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<document id="76DED84F0AAE13938E65D0909EEFB9C5" ID-DOI="10.3897/zookeys.1210.124121" ID-ZooBank="BDD76A5D-A8B4-4F01-9D1B-4BDC9795876D" ID-publisher-id="124121" URI-arpha="57CE6668-C138-5D0B-BE14-880C6C782B2E" URI-zoobank="http://zoobank.org/BDD76A5D-A8B4-4F01-9D1B-4BDC9795876D" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1724943203130" checkinUser="pensoft" docAuthor="Miller, Kelly B., Michat, Mariano C. &amp; Ferreira Jr, Nelson" docDate="2024" docId="946585B2F35C5E65B3C0BCB61748D5D3" docLanguage="en" docName="ZooKeys 1210: 325-332" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/124121/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nilssondytes diversus Miller, Michat &amp; Jr, 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="332" masterDocId="57CE6668C1385D0BBE14880C6C782B2E" masterDocTitle="Description of Nilssondytes diversus gen. et sp. nov. from Venezuela (Coleoptera, Adephaga, Dytiscidae, Cybistrinae)" masterLastPageNumber="332" masterPageNumber="325" pageNumber="325" updateTime="1724944862994" updateUser="admin">
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<mods:title id="73E048149805E5E7298DAC34C201D1F5">Description of Nilssondytes diversus gen. et sp. nov. from Venezuela (Coleoptera, Adephaga, Dytiscidae, Cybistrinae)</mods:title>
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<mods:namePart id="EE7FAAD48AD597D6E5D6D5F299E8D1C7">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="2C99468F2C6B76B9502BD26B51183E22">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
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<mods:namePart id="FFF990418709949432A18888B24D7F86">Michat, Mariano C.</mods:namePart>
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<mods:affiliation id="34D85BDC66F853FABAB1C42E888D1193">Instituto de Biodiversidad y Biología Experimental y Aplicada, CONICET-Universidad de Buenos Aires, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="E0A6A036A24A0BCBC8DDD154F017295D">Ferreira Jr, Nelson</mods:namePart>
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<mods:affiliation id="9BF38ED8418BB7562D40F50C8B2074E3">Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:date id="B5C4900013802C345E8A3407158FAA25">2024</mods:date>
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<subSubSection id="CC7A1A7045D39C979AA60E9849DF9460" type="nomenclature">
<paragraph id="358345AEA6A7C36F589B703885281C7A">
<taxonomicName id="14390727C3E43926539D8B06A2F20929" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="diversus" status="sp. nov.">
<emphasis id="9161BE5EB4D592A6C4B37F8382788A4F" italics="true">Nilssondytes diversus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58113EFF563AAC39D6110DC2DA23D816" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="670327691EE850DA9352E08103B5DC42" type="description">
<paragraph id="D2D194DF4C92D39F13439E483BA3B7E7">
<figureCitation id="5D77982BB8A8851622E50CA8430F4B5C" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">Figs 110</figureCitation>
,
<figureCitation id="49A8BC5B72CF89F104172B2F5B2B32DB" captionStart="Figure 11" captionStartId="F2" captionText="Figure 11. Distribution of Nilssondytes diversus sp. nov. in northern South America." figureDoi="10.3897/zookeys.1210.124121.figure11" httpUri="https://binary.pensoft.net/fig/1123262">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKAAE" type="remarks">
<paragraph id="2F3775C5DFF2C0025EFFC924B03C500E">
<heading id="81805844EC015CF2DBE582DE09B4938B" reason="title">Remark.</heading>
</paragraph>
<paragraph id="BD09D252C0A1CE94CDDF277F7898926F">
The following description is reproduced with some modifications from
<bibRefCitation id="DDF3BB23535B52430D9E1DC7B4310E4A" DOI="10.3897/zookeys.1188.110081" author="Miller KB &amp; Michat MC &amp; Ferreira Jr N" etAl="et al." firstAuthor="Miller" journalOrPublisher="ZooKeys" pagination="125-168" refId="B7" refString="Miller KB, Michat MC, Ferreira Jr N (2024) Reclassification of Cybistrinae Sharp, 1880 in the Neotropical Region (Coleoptera, Adephaga, Dytiscidae), with description of new taxa. ZooKeys 1188: 125168. https://doi.org/10.3897/zookeys.1188.110081" title="Reclassification of Cybistrinae Sharp, 1880 in the Neotropical Region (Coleoptera, Adephaga, Dytiscidae), with description of new taxa." volume="1188" year="2024">Miller et al. (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUAAE" type="type locality">
<paragraph id="C9F0B9C0AD7B9B42374B699822E96E55">
<heading id="5AAD1E01C47C5E357C39666D83825305" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="E251A8A73EDCB6E005ABF8B85E098031">
<collectingCountry id="363F1BE7CD71D4DED5B4A13D9E6F4975" name="Venezuela">Venezuela</collectingCountry>
,
<collectingRegion id="0C69008AE958E74997F654F930F3C933" country="Venezuela" name="Amazonas">Amazonas State</collectingRegion>
, roadside pond ca.
<quantity id="BACFEEE5DEECE06A217E637F2748B99E" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
S Samariapo
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<geoCoordinate id="68909E681C6759CF75B808212CDA5C2C" degrees="5" direction="north" minutes="10.900" orientation="latitude" precision="1" value="5.181667">5 ° 10.900 ' N</geoCoordinate>
,
<geoCoordinate id="9656519EEC4827EAAB0A343B18240ED2" degrees="67" direction="west" minutes="46.078" orientation="longitude" precision="1" value="-67.76797">67 ° 46.078 ' W</geoCoordinate>
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,
<quantity id="BE2B6F55779B9766D17F47A22E23F6E5" metricMagnitude="1" metricUnit="m" metricValue="9.5" unit="m" value="95.0">95 m</quantity>
elev.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECBAE" type="diagnosis">
<paragraph id="25F8549894DD6F564F31021E90F66290">
<heading id="2B19E339442501905D978852FF38A54A" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="AFAAF5AC563C982DF87F7E0C2A244A8F">
This is the only species in the genus and is characterized by its diagnostic combination (see above). Typically, species-level features include the shape of the male median lobe which is unique. In ventral aspect the apex is abruptly constricted with the apex narrowly truncate with laterally pointed processes (Fig.
<figureCitation id="FE6DE81752CCDC6E83775552806785EA" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">6</figureCitation>
). In lateral aspect, the median lobe is moderately evenly curved on the dorsal margin, apically abruptly narrowed with the apex elongate and slender, apex narrowly rounded (Fig.
<figureCitation id="78F9780DBB8FA83C2C073E5A3F9B64EC" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQBAE" type="description">
<paragraph id="5408FFCFE10087DE404A6C011AA13633">
<heading id="9F38425855F91BC245236C9E5C32DF6B" reason="title">Description.</heading>
</paragraph>
<paragraph id="9559000F2B91E6D7D65903ECE972F073">
<emphasis id="D09401ED4D8186AD141054E548CBD6F7" bold="true" italics="true">Measurements.</emphasis>
<abbrev id="ABBRID0E2BAE" xlink_title="total length">TL</abbrev>
=
<quantity id="9C20425E7E8B6A8AB93E2552513659FA" metricMagnitude="-2" metricUnit="m" metricValue="1.8049999999999997" metricValueMax="1.94" metricValueMin="1.67" unit="mm" value="18.049999999999997" valueMax="19.4" valueMin="16.7">16.719.4 mm</quantity>
,
<abbrev id="ABBRID0E6BAE" xlink_title="greatest width across elytra">GW</abbrev>
=
<quantity id="B27E07660CD8A1C4D2719EC89C3CDC68" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.0699999999999998" metricValueMin="0.96" unit="mm" value="10.149999999999999" valueMax="10.7" valueMin="9.6">9.610.7 mm</quantity>
,
<abbrev id="ABBRID0EDCAE" xlink_title="greatest pronotal width">PW</abbrev>
= 7.0
<quantity id="723804EDDE7416B5EF3B2058F91E10D1" metricMagnitude="-3" metricUnit="m" metricValue="8.1" unit="mm" value="8.1">8.1 mm</quantity>
,
<abbrev id="ABBRID0EHCAE" xlink_title="greatest width of the head">HW</abbrev>
=
<quantity id="14A6A6A196EE44DB16285C2D2AD4087C" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.7" metricValueMin="4.2" unit="mm" value="4.45" valueMax="4.7" valueMin="4.2">4.24.7 mm</quantity>
,
<abbrev id="ABBRID0ELCAE" xlink_title="distance between the eyes">EW</abbrev>
=
<quantity id="0CA83C22B38432705546C867ADE1CCDB" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
,
<abbrev id="ABBRID0EPCAE" xlink_title="total length">TL</abbrev>
/
<abbrev id="ABBRID0ETCAE" xlink_title="greatest width across elytra">GW</abbrev>
= 1.71.8,
<abbrev id="ABBRID0EXCAE" xlink_title="greatest width of the head">HW</abbrev>
/
<abbrev id="ABBRID0E2CAE" xlink_title="distance between the eyes">EW</abbrev>
= 1.61.7, WC / WV = 3.13.2. Body shape suboval, slightly expanded posteriorly, widest at ~ 3 / 5 of length (Fig.
<figureCitation id="AEEA59268E554A75ECFE86F6332C4D67" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">1</figureCitation>
); lateral margins evenly, continuously curved between pronotum and elytron. Depressed and somewhat flattened in lateral aspect.
</paragraph>
<paragraph id="15A525F4B462ECFC8A1F6863638C1E05">
<emphasis id="E11286BF758E4996A90218AE5FBC9E04" bold="true" italics="true">Coloration</emphasis>
(Fig.
<figureCitation id="AFC57162242E7211666DF61A9082F050" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">1</figureCitation>
). Head dark green, anterior clypeal margin yellow, more so laterally, testaceous near eyes. Pronotum dark green with broad lateral yellow margin, posteriorly interrupted and green in three of the four examined specimens, in other specimen yellow extending to posterior angle. Elytron dark green with broad lateral yellow band, separated narrowly from lateral margin, slightly expanded near apex. Ventral surfaces largely black, testaceous on head, basal leg segments and elytral epipleuron.
</paragraph>
<paragraph id="160D6E676AAF76ED5BB798E445B808CF">
<emphasis id="3C50349F4B401958468CD132AC895884" bold="true">
<emphasis id="F52B8D6DACEEB7D6AAAAC3649F6F0C5E" bold="true" italics="true">Sculpture and structure</emphasis>
.
</emphasis>
Head broad, frontoclypeal lines elongate, straight, strongly oblique; anterior clypeal margin broadly, shallowly and evenly concave; dorsal surface evenly covered with fine microsculpture and micropunctures. Pronotum with lateral margins evenly and broadly curved; surface similar to surface of head in microsculpture and micropunctation. Elytral lateral margin evenly and slightly curved for most of length, apically broadly curved; surface of elytron similar to surface of head in microsculpture and micropunctation. Prosternal process apically rounded, ventral surface distinctly sulcate (Fig.
<figureCitation id="16694BD61E84C6069E488308CEE15AF3" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">4</figureCitation>
), apex robust, acutely pointed. Metaventral wing broad, slightly less than 1 / 3 width of lateral portion of metacoxa (WC / WV = 3.13.2); surface smooth, without sculpturing. Lateral portion of metacoxa large, broad, surface smooth, without sculpturing; metacoxal lines short, extending less than half distance across metacoxa. Abdominal ventrites smooth, unsculptured.
</paragraph>
<paragraph id="FFBBE214DC965EAC30CB92934E56256C">
<emphasis id="BD18CB3DFD574854A4F4E8E95B7AEBDC" bold="true">
<emphasis id="C01E396B2EEA3475330170BB86E521A4" bold="true" italics="true">Male genitalia</emphasis>
.
</emphasis>
Male median lobe in ventral aspect broad throughout most of length, apically abruptly narrowed, apex laterally produced, submedially with broad, elongate lobes on each side, ventral sclerite short, apically sharp, acuminate, extending to 3 / 5 length of median lobe, apex sharply pointed (Fig.
<figureCitation id="76D6D8B06292C58F7049AC8540A1B818" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">6</figureCitation>
). In lateral aspect shallowly curved, apically abruptly narrowed, apex narrowed, slightly curved, apically narrowly rounded, broad medially (Fig.
<figureCitation id="EAC92098F81E66AEBDA92854A66B946B" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">7</figureCitation>
). Lateral lobe broad in basal half, apically distinctly narrowed, apex narrowly rounded, with series of elongate setae along more than apical half of dorsal margin of lateral lobe (Fig.
<figureCitation id="1BBCF0375E99AA5215D6F0B7D106102D" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">8</figureCitation>
).
</paragraph>
<caption id="38D9468D03AAB77B33AC6101B7EFEE39" ID-DOI="10.3897/zookeys.1210.124121.figures1-10" ID-arpha="4348A308-B567-5496-932B-D21B3E61F535" httpUri="https://binary.pensoft.net/fig/1123261" startId="F1">
<paragraph id="26E59B2260B6A273B8C5D92403DAAD67">
<label id="4F94A5910C6DFB4FAA8F0117B181EF95">Figures 110.</label>
</paragraph>
<paragraph id="659EC2EE77747E166BDC7095E7CEA110">
<taxonomicName id="88C0962DE91C8F978EF9C1CE558BACC9" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="diversus" status="sp. nov.">
<emphasis id="40A418EAB51FDAA6DAE366C6FB0F153F" italics="true">Nilssondytes diversus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51EE3B1008F19EEEA40CE06B795E1271" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="611BFE57C6AFF0745A88F4941E4F022B" bold="true">1</emphasis>
dorsal habitus
<emphasis id="8202892CCE0CCC0AB1C8255B44579A6C" bold="true">2, 3</emphasis>
metatarsal claws, posterior aspect:
<emphasis id="3B4F286D729CD8C2A3F379C0FEDF28D6" bold="true">2</emphasis>
male
<emphasis id="083CADF948E2FDDABEE16DC37F21E309" bold="true">3</emphasis>
female
<emphasis id="925D222BA296A27C6C802DD1CE031C96" bold="true">4</emphasis>
prosternal process, ventral aspect
<emphasis id="3F3A1A219D66E7743244CFFDE8BE5D66" bold="true">5</emphasis>
right mesothoracic leg, posterior aspect
<emphasis id="8020149B8AB12C49FBCEEAB98383FFFE" bold="true">68</emphasis>
male genitalia:
<emphasis id="81DF1CC4E5EE45422F288BC6149E4EBB" bold="true">6</emphasis>
median lobe, ventral aspect
<emphasis id="AB17D796441C20F4ABDF6B09BBEB43C9" bold="true">7</emphasis>
median lobe, right lateral aspect
<emphasis id="D932C9EB2879F1F72AF4A880F0BA338D" bold="true">8</emphasis>
right lateral lobe, right lateral aspect
<emphasis id="07D80B3D8284DCFBE8DE7DB649D5CE48" bold="true">9</emphasis>
female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect
<emphasis id="31041BDF35D901255B805D684967AF92" bold="true">10</emphasis>
male sternite IX, ventral aspect. Scale bars: 5.0 mm (
<emphasis id="0FF9EFA5DAFEB024F8EC4ADA37BF85C1" bold="true">1</emphasis>
); 1.0 mm (
<emphasis id="2F6C8053358960101F1DBDACA426C5B5" bold="true">25, 9, 10</emphasis>
)
</paragraph>
</caption>
<caption id="BB8D1FFD5A3795A6D96CA67C2C4CEDB8" ID-DOI="10.3897/zookeys.1210.124121.figure11" ID-arpha="A1AEF094-ED38-52BB-BFBE-DADEA133F508" httpUri="https://binary.pensoft.net/fig/1123262" startId="F2">
<paragraph id="9E683F9E3A7AAFC0A6BD9765BA88E08A">
<label id="406DE4E5FA362E0A6C4149ADB5033945">Figure 11.</label>
</paragraph>
<paragraph id="6AABA214BA17E0C9D62FA505319FCA8A">
Distribution of
<taxonomicName id="E23B922F606A20FA302ADDA8C09C4DAB" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="diversus" status="sp. nov.">
<emphasis id="91F8B989A5215D01E8786315561472AC" italics="true">Nilssondytes diversus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5408348C40DB7B7BE8467473F841DEA" rank="species">sp. nov.</taxonomicNameLabel>
in northern South America.
</paragraph>
</caption>
<paragraph id="AC8983FBE6AEFCA1376B5C7BEF0B6259">
<emphasis id="1232B32CCD56DA6489DC3103999C3A8F" bold="true">
<emphasis id="76F09C98AE7EE4D3075FF7D1344841D3" bold="true" italics="true">Female genitalia</emphasis>
.
</emphasis>
With a single genital opening, vagina elongate, slender, with enlarged, bulbous region at base of common oviduct; spermatheca short, curved, at apex of enlarged region, with soft tissue region on each side of enlarged region (Fig.
<figureCitation id="BB8BB6A57B65DC62720E8116723A3D36" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">9</figureCitation>
); gonocoxae together broad, apically broadly pointed (Fig.
<figureCitation id="54D94D8EADB9F9E38BE86E3AE005DD82" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">9</figureCitation>
); gonocoxosternite broad, with elongate anterolateral lobe, with medial margin sublinear, without conspicuous setae (Fig.
<figureCitation id="8C75E7B2217B8AA5DC3A530D6B7A0946" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">9</figureCitation>
).
</paragraph>
<paragraph id="05F85DEC24CE56ED75E603172A644B33">
<emphasis id="E8E7EAED838381A9EFCCE7A968585C6E" bold="true">
<emphasis id="2A7419CE6E7BADF8CE6D8A96C96C00D7" bold="true" italics="true">Sexual dimorphism</emphasis>
.
</emphasis>
Males have a characteristic broad protarsal palette with ventral adhesive setae. Males also have mesotarsomeres with clumps of posteroventral setae. Females lack pro- and mesotarsal expansions or adhesive setae. Both males and females have two metatarsal claws with the posterior shorter than the anterior (Figs
<figureCitation id="1BF24B12202C5D9AFF7C5043A489588E" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">2</figureCitation>
,
<figureCitation id="D3145992C0155ED6C2355C1BD2171517" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">3</figureCitation>
), but females have the posterior somewhat more curved than in males (Fig.
<figureCitation id="673D7DC7D9BF71DE419B262508637845" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">3</figureCitation>
). Females have distinctive microsculpture on the surface of the elytron anteriorly in the form of a field of short striae which is absent in males.
</paragraph>
<paragraph id="25743E6A937F779797FA6980AC08B9BA">
<emphasis id="09F4983CDE188BCA3CA424E3B7214267" bold="true">
<emphasis id="2BC24F52715C505217BD8B458C89D4A4" bold="true" italics="true">Variation</emphasis>
.
</emphasis>
<specimenCount id="4CC79E1628417BA8BCC5365458ECF696" count="5" type="generic">Five specimens</specimenCount>
were examined.
<specimenCount id="F6996CBAA774412BC69A048969C328DC" count="1" type="generic">One specimen</specimenCount>
has the lateral pronotal yellow band extending to the posterior margin of the pronotum, the others have a narrow dark green separation from the posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFIAE" type="distribution">
<paragraph id="08C83C11DB464207FE9F7565C0742D5B">
<heading id="46856CD025AF788EA83ACFF50A1AC9B7" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="432401F0CCC6D7C6183CF9E1BE8FB66E">
This species is known from few localities in
<collectingCountry id="32D2BE6E639B10149DB97BDC6A845CCA" name="Venezuela">Venezuela</collectingCountry>
along the northwestern margins of the Guiana Shield craton (Fig.
<figureCitation id="24EC14A0160EAD845CB7FB3445C4DF14" captionStart="Figure 11" captionStartId="F2" captionText="Figure 11. Distribution of Nilssondytes diversus sp. nov. in northern South America." figureDoi="10.3897/zookeys.1210.124121.figure11" httpUri="https://binary.pensoft.net/fig/1123262">11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPIAE" type="natural history">
<paragraph id="7C77E32BFB8D7F63FA7AD2B777CE53B0">
<heading id="9CEC63ABD22BB2D6DAB804BF486CB1D7" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="D0C9A4C8A6761894C471A54730F7BDC8">The only natural history information available from labels is “ roadside pond, ” “ river margin, ” and “ rock outcropping. ”</paragraph>
</subSubSection>
<subSubSection id="SECID0EUIAE" type="etymology">
<paragraph id="086AC28787069276FDCE225FE0CEF49A">
<heading id="FC12DD1ECD707EBD51C52FB77F504EAB" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="6F44D7430E5C660DC2080E784FCDAEA7">
The species is named from the Latin
<taxonomicName id="3012747D4A110912EE882073FC75BB65" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="86B0B7958460E881E0464779212FF76C" italics="true">diversus</emphasis>
</taxonomicName>
, meaning “ different, ” in recognition of the different lengths of the metatarsal claws in both males and females (Figs
<figureCitation id="538F44AC5CFDA9D923066681F5B0D728" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">3</figureCitation>
,
<figureCitation id="83E65244BD3774A5A9D5FC1A4D1D11BE" captionStart="Figures 110" captionStartId="F1" captionText="Figures 110. Nilssondytes diversus sp. nov. 1 dorsal habitus 2, 3 metatarsal claws, posterior aspect: 2 male 3 female 4 prosternal process, ventral aspect 5 right mesothoracic leg, posterior aspect 68 male genitalia: 6 median lobe, ventral aspect 7 median lobe, right lateral aspect 8 right lateral lobe, right lateral aspect 9 female genitalia (gonocoxae, laterotergites, right gonocoxosternite and internal genitalia), ventral aspect 10 male sternite IX, ventral aspect. Scale bars: 5.0 mm (1); 1.0 mm (25, 9, 10)" figureDoi="10.3897/zookeys.1210.124121.figures1-10" httpUri="https://binary.pensoft.net/fig/1123261">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMJAE" type="material">
<paragraph id="75887F7F817E8B6E87E9B1B8CD585603">
<heading id="7A3A5A54EC5B20602E9F8DD2B35ABF88" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="699AF7497C17B54505DF6C1AEA6BB073">
<materialsCitation id="C5DF5596BB6790AEDF2539C315D57D3F" collectedFrom="roadside pond" collectingDate="2009-01-15" collectionCode="MIZA, KUNHM-ENT" collectorName="Short &amp; Miller &amp; Garcia &amp; Camacho &amp; Joly" country="Venezuela" elevation="95" latitude="5.181667" location="Amazonas State" longLatPrecision="1" longitude="-67.76797" specimenCode="SM 0846115" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas State" typeStatus="holotype">
<emphasis id="B1CF826F2BB0F335C10BBF023A363205" bold="true" italics="true">
<typeStatus id="DCCA95B69C04A063E7E0E57D32FACBF8" type="holotype">Holotype</typeStatus>
</emphasis>
,
<specimenCount id="F86829BFD2AC317BC5A5BE41F2366E0C" count="1" type="male">male</specimenCount>
deposited in
<named-content id="E133A727778B1C444DD08679A76EC93C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-central-de-venezuela" xlink_title="Museo del Instituto de Zoología Agrícola Francisco Fernández Yépez, Universidad Central de Venezuela, Maracay, Venezuela">
<collectionCode id="35E7F8C7454E6C1F138C58B1C0A42818" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" type="Museum">MIZA</collectionCode>
</named-content>
(see above) labeled, “
<collectingCountry id="22D932F3FB55B492E388F75BE31D8427" name="Venezuela">Venezuela</collectingCountry>
:
<collectingRegion id="EA9D2A0620CF0D42629AE0976BDB7D85" country="Venezuela" name="Amazonas">Amazonas State</collectingRegion>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5JAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-67.767967,5.181667]}">
<geoCoordinate id="861FBC44FC0D22D8C538CCA60C21E3A0" degrees="5" direction="north" minutes="10.900" orientation="latitude" precision="1" value="5.181667">5 ° 10.900 ' N</geoCoordinate>
,
<geoCoordinate id="96212033BC1C57D5B7097D9A57E5D240" degrees="67" direction="west" minutes="46.078" orientation="longitude" precision="1" value="-67.76797">67 ° 46.078 ' W</geoCoordinate>
</named-content>
,
<quantity id="20E9404AB577A58D176CB64F8197062C" metricMagnitude="1" metricUnit="m" metricValue="9.5" unit="m" value="95.0">
<elevation id="6CE28F0878F379AE016E0E5BE7062A31" metricMagnitude="1" metricUnit="m" metricValue="9.5" unit="m" value="95.0">95 m</elevation>
</quantity>
ca.
<locationDeviation id="40A57A790A43615F37DF12F1612FA617">
<quantity id="A2D33BD94F5D6E30F3BFD8B404A62083" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
S. Samariapo
</locationDeviation>
<date id="94489DCE608E83BC039DDD63A8D2AB5A" value="2009-01-15">
<collectingDate id="22F5F91EB24DB55C0FF80CACCA386133" value="2009-01-15">15. i. 2009</collectingDate>
</date>
; leg.
<collectorName id="77BF1DDB5D0ACD2AEA1004D6BD858E30">Short</collectorName>
,
<collectorName id="4A998932278769FA1C2DFDFCE5018D50">Miller</collectorName>
,
<collectorName id="495CC6440AC5A939042BABD847EAB74A">García</collectorName>
,
<collectorName id="C55E1730B0AA931BF9891D77C9BC2E58">Camacho</collectorName>
,
<collectorName id="692C9FA6FBB946113F8B7044D72491CF">Joly</collectorName>
VZ 09-0115 - 02 X:
<collectedFrom id="7C78EA6F962606A4A16D63CFDCF6E591">roadside pond</collectedFrom>
/
<specimenCode id="4E23AB175557B35F7F128C3C603EB9C7">SM 0846115</specimenCode>
<collectionCode id="5111C71AA77EEB4E6DA7A0C3A9A484C1">KUNHM-ENT</collectionCode>
[barcode label] / Holotype:
<originalDetermination id="032DA1A7BDEC70B07CDD11F58D2F748B">
<taxonomicName id="E0F8CE96A432671CCEB33A6DD8CB0BAD" authority="Miller, Michat" authorityName="Miller, Michat" class="Insecta" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="7E161D668F144F3EBE3C4661E9CAC7C0" italics="true">Nilssondytes diversus</emphasis>
Miller, Michat
</taxonomicName>
&amp; Ferreira-Jr., 2024
</originalDetermination>
[red label with double black line border]. ”
</materialsCitation>
<materialsCitation id="8F39B3B4D3DE3D8E0B7462B921FF958D" collectingDate="1900-04-20" collectionCode="SEMC" collectorName="E. A. Klages" location="Suapure Venez" municipality="Caura River" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="304C77DF30AB5CF4D1F1A47200C72785" bold="true" italics="true">
<typeStatus id="CAF385124B2386CA5F80A678449ECF31" type="paratype">Paratypes</typeStatus>
</emphasis>
,
<specimenCount id="C0FE36F0E5714888704EADDB9F50FD08" count="1" type="male">1 male</specimenCount>
(
<named-content id="502875090CE4A4F252DCE3BAE73E69A0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-kansas-biodiversity-institute" xlink_title="Snow Entomological Collection, University of Kansas, Lawrence, KS, USA">
<collectionCode id="69C40EFA197D482424882A658A7F978A" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" type="Museum">SEMC</collectionCode>
</named-content>
) labeled, “
<location id="212F8233195F8589DE7D7B728FB2F491" LSID="urn:lsid:plazi:treatment:946585B2F35C5E65B3C0BCB61748D5D3:212F8233195F8589DE7D7B728FB2F491" municipality="Caura River" name="Suapure Venez">Suapure Venez</location>
.
<collectingMunicipality id="015EE8B1189542168DBDED538929AAC2">Caura River</collectingMunicipality>
<date id="38B91803D80CEF777C03121CF8C534F4" value="1900-04-20">
<collectingDate id="D403A4DD0F71A4F6829C117E8C6918A5" value="1900-04-20">4.20. 1900</collectingDate>
</date>
[handwritten]
<collectorName id="5E250FBBA035B9BD94293F05AD82E9DE">E. A. Klages</collectorName>
. ”
</materialsCitation>
,
<materialsCitation id="9A5CE564B4A5A4E07EF7431433046F47" collectedFrom="rock outcropping" collectingDate="2008-08-05" collectionCode="MIZA, KUNMH-ENT" collectorName="A. Short &amp; M. Garcia" country="Venezuela" elevation="134" latitude="7.689889" location="Rio Aro" longLatPrecision="1" longitude="-64.03223" specimenCode="SM 0829328" specimenCount="1" specimenCount-female="1" stateProvince="Bolivar State">
<specimenCount id="ADA1EE45A2D4D672F0FACF99E5EB0CB2" count="1" type="female">1 female</specimenCount>
(
<named-content id="D8C84751698AC51665B5F08F308C81C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-central-de-venezuela" xlink_title="Museo del Instituto de Zoología Agrícola Francisco Fernández Yépez, Universidad Central de Venezuela, Maracay, Venezuela">
<collectionCode id="2C9D9D1E347F5A53B1AE9C0C1EBC28F0" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" type="Museum">MIZA</collectionCode>
</named-content>
) labeled “
<collectingCountry id="C8F027E9ADEDAEA125D7A7A5CAA52DCB" name="Venezuela">Venezuela</collectingCountry>
:
<collectingRegion id="040FCB47F9448F6C8BA406B3F11922EB" country="Venezuela" name="Bolivar">Bolivar State</collectingRegion>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3KAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-64.032222,7.689889]}">
<geoCoordinate id="CF0631602002C88A180223E2870339BA" degrees="7" direction="north" minutes="41" orientation="latitude" precision="1" seconds="23.6" value="7.689889">7 ° 41 ' 23.6 &quot; N</geoCoordinate>
,
<geoCoordinate id="1D4AEB1BDD499B711264E925938FAB29" degrees="64" direction="west" minutes="1" orientation="longitude" precision="1" seconds="56.0" value="-64.03223">64 ° 1 ' 56.0 &quot; W</geoCoordinate>
</named-content>
,
<quantity id="4482D1BE94434672F5B2A493E9844C0B" metricMagnitude="2" metricUnit="m" metricValue="1.34" unit="m" value="134.0">
<elevation id="5227340E70D0E1C2EECAE0651B912121" metricMagnitude="2" metricUnit="m" metricValue="1.34" unit="m" value="134.0">134 m</elevation>
</quantity>
ca.
<locationDeviation id="96368F59760D3149281231AEA2675AC1" bearing="90.0" distance="14000.0">
<quantity id="2067DA3C4B6F20FDCCE0A2731DAA44F5" metricMagnitude="4" metricUnit="m" metricValue="1.4" unit="km" value="14.0">14 km</quantity>
E
</locationDeviation>
<location id="2686FBAD20663A25D848D06717F5A4E1" LSID="urn:lsid:plazi:treatment:946585B2F35C5E65B3C0BCB61748D5D3:2686FBAD20663A25D848D06717F5A4E1" country="Venezuela" latitude="7.689889" longLatPrecision="1" longitude="-64.03223" name="Rio Aro" stateProvince="Bolivar State">Rio Aro</location>
;
<date id="083044564F0807224056B43078DB389D" value="2008-08-05">
<collectingDate id="7AB8B9EACA8056E37D778708F3FFB715" value="2008-08-05">5. viii. 2008</collectingDate>
</date>
leg.
<collectorName id="4F5425074663B36142181A0DDEA6CCAF">A. Short</collectorName>
$
<collectorName id="A86DD77D8B50DBFEEAA942D6FC2EA5B9">M. García</collectorName>
AS- 08-073;
<collectedFrom id="1DDF23B57B3BB5C525812433D124329B">rock outcropping</collectedFrom>
/
<specimenCode id="426EF1952EC9E4F486797B36697C8AA8">SM 0829328</specimenCode>
<collectionCode id="F0F93C95152C5B3D9861FE2C23DEA6FD">KUNMH-ENT</collectionCode>
[barcode label], ”
</materialsCitation>
<materialsCitation id="87FD4F014556DB45E66B5ABB32D1DCD0" collectingDate="2009-01-10" collectionCode="SEMC, KUNHM-ENT" collectorName="Short &amp; Miller &amp; Joly &amp; Garcia &amp; Camacho" country="Venezuela" elevation="52" latitude="8.103766" location="Rio Aguaro" longLatPrecision="1" longitude="-66.43713" municipality="San Nicolasito Field Station" specimenCode="SEMC 0852602" specimenCount="1" specimenCount-female="1" stateProvince="Guarico State">
<specimenCount id="26ADDD9D7D5EB8A00FD2807FE2903EB2" count="1" type="female">1 female</specimenCount>
(
<named-content id="BB960EB7E11B35620898AFB978E3F601" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-kansas-biodiversity-institute" xlink_title="Snow Entomological Collection, University of Kansas, Lawrence, KS, USA">
<collectionCode id="212E901AED1C3DC29119D2270EB6CD7A" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" type="Museum">SEMC</collectionCode>
</named-content>
) labeled “
<collectingCountry id="934D4057DCC8772D67379D9D404F5DBE" name="Venezuela">Venezuela</collectingCountry>
:
<collectingRegion id="F07C992B60712A9813A68F3FA3BC3170" country="Venezuela" name="Guarico">Guárico State</collectingRegion>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EILAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-66.437133,8.103767]}">
<geoCoordinate id="A18C9D49177EF3DF37F345D59E324985" degrees="8" direction="north" minutes="6.226" orientation="latitude" precision="1" value="8.103766">8 ° 6.226 ' N</geoCoordinate>
,
<geoCoordinate id="AC09806A351D7319E3C1044BCFCB5D79" degrees="66" direction="west" minutes="26.228" orientation="longitude" precision="1" value="-66.43713">66 ° 26.228 ' W</geoCoordinate>
</named-content>
,
<quantity id="F0D5B8EDFF3DAB1E13237363D937468C" metricMagnitude="1" metricUnit="m" metricValue="5.2" unit="m" value="52.0">
<elevation id="118FA1FB7A51885C25C871EBDDE7301D" metricMagnitude="1" metricUnit="m" metricValue="5.2" unit="m" value="52.0">52 m</elevation>
</quantity>
UCV
<collectingMunicipality id="A42D64D6E4E2DC4EE7DE43980D9AB294">San Nicolasito Field Station</collectingMunicipality>
:
<location id="BDB64C0001BC78FE0D92D6A0B8E4CED6" LSID="urn:lsid:plazi:treatment:946585B2F35C5E65B3C0BCB61748D5D3:BDB64C0001BC78FE0D92D6A0B8E4CED6" country="Venezuela" latitude="8.103766" longLatPrecision="1" longitude="-66.43713" municipality="San Nicolasito Field Station" name="Rio Aguaro" stateProvince="Guarico State">Rio Aguaro</location>
;
<date id="D9DF52E081779D3F48DAE885BDD27BF6" value="2009-01-10">
<collectingDate id="892DEEA043E0499373EF3D10CEF89B08" value="2009-01-10">10. i. 2009</collectingDate>
</date>
leg.
<collectorName id="72AD971E4292BAADC5F1F6E2C9D12456">Short</collectorName>
,
<collectorName id="CBC97697F6CF0EA303056482617A4600">Miller</collectorName>
,
<collectorName id="02B8F30C2B6D9CABCB00858B6DC62CDF">Joly</collectorName>
,
<collectorName id="BF44F2458D4FEEC017F91BDC354A4967">García</collectorName>
,
<collectorName id="6B95B1B71E1A6B0E394313ABBDE38750">Camacho</collectorName>
; VZ 09-0110 - 01 A /
<specimenCode id="FE2EBEC4C4087AA9A671544F5AD81693">SEMC 0852602</specimenCode>
<collectionCode id="43F5D9AC46040542E0EBFDFB3D1FFD1D">KUNHM-ENT</collectionCode>
, ”
</materialsCitation>
<materialsCitation id="44E385C8D74A0ED3868B377EA66EA7AD" collectedFrom="river margin" collectingDate="2009-01-12" collectionCode="MSBA, KUNHM-ENT" collectorName="Short &amp; Miller" country="Venezuela" latitude="6.58694" location="Rio Caripito" longLatPrecision="1" longitude="-67.02912" specimenCode="SM 0844405" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar State">
<specimenCount id="F15C5E23F35395C87CF5B9E59DED9468" count="1" type="male">1 male</specimenCount>
(
<named-content id="3E9D1977716640F2BF962E85CC649559" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-new-mexico" xlink_title="Museum of Southwestern Biology, Division of Arthropods, University of New Mexico, Albuquerque, NM, USA">
<collectionCode id="24EB8025DA6B9BC87230984646E187CC">MSBA</collectionCode>
</named-content>
) labeled “
<collectingCountry id="E0A8241DE7FA71465A3CC4ED5AB850E0" name="Venezuela">Venezuela</collectingCountry>
:
<collectingRegion id="724117AB3DB63B6BD151A33C4D63E59E" country="Venezuela" name="Bolivar">Bolivar State</collectingRegion>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EULAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-67.029120,6.586940]}">
<geoCoordinate id="6890DA32AF057C17AA13542C0EAAAE59" degrees="6.58694" direction="north" orientation="latitude" precision="1" value="6.58694">6.58694 ° N</geoCoordinate>
,
<geoCoordinate id="7964D7EEBA4924C50E2E5F32BA7D1AD0" degrees="67.02912" direction="west" orientation="longitude" precision="1" value="-67.02912">67.02912 ° W</geoCoordinate>
</named-content>
<location id="9D34FA1BF530A14F56EBF570CD20178D" LSID="urn:lsid:plazi:treatment:946585B2F35C5E65B3C0BCB61748D5D3:9D34FA1BF530A14F56EBF570CD20178D" country="Venezuela" latitude="6.58694" longLatPrecision="1" longitude="-67.02912" name="Rio Caripito" stateProvince="Bolivar State">Rio Caripito</location>
<date id="7A5F4C6F5985E80A6E18BA7FFB7ECEBE" value="2009-01-12">
<collectingDate id="E1BBCFC51A7FECD6C7EB1B3D3BAE99E0" value="2009-01-12">12. i. 2009</collectingDate>
</date>
; leg.
<collectorName id="5B78A7B72CE87DCB976C86FDF449AE7F">Short</collectorName>
<collectorName id="8C27A5FCC6065B8416597264B985DE54">Miller</collectorName>
VZ 09-0112 - 02 A:
<collectedFrom id="AA20FDB93860B44C2CC11F15630E2C60">river margin</collectedFrom>
/
<specimenCode id="72B174CE60DD5C29A0B16C27B0FA7B3F">SM 0844405</specimenCode>
<collectionCode id="8C93D235F86FF3BA71D4EE0DEB802A86">KUNHM-ENT</collectionCode>
[barcode label]
</materialsCitation>
. ” All
<typeStatus id="890736E8B7AF3D90520942419CBA4C79" type="paratype">paratypes</typeStatus>
with, “ …
<typeStatus id="99AE35094DCA367884748AC7C3A9FE16" type="paratype">Paratype</typeStatus>
<taxonomicName id="02661A6F7A81993BF16EC9440DEED95F" authority="Miller, Michat" authorityName="Miller, Michat" class="Insecta" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="E21E636718FBD99FD2CD97BF1D07DD76" italics="true">Nilssondytes diversus</emphasis>
Miller, Michat
</taxonomicName>
and Ferreira-Jr., 2024 [blue label with black line border].
</paragraph>
</subSubSection>
</treatment>
</document>