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<document id="73092A28942DDC70EF89778CFE34499E" ID-DOI="10.11646/zootaxa.5569.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734364" ID-ZooBank="C3CFD3D4-43E6-4AAC-8A4C-220BDF952E01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737735265714" checkinUser="plazi" docAuthor="Medina, Milton Norman, Lam, Athena W., Barsevskis, Arvids, Cabras, Analyn A. &amp; Van Dam, Matthew H." docDate="2024" docId="F65B879C062D8319FF6EF8AEB5685E37" docLanguage="en" docName="zootaxa.5569.3.5.pdf" docOrigin="Zootaxa 5569 (3)" docSource="https://doi.org/10.11646/zootaxa.5569.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyriotasiastes rhetenor subsp. solsonanensis Medina 2023" docType="treatment" docVersion="2" lastPageNumber="500" masterDocId="0A62FFE4062E831EFFF9FFE1B7155C2E" masterDocTitle="Integrative taxonomy of the genus Cyriotasiastes (Newman) (Cerambycidae: Lamiinae) with the description of a new species from Mindanao Island Philippines" masterLastPageNumber="508" masterPageNumber="493" pageNumber="496" updateTime="1737735716169" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="233518071F87DE8D5F6F2A670A094E41">Integrative taxonomy of the genus Cyriotasiastes (Newman) (Cerambycidae: Lamiinae) with the description of a new species from Mindanao Island Philippines</mods:title>
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<mods:namePart id="54E644D0313B5DC21BC9D41F11779E0A">Medina, Milton Norman</mods:namePart>
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<mods:namePart id="BA6249F042FCF6357F8BAF9DBE7C02CF">Lam, Athena W.</mods:namePart>
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<mods:affiliation id="BA8435F92EF7928C3F19B91FE6EC353D">Daugavpils University, Institute of Life Sciences and Technology, Coleopterological Research Center, Vienîbas Str. 13, Daugavpils, LV- 5401, Latvia.</mods:affiliation>
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<mods:namePart id="8C84469E1E282C4B24BE1405A5A619CF">Cabras, Analyn A.</mods:namePart>
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<mods:affiliation id="F9A588B658550E6FE276A1F78EEFBD1D">Tropical Genomics Laboratory-URESCOM, and Faculty of Agriculture and Life Sciences, Davao Oriental State University, Mati City, 8200 Philippines. &amp; National Museum of Natural History, Ermita, 1000 Philippines.</mods:affiliation>
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<mods:namePart id="632E8CD6EB86821C1859E867E14429AD">Van Dam, Matthew H.</mods:namePart>
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<mods:affiliation id="5065DD95FDBD66F5BA3C8B55611F9317">Center for Comparative Genomics, California Academy of Sciences, San Francesco, USA.</mods:affiliation>
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<paragraph id="7E4D368A062D831DFF6EF8AEB45F5B47" blockId="3.[151,842,1871,1933]" box="[151,842,1871,1898]" pageId="3" pageNumber="496">
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<taxonomicName id="B9F24D09062D831DFF6EF8AEB45F5B47" authority="Medina et al. 2023" authorityName="Medina" authorityYear="2023" box="[151,842,1871,1898]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA98062D831DFF6EF8AEB45F5B47" bold="true" box="[151,842,1871,1898]" pageId="3" pageNumber="496">
<emphasis id="4C86EA98062D831DFF6EF8AEB54B5B44" bold="true" box="[151,606,1871,1898]" italics="true" pageId="3" pageNumber="496">Cyriotasiastes rhetenor solsonanensis</emphasis>
<bibRefCitation id="1A634B7B062D831DFD9CF8AEB45F5B47" author="Medina, M. N. D. &amp; Cabras, A. A. &amp; Van Dam, M." box="[613,842,1871,1898]" pageId="3" pageNumber="496" pagination="177 - 184" refId="ref9050" refString="Medina, M. N. D., Cabras, A. A. &amp; Van Dam, M. (2023) Two new species of the genus Glenea Newman, 1842 (Coleoptera: Cerambycidae: Lamiinae: Saperdini) from Mindanao and Luzon Islands Philippines. Zootaxa, 5284 (1), 177 - 184. https: // doi. org / 10.11646 / zootaxa. 5284.1.8" type="journal article" year="2023">
Medina
<emphasis id="4C86EA98062D831DFD34F8AEB41D5B44" bold="true" box="[717,776,1871,1898]" italics="true" pageId="3" pageNumber="496">et al.</emphasis>
2023
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</emphasis>
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</paragraph>
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<figureCitation id="E6C92A0F062D831DFF6EF892B6395BA3" box="[151,300,1907,1933]" captionStart="FIGURE 1" captionStartId="4.[152,255,1923,1947]" captionTargetBox="[221,1409,348,1894]" captionTargetId="figure-60@4.[177,1411,306,1896]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus: Cyriotasiastes rhetenor solsonanensis, male, Holotype (A), C. r. solsonanensis, female, paratype (B), C. r. mindorensis, male, (C), C. r. mindorensis, female (D), C. r. panayensis, male, paratype (E), C. r. panayensis, female, panayensis (F), C. r. rhetenor, male (G), C. mindanaonis sp. nov., female, holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.14734366" httpUri="https://zenodo.org/record/14734366/files/figure.png" pageId="3" pageNumber="496">Figure 1AB</figureCitation>
,
<figureCitation id="E6C92A0F062D831DFEC1F892B6DB5BA3" box="[312,462,1907,1933]" captionStart="FIGURE 2" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[360,1228,206,1916]" captionTargetId="figure-18@5.[356,1232,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Lateral habitus: Cyriotasiastes rhetenor solsonanensis, male, Holotype (A), C. r. solsonanensis, female, paratype (B), C. r. mindorensis, male, (C), C. r. mindorensis, female (D), C. r. panayensis, male, paratype (E), C. r. panayensis, female, paratype (F), C. r. rhetenor, male (G), C. agusanensis sp. nov., female, holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.14734370" httpUri="https://zenodo.org/record/14734370/files/figure.png" pageId="3" pageNumber="496">Figure 2AB</figureCitation>
,
<figureCitation id="E6C92A0F062D831DFE20F892B5795BA3" box="[473,620,1907,1933]" captionStart="FIGURE 3" captionStartId="6.[152,255,1000,1024]" captionTargetBox="[171,1416,188,970]" captionTargetId="figure-20@6.[164,1423,181,977]" captionTargetPageId="6" captionText="FIGURE 3. Frons: Cyriotasiastes rhetenor solsonanensis, male, Holotype (A), C. r. solsonanensis, female, paratype (B), C. r. mindorensis, male, (C), C. r. mindorensis, female (D), C. r. panayensis, male, paratype (E), C. r. panayensis, female, paratype (F), C. r. rhetenor, male (G), C. agusanensis sp. nov., female, holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.14734372" httpUri="https://zenodo.org/record/14734372/files/figure.png" pageId="3" pageNumber="496">Figure 3AB</figureCitation>
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<emphasis id="4C86EA98062D831DFF6EF854B6695BE1" bold="true" box="[151,380,1973,1999]" pageId="3" pageNumber="496">Material examined</emphasis>
:
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leg.
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, to be deposited at
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;
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female,
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the same label as
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;
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,
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,
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,
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,
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</date>
, loc.
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;
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</date>
, loc.
<collectorName id="D307535C062A831AFD31FF5AB4155CFB" box="[712,768,187,213]" pageId="4" pageNumber="497">Coll.</collectorName>
<collectionCode id="18E3AE4F062A831AFCF4FF5AB44B5CFB" box="[781,862,187,213]" pageId="4" pageNumber="497">DUBC</collectionCode>
</materialsCitation>
;
<materialsCitation id="CE9A3CD7062A831AFC89FF5AB6DC5CD7" collectingDate="2013-07" collectionCode="DUBC" collectorName="Coll." country="Philippines" location="Eastern Luzon" pageId="4" pageNumber="497" specimenCount="1" specimenCount-male="1" stateProvince="Isabela" typeStatus="paratype">
<specimenCount id="68F4FD03062A831AFC89FF5AB4D15CFB" box="[880,964,187,213]" count="1" pageId="4" pageNumber="497" type="male">1 male</specimenCount>
,
<collectingCountry id="06E5761A062A831AFC2FFF5AB36E5CFB" box="[982,1147,187,213]" name="Philippines" pageId="4" pageNumber="497">PHILIPPINES</collectingCountry>
<location id="7B2D6051062A831AFB70FF5AB2215CFB" LSID="urn:lsid:plazi:treatment:F65B879C062D8319FF6EF8AEB5685E37:7B2D6051062A831AFB70FF5AB2215CFB" box="[1161,1332,187,213]" country="Philippines" name="Eastern Luzon" pageId="4" pageNumber="497" stateProvince="Isabela">Eastern Luzon</location>
,
<collectingRegion id="BC36F868062A831AFABFFF5AB28D5CFB" box="[1350,1432,187,213]" country="Philippines" name="Isabela" pageId="4" pageNumber="497">Isabela</collectingRegion>
,
<date id="0A4C104A062A831AFF6EFF3EB7E35CD7" box="[151,246,223,249]" pageId="4" pageNumber="497" value="2013-07">
<collectingDate id="1A08E9A2062A831AFF6EFF3EB7E35CD7" box="[151,246,223,249]" pageId="4" pageNumber="497" value="2013-07">vii.2013</collectingDate>
</date>
, loc.
<collectorName id="D307535C062A831AFECEFF3EB67A5CD7" box="[311,367,223,249]" pageId="4" pageNumber="497">Coll.</collectorName>
<collectionCode id="18E3AE4F062A831AFE81FF3EB6DC5CD7" box="[376,457,223,249]" pageId="4" pageNumber="497">DUBC</collectionCode>
</materialsCitation>
;
<materialsCitation id="CE9A3CD7062A831AFE21FF3EB6335D33" collectingDate="2014-06" collectionCode="DUBC" collectorName="Coll." country="Philippines" county="Northern Luzon" location="Santa Ana" municipality="Valley" pageId="4" pageNumber="497" specimenCount="2" specimenCount-female="2" stateProvince="Cagayan" typeStatus="paratype">
<specimenCount id="68F4FD03062A831AFE21FF3EB52B5CD7" box="[472,574,223,249]" count="2" pageId="4" pageNumber="497" type="female">2 female</specimenCount>
,
<collectingCountry id="06E5761A062A831AFDB5FF3EB5E55CD7" box="[588,752,223,249]" name="Philippines" pageId="4" pageNumber="497">PHILIPPINES</collectingCountry>
<collectingCounty id="972C4E06062A831AFD06FF3EB4AD5CD7" box="[767,952,223,249]" pageId="4" pageNumber="497">Northern Luzon</collectingCounty>
,
<collectingRegion id="BC36F868062A831AFC3FFF3EB33D5CD7" box="[966,1064,223,249]" country="Philippines" name="Cagayan" pageId="4" pageNumber="497">Cagayan</collectingRegion>
<collectingMunicipality id="9E29ACF0062A831AFBC8FF3EB36F5CD7" box="[1073,1146,223,249]" pageId="4" pageNumber="497">Valley</collectingMunicipality>
,
<location id="7B2D6051062A831AFB71FF3EB3EA5CD7" LSID="urn:lsid:plazi:treatment:F65B879C062D8319FF6EF8AEB5685E37:7B2D6051062A831AFB71FF3EB3EA5CD7" box="[1160,1279,223,249]" country="Philippines" county="Northern Luzon" municipality="Valley" name="Santa Ana" pageId="4" pageNumber="497" stateProvince="Cagayan">Santa Ana</location>
,
<date id="0A4C104A062A831AFAF4FF3EB2705CD7" box="[1293,1381,223,249]" pageId="4" pageNumber="497" value="2014-06">
<collectingDate id="1A08E9A2062A831AFAF4FF3EB2705CD7" box="[1293,1381,223,249]" pageId="4" pageNumber="497" value="2014-06">vi.2014</collectingDate>
</date>
, loc.
<collectorName id="D307535C062A831AFF6EFEE2B7DA5D33" box="[151,207,259,285]" pageId="4" pageNumber="497">Coll.</collectorName>
<collectionCode id="18E3AE4F062A831AFF2CFEE2B6335D33" box="[213,294,259,285]" pageId="4" pageNumber="497">DUBC</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="2A8D6602062A831AFF61F862B3055BCD" ID-DOI="http://doi.org/10.5281/zenodo.14734366" ID-Zenodo-Dep="14734366" httpUri="https://zenodo.org/record/14734366/files/figure.png" pageId="4" pageNumber="497" startId="4.[152,255,1923,1947]" targetBox="[221,1409,348,1894]" targetPageId="4" targetType="figure">
<paragraph id="7E4D368A062A831AFF61F862B3055BCD" blockId="4.[151,1437,1923,2019]" pageId="4" pageNumber="497">
<emphasis id="4C86EA98062A831AFF61F862B6065BB5" bold="true" box="[152,275,1923,1947]" pageId="4" pageNumber="497">FIGURE 1</emphasis>
. Dorsal habitus:
<taxonomicName id="B9F24D09062A831AFE3BF862B4505BB5" box="[450,837,1923,1947]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA98062A831AFE3BF862B4505BB5" box="[450,837,1923,1947]" italics="true" pageId="4" pageNumber="497">Cyriotasiastes rhetenor solsonanensis</emphasis>
</taxonomicName>
, male, Holotype (A),
<taxonomicName id="B9F24D09062A831AFBDFF862B3F35BB5" box="[1062,1254,1923,1947]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA98062A831AFBDFF862B3F35BB5" box="[1062,1254,1923,1947]" italics="true" pageId="4" pageNumber="497">C. r. solsonanensis</emphasis>
</taxonomicName>
, female, paratype (B),
<taxonomicName id="B9F24D09062A831AFF3FF846B6615B91" authorityName="Vives" authorityYear="2009" box="[198,372,1959,1983]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="mindorensis">
<emphasis id="4C86EA98062A831AFF3FF846B6615B91" box="[198,372,1959,1983]" italics="true" pageId="4" pageNumber="497">C. r. mindorensis</emphasis>
</taxonomicName>
, male, (C),
<taxonomicName id="B9F24D09062A831AFE16F846B5895B91" authorityName="Vives" authorityYear="2009" box="[495,668,1959,1983]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="mindorensis">
<emphasis id="4C86EA98062A831AFE16F846B5895B91" box="[495,668,1959,1983]" italics="true" pageId="4" pageNumber="497">C. r. mindorensis</emphasis>
</taxonomicName>
, female (D),
<taxonomicName id="B9F24D09062A831AFCDFF846B4DC5B91" authorityName="Barsevskis" authorityYear="2018" box="[806,969,1959,1983]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="panayensis">
<emphasis id="4C86EA98062A831AFCDFF846B4DC5B91" box="[806,969,1959,1983]" italics="true" pageId="4" pageNumber="497">C. r. panayensis</emphasis>
</taxonomicName>
, male, paratype (E),
<taxonomicName id="B9F24D09062A831AFB58F846B2515B91" authorityName="Barsevskis" authorityYear="2018" box="[1185,1348,1959,1983]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="panayensis">
<emphasis id="4C86EA98062A831AFB58F846B2515B91" box="[1185,1348,1959,1983]" italics="true" pageId="4" pageNumber="497">C. r. panayensis</emphasis>
</taxonomicName>
, female, panayensis (F),
<taxonomicName id="B9F24D09062A831AFEC1F82AB6D45BCD" baseAuthorityName="Newman" baseAuthorityYear="1842" box="[312,449,1995,2019]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="rhetenor">
<emphasis id="4C86EA98062A831AFEC1F82AB6D45BCD" box="[312,449,1995,2019]" italics="true" pageId="4" pageNumber="497">C. r. rhetenor</emphasis>
</taxonomicName>
, male (G),
<taxonomicName id="B9F24D09062A831AFDCDF82AB5C05BCD" authorityName="Medina &amp; Lam &amp; Barsevskis &amp; Cabras &amp; Van Dam" authorityYear="2024" box="[564,725,1995,2019]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="mindanaonis" status="sp. nov.">
<emphasis id="4C86EA98062A831AFDCDF82AB5C05BCD" box="[564,725,1995,2019]" italics="true" pageId="4" pageNumber="497">C. mindanaonis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57B557E3062A831AFD22F82AB4395BCC" box="[731,812,1995,2018]" pageId="4" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype (H).
</paragraph>
</caption>
<caption id="2A8D6602062B831BFF61F876B4E45BD6" ID-DOI="http://doi.org/10.5281/zenodo.14734370" ID-Zenodo-Dep="14734370" httpUri="https://zenodo.org/record/14734370/files/figure.png" pageId="5" pageNumber="498" startId="5.[152,255,1943,1967]" targetBox="[360,1228,206,1916]" targetPageId="5" targetType="figure">
<paragraph id="7E4D368A062B831BFF61F876B4E45BD6" blockId="5.[151,1437,1943,2040]" pageId="5" pageNumber="498">
<emphasis id="4C86EA98062B831BFF61F876B6065B81" bold="true" box="[152,275,1943,1967]" pageId="5" pageNumber="498">FIGURE 2</emphasis>
. Lateral habitus:
<taxonomicName id="B9F24D09062B831BFE3CF879B4525B9E" box="[453,839,1944,1968]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA98062B831BFE3CF879B4525B9E" box="[453,839,1944,1968]" italics="true" pageId="5" pageNumber="498">Cyriotasiastes rhetenor solsonanensis</emphasis>
</taxonomicName>
, male, Holotype (A),
<taxonomicName id="B9F24D09062B831BFBDEF879B3F35B9E" box="[1063,1254,1944,1968]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA98062B831BFBDEF879B3F35B9E" box="[1063,1254,1944,1968]" italics="true" pageId="5" pageNumber="498">C. r. solsonanensis</emphasis>
</taxonomicName>
, female, paratype (B),
<taxonomicName id="B9F24D09062B831BFF3FF85DB6615BFA" authorityName="Vives" authorityYear="2009" box="[198,372,1980,2004]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="mindorensis">
<emphasis id="4C86EA98062B831BFF3FF85DB6615BFA" box="[198,372,1980,2004]" italics="true" pageId="5" pageNumber="498">C. r. mindorensis</emphasis>
</taxonomicName>
, male, (C),
<taxonomicName id="B9F24D09062B831BFE16F85DB5895BFA" authorityName="Vives" authorityYear="2009" box="[495,668,1980,2004]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="mindorensis">
<emphasis id="4C86EA98062B831BFE16F85DB5895BFA" box="[495,668,1980,2004]" italics="true" pageId="5" pageNumber="498">C. r. mindorensis</emphasis>
</taxonomicName>
, female (D),
<taxonomicName id="B9F24D09062B831BFCDFF85DB4DC5BFD" authorityName="Barsevskis" authorityYear="2018" box="[806,969,1980,2003]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="panayensis">
<emphasis id="4C86EA98062B831BFCDFF85DB4DC5BFD" box="[806,969,1980,2003]" italics="true" pageId="5" pageNumber="498">C. r. panayensis</emphasis>
</taxonomicName>
, male, paratype (E),
<taxonomicName id="B9F24D09062B831BFB58F85DB2515BFD" authorityName="Barsevskis" authorityYear="2018" box="[1185,1348,1980,2003]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="panayensis">
<emphasis id="4C86EA98062B831BFB58F85DB2515BFD" box="[1185,1348,1980,2003]" italics="true" pageId="5" pageNumber="498">C. r. panayensis</emphasis>
</taxonomicName>
, female, paratype (F),
<taxonomicName id="B9F24D09062B831BFED9F801B6BC5BD6" baseAuthorityName="Newman" baseAuthorityYear="1842" box="[288,425,2016,2040]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="rhetenor">
<emphasis id="4C86EA98062B831BFED9F801B6BC5BD6" box="[288,425,2016,2040]" italics="true" pageId="5" pageNumber="498">C. r. rhetenor</emphasis>
</taxonomicName>
, male (G),
<taxonomicName id="B9F24D09062B831BFDE5F801B5A35BD9" authorityName="Medina &amp; Lam &amp; Barsevskis &amp; Cabras &amp; Van Dam" authorityYear="2024" box="[540,694,2016,2039]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="agusanensis" status="sp. nov.">
<emphasis id="4C86EA98062B831BFDE5F801B5A35BD9" box="[540,694,2016,2039]" italics="true" pageId="5" pageNumber="498">C. agusanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57B557E3062B831BFD45F801B4185BD9" box="[700,781,2016,2039]" pageId="5" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype (H).
</paragraph>
</caption>
<caption id="2A8D660206288318FF61FC09B4815867" ID-DOI="http://doi.org/10.5281/zenodo.14734372" ID-Zenodo-Dep="14734372" httpUri="https://zenodo.org/record/14734372/files/figure.png" pageId="6" pageNumber="499" startId="6.[152,255,1000,1024]" targetBox="[171,1416,188,970]" targetPageId="6" targetType="figure">
<paragraph id="7E4D368A06288318FF61FC09B4815867" blockId="6.[151,1437,1000,1097]" pageId="6" pageNumber="499">
<emphasis id="4C86EA9806288318FF61FC09B606582F" bold="true" box="[152,275,1000,1025]" pageId="6" pageNumber="499">FIGURE 3</emphasis>
. Frons:
<taxonomicName id="B9F24D0906288318FE9FFC08B5FD582F" box="[358,744,1001,1025]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA9806288318FE9FFC08B5FD582F" box="[358,744,1001,1025]" italics="true" pageId="6" pageNumber="499">Cyriotasiastes rhetenor solsonanensis</emphasis>
</taxonomicName>
, male, Holotype (A),
<taxonomicName id="B9F24D0906288318FC31FC08B393582F" box="[968,1158,1001,1025]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="solsonanensis">
<emphasis id="4C86EA9806288318FC31FC08B393582F" box="[968,1158,1001,1025]" italics="true" pageId="6" pageNumber="499">C. r. solsonanensis</emphasis>
</taxonomicName>
, female, paratype (B),
<taxonomicName id="B9F24D0906288318FA88FC08B607580B" authorityName="Vives" authorityYear="2009" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="mindorensis">
<emphasis id="4C86EA9806288318FA88FC08B607580B" italics="true" pageId="6" pageNumber="499">C. r. mindorensis</emphasis>
</taxonomicName>
, male, (C),
<taxonomicName id="B9F24D0906288318FE77FBECB529580B" authorityName="Vives" authorityYear="2009" box="[398,572,1037,1061]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="mindorensis">
<emphasis id="4C86EA9806288318FE77FBECB529580B" box="[398,572,1037,1061]" italics="true" pageId="6" pageNumber="499">C. r. mindorensis</emphasis>
</taxonomicName>
, female (D),
<taxonomicName id="B9F24D0906288318FD3FFBECB47F580A" authorityName="Barsevskis" authorityYear="2018" box="[710,874,1037,1060]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="panayensis">
<emphasis id="4C86EA9806288318FD3FFBECB47F580A" box="[710,874,1037,1060]" italics="true" pageId="6" pageNumber="499">C. r. panayensis</emphasis>
</taxonomicName>
, male, paratype (E),
<taxonomicName id="B9F24D0906288318FBBBFBECB3F3580A" authorityName="Barsevskis" authorityYear="2018" box="[1090,1254,1037,1060]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="panayensis">
<emphasis id="4C86EA9806288318FBBBFBECB3F3580A" box="[1090,1254,1037,1060]" italics="true" pageId="6" pageNumber="499">C. r. panayensis</emphasis>
</taxonomicName>
, female, paratype (F),
<taxonomicName id="B9F24D0906288318FF3AFBD0B65E5867" baseAuthorityName="Newman" baseAuthorityYear="1842" box="[195,331,1073,1097]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="rhetenor" subSpecies="rhetenor">
<emphasis id="4C86EA9806288318FF3AFBD0B65E5867" box="[195,331,1073,1097]" italics="true" pageId="6" pageNumber="499">C. r. rhetenor</emphasis>
</taxonomicName>
, male (G),
<taxonomicName id="B9F24D0906288318FE46FBD0B54C5866" authorityName="Medina &amp; Lam &amp; Barsevskis &amp; Cabras &amp; Van Dam" authorityYear="2024" box="[447,601,1073,1096]" class="Insecta" family="Cerambycidae" genus="Cyriotasiastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="agusanensis" status="sp. nov.">
<emphasis id="4C86EA9806288318FE46FBD0B54C5866" box="[447,601,1073,1096]" italics="true" pageId="6" pageNumber="499">C. agusanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57B557E306288318FDA6FBD0B5A55866" box="[607,688,1073,1096]" pageId="6" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype (H).
</paragraph>
</caption>
<paragraph id="7E4D368A06288318FF3EFB7AB5FB590F" blockId="6.[151,1437,1179,2033]" pageId="6" pageNumber="499">
Measurements of the various body parts (in range) are as follows: Males (n=3): LB (
<collectionCode id="18E3AE4F06288318FB85FB7AB3B6589B" box="[1148,1187,1179,1205]" pageId="6" pageNumber="499">LH</collectionCode>
+LP+
<collectionCode id="18E3AE4F06288318FB1AFB7AB212589B" box="[1251,1287,1179,1205]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="6" pageNumber="499">LE</collectionCode>
) =
<quantity id="B90A9B6F06288318FAD4FB7AB7D358F7" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.55" metricValueMin="1.15" pageId="6" pageNumber="499" unit="mm" value="13.5" valueMax="15.5" valueMin="11.5">11.515.5 mm</quantity>
; LH = 1.52.0 mm; WH = 2.03.0 mm; LP = 2.03.0 mm; WP =
<quantity id="B90A9B6F06288318FC39FB5EB35958F7" box="[960,1100,1215,1242]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="6" pageNumber="499" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.54.5 mm</quantity>
; LE = 8.0
<quantity id="B90A9B6F06288318FB29FB5EB22258F7" box="[1232,1335,1215,1242]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="499" unit="mm" value="10.5">10.5 mm</quantity>
; WEH = 4.05.0 mm. Females (n=4): LB (
<collectionCode id="18E3AE4F06288318FDE8FB02B52D58D3" box="[529,568,1251,1277]" pageId="6" pageNumber="499">LH</collectionCode>
+LP+
<collectionCode id="18E3AE4F06288318FD80FB02B58858D3" box="[633,669,1251,1277]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="6" pageNumber="499">LE</collectionCode>
) = 18.019.0 mm; LH = 3.0 mm; WH = 3.0
<quantity id="B90A9B6F06288318FB65FB02B3E058D3" box="[1180,1269,1251,1278]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="499" unit="mm" value="3.5">3.5 mm</quantity>
; LP = 3.0 mm; WP = 5.0 mm; LE = 12.013.0 mm; WEH = 6.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="36E8650106288319FF3EFACAB6F85DFF" lastPageId="7" lastPageNumber="500" pageId="6" pageNumber="499" type="description">
<paragraph id="7E4D368A06288318FF3EFACAB63459D7" blockId="6.[151,1437,1179,2033]" pageId="6" pageNumber="499">
<emphasis id="4C86EA9806288318FF3EFACAB660596B" bold="true" box="[199,373,1323,1349]" pageId="6" pageNumber="499">Redescription.</emphasis>
Adult male. Teguments: head, prothorax, abdomen, and legs dark brown to matte black, antennae brown to dark brown. Head densely covered with tomentose white to yellowish pubescence, denser at the frons; long erect black setae randomly scattered at frons, denser at the posterior margin. Antennae dark brown, more than twice longer than body length, densely covered with very fine whitish recumbent pubescence; scape slightly robust, with apical cicatrix; antennomeres IIIV of the same length. Eyes dark brown and mandible dark brown to matte black.
</paragraph>
<paragraph id="7E4D368A06288318FF3EF9E2B40B5A83" blockId="6.[151,1437,1179,2033]" pageId="6" pageNumber="499">Elytra twice longer than wide, tomentose, coarsely punctate near the base up to apical third, finer towards apical end; densely covered with light to dark brown recumbent setae; with three vertical bands of white to yellowish recumbent setae: slender S-shaped at the basal third, broad C-shaped at postmedian from elytral margin up to near suture, and a small band at the apical end. Elytra apical margin flat, elytral suture miniscule. Scutellum rounded, matte black, densely covered with recumbent white setae.</paragraph>
<paragraph id="7E4D368A06288318FF3EF956B6DB5B37" blockId="6.[151,1437,1179,2033]" pageId="6" pageNumber="499">Prothorax armed at each side, slightly recurved white band of longitudinal recumbent setae at each sides. Basal margin of pronotum lined with white recumbent pubescence, minute band of white pubescence near the basal margin, disc slightly raised.</paragraph>
<paragraph id="7E4D368A06288318FF3EF8C2B6E65BAB" blockId="6.[151,1437,1179,2033]" pageId="6" pageNumber="499">Legs dark brown to matte black. Coxae, trochanters black densely covered with white recumbent pubescence; femur and tibiae dark brown, densely covered with minute recumbent pubescence with erect golden-brown bristles at the apex. Claws dark brown.</paragraph>
<paragraph id="7E4D368A06288318FF3EF86EB26D5BDF" blockId="6.[151,1437,1179,2033]" pageId="6" pageNumber="499">Prosternum, mesosternum, metasternum, abdominal sternites matte black covered with white recumbent pubescence. Metepisternum paralleled, lanceolate at the apical end, covered with whitish and yellowish recumbent pubescence. Pygidium matte black, widely ovate at apex, densely covered with yellowish and black erect setae.</paragraph>
<paragraph id="7E4D368A06298319FF3EFF76B53F5D33" blockId="7.[151,1437,151,537]" pageId="7" pageNumber="500">
Genitalia. Endophallus approximately twice as long as the whole system of genitalia (
<figureCitation id="E6C92A0F06298319FB33FF76B23F5C9F" box="[1226,1322,151,177]" captionStart="FIGURE 4" captionStartId="8.[152,255,1903,1927]" captionTargetBox="[155,1430,186,1875]" captionTargetId="figure-20@8.[151,1435,181,1879]" captionTargetPageId="8" captionText="FIGURE 4. AC Whole system of genitalia: A. Cyriotasiastes rhetenor solsonanensis, holotype, B. C. r. mindorensis, C. C. r. panayensis, paratype. DF Aedeagus, ventral aspect: D. C. r. mindorensis, E. C. r. panayensis, paratype, F. C. r. solsonanensis. GI Aedeagus, lateral aspect: G. C. r. mindorensis, H. C. r. panayensis, paratype, I. C. r. solsonanensis. JL Tegmen, ventral aspect: J. C. r. mindorensis, K. C. r. panayensis, paratype, L. C. r. solsonanensis." figureDoi="http://doi.org/10.5281/zenodo.14734378" httpUri="https://zenodo.org/record/14734378/files/figure.png" pageId="7" pageNumber="500">Fig. 4A</figureCitation>
). Tegmen relatively slender, parameres broad; apex of parameres lined with long erect yellowish setae. Aedeagus strongly recurved when viewed laterally (
<figureCitation id="E6C92A0F06298319FDF0FF3EB5425CD7" box="[521,599,223,249]" captionStart="FIGURE 4" captionStartId="8.[152,255,1903,1927]" captionTargetBox="[155,1430,186,1875]" captionTargetId="figure-20@8.[151,1435,181,1879]" captionTargetPageId="8" captionText="FIGURE 4. AC Whole system of genitalia: A. Cyriotasiastes rhetenor solsonanensis, holotype, B. C. r. mindorensis, C. C. r. panayensis, paratype. DF Aedeagus, ventral aspect: D. C. r. mindorensis, E. C. r. panayensis, paratype, F. C. r. solsonanensis. GI Aedeagus, lateral aspect: G. C. r. mindorensis, H. C. r. panayensis, paratype, I. C. r. solsonanensis. JL Tegmen, ventral aspect: J. C. r. mindorensis, K. C. r. panayensis, paratype, L. C. r. solsonanensis." figureDoi="http://doi.org/10.5281/zenodo.14734378" httpUri="https://zenodo.org/record/14734378/files/figure.png" pageId="7" pageNumber="500">Fig. 4I</figureCitation>
), slightly bottle-shaped ventrally (
<figureCitation id="E6C92A0F06298319FC23FF3EB3255CD7" box="[986,1072,223,249]" captionStart="FIGURE 4" captionStartId="8.[152,255,1903,1927]" captionTargetBox="[155,1430,186,1875]" captionTargetId="figure-20@8.[151,1435,181,1879]" captionTargetPageId="8" captionText="FIGURE 4. AC Whole system of genitalia: A. Cyriotasiastes rhetenor solsonanensis, holotype, B. C. r. mindorensis, C. C. r. panayensis, paratype. DF Aedeagus, ventral aspect: D. C. r. mindorensis, E. C. r. panayensis, paratype, F. C. r. solsonanensis. GI Aedeagus, lateral aspect: G. C. r. mindorensis, H. C. r. panayensis, paratype, I. C. r. solsonanensis. JL Tegmen, ventral aspect: J. C. r. mindorensis, K. C. r. panayensis, paratype, L. C. r. solsonanensis." figureDoi="http://doi.org/10.5281/zenodo.14734378" httpUri="https://zenodo.org/record/14734378/files/figure.png" pageId="7" pageNumber="500">Fig. 4F</figureCitation>
), and apical third steeply angled towards a relatively blunt apical tip.
</paragraph>
<paragraph id="7E4D368A06298319FF3EFEC6B6F85DFF" blockId="7.[151,1437,151,537]" pageId="7" pageNumber="500">Adult Female. Morphologically similar to males except females are broader and larger.Antennae slightly longer than body length but much shorter compared to males. Females have two-toned (basal third or basal half with whitish or yellowish recumbent pubescence, the other half covered with black recumbent pubescence) antennomeres IIIVI, males do not have two-toned antennomeres. The S-shaped band of white pubescence at the basal third is much broader compared to the male.</paragraph>
</subSubSection>
<subSubSection id="36E8650106298319FF3EFE3BB5685E37" pageId="7" pageNumber="500" type="distribution">
<paragraph id="7E4D368A06298319FF3EFE3BB5685E37" blockId="7.[151,1437,151,537]" pageId="7" pageNumber="500">
<emphasis id="4C86EA9806298319FF3EFE3BB6765DDA" bold="true" box="[199,355,474,500]" pageId="7" pageNumber="500">Distribution:</emphasis>
<collectingCountry id="06E5761A06298319FE9EFE3AB6F05DDB" box="[359,485,475,501]" name="Philippines" pageId="7" pageNumber="500">Philippines</collectingCountry>
(Northern Luzon:
<collectingRegion id="BC36F86806298319FD48FE3AB4285DDB" box="[689,829,475,501]" country="Philippines" name="Ilocos Norte" pageId="7" pageNumber="500">Ilocos Norte</collectingRegion>
, Solsona;
<collectingRegion id="BC36F86806298319FC52FE3AB3135DDB" box="[939,1030,475,501]" country="Philippines" name="Kalinga" pageId="7" pageNumber="500">Kalinga</collectingRegion>
, Pinukpuk;
<collectingRegion id="BC36F86806298319FB7EFE3AB2275DDB" box="[1159,1330,475,501]" country="Philippines" name="Nueva Vizcaya" pageId="7" pageNumber="500">Nueva Vizcaya</collectingRegion>
;
<collectingRegion id="BC36F86806298319FAC3FE3AB2895DDB" box="[1338,1436,475,501]" country="Philippines" name="Cagayan" pageId="7" pageNumber="500">Cagayan</collectingRegion>
Valley, Santa Ana. Eastern Luzon:
<collectingRegion id="BC36F86806298319FDE5FE1EB5655E37" box="[540,624,511,537]" country="Philippines" name="Isabela" pageId="7" pageNumber="500">Isabela</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>