4FF2E" box="[972,1042,160,185]" italics="true" pageId="9" pageNumber="249">Empis</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C2FB1839977BF612300B1D8FB26FF2E" box="[1091,1216,160,185]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="family">Empididae</taxonomicName>
;
<figureCitation id="1314D6859977BF61238AB1D8FAEFFF2E" box="[1225,1289,160,185]" captionStart="FIGURE 7" captionStartId="8.[136,229,1066,1087]" captionTargetBox="[151,1436,190,1041]" captionTargetId="figure-14@8.[151,1436,190,1041]" captionTargetPageId="8" captionText="FIGURE 7. Pollinator (Empis sp.) of Impatiens pinganoensis, phot. B. Ditsch." figureDoi="http://doi.org/10.5281/zenodo.13671422" httpUri="https://zenodo.org/record/13671422/files/figure.png" pageId="9" pageNumber="249">Fig. 7</figureCitation>
) was observed legitimately visiting and pollinating the flowers of
<taxonomicName id="4C2FB1839977BF612593B1BCFC3AFF4A" box="[720,988,196,221]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="249" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B16129977BF612593B1BCFC3AFF4A" box="[720,988,196,221]" italics="true" pageId="9" pageNumber="249">Impatiens pinganoensis</emphasis>
</taxonomicName>
. Additionally, pollination by lepidoptera seems also possible, since both pollinator groups visit flowers with the same flower morphology, which has already been observed for
<taxonomicName id="4C2FB1839977BF61261BB074FDB7FEB2" box="[344,593,268,293]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="249" phylum="Tracheophyta" rank="species" species="hochstetteri">
<emphasis id="B95B16129977BF61261BB074FDB7FEB2" box="[344,593,268,293]" italics="true" pageId="9" pageNumber="249">Impatiens hochstetteri</emphasis>
</taxonomicName>
in
<collectingCountry id="F3388A909977BF612536B074FCE2FEB2" box="[629,772,268,293]" name="South Africa" pageId="9" pageNumber="249">South Africa</collectingCountry>
(
<bibRefCitation id="EFBEB7F19977BF612451B074FBA5FEB2" author="Potgieter, C. J. &amp; Edwards, T. J." box="[786,1091,268,293]" pageId="9" pageNumber="249" pagination="254 - 267" refId="ref3941" refString="Potgieter, C. J. &amp; Edwards, T. J. (2005) The Stenobasipteron wiedemanni (Diptera, Nemestrinidae) pollination guild in eastern southern Africa. Annals of the Missouri Botanical Garden 92: 254 - 267." type="journal article" year="2005">Potgieter &amp; Edwards 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8B90CA009977BF6127FEB048FF3FFE22" blockId="9.[136,1452,160,437]" pageId="9" pageNumber="249">
Cultivating the plants, the hairs at the hilum and at the narrow side of the seeds were observed to be tearing along the spirally thickened cell walls developing into relatively tear-resistant, spiral fibres (
<figureCitation id="1314D6859977BF612305B02CFB6FFEFA" box="[1094,1161,340,365]" captionStart="FIGURE 8" captionStartId="8.[136,229,1630,1651]" captionTargetBox="[466,1121,1113,1605]" captionTargetId="figure-29@8.[466,1121,1113,1605]" captionTargetPageId="8" captionText="FIGURE 8. Seed hairs with spirally thickened cell walls developing into relatively tear-resistant, spiral fibres, phot. M. Günther." figureDoi="http://doi.org/10.5281/zenodo.13671426" httpUri="https://zenodo.org/record/13671426/files/figure.png" pageId="9" pageNumber="249">Fig. 8</figureCitation>
), which probably serve to attach the seeds to passing distributers (
<bibRefCitation id="EFBEB7F19977BF61257EB000FD23FE06" author="Hilger, H. H." box="[573,709,376,401]" pageId="9" pageNumber="249" pagination="333 - 356" refId="ref3410" refString="Hilger, H. H. (1983) Epizoochory dominates in ground layer of grazed evergreen mountain rain forest of mount Kulal north Kenya. Beitrage zur Biologie der Pflanzen 1983: 333 - 356." type="journal article" year="1983">Hilger 1983</bibRefCitation>
) as well as to the stones of the cliffs where
<taxonomicName id="4C2FB1839977BF6123E0B000FA4AFE06" box="[1187,1452,376,401]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="249" phylum="Tracheophyta" rank="species" species="pinganoensis">
<emphasis id="B95B16129977BF6123E0B000FA4AFE06" box="[1187,1452,376,401]" italics="true" pageId="9" pageNumber="249">Impatiens pinganoensis</emphasis>
</taxonomicName>
occurs.
</paragraph>
</subSubSection>
</treatment>
</document>