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<document id="206197AC73E97F6966C1B7AC8727F22C" ID-CLB-Dataset="21285" ID-DOI="10.1080/00222933.2015.1104393" ID-GBIF-Dataset="6d59feb3-23d2-4911-88b2-0db9163acf76" ID-ISSN="1464-5262" ID-Zenodo-Dep="3990373" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597772314732" checkinUser="carolina" docAuthor="Yamada, Kazutaka, Yasunaga, Tomohide &amp; Artchawakom, Taksin" docDate="2015" docId="EA37063720440D3E41403239FEC8FCDD" docLanguage="en" docName="JNATHIST.50.17-18.1103-1157.pdf.imf" docOrigin="Journal of Natural History 50" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Orius (Orius) inthanonus Yamada and Yasunaga, sp. nov." docType="treatment" docVersion="6" lastPageNumber="1133" masterDocId="160E7E4F20580D2040773102FFDBFE7C" masterDocTitle="The flower bug genus Orius Wolff, 1811 (Hemiptera: Heteroptera: Anthocoridae: Oriini) of Thailand" masterLastPageNumber="1157" masterPageNumber="1103" pageNumber="1131" updateTime="1698844749935" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="28860A22269FB35D15D5D95E550158DF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3742C2D6D5F6370C6C3AD3174F4C9302">
<mods:title id="ED094D81C7008C52D062D749897C446E">The flower bug genus Orius Wolff, 1811 (Hemiptera: Heteroptera: Anthocoridae: Oriini) of Thailand</mods:title>
</mods:titleInfo>
<mods:name id="3944B834CF1AAB3F70F9E83036AC9519" type="personal">
<mods:role id="308B255E355558185907ECE8200DE8F7">
<mods:roleTerm id="613AE7EC31BB131195AECF967F2FFFB4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3B27E01D1BE5F04B5707F41DF1E7CBA">Yamada, Kazutaka</mods:namePart>
<mods:affiliation id="02CC0D0CDA6CB4C63C24A2464C8DB424">Tokushima Prefectural Museum, Bunka-no-Mori Park, Tokushima, Japan;</mods:affiliation>
</mods:name>
<mods:name id="A053D841FA53D8DE220FC07F2A89BDA0" type="personal">
<mods:role id="687D88A83E1371C317BCAE8776743A74">
<mods:roleTerm id="FEBC04584417ACD713810DB6232EF003">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D6C00961CEE78CEA0E9C6BF78C9549E">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="092770D41F435C4385567F0913197084">American Museum of Natural History, New York, NY, USA; &amp; Plant Protection Division, Myanmar Ministry of Agriculture &amp; Irrigation, c / o Japan International Cooperation Agency (JICA), Yangon, Myanmar;</mods:affiliation>
</mods:name>
<mods:name id="07CA72DD3767378990962B2B2EFC013F" type="personal">
<mods:role id="82735EAF63E8511CDB550B2F733FA1E5">
<mods:roleTerm id="B9042EB921BCE43048858F774767D6D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4FB8B8E83DF093FDA39DF6617CD3FE1">Artchawakom, Taksin</mods:namePart>
<mods:affiliation id="189A6448937E389F4D9CFD3E9BCFE062">Sakaerat Environmental Research Station (SERS), Thailand Institute of Scientific and Technological Research (TISTR), Ministry of Science and Technology, Nakhon Ratchasima, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BEB5231D5F0ED7757CD6CCBD500C600F">text</mods:typeOfResource>
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<mods:title id="9362A60D03D6155FC63D6A96370E41E6">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="4E7DC96A7EF8D0621544D7DD30B48DF1">
<mods:date id="079C550FB7209C6BDF8F09FDF1D4C2E0">2015</mods:date>
<mods:detail id="9C0BCA0984E5F739C026224014578829" type="pubDate">
<mods:number id="7CE5913BAD01A79AD3E951C714F41E05">2015-09-30</mods:number>
</mods:detail>
<mods:detail id="64677DD43148D6D1FA62DF583AF4B172" type="volume">
<mods:number id="A0B2D8C4D1ED8A791516B7DDA215FF8F">50</mods:number>
</mods:detail>
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<mods:start id="407A7C4F12FA80E526F62DC6FBCACC9A">1103</mods:start>
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<mods:classification id="1B81A9163C287EECA37929616D8F8679">journal article</mods:classification>
<mods:identifier id="EF11A0E03B3A7600325B790A9DBA06BB" type="CLB-Dataset">21285</mods:identifier>
<mods:identifier id="F75591B83A82F9EE788E928700E77796" type="DOI">10.1080/00222933.2015.1104393</mods:identifier>
<mods:identifier id="098F77C3BF26A493EA29AD63209A305A" type="GBIF-Dataset">6d59feb3-23d2-4911-88b2-0db9163acf76</mods:identifier>
<mods:identifier id="0BFAA6AA5F08840CC510968C0C8A12D3" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="8F74328C93360DFE662FD97888B7329C" type="Zenodo-Dep">3990373</mods:identifier>
</mods:mods>
<treatment id="EA37063720440D3E41403239FEC8FCDD" ID-DOI="http://doi.org/10.5281/zenodo.4332769" ID-GBIF-Taxon="166254624" ID-Zenodo-Dep="4332769" LSID="urn:lsid:plazi:treatment:EA37063720440D3E41403239FEC8FCDD" httpUri="http://treatment.plazi.org/id/EA37063720440D3E41403239FEC8FCDD" lastPageId="30" lastPageNumber="1133" pageId="28" pageNumber="1131">
<subSubSection id="2A84E4AA20440D3C41403239FCB3FD0B" pageId="28" pageNumber="1131" type="nomenclature">
<paragraph id="6221B72120440D3C41403239FCB3FD0B" blockId="28.[311,1005,827,887]" pageId="28" pageNumber="1131">
<heading id="3969004D20440D3C41403239FCB3FD0B" centered="true" fontSize="11" level="2" pageId="28" pageNumber="1131" reason="2">
<taxonomicName id="A59ECCA220440D3C41403239FC58FD28" authority="Yamada and Yasunaga" authorityName="Yamada and Yasunaga" box="[311,899,827,852]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1131" phylum="Arthropoda" rank="species" species="inthanonus" status="sp. nov." subGenus="Orius">
<emphasis id="50EA6B3320440D3C41403239FDB3FD28" bold="true" box="[311,616,827,852]" inLineHeading="true" pageId="28" pageNumber="1131" reason="3">
<emphasis id="50EA6B3320440D3C41403239FEA3FD28" bold="true" box="[311,376,827,852]" italics="true" pageId="28" pageNumber="1131">Orius</emphasis>
(
<emphasis id="50EA6B3320440D3C41FD3239FE10FD28" bold="true" box="[394,459,827,852]" italics="true" pageId="28" pageNumber="1131">Orius</emphasis>
)
<emphasis id="50EA6B3320440D3C41AA3239FDB3FD28" bold="true" box="[477,616,827,852]" italics="true" pageId="28" pageNumber="1131">inthanonus</emphasis>
</emphasis>
Yamada and Yasunaga
</taxonomicName>
,
<taxonomicNameLabel id="4BD9D64820440D3C43E73239FC36FD28" box="[912,1005,827,852]" pageId="28" pageNumber="1131" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="FAA5ABA420440D3C41B4325CFD9CFD0B" box="[451,583,862,887]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figures 3I,J</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C4221325CFDAFFD0B" box="[598,628,862,887]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="28" pageNumber="1131">5K</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C42F4325CFD44FD0B" box="[643,671,862,887]" captionStart="Figure 6" captionStartId="7.[160,225,1375,1398]" captionTargetBox="[178,1138,142,1344]" captionTargetId="figure@7.[178,1138,142,1345]" captionTargetPageId="7" captionText="Figure 6. SEM images of ostiolar peritreme and evaporatorium, left lateroventral view. (A) O. sakaerat, holotype, male; (B) O. taksini, holotype, male; (C) O. tomokunii, paratype, female;(D) O. filiferus, paratype, female;(E) O. machaerus, holotype, male; (F) O. inthanonus, holotype,male. Abbreviations: eva = evaporatorium; mf = median furrow; op = ostiolar peritreme; sca = supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.3990385" httpUri="https://zenodo.org/record/3990385/files/figure.png" pageId="28" pageNumber="1131">6F</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C42D9325CFD11FD0B" box="[686,714,862,887]" captionStart="Figure 9" captionStartId="12.[160,225,1513,1536]" captionTargetBox="[178,1138,142,1482]" captionTargetId="figure@12.[178,1138,142,1482]" captionTargetPageId="12" captionText="Figure 9. Head and pronotum, male, dorsal view. (A) O. sakaerat, holotype; (B) O. taksini, holotype; (C) O. tomokunii, holotype; (D) O. filiferus, paratype; (E) O. machaerus, paratype; (F) O. inthanonus, holotype. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990391" httpUri="https://zenodo.org/record/3990391/files/figure.png" pageId="28" pageNumber="1131">9F</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C42AE325CFCDDFD0B" box="[729,774,862,887]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="28" pageNumber="1131">12C</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C4362325CFCBBFD0B" box="[789,864,861,887]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="28" pageNumber="1131">
16D
<emphasis id="50EA6B3320440D3C4333325FFC89FD0A" box="[836,850,861,886]" italics="true" pageId="28" pageNumber="1131">–</emphasis>
F
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20440D3C40D732A4FC5FFA35" pageId="28" pageNumber="1131" type="materials_examined">
<paragraph id="6221B72120440D3C40D732A4FE9EFDC3" blockId="28.[160,1156,934,1097]" box="[160,325,934,959]" pageId="28" pageNumber="1131">
<heading id="3969004D20440D3C40D732A4FE9EFDC3" box="[160,325,934,959]" fontSize="11" level="3" pageId="28" pageNumber="1131" reason="5">
<emphasis id="50EA6B3320440D3C40D732A4FE9EFDC3" bold="true" box="[160,325,934,959]" inLineHeading="true" italics="true" pageId="28" pageNumber="1131" reason="3">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120440D3C40D732CBFC5FFA35" blockId="28.[160,1156,934,1097]" pageId="28" pageNumber="1131">
<materialsCitation id="D2F6BD7C20440D3C40D732CBFC5FFA35" ID-GBIF-Occurrence="2831862356" collectingDate="1983-10-20" collectionCode="NSMT" collectorName="M. Sakai &amp; Right" country="Thailand" elevation="1300" location="Maeo Khun Klang" pageId="28" pageNumber="1131" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="50EA6B3320440D3C40D732CBFED1FD9D" bold="true" box="[160,266,969,993]" italics="true" pageId="28" pageNumber="1131">
<typeStatus id="BD25098320440D3C40D732CBFED1FD9D" box="[160,266,969,993]" pageId="28" pageNumber="1131" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="74987CA820440D3C416F32CBFEF7FD9D" box="[280,300,969,993]" pageId="28" pageNumber="1131" type="male">♂</specimenCount>
(NSMT-I-He 74287;
<figureCitation id="FAA5ABA420440D3C427A32CBFD48FD9D" box="[525,659,969,993]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figures 3I, J</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C42D632CBFD66FD9D" box="[673,701,969,993]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="28" pageNumber="1131">5K</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C42BC32CBFD3DFD9D" box="[715,742,969,993]" captionStart="Figure 6" captionStartId="7.[160,225,1375,1398]" captionTargetBox="[178,1138,142,1344]" captionTargetId="figure@7.[178,1138,142,1345]" captionTargetPageId="7" captionText="Figure 6. SEM images of ostiolar peritreme and evaporatorium, left lateroventral view. (A) O. sakaerat, holotype, male; (B) O. taksini, holotype, male; (C) O. tomokunii, paratype, female;(D) O. filiferus, paratype, female;(E) O. machaerus, holotype, male; (F) O. inthanonus, holotype,male. Abbreviations: eva = evaporatorium; mf = median furrow; op = ostiolar peritreme; sca = supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.3990385" httpUri="https://zenodo.org/record/3990385/files/figure.png" pageId="28" pageNumber="1131">6F</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C428332CBFCD4FD9D" box="[756,783,969,993]" captionStart="Figure 9" captionStartId="12.[160,225,1513,1536]" captionTargetBox="[178,1138,142,1482]" captionTargetId="figure@12.[178,1138,142,1482]" captionTargetPageId="12" captionText="Figure 9. Head and pronotum, male, dorsal view. (A) O. sakaerat, holotype; (B) O. taksini, holotype; (C) O. tomokunii, holotype; (D) O. filiferus, paratype; (E) O. machaerus, paratype; (F) O. inthanonus, holotype. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990391" httpUri="https://zenodo.org/record/3990391/files/figure.png" pageId="28" pageNumber="1131">9F</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C436B32CBFC9CFD9D" box="[796,839,969,993]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="28" pageNumber="1131">12C</figureCitation>
,
<figureCitation id="FAA5ABA420440D3C432232CBFC46FD9D" box="[853,925,969,993]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="28" pageNumber="1131">
16D
<emphasis id="50EA6B3320440D3C43F532CBFC54FD9D" box="[898,911,969,993]" italics="true" pageId="28" pageNumber="1131">–</emphasis>
F
</figureCitation>
), (
<collectingCountry id="1A89F7B120440D3C43CF32CBFBEFFD9D" box="[952,1076,969,993]" name="Thailand" pageId="28" pageNumber="1131">THAILAND</collectingCountry>
),
<location id="6741E1FA20440D3C443232CBFEFFFA78" LSID="urn:lsid:plazi:treatment:EA37063720440D3E41403239FEC8FCDD:6741E1FA20440D3C443232CBFEFFFA78" country="Thailand" name="Maeo Khun Klang" pageId="28" pageNumber="1131">Maeo Khun Klang</location>
,
<quantity id="A5661AC420440D3C415A32EEFE58FA78" box="[301,387,1004,1028]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="28" pageNumber="1131" unit="m" value="1300.0">
<elevation id="E9B3501220440D3C415A32EEFE58FA78" box="[301,387,1004,1028]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="28" pageNumber="1131" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, Doi Inthanon,
<date id="162091E120440D3C425932EEFD30FA78" box="[558,747,1004,1028]" pageId="28" pageNumber="1131" value="1983-10-20">
<collectingDate id="0664680920440D3C425932EEFD30FA78" box="[558,747,1004,1028]" pageId="28" pageNumber="1131" value="1983-10-20">20 October 1983</collectingDate>
</date>
,
<collectorName id="CF6BD2F720440D3C428232EEFC8AFA78" box="[757,849,1004,1028]" pageId="28" pageNumber="1131">M. Sakai</collectorName>
(
<collectionCode id="048F2FE420440D3C432A32EEFC7EFA78" box="[861,933,1004,1028]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="28" pageNumber="1131">NSMT</collectionCode>
).
<collectorName id="CF6BD2F720440D3C43C332EEFC34FA78" box="[948,1007,1004,1028]" pageId="28" pageNumber="1131">Right</collectorName>
antenna, left antennal segments III and IV, pronotal corner setae excepting left posterolateral one, and long, erect setae between each eye and ocellus are missing.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20440D3C40D7357BFBE3FB1E" pageId="28" pageNumber="1131" type="diagnosis">
<paragraph id="6221B72120440D3C40D7357BFECFFAEE" blockId="28.[160,1155,1145,1378]" box="[160,276,1145,1170]" pageId="28" pageNumber="1131">
<heading id="3969004D20440D3C40D7357BFECFFAEE" box="[160,276,1145,1170]" fontSize="11" level="3" pageId="28" pageNumber="1131" reason="5">
<emphasis id="50EA6B3320440D3C40D7357BFECFFAEE" bold="true" box="[160,276,1145,1170]" inLineHeading="true" italics="true" pageId="28" pageNumber="1131" reason="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120440D3C40D7359EFBE3FB1E" blockId="28.[160,1155,1145,1378]" pageId="28" pageNumber="1131">
Recognised by the following characters: body generally blackish brown (
<figureCitation id="FAA5ABA420440D3C4382359EFBADFAC8" box="[1013,1142,1180,1204]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3I, J</figureCitation>
); hemelytra brown to greyish brown, embolium posteriorly darkened, cuneus darker (
<figureCitation id="FAA5ABA420440D3C40D035E0FED6FA86" box="[167,269,1250,1274]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3I</figureCitation>
); legs yellowish brown, meso- and metafemora dark brown (
<figureCitation id="FAA5ABA420440D3C43B935E0FBEDFA86" box="[974,1078,1250,1274]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3J</figureCitation>
); male profemora with six fuscous small teeth on ventral side; cone thin, acute apicad, elevated dorsally with a projection (
<figureCitation id="FAA5ABA420440D3C41AC3425FDAFFB43" box="[475,628,1319,1343]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="28" pageNumber="1131">
Figure 16D
<emphasis id="50EA6B3320440D3C422D3425FDBCFB43" box="[602,615,1319,1343]" italics="true" pageId="28" pageNumber="1131">–</emphasis>
F
</figureCitation>
); denticule lacking; flagellum robust, roundly curved, noticeably expanded from base to middle in lateral view (
<figureCitation id="FAA5ABA420440D3C43E33448FBF0FB1E" box="[916,1067,1353,1378]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="28" pageNumber="1131">
Figure 16D
<emphasis id="50EA6B3320440D3C4467344BFBC6FB1D" box="[1040,1053,1353,1377]" italics="true" pageId="28" pageNumber="1131">–</emphasis>
F
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20440D3D40D73493FE3DF8C3" lastPageId="29" lastPageNumber="1132" pageId="28" pageNumber="1131" type="description">
<paragraph id="6221B72120440D3C40D73493FEFDFBD6" blockId="28.[160,1155,1425,1727]" box="[160,294,1425,1450]" pageId="28" pageNumber="1131">
<heading id="3969004D20440D3C40D73493FEFDFBD6" box="[160,294,1425,1450]" fontSize="11" level="3" pageId="28" pageNumber="1131" reason="5">
<emphasis id="50EA6B3320440D3C40D73493FEFDFBD6" bold="true" box="[160,294,1425,1450]" inLineHeading="true" italics="true" pageId="28" pageNumber="1131" reason="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120440D3D40D734B6FD23FB3D" blockId="28.[160,1155,1425,1727]" lastBlockId="29.[160,1156,142,1345]" lastPageId="29" lastPageNumber="1132" pageId="28" pageNumber="1131">
<emphasis id="50EA6B3320440D3C40D734B6FEB3FBB1" bold="true" box="[160,360,1460,1485]" inLineHeading="true" italics="true" pageId="28" pageNumber="1131" reason="3">
Male (
<typeStatus id="BD25098320440D3C408534B6FE83FBB1" box="[242,344,1460,1485]" pageId="28" pageNumber="1131" type="holotype">holotype</typeStatus>
).
</emphasis>
Colouration: body generally blackish brown (
<figureCitation id="FAA5ABA420440D3C43D634B7FBF0FBB1" box="[929,1067,1461,1485]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3I, J</figureCitation>
). Head brown to blackish brown, with paler apex (
<figureCitation id="FAA5ABA420440D3C42DA34D5FCC5FB93" box="[685,798,1495,1519]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="28" pageNumber="1131">Figure 5K</figureCitation>
); eyes reddish brown, ocellus and its surrounding area red to reddish brown. Antennal segments I and II yellowish brown (
<figureCitation id="FAA5ABA420440D3C408B371FFEB1F849" box="[252,362,1565,1589]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="28" pageNumber="1131">Figure 5K</figureCitation>
). Labium uniformly pale yellow, but segments I and II with fuscous tinge (
<figureCitation id="FAA5ABA420440D3C4085373DFEBAF82B" box="[242,353,1599,1623]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3J</figureCitation>
). Pronotum and scutellum uniformly blackish brown (
<figureCitation id="FAA5ABA420440D3C4475373DFBADF82B" box="[1026,1142,1599,1623]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="28" pageNumber="1131">Figure 5K</figureCitation>
). Hemelytra brown to greyish brown; embolium posteriorly darkened; cuneus darker than remainder area of corium; membrane uniformly somber dark brown (
<figureCitation id="FAA5ABA420440D3C44783787FBADF8E1" box="[1039,1142,1669,1693]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3I</figureCitation>
). Legs pale yellow to yellowish brown; meso- and metafemora dark brown (
<figureCitation id="FAA5ABA420440D3C447B37A5FBADF8C3" box="[1036,1142,1703,1727]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="28" pageNumber="1131">Figure 3J</figureCitation>
). Venter of thorax and abdomen dark brown, tinged with darker orange brown (
<figureCitation id="FAA5ABA420450D3D40DF31B3FECCFEB5" box="[168,279,177,201]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="29" pageNumber="1132">Figure 3J</figureCitation>
). Structure: body elongate oval (
<figureCitation id="FAA5ABA420450D3D42C531B3FCC5FEB5" box="[690,798,177,201]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="29" pageNumber="1132">Figure 3I</figureCitation>
). Head smooth, shiny, about 0.66 times as long as width across eyes, and with a longer erect seta on each side of clypeus, and near anteromedial margin of each eye; ante-ocular portion 0.64 times as long as length of eye in dorsal view; vertex about 1.8 times as wide as eye in dorsal view; eye 1.4 times as long as eye width in dorsal view; postocular portion constricted; neck distinct (
<figureCitation id="FAA5ABA420450D3D413E305CFE63FF0A" box="[329,440,350,374]" captionStart="Figure 9" captionStartId="12.[160,225,1513,1536]" captionTargetBox="[178,1138,142,1482]" captionTargetId="figure@12.[178,1138,142,1482]" captionTargetPageId="12" captionText="Figure 9. Head and pronotum, male, dorsal view. (A) O. sakaerat, holotype; (B) O. taksini, holotype; (C) O. tomokunii, holotype; (D) O. filiferus, paratype; (E) O. machaerus, paratype; (F) O. inthanonus, holotype. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990391" httpUri="https://zenodo.org/record/3990391/files/figure.png" pageId="29" pageNumber="1132">Figure 9F</figureCitation>
). Antennal segment I stout, much exceeding apex of head, sparsely covered with short suberect setae; segment II 0.63 times as long as head width across eyes, densely covered with suberect setae which are shorter than width of the segment (
<figureCitation id="FAA5ABA420450D3D411F30C4FE39FFA2" box="[360,482,454,478]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="29" pageNumber="1132">Figures 5K</figureCitation>
,
<figureCitation id="FAA5ABA420450D3D418430C4FDD5FFA2" box="[499,526,454,478]" captionStart="Figure 9" captionStartId="12.[160,225,1513,1536]" captionTargetBox="[178,1138,142,1482]" captionTargetId="figure@12.[178,1138,142,1482]" captionTargetPageId="12" captionText="Figure 9. Head and pronotum, male, dorsal view. (A) O. sakaerat, holotype; (B) O. taksini, holotype; (C) O. tomokunii, holotype; (D) O. filiferus, paratype; (E) O. machaerus, paratype; (F) O. inthanonus, holotype. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990391" httpUri="https://zenodo.org/record/3990391/files/figure.png" pageId="29" pageNumber="1132">9F</figureCitation>
). Labium sparsely covered with short suberect setae. Pronotum shiny, smooth, sparsely covered with short decumbent setae and tiny punctures; anterior margin slightly concave, width about as wide as mesal length; lateral margin moderately curved; lateral carinae distinctly expanded; posterior margin concave, width 2.5 times as wide as anterior pronotal width; collar distinct, strigose, demarcated by shallow transverse impression from callus; callus strongly convex, with scattered short setae, demarcated posteriorly by deep sinuate transverse impression (
<figureCitation id="FAA5ABA420450D3D40DF33BBFEF1FCAD" box="[168,298,697,721]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="29" pageNumber="1132">Figures 5K</figureCitation>
,
<figureCitation id="FAA5ABA420450D3D413633BBFE87FCAD" box="[321,348,697,721]" captionStart="Figure 9" captionStartId="12.[160,225,1513,1536]" captionTargetBox="[178,1138,142,1482]" captionTargetId="figure@12.[178,1138,142,1482]" captionTargetPageId="12" captionText="Figure 9. Head and pronotum, male, dorsal view. (A) O. sakaerat, holotype; (B) O. taksini, holotype; (C) O. tomokunii, holotype; (D) O. filiferus, paratype; (E) O. machaerus, paratype; (F) O. inthanonus, holotype. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990391" httpUri="https://zenodo.org/record/3990391/files/figure.png" pageId="29" pageNumber="1132">9F</figureCitation>
). Scutellum nearly equilateral, shorter than basal width, deeply depressed through middle, sparsely covered with short decumbent setae and tiny punctures. Hemelytra overall sparsely covered with short decumbent setae and tiny punctures; maximum width of endocorium twice as wide as embolium; cuneal margin 0.5 times as long as embolial margin; membrane with one indistinct vein located near costal margin. Ostiolar peritreme similar in general shape to
<emphasis id="50EA6B3320450D3D43D03264FBC4FD02" box="[935,1055,870,894]" italics="true" pageId="29" pageNumber="1132">
O.
<emphasis id="50EA6B3320450D3D43A73264FC06FD02" bold="true" box="[976,989,870,894]" italics="true" pageId="29" pageNumber="1132">fi</emphasis>
liferus
</emphasis>
and
<taxonomicName id="A59ECCA220450D3D441C3264FEC2FDDD" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="1132" phylum="Arthropoda" rank="species" species="machaerus">
<emphasis id="50EA6B3320450D3D441C3264FEC2FDDD" italics="true" pageId="29" pageNumber="1132">O. machaerus</emphasis>
</taxonomicName>
; anterior area to median furrow in ostiolar peritreme smooth, about as wide as maximum width of posterior area to median furrow; posterior area squamous entirely; supracoxal area transversely rugose (
<figureCitation id="FAA5ABA420450D3D42C832CCFCF5FD9A" box="[703,814,974,998]" captionStart="Figure 6" captionStartId="7.[160,225,1375,1398]" captionTargetBox="[178,1138,142,1344]" captionTargetId="figure@7.[178,1138,142,1345]" captionTargetPageId="7" captionText="Figure 6. SEM images of ostiolar peritreme and evaporatorium, left lateroventral view. (A) O. sakaerat, holotype, male; (B) O. taksini, holotype, male; (C) O. tomokunii, paratype, female;(D) O. filiferus, paratype, female;(E) O. machaerus, holotype, male; (F) O. inthanonus, holotype,male. Abbreviations: eva = evaporatorium; mf = median furrow; op = ostiolar peritreme; sca = supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.3990385" httpUri="https://zenodo.org/record/3990385/files/figure.png" pageId="29" pageNumber="1132">Figure 6F</figureCitation>
). Legs densely covered with stramineous decumbent setae; male protrochanters bearing one fuscous small tooth on ventral side; male profemora with six fuscous small teeth on ventral side; male protibiae with a row of 16 fuscous small teeth on ventral side. Male genitalia (
<figureCitation id="FAA5ABA420450D3D44473534FF10FA0D" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="29" pageNumber="1132">Figures 12C</figureCitation>
,
<figureCitation id="FAA5ABA420450D3D40AD355BFEF8FA0D" box="[218,291,1113,1137]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="29" pageNumber="1132">
16D
<emphasis id="50EA6B3320450D3D417F355BFECEFA0D" box="[264,277,1113,1137]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
F
</figureCitation>
): pygophore ovoid shaped, posteroventrally covered with four long, stout setae which are shorter than length of pygophore (
<figureCitation id="FAA5ABA420450D3D43743579FC5AFAEF" box="[771,897,1147,1171]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="29" pageNumber="1132">Figure 12C</figureCitation>
); mediodorsal surface densely distributed with short, suberect setae; cone thin, acute apicad in dorsal view, elevated dorsally with a projection (
<figureCitation id="FAA5ABA420450D3D423735C3FD02FAA5" box="[576,729,1217,1241]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="29" pageNumber="1132">
Figure 16D
<emphasis id="50EA6B3320450D3D42C835C3FD17FAA5" box="[703,716,1217,1241]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
F
</figureCitation>
); denticule lacking; flagellum robust, roundly curved, longer than twice of maximum width of cone, basally thickened in dorsal view, noticeably expanded from base to middle in lateral view, extending well beyond the left edge of pygophore (
<figureCitation id="FAA5ABA420450D3D4226342BFD30FB3D" box="[593,747,1321,1345]" captionStart="Figure 16" captionStartId="38.[160,225,1534,1557]" captionTargetBox="[178,1130,142,1503]" captionTargetId="figure@38.[178,1138,142,1503]" captionTargetPageId="38" captionText="Figure 16. Paramere, three different views. (A–C) O. machaerus, holotype; (D–F) O. inthanonus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990407" httpUri="https://zenodo.org/record/3990407/files/figure.png" pageId="29" pageNumber="1132">
Figure 16D
<emphasis id="50EA6B3320450D3D42A6342BFD05FB3D" box="[721,734,1321,1345]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
F
</figureCitation>
).
</paragraph>
<paragraph id="6221B72120450D3D40D7347DFEA4FBE4" blockId="29.[160,383,1407,1432]" box="[160,383,1407,1432]" pageId="29" pageNumber="1132">
<emphasis id="50EA6B3320450D3D40D7347DFF27FBE4" bold="true" box="[160,252,1407,1432]" inLineHeading="true" italics="true" pageId="29" pageNumber="1132" reason="3">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="6221B72120450D3D40D734D4FE4FFB93" blockId="29.[160,1155,1494,1727]" box="[160,404,1494,1519]" pageId="29" pageNumber="1132">
<heading id="3969004D20450D3D40D734D4FE4FFB93" box="[160,404,1494,1519]" fontSize="11" level="3" pageId="29" pageNumber="1132" reason="5">
<emphasis id="50EA6B3320450D3D40D734D4FE4FFB93" bold="true" box="[160,404,1494,1519]" inLineHeading="true" italics="true" pageId="29" pageNumber="1132" reason="3">Measurements (mm)</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120450D3D40D734F8FE3DF8C3" blockId="29.[160,1155,1494,1727]" pageId="29" pageNumber="1132">
[
<specimenCount id="74987CA820450D3D40D034F8FF60F86E" box="[167,187,1530,1554]" pageId="29" pageNumber="1132" type="male">♂</specimenCount>
(n = 1)]. Body length 2.05; head length (excluding neck) 0.25; head width across eyes 0.38; vertex width 0.18; width between ocelli 0.15; lengths of antennal segments I
<emphasis id="50EA6B3320450D3D4437371EFB96F848" box="[1088,1101,1564,1588]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
II: I
<emphasis id="50EA6B3320450D3D4401371EFB58F848" box="[1142,1155,1564,1588]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
0.09, II
<emphasis id="50EA6B3320450D3D4083373DFEDAF82B" box="[244,257,1599,1623]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
0.24, III
<emphasis id="50EA6B3320450D3D4113373DFEAAF82B" box="[356,369,1599,1623]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
missing, IV
<emphasis id="50EA6B3320450D3D4189373DFDD0F82B" box="[510,523,1599,1623]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
missing; lengths of labial segments II
<emphasis id="50EA6B3320450D3D43C9373DFC10F82B" box="[958,971,1599,1623]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
IV: II
<emphasis id="50EA6B3320450D3D4472373DFBC9F82B" box="[1029,1042,1599,1623]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
0.09, III
<emphasis id="50EA6B3320450D3D4401373DFB58F82B" box="[1142,1155,1599,1623]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
0.25, IV
<emphasis id="50EA6B3320450D3D408C3760FED3F806" box="[251,264,1634,1658]" italics="true" pageId="29" pageNumber="1132">–</emphasis>
0.22; anterior pronotal width 0.30; mesal pronotal length 0.30; basal pronotal width 0.76; length of embolial margin 0.68; length of cuneal margin 0.35; maximum width across hemelytra 0.84.
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20460D3E40D7318CFD05FE90" pageId="30" pageNumber="1133" type="etymology">
<paragraph id="6221B72120460D3E40D7318CFEC6FEDB" blockId="30.[160,1155,142,236]" box="[160,285,142,167]" pageId="30" pageNumber="1133">
<heading id="3969004D20460D3E40D7318CFEC6FEDB" box="[160,285,142,167]" fontSize="11" level="3" pageId="30" pageNumber="1133" reason="5">
<emphasis id="50EA6B3320460D3E40D7318CFEC6FEDB" bold="true" box="[160,285,142,167]" inLineHeading="true" italics="true" pageId="30" pageNumber="1133" reason="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120460D3E40D731B0FD05FE90" blockId="30.[160,1155,142,236]" pageId="30" pageNumber="1133">
<materialsCitation id="D2F6BD7C20460D3E40D731B0FD01FE90" ID-GBIF-Occurrence="2831862402" country="Thailand" location="Doi Inthanon in Chom Thong District" pageId="30" pageNumber="1133" specimenCount="1" stateProvince="Chiang Mai">
Named after the
<typeStatus id="BD25098320460D3E410F31B0FE71FEB6" box="[376,426,178,202]" pageId="30" pageNumber="1133">type</typeStatus>
locality,
<location id="6741E1FA20460D3E426931B0FC31FEB6" LSID="urn:lsid:plazi:treatment:EA37063720440D3E41403239FEC8FCDD:6741E1FA20460D3E426931B0FC31FEB6" box="[542,1002,178,202]" country="Thailand" name="Doi Inthanon in Chom Thong District" pageId="30" pageNumber="1133" stateProvince="Chiang Mai">Doi Inthanon in Chom Thong District</location>
,
<collectingRegion id="A05A79C320460D3E438B31B0FED8FE90" country="Thailand" name="Chiang Mai" pageId="30" pageNumber="1133">Chiang Mai Province</collectingRegion>
, northern
<collectingCountry id="1A89F7B120460D3E410C31D6FE05FE90" box="[379,478,212,236]" name="Thailand" pageId="30" pageNumber="1133">Thailand</collectingCountry>
; a noun in apposition
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20460D3E40D73016FDD6FF2C" box="[160,525,276,336]" pageId="30" pageNumber="1133" type="distribution">
<paragraph id="6221B72120460D3E40D73016FEF7FF51" blockId="30.[160,525,276,336]" box="[160,300,276,301]" pageId="30" pageNumber="1133">
<heading id="3969004D20460D3E40D73016FEF7FF51" box="[160,300,276,301]" fontSize="11" level="3" pageId="30" pageNumber="1133" reason="5">
<emphasis id="50EA6B3320460D3E40D73016FEF7FF51" bold="true" box="[160,300,276,301]" inLineHeading="true" italics="true" pageId="30" pageNumber="1133" reason="3">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120460D3E40D7303AFDD6FF2C" blockId="30.[160,525,276,336]" box="[160,525,312,336]" pageId="30" pageNumber="1133">
Northern
<collectingCountry id="1A89F7B120460D3E4179303AFEB5FF2C" box="[270,366,312,336]" name="Thailand" pageId="30" pageNumber="1133">Thailand</collectingCountry>
(
<collectingRegion id="A05A79C320460D3E410A303AFDD8FF2C" box="[381,515,312,336]" country="Thailand" name="Chiang Mai" pageId="30" pageNumber="1133">Chiang Mai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20460D3E40D7307AFEC8FCDD" pageId="30" pageNumber="1133" type="discussion">
<paragraph id="6221B72120460D3E40D7307AFEDDFFED" blockId="30.[160,1156,376,574]" box="[160,262,376,401]" pageId="30" pageNumber="1133">
<heading id="3969004D20460D3E40D7307AFEDDFFED" box="[160,262,376,401]" fontSize="11" level="3" pageId="30" pageNumber="1133" reason="5">
<emphasis id="50EA6B3320460D3E40D7307AFEDDFFED" bold="true" box="[160,262,376,401]" inLineHeading="true" italics="true" pageId="30" pageNumber="1133" reason="3">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120460D3E40D73099FB5FFC42" blockId="30.[160,1156,376,574]" pageId="30" pageNumber="1133">
<taxonomicName id="A59ECCA220460D3E40D73099FE88FFCF" box="[160,339,411,435]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1133" phylum="Arthropoda" rank="species" species="inthanonus">
<emphasis id="50EA6B3320460D3E40D73099FE88FFCF" box="[160,339,411,435]" italics="true" pageId="30" pageNumber="1133">Orius inthanonus</emphasis>
</taxonomicName>
is distinctive among the species of the subgenus
<taxonomicName id="A59ECCA220460D3E43033099FC71FFCF" authorityName="Wolff" authorityYear="1811" box="[884,938,411,435]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1133" phylum="Arthropoda" rank="genus">
<emphasis id="50EA6B3320460D3E43033099FC71FFCF" box="[884,938,411,435]" italics="true" pageId="30" pageNumber="1133">Orius</emphasis>
</taxonomicName>
in having brown to greyish brown hemelytra with posteriorly darkened embolium and darker cuneus and the male profemora with six fuscous small teeth on ventral side. The structure of male genitalia of
<taxonomicName id="A59ECCA220460D3E40D73301FEE9FC67" box="[160,306,515,539]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1133" phylum="Arthropoda" rank="species" species="inthanonus">
<emphasis id="50EA6B3320460D3E40D73301FEE9FC67" box="[160,306,515,539]" italics="true" pageId="30" pageNumber="1133">O. inthanonus</emphasis>
</taxonomicName>
somewhat resembles that of
<taxonomicName id="A59ECCA220460D3E421B3301FD20FC67" box="[620,763,515,539]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1133" phylum="Arthropoda" rank="species" species="machaerus">
<emphasis id="50EA6B3320460D3E421B3301FD20FC67" box="[620,763,515,539]" italics="true" pageId="30" pageNumber="1133">O. machaerus</emphasis>
</taxonomicName>
; however, it differs from the latter by the roundly curved flagellum being noticeably expanded from base to middle in lateral view.
</paragraph>
<paragraph id="6221B72120460D3E40D73364FF22FC03" blockId="30.[160,275,614,673]" box="[160,249,614,639]" pageId="30" pageNumber="1133">
<heading id="3969004D20460D3E40D73364FF22FC03" box="[160,249,614,639]" fontSize="11" level="3" pageId="30" pageNumber="1133" reason="5">
<emphasis id="50EA6B3320460D3E40D73364FF22FC03" bold="true" box="[160,249,614,639]" inLineHeading="true" italics="true" pageId="30" pageNumber="1133" reason="3">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120460D3E40D7338BFEC8FCDD" blockId="30.[160,275,614,673]" box="[160,275,649,673]" pageId="30" pageNumber="1133">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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