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<document ID-CLB-Dataset="268381" ID-DOI="http://dx.doi.org/10.3897/vz.73.e109854" ID-GBIF-Dataset="83e9e5f1-7a0c-4650-aa6f-3df62c9a9a0f" ID-Pensoft-Pub="2625-8498-73-697" ID-Pensoft-UUID="7955E10D927A5196997DE0497DD45CCB" ID-ZooBank="CA5997B2B4624B40B861EFF08FC1768D" ModsDocID="2625-8498-73-697" checkinTime="1695177574508" checkinUser="pensoft" docAuthor="Idiiatullina, Sabira S., Pawangkhanant, Parinya, Tawan, Tanapong, Worranuch, Thanawut, Dechochai, Bunyarit, Suwannapoom, Chatmongkon, Nguyen, Tan Van, Chanhome, Lawan &amp; Poyarkov, Nikolay A." docDate="2023" docId="A03C6F98B1935B4092551746C8396712" docLanguage="en" docName="VertZool 73: 697-716" docOrigin="Vertebrate Zoology 73" docPubDate="2023-09-19" docSource="http://dx.doi.org/10.3897/vz.73.e109854" docTitle="Trimeresurus ciliaris Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov 2023, sp. nov." docType="treatment" docUuid="898F9CF2-0457-4F77-B53A-BB8D1E8AB3F2" docUuidSource="ZooBank" docVersion="5" id="7955E10D927A5196997DE0497DD45CCB" lastPageNumber="697" masterDocId="7955E10D927A5196997DE0497DD45CCB" masterDocTitle="Limestone jewel: A new colourful karst-dwelling pitviper (Serpentes: Viperidae: Trimeresurus) from the poorly explored borderlands of southern peninsular Thailand" masterLastPageNumber="716" masterPageNumber="697" pageNumber="697" updateTime="1732832345199" updateUser="ExternalLinkService">
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<mods:title id="54D076316C770A1FAF27F7823FDBE07F">Limestone jewel: A new colourful karst-dwelling pitviper (Serpentes: Viperidae: Trimeresurus) from the poorly explored borderlands of southern peninsular Thailand</mods:title>
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<mods:namePart id="2203646EA768FD1BE8EC7A7BE9CE7C8F">Idiiatullina, Sabira S.</mods:namePart>
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<mods:affiliation id="18E709DD1F59B362108D8E1FC1309907">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia</mods:affiliation>
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<mods:namePart id="8CC4217C6888AF7A64F619C73037AB30">Pawangkhanant, Parinya</mods:namePart>
<mods:nameIdentifier id="EBB369752D4F2B0E9AF266B45711A939" type="ORCID">https://orcid.org/0000-0002-0947-5729</mods:nameIdentifier>
<mods:affiliation id="C400ADE9DD6930EDA920A5B9F857F4EB">Rabbit in the Moon Foundation, Suanphueng, Ratchaburi 70180, Thailand &amp; Division of Fishery, School of Agriculture and Natural Resources, University of Phayao, Phayao 56000, Thailand</mods:affiliation>
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<mods:namePart id="F0C69519BB12132FF3EBA47CC598D141">Tawan, Tanapong</mods:namePart>
<mods:nameIdentifier id="B210F003D8235764906D5A27BF2BA2C7" type="ORCID">https://orcid.org/0000-0003-3448-1132</mods:nameIdentifier>
<mods:affiliation id="2446B1DB570A620385A20573108FA625">Queen Saovabha Memorial Institute, Thai Red Cross Society, Pathumwan, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="3C0D37EEE46D08CA8B75B5F69F63A9BA">Worranuch, Thanawut</mods:namePart>
<mods:affiliation id="03A9D3F04680BE05CE12B8361D914163">Tontan Travel Wildlife Tours, Phetchaburi 91160, Thailand</mods:affiliation>
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<mods:namePart id="BA5B0C7E6D606E5F62157B19AD906B08">Dechochai, Bunyarit</mods:namePart>
<mods:affiliation id="2F6C7EC9C6747C14AFD9B720C14DA738">Thaleban National Park, Wang Pra Chan, Kuan Don, Satun, Thailand</mods:affiliation>
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<mods:namePart id="D0B7BA226FC8A98680BDF69B474BD1E4">Suwannapoom, Chatmongkon</mods:namePart>
<mods:nameIdentifier id="C061CED4FD17C351342AC68A260EE4DE" type="ORCID">https://orcid.org/0000-0002-3342-1464</mods:nameIdentifier>
<mods:affiliation id="7B1628C3481701D0A9EF213140D87667">Division of Fishery, School of Agriculture and Natural Resources, University of Phayao, Phayao 56000, Thailand</mods:affiliation>
</mods:name>
<mods:name id="52119567A3CEA419B0AA3E07963BC24B" type="personal">
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<mods:namePart id="252D9136EEC3EA5F287E24DEA375EB06">Nguyen, Tan Van</mods:namePart>
<mods:nameIdentifier id="FE9E1DC0C141121D3FCD2DC35CE5060D" type="ORCID">https://orcid.org/0000-0001-5413-968X</mods:nameIdentifier>
<mods:affiliation id="5395A7FE07022431D7C870319E1BE65E">Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam</mods:affiliation>
<mods:nameIdentifier id="F9D562B625B7BCE72CEFB4663EB6E623" type="email">tan.sifasv@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0A84CB7D86F110EDB0B04BE41D1D4F52">Chanhome, Lawan</mods:namePart>
<mods:nameIdentifier id="E220CBF2ED455D04AAEF5C5CBCC825ED" type="ORCID">https://orcid.org/0000-0003-2425-5000</mods:nameIdentifier>
<mods:affiliation id="5E9AA1974FCF6A5522337DE24DEACC8D">Queen Saovabha Memorial Institute, Thai Red Cross Society, Pathumwan, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="87C1987CD3D6ED8C8AD3EBA670EE91A8" type="email">lchanhome@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="335B14568600FC8D27513153088384E7">Poyarkov, Nikolay A.</mods:namePart>
<mods:nameIdentifier id="9C8D2E91949729EB10B65329E500370C" type="ORCID">https://orcid.org/0000-0002-7576-2283</mods:nameIdentifier>
<mods:affiliation id="3F214BE141AE04AE46AEDFD4CF4E034B">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia &amp; College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, 550000, Vietnam</mods:affiliation>
<mods:nameIdentifier id="0F9168F6DAC2A2CEB270E7071402C6BE" type="email">n.poyarkov@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="90D54A07511327C2AA86DAB675C5569B">text</mods:typeOfResource>
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<mods:title id="F40200E0FC04D583473ABF7376A46684">Vertebrate Zoology</mods:title>
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<mods:part id="55FB3B4E8FD737E2D26682319989B5D4">
<mods:date id="B6B19333EAD0174A625215D94618482C">2023</mods:date>
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<mods:number id="A7083F1576AF19C2AF97DECC93B0007A">73</mods:number>
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<mods:classification id="C6CE4FBE7D8218D8257845B1C8A22FD9">journal article</mods:classification>
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<treatment id="A03C6F98B1935B4092551746C8396712" ID-GBIF-Taxon="213212069" LSID="urn:lsid:zoobank.org:act:898F9CF2-0457-4F77-B53A-BB8D1E8AB3F2" httpUri="http://treatment.plazi.org/id/A03C6F98B1935B4092551746C8396712" lastPageNumber="697" pageId="0" pageNumber="697" scope_family="Viperidae">
<subSubSection id="A6B43B2E03462B9A839243524F6B25E9" pageId="0" pageNumber="697" type="nomenclature">
<paragraph id="2D1A070C1ECD519D6D16BF79D4A96B06" pageId="0" pageNumber="697">
<taxonomicName id="EAD673914270C72A841407463BED9824" LSID="https://zoobank.org/898F9CF2-0457-4F77-B53A-BB8D1E8AB3F2" authority="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov, 2023" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris" status="sp. nov.">Trimeresurus ciliaris</taxonomicName>
<taxonomicNameLabel id="740341D83CD93AF72C024078C2A23F3C" pageId="0" pageNumber="697">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A340042E33CE111EC039E1B8938F1DDC" pageId="0" pageNumber="697" type="description">
<paragraph id="E4A28D3522F5DBE4EBCEF189BA855A92" pageId="0" pageNumber="697">
<figureCitation id="27702B48A9A52C9AEEAA06F620D5CF08" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">Figures 3</figureCitation>
<figureCitation id="EB1DCFC5586E7A515DFBDCD696357C01" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in head scalation and coloration in male paratypes of Trimeresurus ciliaris sp. nov.: A ZMMU Re- 17661; B ZMMU Re- 17662; C AUP- 02011; and D ZMMU Re- 17663. See Table 4 for details. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure4" httpUri="https://binary.pensoft.net/fig/908086" pageId="0" pageNumber="697">, 4</figureCitation>
<figureCitation id="3573396ECF050CF3AFD4DFEFDB3C95C1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="01C5D068A9D83D25FB356AF93F6445A5" pageId="0" pageNumber="697" type="chresonymy">
<paragraph id="47A8BD3BF58DCC79415D82914AD4A4CC" pageId="0" pageNumber="697">Chresonymy.</paragraph>
<paragraph id="CA5DE7B47867500AD6A14CA1244D3A12" pageId="0" pageNumber="697">
<taxonomicName id="45221A7ABE2157DA10972225D7B01F52" baseAuthorityName="Vogel" baseAuthorityYear="1991" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptelytrops venustus" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="venustus">
<emphasis id="55A042D9A07C2FCBBBE583E8D1EA8AE2" bold="true" pageId="0" pageNumber="697">Cryptelytrops cf. venustus</emphasis>
</taxonomicName>
(partim):
<bibRefCitation id="5F4F28CAEFC65ABF518846E1A22E6F24" author="Chan, KO" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="697" pagination="253 - 259" refId="B3" refString="Chan, KO, Grismer, LL, Anuar, S, Quah, E, Grismer, JL, Jr, PLW, Muin, MA, Ahmad, N, 2011. A new species of Chiromantis Peters 1854 (Anura: Rhacophoridae) from Perlis State Park in extreme northern peninsular Malaysia with additional herpetofaunal records for the park. Russian Journal of Herpetology 18: 253 - 259" title="A new species of Chiromantis Peters 1854 (Anura: Rhacophoridae) from Perlis State Park in extreme northern peninsular Malaysia with additional herpetofaunal records for the park." volume="18" year="2011">Chan et al. (2011</bibRefCitation>
: 258).
</paragraph>
</subSubSection>
<subSubSection id="342E9CFA805F26D4ECBF25C75D88F73E" pageId="0" pageNumber="697" type="holotype">
<paragraph id="8F5E96A72B8574454FB8D030F46C08E1" pageId="0" pageNumber="697">Holotype.</paragraph>
<paragraph id="29BA8BFE6D675BDE1F8141F2D8EF0D6D" pageId="0" pageNumber="697">
ZMMU Re-17661, adult male collected by S. Idiiatullina, P. Pawangkhanant and T. Woranuch on 23 January 2023 near Thum Khao Ting, Palian District, Trang Province, Thailand (
<geoCoordinate id="8A49FC8A46E5439CF6C14E0F1082640D" degrees="07" direction="north" minutes="09.943" orientation="latitude" precision="1" value="7.1657166">07°09.943N</geoCoordinate>
,
<geoCoordinate id="EC2445D8CB7E271A235083234C990F95" degrees="99" direction="east" minutes="48.142" orientation="longitude" precision="1" value="99.80237">99°48.142E</geoCoordinate>
; elevation 28 m a.s.l).
</paragraph>
</subSubSection>
<subSubSection id="BBD6AD085D5192EBDBC808B98D106DAA" pageId="0" pageNumber="697" type="paratypes (n = 4)">
<paragraph id="15433886D0CDE3A74847B0C408D3C70A" pageId="0" pageNumber="697">Paratypes (n = 4).</paragraph>
<paragraph id="0CA608035CABE87082D5F06C32BFC804" pageId="0" pageNumber="697">QSMI 1538, adult female collected by P. Pawangkhanant on 18 June 2016 from the same locality as the holotype; AUP-02011, ZMMU Re-17662, ZMMU Re-17663, three adult males collected on 23 January 2023 from the same locality and by the same collectors as for the holotype.</paragraph>
</subSubSection>
<subSubSection id="8E71582BF1D6401D3CA79E816452C37F" pageId="0" pageNumber="697" type="diagnosis">
<paragraph id="7C6FE2DA03845A781393DC3AFA5EFFD1" pageId="0" pageNumber="697">Diagnosis.</paragraph>
<paragraph id="C9EC5873AC1AD86F367495AEA9BC4369" pageId="0" pageNumber="697">
A species of the genus
<taxonomicName id="FFDF9C73828D342EFECEB67AD4863975" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus" order="Squamata" pageId="0" pageNumber="697" rank="genus">Trimeresurus</taxonomicName>
which is assigned to the subgenus
<taxonomicName id="D00071468314B5B0BC606AD066B724AA" authority="Trimeresurus" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus (Trimeresurus)" order="Squamata" pageId="0" pageNumber="697" rank="subGenus" subGenus="Trimeresurus">Trimeresurus Trimeresurus</taxonomicName>
based on the following morphological attributes: a long papillose or calyculate hemipenis and partially fused first supralabial and nasal scales (
<bibRefCitation id="37364CFC1164C62F08F211FEFC6B86A3" DOI="https://doi.org/10.1016/j.ympev.2004.02.008" author="Malhotra, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="697" pagination="83 - 100" refId="B35" refString="Malhotra, A, Thorpe, RS, 2004a. A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution 32: 83 - 100, DOI: https://doi.org/10.1016/j.ympev.2004.02.008" title="A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis)." url="https://doi.org/10.1016/j.ympev.2004.02.008" volume="32" year="2004 a">Malhotra and Thorpe 2004a</bibRefCitation>
;
<bibRefCitation id="0C207971347C1605DEFEDB31EA9674B2" author="David, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="697" pagination="1 - 51" refId="B14" refString="David, P, Vogel, G, Dubois, A, 2011. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae). Zootaxa 2992: 1 - 51" title="On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae)." volume="2992" year="2011">David et al. 2011</bibRefCitation>
). The new species
<taxonomicName id="BBA3BDF9FE5A23D9C6FD5318F750D178" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="BF0889135675A2833BB74FD510A08552" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="A99F10374DBEA0EA5145ABCCC4005424" bold="true" pageId="0" pageNumber="697">sp. nov</emphasis>
. can be distinguished from all other congeneric species by the following combination of morphological characters: three or four small convex supraocular scales; internasals not in contact; small scale between nasal and the scale formed by the fused second supralabial and loreal present; dorsal scales in 17-17-15 rows; ventral scales 172-175 in males, 171 in a single female; subcaudal scales 59-63 in males, 52 in a single female, all paired; in life an emerald-green dorsum with reddish-brown cross-bands; creamy-white venter lacking dark dots or stripes on the lateral sides of the ventrals; iris olive-green with faded-brown horizontal stripe; tail dark brown mottled with rusty spots; white vertebral spots present in both sexes located approximately every two or three dorsal scales; dark brown spots forming discontinuous pattern present on 1-3 lateral dorsal scale rows; males having a reddish-brown postocular stripe.
</paragraph>
</subSubSection>
<subSubSection id="E180E2B48CA152BCBAEA1FFEFE8292B9" pageId="0" pageNumber="697" type="description">
<paragraph id="DFFBD0EEF6D120E92EC9EA630EE817C8" pageId="0" pageNumber="697">Description of the holotype.</paragraph>
<paragraph id="EFED781CF6B023EA6950E2DE92AE2CA3" pageId="0" pageNumber="697">
Adult male (Fig.
<figureCitation id="C455ABD290B7261243B4574F431FF121" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3</figureCitation>
), specimen in a good state of preservation. Body cylindrical, long and thin (SVL 345 mm, TaL 72 mm, TL 417 mm, TaL/TL 0.17) (Fig.
<figureCitation id="8DE2252979F3465C77BBA36558FD0617" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3A,B</figureCitation>
). Head triangular in dorsal view (Fig.
<figureCitation id="F862A7C579FE7B85AD5B775F01417D4D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3C</figureCitation>
), elongate, clearly distinct from the neck (HL 16.5 mm, HW 11.4 mm, HW/HL 0.69). Snout elongate, flattened and rounded in dorsal view (Fig.
<figureCitation id="F4AEE77438F434F2353EC2AF332953B9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3C</figureCitation>
), rather rectangular in lateral view (Fig.
<figureCitation id="7ACA4F6A5354B74B4EAC31854510DED3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E,F</figureCitation>
), with a very distinct and sharp canthus rostralis (SnL/HL 0.30, SnL/ED 1.92). Distance between eye and nostril 3.9 mm on both sides. Rostral slightly visible in dorsal aspect, triangular (Fig.
<figureCitation id="5F5C2A672EDCFAC77A772EC112C10252" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3C</figureCitation>
). Pupil vertically elliptical, loreal pit present, triangular in shape (Fig.
<figureCitation id="EEAA913094BAC18C27D065D7654FB7B3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E,F</figureCitation>
). Nostril completely enclosed in nasal scale; nasal scale partially fused with first supralabial (Fig.
<figureCitation id="3EB9F33E86B354A9FB45D29A91A16D3C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E</figureCitation>
). Shield bordering the anterior edge of loreal pit fused with second supralabial. Anterior subocular long, thin, and crescent-like, separated from the 4th and 5th supralabials by one row of scales; posterior subocular ovoid, separated from the 6th supralabial by 2/1 scales (Fig.
<figureCitation id="115203861E11D6880B3386FAF0270679" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E, F</figureCitation>
). Three preoculars on each side of the head; two upper preoculars located above the loreal pit, elongated, in contact with the single loreal which separates them from the nasal; lower preocular forming the lower margin of the loreal pit, lower preocular in contact with third supralabial (Fig.
<figureCitation id="ADFF51B1847861D7B071712EE587D456" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E, F</figureCitation>
). A small scale between the nasal and the scale formed by the fused second supralabial and loreal; 2/2 postoculars; 9/9 supralabials, third the largest (Fig.
<figureCitation id="5333DE393FFB3C57F50C96E1B121AF55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E,F</figureCitation>
); 11/10 infralabials, those of the first pair in contact with each other behind the mental; the first three pairs of infralabials in contact with the single pair of chin shields (Fig.
<figureCitation id="BD47863A1D063D003E4A6C3C1BB97A85" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3D</figureCitation>
). Six pairs of gulars aligned between the chin shields and the first preventral (Fig.
<figureCitation id="C8FE39E9FA07256D8A5428036DB8BA39" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3D</figureCitation>
). One large pair of enlarged internasals, separated by three small scales. Three small slightly convex supraoculars, same size as or smaller than preoculars and postoculars (Fig.
<figureCitation id="66A08F0499D416897D393812C9BD95AA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3C</figureCitation>
). Scales on snout and in the interorbital region smooth, irregular, subimbricate; temporal and occipital scales moderately keeled (Fig.
<figureCitation id="E9FE66AE6B4278DD843043EA257C8EF3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3C,E,F</figureCitation>
). Dorsal scales in 17-17-15 rows. Dorsal scales all strongly keeled, except the first row, which is smooth (Fig.
<figureCitation id="42DE0C80E581A91289427D2CA0267926" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3H</figureCitation>
). One preventral + 172 ventrals. Cloacal plate single; 61 subcaudals, all divided. Hemipenes long, papillose, and deeply forked with small, soft, basal spines (Fig.
<figureCitation id="D0A16AD8B20979670CCE5824E2CAB855" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3G</figureCitation>
). Each maxilla bearing a single large venom tooth; three teeth on palatine, 11 on pterygoid, and 12 on dentary.
</paragraph>
<caption id="F237FADBB83A844352C08A9F6918F75C" doi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697" start="Figure 3" startId="F3">
<paragraph id="5325AE965E412148FD8228BF8C79A3E1" pageId="0" pageNumber="697">
<emphasis id="74199B834B0CD1F34098ED1E57239AD4" bold="true" pageId="0" pageNumber="697">Figure 3.</emphasis>
The holotype of
<taxonomicName id="4FD0815550B44D48070BF312EA7DB912" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="82E27B59A8BB60B7D7B916684DD6AF5A" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="71406C7AC3B1988BA6343ECFC97EC352" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
in life (ZMMU Re-17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand.
<emphasis id="ED857272F41E8ECE658C7D1BB0292545" bold="true" pageId="0" pageNumber="697">A</emphasis>
dorsolateral and
<emphasis id="C3B8CBAD71387F94BC39C7331D19315D" bold="true" pageId="0" pageNumber="697">B</emphasis>
ventral view of body;
<emphasis id="DA080758CFB8D84A18E55F32635109A6" bold="true" pageId="0" pageNumber="697">C</emphasis>
head in dorsal,
<emphasis id="BC1ED38EEB84F409380B04A6B44B93F6" bold="true" pageId="0" pageNumber="697">D</emphasis>
in ventral,
<emphasis id="011905D878C4FA8AD49F87BA5E78A8E4" bold="true" pageId="0" pageNumber="697">E</emphasis>
in lateral right, and
<emphasis id="AD080148D9B8D9D357A7B68753B9F236" bold="true" pageId="0" pageNumber="697">F</emphasis>
in lateral left aspects;
<emphasis id="4D7603E72CFA6F3374CBC87C4AF8047E" bold="true" pageId="0" pageNumber="697">G</emphasis>
right hemipenis, partially everted;
<emphasis id="37CE9B71A0F034687D6445F03D6143E4" bold="true" pageId="0" pageNumber="697">H</emphasis>
close-up of dorsal scales. Photographs by P. Pawangkhanant.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9F99BFB61A9ACDE736AA4C42D4FA7033" pageId="0" pageNumber="697" type="coloration in life (figs 3–5)">
<paragraph id="31EDD70F2710A7BD2AE0F9359F8A48BF" pageId="0" pageNumber="697">
Coloration in life (Figs
<figureCitation id="829137AC00446677C0BEBA5E7E0E94B1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3</figureCitation>
-
<figureCitation id="DE988ECA26EA51FADF6B37C2D63F00CA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5</figureCitation>
).
</paragraph>
<paragraph id="0792F500B4A1085988C7931D80CFAFBC" pageId="0" pageNumber="697">
Dorsal surface of the head emerald-green with many scales partly or entirely dark red, especially on the snout, and on the interorbital, temporal and occipital regions (Fig.
<figureCitation id="F17F5D3612AEB6EE9EDFBF554C6A0E96" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3C</figureCitation>
). A reddish-brown postocular stripe extends from postocular scales to the neck (Fig.
<figureCitation id="DD3279FD32EAD05FFCBCC61832A87AE7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E,F</figureCitation>
). Black vertical pupil; iris olive-green with faded brown horizontal stripe (Fig.
<figureCitation id="0FC82B4CC1BDB64825F5EBAB09748C9E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E,F</figureCitation>
). The background color of the dorsum emerald-green, similar to that of the head, with about 75 irregular, dark red cross-bands (Fig.
<figureCitation id="F75F04EB093A15B313B1C36C909E61C4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3A</figureCitation>
). These red bands are about two dorsal scales long mediodorsally, but getting narrower on the lower flanks where they are about 1-2 dorsal scale in length. At the level of the vertebral row, the red bands are generally separated by one dorsal scale. White vertebral spots present along the vertebral scale row, approximately every two or three scales (Fig.
<figureCitation id="C43F840066C1C59B1101718438891E96" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3A</figureCitation>
). Ventrolateral stripe absent. Dark brown spots forming discontinuous pattern present on first to third lateral dorsal scale rows (Fig.
<figureCitation id="5D07189445B4CF1C2BB788D4E43FF09E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3H</figureCitation>
). Tail dark brown mottled with rusty spots. Ventral surface of the tail dark brown with white irregular bands, tail tip completely black. Infralabials greenish-white with faint brown spots (Fig.
<figureCitation id="1BB53DFC8B23E71B7A302E13365E47CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3E,F</figureCitation>
); the ventral surfaces of the head creamy white with bluish tint (Fig.
<figureCitation id="F0747B773F85097A59AC6BA9AD92A4BB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3D</figureCitation>
); the ventral surfaces of body creamy white (Fig.
<figureCitation id="5E9C4C810CD41627D42FEBCBF29F4F8A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The holotype of Trimeresurus ciliaris sp. nov. in life (ZMMU Re- 17661, adult male) from Thum Khao Ting, Palian District, Trang Province, Thailand. A dorsolateral and B ventral view of body; C head in dorsal, D in ventral, E in lateral right, and F in lateral left aspects; G right hemipenis, partially everted; H close-up of dorsal scales. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure3" httpUri="https://binary.pensoft.net/fig/908085" pageId="0" pageNumber="697">3B</figureCitation>
).
</paragraph>
<paragraph id="475A9E96D8B148989001A0C63F2E8717" pageId="0" pageNumber="697">In preservative the background dorsal color faded to greyish-brown, with less contrasting dark marks on the head and bands on the dorsum; the ventral color became whitish grey.</paragraph>
</subSubSection>
<subSubSection id="9CCCC86D5B602291064C3D9BB1DF3C6F" pageId="0" pageNumber="697" type="variation">
<paragraph id="E7A19CED15BAEB324171C34CB9533AB1" pageId="0" pageNumber="697">Variation.</paragraph>
<paragraph id="0083825E716833B10F09D17E7D28AFE1" pageId="0" pageNumber="697">
The main meristic and morphometric characters of the type series of
<taxonomicName id="1D9AAF04475A6C33B223DB82CC1AAA75" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="7DD8448F2816EB4964B3DB4F5128674C" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="025E897AFE17040294191C207EB472DE" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
are summarized in Table
<tableCitation id="8BD5B03F33B48AFF7CE535083B2D94E1" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological data on the type series of Trimeresurus ciliaris sp. nov. Notes: For abbreviations see Materials and methods section." httpUri="http://table.plazi.org/id/3DFF69BB9E3C24255F0BF12110F8CA24" pageId="0" pageNumber="697" tableUuid="3DFF69BB9E3C24255F0BF12110F8CA24">4</tableCitation>
; color variation of the type series is presented in Figures
<figureCitation id="012CAF31AF509B8433F68450D5247D99" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in head scalation and coloration in male paratypes of Trimeresurus ciliaris sp. nov.: A ZMMU Re- 17661; B ZMMU Re- 17662; C AUP- 02011; and D ZMMU Re- 17663. See Table 4 for details. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure4" httpUri="https://binary.pensoft.net/fig/908086" pageId="0" pageNumber="697">4</figureCitation>
,
<figureCitation id="8C4173225F86606DE8CAF74466B07BCC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5A</figureCitation>
,
<figureCitation id="27C374BF99C1A4C2EB1D7D57BDA9C60C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6A-B</figureCitation>
,
<figureCitation id="F86A24EEFCE87882F3C125CC6AEACFF3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitat of Trimeresurus ciliaris sp. nov. A microhabitat of the new species near the Thum Khao Ting Cave, Palian District, Trang Province, Thailand; B holotype ZMMU Re- 17661, adult male, in life in situ; C paratype QSMI 1538, adult female, in life in situ; D - E male specimen from Wang Kelian, Perlis State, Malaysia (not collected). Photographs by P. Pawangkhanant (A-C) and Azrul Azrizal Azmy (D-E)." figureDoi="10.3897/vz.73.e109854.figure7" httpUri="https://binary.pensoft.net/fig/908089" pageId="0" pageNumber="697">7B-E</figureCitation>
. All members of the type series, similarly to the holotype, show a nasal scale partially fused with the first supralabial. Head scalation of the paratypes generally agrees with that of the holotype, however male ZMMU Re-17662 has a suboculars fused in a single long crescent-shaped scale (Fig.
<figureCitation id="76AA67908DC0DA14DAA4D63AB6BD29C3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in head scalation and coloration in male paratypes of Trimeresurus ciliaris sp. nov.: A ZMMU Re- 17661; B ZMMU Re- 17662; C AUP- 02011; and D ZMMU Re- 17663. See Table 4 for details. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure4" httpUri="https://binary.pensoft.net/fig/908086" pageId="0" pageNumber="697">4B</figureCitation>
), while the two males AUP-02011 (Fig.
<figureCitation id="9130C000774A88BADCF9B36FF080C1AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in head scalation and coloration in male paratypes of Trimeresurus ciliaris sp. nov.: A ZMMU Re- 17661; B ZMMU Re- 17662; C AUP- 02011; and D ZMMU Re- 17663. See Table 4 for details. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure4" httpUri="https://binary.pensoft.net/fig/908086" pageId="0" pageNumber="697">4C</figureCitation>
) and ZMMU Re-17663 (Fig.
<figureCitation id="EFA301370AD19C8D49AE28A894A31AA0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in head scalation and coloration in male paratypes of Trimeresurus ciliaris sp. nov.: A ZMMU Re- 17661; B ZMMU Re- 17662; C AUP- 02011; and D ZMMU Re- 17663. See Table 4 for details. Photographs by P. Pawangkhanant." figureDoi="10.3897/vz.73.e109854.figure4" httpUri="https://binary.pensoft.net/fig/908086" pageId="0" pageNumber="697">4D</figureCitation>
) have four small supraoculars on each side of the head instead of three as in the holotype. Coloration of paratypes is very similar to that of the holotype; all specimens have a creamy-white belly and lack ventrolateral stripe. Female specimen QSMI 1538 has slightly lower number of subcaudals than in males (SC 52 vs. 59-63). The white vertebral spots are present both in males and in females; they are regularly spaced by two or three vertebral scales. The number of dark bands on dorsum varies from 70 to 81 among the members of the type series, without apparent sexual dimorphism. Coloration of female specimen QSMI 1538 largely faded due to preservation in formalin for seven years.
</paragraph>
<caption id="EC16500099EBF6285BC87ADFC92A0D3F" doi="10.3897/vz.73.e109854.figure4" httpUri="https://binary.pensoft.net/fig/908086" pageId="0" pageNumber="697" start="Figure 4" startId="F4">
<paragraph id="2480097FB462C0E6458E53303190105D" pageId="0" pageNumber="697">
<emphasis id="8DEED0A7843B9062BAB929FAA9DC29F1" bold="true" pageId="0" pageNumber="697">Figure 4.</emphasis>
Variation in head scalation and coloration in male paratypes of
<taxonomicName id="5B89D036729954027379A0BE69897B74" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="1525312D8293C038502E697A5F818BE4" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="44C74B6219BA043D09D6780B58DF3B94" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
:
<emphasis id="1A477C773E2847731EA97D83F0203865" bold="true" pageId="0" pageNumber="697">A</emphasis>
ZMMU Re-17661;
<emphasis id="20FCE0BB6976473F33277B9BC8CA8ACC" bold="true" pageId="0" pageNumber="697">B</emphasis>
ZMMU Re-17662;
<emphasis id="4E2D16E5108BE08D6F2E86054E207092" bold="true" pageId="0" pageNumber="697">C</emphasis>
AUP-02011; and
<emphasis id="012CC10B8B37E78E14287D93E88B57CF" bold="true" pageId="0" pageNumber="697">D</emphasis>
ZMMU Re-17663. See Table
<tableCitation id="02D8870387E909A472EE8A78893A8262" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological data on the type series of Trimeresurus ciliaris sp. nov. Notes: For abbreviations see Materials and methods section." httpUri="http://table.plazi.org/id/3DFF69BB9E3C24255F0BF12110F8CA24" pageId="0" pageNumber="697" tableUuid="3DFF69BB9E3C24255F0BF12110F8CA24">4</tableCitation>
for details. Photographs by P. Pawangkhanant.
</paragraph>
</caption>
<caption id="5C2FA74F9BE65CF1B56E8B4EB9043824" doi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697" start="Figure 5" startId="F5">
<paragraph id="002B8D33F4B0D71A037DD769707F68D6" pageId="0" pageNumber="697">
<emphasis id="B4DF3CD76418AF64C3525163B7090209" bold="true" pageId="0" pageNumber="697">Figure 5.</emphasis>
Comparison of body coloration between males of
<emphasis id="9DC30B3FBC1185EAAE6919E12817BBD9" bold="true" pageId="0" pageNumber="697">A</emphasis>
<taxonomicName id="19132BB0F699889473A5A94D84E13AE1" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="E244B0183D1191CA8B531FA184F9F40A" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="887EBB225376F2529593430D0237DB12" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
;
<emphasis id="685A02E8F2DE1FF5403EE0BEF0445F3C" bold="true" pageId="0" pageNumber="697">B</emphasis>
<taxonomicName id="ADEF29FE76BE39EAA8176ADEA818C536" lsidName="T. kanburiensis" pageId="0" pageNumber="697" rank="species" species="kanburiensis">
<emphasis id="D9B689F6ECDBFEE4B7CCA56FCDC1D79F" italics="true" pageId="0" pageNumber="697">T. kanburiensis</emphasis>
</taxonomicName>
;
<emphasis id="BDA69512E60678D2549EFF061FB5730F" bold="true" pageId="0" pageNumber="697">C</emphasis>
<taxonomicName id="4BDB02D45F27874F05E524D7B030EB12" lsidName="T. kuiburi" pageId="0" pageNumber="697" rank="species" species="kuiburi">
<emphasis id="B391BDBEFE522B68991C9FE370F1D8D8" italics="true" pageId="0" pageNumber="697">T. kuiburi</emphasis>
</taxonomicName>
; and
<emphasis id="A9621729279E29A8314E2C00636990D2" bold="true" pageId="0" pageNumber="697">D</emphasis>
<taxonomicName id="17CF9B2071A39193B1B50E3F0CC9B56A" lsidName="T. venustus" pageId="0" pageNumber="697" rank="species" species="venustus">
<emphasis id="9BEB69C448A0CF1D456BA5DC4611C580" italics="true" pageId="0" pageNumber="697">T. venustus</emphasis>
</taxonomicName>
s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C).
</paragraph>
</caption>
<caption id="F126F6C88F06C1517274568E1BAD12DD" doi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697" start="Figure 6" startId="F6">
<paragraph id="9B8EE56E7A2CA51A851A5BAB87F51694" pageId="0" pageNumber="697">
<emphasis id="61E707D87918EACF26870A8219C91ED5" bold="true" pageId="0" pageNumber="697">Figure 6.</emphasis>
Comparison of head coloration (left profile and dorsal view of the head) between males of
<emphasis id="1F9573C24C311632873C6DFFB9364324" bold="true" pageId="0" pageNumber="697">A-B</emphasis>
<taxonomicName id="6039FD072DCDBB5D3535A185A3C71CE3" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="4D38B66F8C36E68933FA96ED49E208C5" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="930F3A4D7988D57D413AA1C9120059D9" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
;
<emphasis id="A1D952FFDF159A73B3F4F9DD3B34C7E2" bold="true" pageId="0" pageNumber="697">C</emphasis>
-
<emphasis id="ADAB513DF7FE240FE9F7839F248A569D" bold="true" pageId="0" pageNumber="697">D</emphasis>
<taxonomicName id="AF3F06F621A1752C171CD17F2693EA55" lsidName="T. kanburiensis" pageId="0" pageNumber="697" rank="species" species="kanburiensis">
<emphasis id="E21E1B9CB94BEDADE3E2B305FBCFFEFA" italics="true" pageId="0" pageNumber="697">T. kanburiensis</emphasis>
</taxonomicName>
;
<emphasis id="9E858AAED4DAA81417185141F597DDD0" bold="true" pageId="0" pageNumber="697">E</emphasis>
-
<emphasis id="B13BEF440016774152E3110C20F8EDEA" bold="true" pageId="0" pageNumber="697">F</emphasis>
<taxonomicName id="11878811C2A0CFD7B547301F91398654" lsidName="T. kuiburi" pageId="0" pageNumber="697" rank="species" species="kuiburi">
<emphasis id="B912BED9BD22F98E7039FD9BB5AA44F0" italics="true" pageId="0" pageNumber="697">T. kuiburi</emphasis>
</taxonomicName>
; and
<emphasis id="63F9DC80FA76F7A9C30F4E1CD7C6D964" bold="true" pageId="0" pageNumber="697">G</emphasis>
-
<emphasis id="64A61EA6B6455B2C766392B4D74126EA" bold="true" pageId="0" pageNumber="697">H</emphasis>
<taxonomicName id="B19508DB19CB5870EA91E729E0F046D1" lsidName="T. venustus" pageId="0" pageNumber="697" rank="species" species="venustus">
<emphasis id="C1E305A11D690210701055D1D4B5BBA1" italics="true" pageId="0" pageNumber="697">T. venustus</emphasis>
</taxonomicName>
s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G).
</paragraph>
</caption>
<caption id="9EE9F703B0B249044D29BE1AE977257E" doi="10.3897/vz.73.e109854.figure7" httpUri="https://binary.pensoft.net/fig/908089" pageId="0" pageNumber="697" start="Figure 7" startId="F7">
<paragraph id="04ABB3DCA872E50AA900EF8C3BE85BC8" pageId="0" pageNumber="697">
<emphasis id="4A3E1B49F0ABAFAF87E42CD7AA206BF5" bold="true" pageId="0" pageNumber="697">Figure 7.</emphasis>
Habitat of
<taxonomicName id="538D7B4A266452EABED5620A25421659" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="1DB58CD88CAA86B64846F34A42BFED67" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="D83DD60B38390EDC420049C68829E87C" bold="true" pageId="0" pageNumber="697">sp. nov</emphasis>
.
<emphasis id="6DBA9C2C35FA06F204DAA0DCDF03826D" bold="true" pageId="0" pageNumber="697">A</emphasis>
microhabitat of the new species near the Thum Khao Ting Cave, Palian District, Trang Province, Thailand;
<emphasis id="A5B2ED98D1C403DD7690CCD29CFD7946" bold="true" pageId="0" pageNumber="697">B</emphasis>
holotype ZMMU Re-17661, adult male, in life in situ;
<emphasis id="18DFBAC9103A6C7BEBF55365209FC760" bold="true" pageId="0" pageNumber="697">C</emphasis>
paratype QSMI 1538, adult female, in life in situ;
<emphasis id="A3380758AA7695FD4EC15F1CBE665E24" bold="true" pageId="0" pageNumber="697">D</emphasis>
-
<emphasis id="6060D843C9D14E576BA37F9E81A9F644" bold="true" pageId="0" pageNumber="697">E</emphasis>
male specimen from Wang Kelian, Perlis State, Malaysia (not collected). Photographs by P. Pawangkhanant (A-C) and Azrul Azrizal Azmy (D-E).
</paragraph>
</caption>
<caption id="3DFF69BB9E3C24255F0BF12110F8CA24" ID-Table-UUID="3DFF69BB9E3C24255F0BF12110F8CA24" httpUri="http://table.plazi.org/id/3DFF69BB9E3C24255F0BF12110F8CA24" pageId="0" pageNumber="697" start="Table 4" startId="T4">
<paragraph id="4DAAAF1EA77D9AF9B4B1CB82FED6F3E4" pageId="0" pageNumber="697">
<emphasis id="4B3C0C05EDE6BD73F5F31656E1C385B6" bold="true" pageId="0" pageNumber="697">Table 4.</emphasis>
Morphological data on the type series of
<taxonomicName id="020496A6CA0455B79C0466E5E2885C36" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="17B43010097E3C56D8B4C15488A5AA76" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="3CF86917C7A94F419B2B80D2501A5E28" bold="true" pageId="0" pageNumber="697">sp. nov</emphasis>
. Notes: For abbreviations see Materials and methods section.
</paragraph>
</caption>
<paragraph id="B2F138AB3621B0E9D6A26EAC7D4EC77B" pageId="0" pageNumber="697">
<table id="CE3F841A6B8733FF1F192AF3DBE779EA" pageId="0" pageNumber="697">
<tr id="E18BDD210D2706957D6F7219F797FC3F" pageId="0" pageNumber="697">
<td id="1E85042AC7D72CB571D8ECFB63421F22" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="F01DE7298A811AEBC285D3A60D88743D" bold="true" pageId="0" pageNumber="697">No.</emphasis>
</td>
<td id="BCF7EE0ED57D82C07CA631144F57EC17" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="76EF127D3915EA0BF3D17CBC29CCC18B" bold="true" pageId="0" pageNumber="697">ZMMU Re-17661</emphasis>
</td>
<td id="0001E11C83BD63ED7816EB07CC5B1B3F" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="7FAB2970B2199D874FF678EAAFEF259F" bold="true" pageId="0" pageNumber="697">AUP-02011</emphasis>
</td>
<td id="08600615345D72FF8466F6933A2B5105" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="93D39D4535EF81A8E8402F42494B6402" bold="true" pageId="0" pageNumber="697">ZMMU Re-17662</emphasis>
</td>
<td id="AC2137FB955C345BC7D82EB8DE645DFE" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="0F23143423F79756D8978ED358F3BA99" bold="true" pageId="0" pageNumber="697">ZMMU Re-17663</emphasis>
</td>
<td id="387DA50795CDD93653C3AA475BFF828C" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="0620C99B43AFBED1A2EB8B86823ABAEC" bold="true" pageId="0" pageNumber="697">QSMI 1538</emphasis>
</td>
</tr>
<tr id="9A62E24782847813A791714C7BF53010" pageId="0" pageNumber="697">
<td id="35DE9148DC2B34247309CC5D0EBE4D7E" colspan="1" pageId="0" pageNumber="697" rowspan="1">Type status</td>
<td id="02B762E4E7958B7036B3AC65B402B09C" colspan="1" pageId="0" pageNumber="697" rowspan="1">Holotype</td>
<td id="B1FE8EF1D6E1565E4FF0AC1A41166817" colspan="1" pageId="0" pageNumber="697" rowspan="1">Paratype</td>
<td id="5B4D840413AACFCA96908EF818725E25" colspan="1" pageId="0" pageNumber="697" rowspan="1">Paratype</td>
<td id="45D9F2CB6AC7F2967F8FED49E10A4023" colspan="1" pageId="0" pageNumber="697" rowspan="1">Paratype</td>
<td id="218A9E555A2DD9DD395C7D4DA609523D" colspan="1" pageId="0" pageNumber="697" rowspan="1">Paratype</td>
</tr>
<tr id="BDE34C460CC38BA8A203C2AA8E640209" pageId="0" pageNumber="697">
<td id="7BC3CE3D7141FE25DAF02F8C10FEB754" colspan="1" pageId="0" pageNumber="697" rowspan="1">Sex</td>
<td id="8E4443ECA27757BF493A0D05A758568C" colspan="1" pageId="0" pageNumber="697" rowspan="1">Male</td>
<td id="9E4E63A9889862A90488D2A686082477" colspan="1" pageId="0" pageNumber="697" rowspan="1">Male</td>
<td id="8B9CCD69345AEC9F121AEF5DCF743F30" colspan="1" pageId="0" pageNumber="697" rowspan="1">Male</td>
<td id="D5442A86F61354084C8268130A7EAC96" colspan="1" pageId="0" pageNumber="697" rowspan="1">Male</td>
<td id="AB0822AE9C22D3B2840F1A8A643012A0" colspan="1" pageId="0" pageNumber="697" rowspan="1">Female</td>
</tr>
<tr id="C7FEFCDA953E36C93706E585688E8A4D" pageId="0" pageNumber="697">
<td id="FE53193FEC8E7EDB3C1671376C11A219" colspan="1" pageId="0" pageNumber="697" rowspan="1">SVL (mm)</td>
<td id="CB3C4F80369BADEEDCAFD574D4889133" colspan="1" pageId="0" pageNumber="697" rowspan="1">345</td>
<td id="6156A2A58F79570F2835E540CD56D0DB" colspan="1" pageId="0" pageNumber="697" rowspan="1">357</td>
<td id="5707AA8FBBC4CFD0BC197990B83CCDA9" colspan="1" pageId="0" pageNumber="697" rowspan="1">360</td>
<td id="8D9D88FF5746DDACD86DAF7AD944269C" colspan="1" pageId="0" pageNumber="697" rowspan="1">352</td>
<td id="F5493E59BD6082385B6EB0195168FC34" colspan="1" pageId="0" pageNumber="697" rowspan="1">333</td>
</tr>
<tr id="892CB8BC009C199A2A1C42DC79032600" pageId="0" pageNumber="697">
<td id="3E0F061DD7EB56C09EE11D1B1042C152" colspan="1" pageId="0" pageNumber="697" rowspan="1">TaL (mm)</td>
<td id="A585A877E17EC92ED2D49C0B890A7012" colspan="1" pageId="0" pageNumber="697" rowspan="1">72</td>
<td id="B68A294DFC50807CF923588BE311CAD9" colspan="1" pageId="0" pageNumber="697" rowspan="1">73</td>
<td id="83E0AFF7C2AD11CE8FA9CE4A3025FDC9" colspan="1" pageId="0" pageNumber="697" rowspan="1">72</td>
<td id="A7BE8C896162BBDCDAC4224CDBD943DB" colspan="1" pageId="0" pageNumber="697" rowspan="1">71</td>
<td id="D3925F512065A65CFDEBEED13F0DF886" colspan="1" pageId="0" pageNumber="697" rowspan="1">51</td>
</tr>
<tr id="41E5C32CF31641DD0590DD17DD1D4406" pageId="0" pageNumber="697">
<td id="BFAC27334F6C9D69625FBF86E593FA9E" colspan="1" pageId="0" pageNumber="697" rowspan="1">TL (mm)</td>
<td id="A18AD28B5CA71F79A8DAD1413AB82396" colspan="1" pageId="0" pageNumber="697" rowspan="1">417</td>
<td id="40D2E34E4373701FE3E2BA194A61C0E6" colspan="1" pageId="0" pageNumber="697" rowspan="1">430</td>
<td id="8B5377D50A798A5A6BBCF8707BC4DCB9" colspan="1" pageId="0" pageNumber="697" rowspan="1">432</td>
<td id="C83917E8F3D1DB02FE885C85D24A9641" colspan="1" pageId="0" pageNumber="697" rowspan="1">423</td>
<td id="F383E190C0E00BD3CA53532BAA96E388" colspan="1" pageId="0" pageNumber="697" rowspan="1">384</td>
</tr>
<tr id="F8E8F38F9B8799623429D7F6243A0236" pageId="0" pageNumber="697">
<td id="A62A0DC2A4B29B1E6ADE0951DFD61D78" colspan="1" pageId="0" pageNumber="697" rowspan="1">HL (mm)</td>
<td id="642365EE51A6349D3F64831236A78384" colspan="1" pageId="0" pageNumber="697" rowspan="1">16.5</td>
<td id="A5163DBAD2B0FFDA4FE6D676F59FE158" colspan="1" pageId="0" pageNumber="697" rowspan="1">16.9</td>
<td id="7E10DCD1B6FE60234E16B0A5F3AA5FC1" colspan="1" pageId="0" pageNumber="697" rowspan="1">17.1</td>
<td id="1065307FE2C715D40C47831F8FB928C8" colspan="1" pageId="0" pageNumber="697" rowspan="1">16.8</td>
<td id="3F8A783BB6239B38084ACBB4409ED0F9" colspan="1" pageId="0" pageNumber="697" rowspan="1">16</td>
</tr>
<tr id="AEDE14E3F419CB0B5E396BE8E5EC5C42" pageId="0" pageNumber="697">
<td id="2A2EC6B492CD11856891D9EE76591A4F" colspan="1" pageId="0" pageNumber="697" rowspan="1">HW (mm)</td>
<td id="6AF064361EE8F0FCD64A578AFD5C629A" colspan="1" pageId="0" pageNumber="697" rowspan="1">11.4</td>
<td id="A3975AA8C584F361AADAC58C96F94CA6" colspan="1" pageId="0" pageNumber="697" rowspan="1">12.2</td>
<td id="596D16A2694D3A3A6F491CC43E02DD0A" colspan="1" pageId="0" pageNumber="697" rowspan="1">12.7</td>
<td id="DCD4C9FA60A0BEE5D6BB0F86CE40C44F" colspan="1" pageId="0" pageNumber="697" rowspan="1">11.3</td>
<td id="1EF3E2299BC4FD383C29D92E9137B8C0" colspan="1" pageId="0" pageNumber="697" rowspan="1">11.3</td>
</tr>
<tr id="BB0AE8B1FBA532C5AA25CB3B098EA8B4" pageId="0" pageNumber="697">
<td id="C9117548928F956B90D7B9EA764B7EED" colspan="1" pageId="0" pageNumber="697" rowspan="1">HD (mm)</td>
<td id="76514FCB61EBE3E8E99D0471B461F711" colspan="1" pageId="0" pageNumber="697" rowspan="1">5</td>
<td id="7017DE3AE8E87B9F4AFF9739BA420C5F" colspan="1" pageId="0" pageNumber="697" rowspan="1">5.7</td>
<td id="E3C08138819DAEC65AD5F0C1E67AE125" colspan="1" pageId="0" pageNumber="697" rowspan="1">6.1</td>
<td id="36A38FE51B0C6E9F1FD0E3FCBF8D9994" colspan="1" pageId="0" pageNumber="697" rowspan="1">5.6</td>
<td id="9FB439F5580C7D8DEECD8D64908488F4" colspan="1" pageId="0" pageNumber="697" rowspan="1">5.7</td>
</tr>
<tr id="7BD8B5677C15451213A6B20B82F03C5F" pageId="0" pageNumber="697">
<td id="8D40C349D8571C07E0588A61745D6281" colspan="1" pageId="0" pageNumber="697" rowspan="1">SnL (mm)</td>
<td id="5551C5499CB041AA80B8CC7FCA664C03" colspan="1" pageId="0" pageNumber="697" rowspan="1">5</td>
<td id="5FDD542E09631243CC1DA427F5EDC498" colspan="1" pageId="0" pageNumber="697" rowspan="1">5.1</td>
<td id="A5C68554FF9C028B51A628D62B2A50DF" colspan="1" pageId="0" pageNumber="697" rowspan="1">5.3</td>
<td id="2444D3D6D82CBF05070ACC821191718E" colspan="1" pageId="0" pageNumber="697" rowspan="1">4.9</td>
<td id="6564F0083907650155132FFCD3C66891" colspan="1" pageId="0" pageNumber="697" rowspan="1">5.6</td>
</tr>
<tr id="FD838E585B682AF8B83EC3FA051A2113" pageId="0" pageNumber="697">
<td id="F0D7ACC3D5B7F670CF1CA041474547A7" colspan="1" pageId="0" pageNumber="697" rowspan="1">ED (mm)</td>
<td id="8B42AC80857D4DA37B610491786453FC" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.6</td>
<td id="740D0B79A710CB8FADD20923766941D2" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.8</td>
<td id="AE3F7FC281CBBD2990731EEF327BFB1D" colspan="1" pageId="0" pageNumber="697" rowspan="1">3</td>
<td id="C5AAB52150E4D6E505A3220DE4020198" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.5</td>
<td id="9FF8F3543A4D01C1DE12EAE3A857BA1B" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.7</td>
</tr>
<tr id="483081BA361F1786BAB87CECC152D890" pageId="0" pageNumber="697">
<td id="9295B36AC28905FB4B322C55F22DCF4A" colspan="1" pageId="0" pageNumber="697" rowspan="1">VED (mm)</td>
<td id="A5D924FE2140391E7C4255B32F84D11F" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.3</td>
<td id="4DF5DB92A4DA5CC835438D96F452E98C" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.1</td>
<td id="7C2BA2B045AE7E3276C77FBB1340EAB9" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.5</td>
<td id="7F003D3D4F09564407D51015C8B27535" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.1</td>
<td id="4C3ABB9576DD0DB26253B209274929A9" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.2</td>
</tr>
<tr id="AE228F671B3B1C433AAA5477B5DF28F2" pageId="0" pageNumber="697">
<td id="44A2C40DB240FCC8D1ACB37E3B316355" colspan="1" pageId="0" pageNumber="697" rowspan="1">SOL (mm)</td>
<td id="11FC185FB2364FA43B3F5B1540ADDBB6" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.8</td>
<td id="BA30954E591FADEBE76E27E0CB27F012" colspan="1" pageId="0" pageNumber="697" rowspan="1">3</td>
<td id="B34EA2B9C3F5DF6B06C39DDE2568ECA5" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.8</td>
<td id="64478A4CB7335E3BD624FED45DD3AB71" colspan="1" pageId="0" pageNumber="697" rowspan="1">2.7</td>
<td id="970DCE6453961295798BA49A459BC611" colspan="1" pageId="0" pageNumber="697" rowspan="1">3.4</td>
</tr>
<tr id="B1F2B93BB319FA19D051C5AC5546EC18" pageId="0" pageNumber="697">
<td id="E3B50EE01396FDBD98F17496D76D4465" colspan="1" pageId="0" pageNumber="697" rowspan="1">SOW (mm)</td>
<td id="CCA9DABAC59F920D4D6B5D2C20035C18" colspan="1" pageId="0" pageNumber="697" rowspan="1">0.7</td>
<td id="36CBE79A8F67DD7B5CB17CAC638E0A2B" colspan="1" pageId="0" pageNumber="697" rowspan="1">0.6</td>
<td id="1DD48FA094DF08E5C8CED421F717DACE" colspan="1" pageId="0" pageNumber="697" rowspan="1">0.7</td>
<td id="9F8F376E1F530E8BBCACDEB95EBF0A08" colspan="1" pageId="0" pageNumber="697" rowspan="1">0.7</td>
<td id="6BF7F671D02ED4215FA267E89646D142" colspan="1" pageId="0" pageNumber="697" rowspan="1">0.8</td>
</tr>
<tr id="F3EDA447E3B99CB8083237726BE6CFBD" pageId="0" pageNumber="697">
<td id="E495C17229E6E7A0F2B1EBD59B50A20E" colspan="1" pageId="0" pageNumber="697" rowspan="1">Internasals in contact</td>
<td id="749577EE449CA18CCBAB459993BC5131" colspan="1" pageId="0" pageNumber="697" rowspan="1">no</td>
<td id="A7CE3E618BF9258EB13A187E79441DC6" colspan="1" pageId="0" pageNumber="697" rowspan="1">no</td>
<td id="F5527EEDB286EBBEC9C4119DFF0B507F" colspan="1" pageId="0" pageNumber="697" rowspan="1">no</td>
<td id="AAA500D8E460EB9931AD83E72CE824E8" colspan="1" pageId="0" pageNumber="697" rowspan="1">no</td>
<td id="FE8DBAA6A5BCA86FCD0A769B00859800" colspan="1" pageId="0" pageNumber="697" rowspan="1">no</td>
</tr>
<tr id="0504204B265C966D7D62439A1F3EE04C" pageId="0" pageNumber="697">
<td id="9D0AF78818CD071F56F92DC17B289963" colspan="1" pageId="0" pageNumber="697" rowspan="1">Contact nasal-1st SL</td>
<td id="006F3AB05D8D146DB444799BE30D3919" colspan="1" pageId="0" pageNumber="697" rowspan="1">partially fused</td>
<td id="D4D4985EFB6E1170717CBD1CF23D38C2" colspan="1" pageId="0" pageNumber="697" rowspan="1">partially fused</td>
<td id="7C6FACC9EFD6E0B32D6B4D43BA8D793F" colspan="1" pageId="0" pageNumber="697" rowspan="1">partially fused</td>
<td id="EC6227D55AAD2FA327EAEABFCFE4CCC0" colspan="1" pageId="0" pageNumber="697" rowspan="1">partially fused</td>
<td id="7DA8B10F6B1EFF5739B11100121E3D77" colspan="1" pageId="0" pageNumber="697" rowspan="1">partially fused</td>
</tr>
<tr id="04E981183B7912E66C6D9556AF3D9CA6" pageId="0" pageNumber="697">
<td id="FCB1564A7BDFD3F8ED3595F95D1921C0" colspan="1" pageId="0" pageNumber="697" rowspan="1">SupOc</td>
<td id="B4EBA093647DFE566EE2131FB71B6589" colspan="1" pageId="0" pageNumber="697" rowspan="1">3</td>
<td id="49937EC3E98BEB8694D5EE9101ABD169" colspan="1" pageId="0" pageNumber="697" rowspan="1">3</td>
<td id="99F46DDC944DA7D6AA2F783E5254AF6E" colspan="1" pageId="0" pageNumber="697" rowspan="1">4</td>
<td id="9902591CB5C51254BC9A20D740D46398" colspan="1" pageId="0" pageNumber="697" rowspan="1">4</td>
<td id="168A0FC1E0E9299B2D07657AD29CFB7A" colspan="1" pageId="0" pageNumber="697" rowspan="1">3</td>
</tr>
<tr id="904D03D5BB6E4F6DEC1C89D447692D56" pageId="0" pageNumber="697">
<td id="B1549B753D96BD86E3274064CAFE1420" colspan="1" pageId="0" pageNumber="697" rowspan="1">PosOc</td>
<td id="E4F94C666A849E2A7DD4F2BAD5357227" colspan="1" pageId="0" pageNumber="697" rowspan="1">2/2</td>
<td id="CA792FE664073B3FE404F5D62EDE0850" colspan="1" pageId="0" pageNumber="697" rowspan="1">2/2</td>
<td id="6519BB0C7DDB7F13366A9902272C9C16" colspan="1" pageId="0" pageNumber="697" rowspan="1">2/2</td>
<td id="9C27CE339DDC92100BF1A7DD829D9AB8" colspan="1" pageId="0" pageNumber="697" rowspan="1">2/2</td>
<td id="FD565C2FC7A1B8FC80EEAA5F11AEE459" colspan="1" pageId="0" pageNumber="697" rowspan="1">2/2</td>
</tr>
<tr id="BB59804D2B76B78C64B3E809AABE2DF6" pageId="0" pageNumber="697">
<td id="91A9F86F71BEE36A7508F5A377E49566" colspan="1" pageId="0" pageNumber="697" rowspan="1">IOS</td>
<td id="8DD09EB0451CC3F1228E5F1D26B576DD" colspan="1" pageId="0" pageNumber="697" rowspan="1">11</td>
<td id="BDCDAC5C4F02933C0C25D1828D2C4F9A" colspan="1" pageId="0" pageNumber="697" rowspan="1">14</td>
<td id="6A39464BF37735573AC70320C4367DB8" colspan="1" pageId="0" pageNumber="697" rowspan="1">11</td>
<td id="D26410C61A2F429B2A877A358D99F95A" colspan="1" pageId="0" pageNumber="697" rowspan="1">11</td>
<td id="578B4F8C3C60929BD8903CDD1827B790" colspan="1" pageId="0" pageNumber="697" rowspan="1">12</td>
</tr>
<tr id="A3CE64B8BFCC4D2261596709DA8B43C1" pageId="0" pageNumber="697">
<td id="A09E57826104E71635779DB94A1A4077" colspan="1" pageId="0" pageNumber="697" rowspan="1">SL</td>
<td id="FFB76A1B0FD4995D8EAF58CAA85CECCC" colspan="1" pageId="0" pageNumber="697" rowspan="1">9/9</td>
<td id="B0EDAC362A59D00C9A3D06FCFF667FDD" colspan="1" pageId="0" pageNumber="697" rowspan="1">9/8</td>
<td id="DE54E5F07439C878C077221CAA162809" colspan="1" pageId="0" pageNumber="697" rowspan="1">9/9</td>
<td id="EF912D1D80ED14F08D27636D72CDBFDA" colspan="1" pageId="0" pageNumber="697" rowspan="1">9/9</td>
<td id="A45AF53F89340544FF676AAC684CD6D2" colspan="1" pageId="0" pageNumber="697" rowspan="1">9/9</td>
</tr>
<tr id="8D7270ACCCFD035B72C1546105B78090" pageId="0" pageNumber="697">
<td id="0AA7A798B473CB71E6DBEE4F2FFB7900" colspan="1" pageId="0" pageNumber="697" rowspan="1">IL</td>
<td id="7BA6C018DBF107D12613732AD802907F" colspan="1" pageId="0" pageNumber="697" rowspan="1">11/10</td>
<td id="809C904A181240152EA2F8FEC687FF2F" colspan="1" pageId="0" pageNumber="697" rowspan="1">10/10</td>
<td id="FF08BA37481C3C494D5107DF336832E7" colspan="1" pageId="0" pageNumber="697" rowspan="1">10/10</td>
<td id="1E2E3F2D7E70F4778BC620BE4FC8E779" colspan="1" pageId="0" pageNumber="697" rowspan="1">11/10</td>
<td id="9606AE819FD9E5786FF5569CD771DD2E" colspan="1" pageId="0" pageNumber="697" rowspan="1">10/11</td>
</tr>
<tr id="7AACD21D1BCD2C178D447E61132B4DC2" pageId="0" pageNumber="697">
<td id="25990138C8FC17DA1359E41273085442" colspan="1" pageId="0" pageNumber="697" rowspan="1">ASR</td>
<td id="FD34101442CAFF0159BDE6350CE7D87D" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="A27C29B75D90FD8D62B7528BD892330A" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="7BE3BEE568B1F1637311E191784AAAFC" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="AB921BDFFEF375E027A8B847B9A722D3" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="3EB324A1169AD963E4F96A4AC014C640" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
</tr>
<tr id="CA7D2DAEDC3D07B218139040F7584EC5" pageId="0" pageNumber="697">
<td id="BB10E59CA45A2C3B8392A90A81EA8150" colspan="1" pageId="0" pageNumber="697" rowspan="1">MSR</td>
<td id="4F6FC9F4E9025F8EBE93A658D40ADB58" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="9D1FB016DA500B9D1ED2378891DA0212" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="FD2F81328161D29382479FE393E96AD1" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="F3BE668CD704AC56D003A7AB2AF8DFA6" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="FA474997786F9F109326019558E7F7C2" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
</tr>
<tr id="9430B9A5D592EF35C623508ABA0A574C" pageId="0" pageNumber="697">
<td id="BC78FF27CDECE3FABFB8D3DF475A8E90" colspan="1" pageId="0" pageNumber="697" rowspan="1">PSR</td>
<td id="0A8362CEB1F14538363EA256EF49BCC9" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
<td id="E3527501885B8814794B469F4021AEF2" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
<td id="ABAA7071480E380054BB822104380F4C" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
<td id="3707A8CD705A89E38B9CF545BD695E89" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
<td id="9E4754D20AB72F7F7C9F6E92628FDBE0" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
</tr>
<tr id="4075E948AA6171634CF2AC4E9EFF3563" pageId="0" pageNumber="697">
<td id="EF2BA8946F5C4DF9E9C16E741935D65C" colspan="1" pageId="0" pageNumber="697" rowspan="1">SRR 17 to 15 (V)</td>
<td id="2F5B00FF6BCADC874BABD12EF336ED99" colspan="1" pageId="0" pageNumber="697" rowspan="1">120-121</td>
<td id="068CD1DC0BA47B38AAF40A2FCC56D1DA" colspan="1" pageId="0" pageNumber="697" rowspan="1">123-124</td>
<td id="03CDB61F482F5F70BE41CD9409673EDF" colspan="1" pageId="0" pageNumber="697" rowspan="1">123-124</td>
<td id="12EF011D81D4F812B123D1230D8601AE" colspan="1" pageId="0" pageNumber="697" rowspan="1">125-126</td>
<td id="2FD03078D41E31763AC11F1B5AFE0F24" colspan="1" pageId="0" pageNumber="697" rowspan="1">123</td>
</tr>
<tr id="3EBE34859910132E20632BE648FDABE5" pageId="0" pageNumber="697">
<td id="263716999A1E5FEA7AB6EC5A181B4130" colspan="1" pageId="0" pageNumber="697" rowspan="1">PreVEN</td>
<td id="C59F33121E67D80847FB7C68B4E1B646" colspan="1" pageId="0" pageNumber="697" rowspan="1">2</td>
<td id="8BA873159F0D7FA4308B405864D1AA49" colspan="1" pageId="0" pageNumber="697" rowspan="1">2</td>
<td id="3E76B0436BBCD141E658CCFAEFADB6D9" colspan="1" pageId="0" pageNumber="697" rowspan="1">2</td>
<td id="FEDCB2ECF2CC164F33594D6793EDF7D6" colspan="1" pageId="0" pageNumber="697" rowspan="1">2</td>
<td id="AF1C0977B17C37309DFF8F3A3AA470F0" colspan="1" pageId="0" pageNumber="697" rowspan="1">2</td>
</tr>
<tr id="AD134E140A43347E8633585FEC548E10" pageId="0" pageNumber="697">
<td id="3ECB4DE6602C23B3C06B2BDE7D5AE7B2" colspan="1" pageId="0" pageNumber="697" rowspan="1">VEN</td>
<td id="27B9EB2FDFB8F97EFFDF2D40C5FEA8B1" colspan="1" pageId="0" pageNumber="697" rowspan="1">172</td>
<td id="9C4282484EC9A44DD3E6646E7F546F31" colspan="1" pageId="0" pageNumber="697" rowspan="1">173</td>
<td id="303F0F59167AF9F898063E380B7446A3" colspan="1" pageId="0" pageNumber="697" rowspan="1">175</td>
<td id="6C96511A7AFEC37E794DC7D504B764FD" colspan="1" pageId="0" pageNumber="697" rowspan="1">172</td>
<td id="6F2BAB239EDDC2479ED9CE0A6CF6A9BB" colspan="1" pageId="0" pageNumber="697" rowspan="1">171</td>
</tr>
<tr id="57BA9884802BE26396D2BADCBB10793C" pageId="0" pageNumber="697">
<td id="4B5A15AE9791EDA36423C6DED2227377" colspan="1" pageId="0" pageNumber="697" rowspan="1">CP</td>
<td id="87BE86EC2245FBA1ECAED590232FB14A" colspan="1" pageId="0" pageNumber="697" rowspan="1">1</td>
<td id="567646ED69478B0012DC1E6DD73EA7B8" colspan="1" pageId="0" pageNumber="697" rowspan="1">1</td>
<td id="AFBE37268982A88578C0967CE5128CDC" colspan="1" pageId="0" pageNumber="697" rowspan="1">1</td>
<td id="03A2620759AAC8BFC621E89C630D1E87" colspan="1" pageId="0" pageNumber="697" rowspan="1">1</td>
<td id="EF85FD4A0BE401774E2CACD73AE8F6A2" colspan="1" pageId="0" pageNumber="697" rowspan="1">1</td>
</tr>
<tr id="9D54E0DF12B120161CAB7973A23C2DE2" pageId="0" pageNumber="697">
<td id="28E7D52FA18CB2A8C5E95A8C6025E971" colspan="1" pageId="0" pageNumber="697" rowspan="1">SC</td>
<td id="AF6CF1C2DB407948CFB83776F55F947F" colspan="1" pageId="0" pageNumber="697" rowspan="1">61</td>
<td id="9BCDA38660EADE072D3CAF2831B0F2F3" colspan="1" pageId="0" pageNumber="697" rowspan="1">59</td>
<td id="5A78D8357E1F019B1262FF6D9E89EEA4" colspan="1" pageId="0" pageNumber="697" rowspan="1">63</td>
<td id="DDF53AA29FBDA37857349CB06C1EFCD8" colspan="1" pageId="0" pageNumber="697" rowspan="1">61</td>
<td id="453061EA1C45AC0FBE703A843B713DF3" colspan="1" pageId="0" pageNumber="697" rowspan="1">52</td>
</tr>
<tr id="836753D695E66855B60F275635030589" pageId="0" pageNumber="697">
<td id="2120C0D9677795E0939D07D6592F4D46" colspan="1" pageId="0" pageNumber="697" rowspan="1">Postocular stripe</td>
<td id="0C9AEE889F173729E19B3114DC5BB163" colspan="1" pageId="0" pageNumber="697" rowspan="1">reddish-brown</td>
<td id="F7905B7A023AC7C70E481A59AE8AF98C" colspan="1" pageId="0" pageNumber="697" rowspan="1">reddish-brown</td>
<td id="EF57C5D97847D0B7AF83A22E02FEDD78" colspan="1" pageId="0" pageNumber="697" rowspan="1">reddish-brown</td>
<td id="B3986426021B0D009AA62C31C9E0A8A2" colspan="1" pageId="0" pageNumber="697" rowspan="1">reddish-brown</td>
<td id="CF42CDF6160A47444297EF9193369CCA" colspan="1" pageId="0" pageNumber="697" rowspan="1">Faint, reddish-brown</td>
</tr>
<tr id="9E67489CD148D917295F28623F98773E" pageId="0" pageNumber="697">
<td id="3B24EA15F0B33D23B607401C385A4916" colspan="1" pageId="0" pageNumber="697" rowspan="1">Number of red bands on dorsum</td>
<td id="C2C84AA8EC8175756A9E6BEEA632E6D2" colspan="1" pageId="0" pageNumber="697" rowspan="1">74</td>
<td id="83DF4C08E34A1F82498BB1CB4287901A" colspan="1" pageId="0" pageNumber="697" rowspan="1">81</td>
<td id="6CCED2EF3A5AEFB1349FBF4BECE7147A" colspan="1" pageId="0" pageNumber="697" rowspan="1">73</td>
<td id="821F2E7EFA60F683338A7FC62368A667" colspan="1" pageId="0" pageNumber="697" rowspan="1">70</td>
<td id="3ABE4EDFF8D281A9DAA4E94AD9DAC1E9" colspan="1" pageId="0" pageNumber="697" rowspan="1">80</td>
</tr>
<tr id="5FB4DAE7CFA8A3947E2B0A54F11708AC" pageId="0" pageNumber="697">
<td id="7428AAC57E3C6C5D7969FC5CBCE0B95E" colspan="1" pageId="0" pageNumber="697" rowspan="1">White vertebral dots</td>
<td id="B9AD2BF54F71A709946201B29C0A7492" colspan="1" pageId="0" pageNumber="697" rowspan="1">every 2-3 scales</td>
<td id="9C35B9014A7F7E8A7E03BF35A3E1559C" colspan="1" pageId="0" pageNumber="697" rowspan="1">every 2-3 scales</td>
<td id="B01CFDCB04EEC5FF63E934CFC51D0212" colspan="1" pageId="0" pageNumber="697" rowspan="1">every 2-3 scales</td>
<td id="742B176725ACC011EC3BD1E09DBF36E7" colspan="1" pageId="0" pageNumber="697" rowspan="1">every 2-3 scales</td>
<td id="B8923CC84A50297E3936C4EA60AA8653" colspan="1" pageId="0" pageNumber="697" rowspan="1">every 2-3 scales</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="E32263BB48DFFB1A2958702ED8E36DFD" pageId="0" pageNumber="697" type="distribution">
<paragraph id="485483FCF9BACFE5BA5CBF1493B9A90D" pageId="0" pageNumber="697">Distribution and natural history.</paragraph>
<paragraph id="77EDD059CE7E1BFF505EF287AA4EA624" pageId="0" pageNumber="697">
Currently,
<taxonomicName id="7A3D69F7E6C6A0FF54C26E29CA481726" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="AB3A7C397C046993A5E95C9A783C556B" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="7504AEFD778FD5386D849EB3192EF715" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
is known only from a narrow limestone area in the Nakawan Range spanning the border of Thailand and Malaysia, in particular in limestone forests in Trang (Palian District) and Satun (Tha Le Ban National Park) provinces, Thailand (Fig.
<figureCitation id="BB8BDD96BF1FBE1CDB033B71527DEDF0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Trimeresurus kanburiensis species complex in Thai-Malay Peninsula: Localities. T. kanburiensis (yellow): 1 Kanchanaburi Prov., Sai Yok Dist., Wat Tham, Phom Lo Khao Yai; T. kuiburi (blue): 2 Prachuap Khiri Khan Prov., Kuiburi Dist., Wat Khao Daeng; 3 Prachuap Khiri Khan Prov., Kuiburi Dist., Khao Daeng Beach; 4 Prachuap Khiri Khan Prov., Kuiburi Dist., Khao Daeng, near Ban Thung Noi; T. cf. venustus (1) (green): 5 Chumphon Prov., Wat Tham Sanook; T. venustus s. str. (pink): 6 Krabi Prov., Mueang Krabi Dist., Tiger Cave viewpoint; 7 Thailand, Nakhon Si Thammarat Prov., Khao Luang; 8 Nakhon Si Thammarat Prov., Thung Song; 9 Surat Thani Prov.; Trimeresurus ciliaris sp. nov. (red): 10 Trang Prov., Palian Dist., Thum Khao Ting; 11 Tha Le Ban NP., Khuan Don Dist., Satun Prov.; 12 Perlis State National Park, Perlis State; T. cf. venustus (2) (purple): 13 Pulau Langkawi Island, Kedah State. Remark: stars denote type localities (the type locality is not shown for T. kanburiensis as it was not sampled). Abbreviations: MY - Myanmar, MA - Malaysia, Prov. - Province, Dist. - District." figureDoi="10.3897/vz.73.e109854.figure1" httpUri="https://binary.pensoft.net/fig/908083" pageId="0" pageNumber="697">1</figureCitation>
). It is highly likely that the new species also inhabits the northernmost part of Perlis State of Peninsular Malaysia (Fig.
<figureCitation id="0862F4C4368FD5C15201CB1A9F94014E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Trimeresurus kanburiensis species complex in Thai-Malay Peninsula: Localities. T. kanburiensis (yellow): 1 Kanchanaburi Prov., Sai Yok Dist., Wat Tham, Phom Lo Khao Yai; T. kuiburi (blue): 2 Prachuap Khiri Khan Prov., Kuiburi Dist., Wat Khao Daeng; 3 Prachuap Khiri Khan Prov., Kuiburi Dist., Khao Daeng Beach; 4 Prachuap Khiri Khan Prov., Kuiburi Dist., Khao Daeng, near Ban Thung Noi; T. cf. venustus (1) (green): 5 Chumphon Prov., Wat Tham Sanook; T. venustus s. str. (pink): 6 Krabi Prov., Mueang Krabi Dist., Tiger Cave viewpoint; 7 Thailand, Nakhon Si Thammarat Prov., Khao Luang; 8 Nakhon Si Thammarat Prov., Thung Song; 9 Surat Thani Prov.; Trimeresurus ciliaris sp. nov. (red): 10 Trang Prov., Palian Dist., Thum Khao Ting; 11 Tha Le Ban NP., Khuan Don Dist., Satun Prov.; 12 Perlis State National Park, Perlis State; T. cf. venustus (2) (purple): 13 Pulau Langkawi Island, Kedah State. Remark: stars denote type localities (the type locality is not shown for T. kanburiensis as it was not sampled). Abbreviations: MY - Myanmar, MA - Malaysia, Prov. - Province, Dist. - District." figureDoi="10.3897/vz.73.e109854.figure1" httpUri="https://binary.pensoft.net/fig/908083" pageId="0" pageNumber="697">1</figureCitation>
, see Discussion for details). The new species occurs in lowland dipterocarp forest on limestone rocks and appears to be strongly associated with a karst landscape. The new species was recorded for the first time in limestone formations within the Tha Le Ban National Park, Satun Province of Thailand, in June 2016 (P. Pawangkhanant pers. obser.), where it was observed in a lowland dipterocarp forest with numerous large karst rocks. Almost all specimens of
<taxonomicName id="7E8B94BD5246DEEA5B204B8F956A3173" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="79164CDD667E29F524A3152DEFCA687D" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="C0D651E3568E5E37FC3D62EA2D9DBCD2" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
were collected during daytime (except ZMMU Re-17663 that was collected at night around 20h50). All specimens were found near large boulders of karst rocks covered with dense vegetation and leaf litter (Fig.
<figureCitation id="70F0229B592B110657CF2E90CB4A3206" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitat of Trimeresurus ciliaris sp. nov. A microhabitat of the new species near the Thum Khao Ting Cave, Palian District, Trang Province, Thailand; B holotype ZMMU Re- 17661, adult male, in life in situ; C paratype QSMI 1538, adult female, in life in situ; D - E male specimen from Wang Kelian, Perlis State, Malaysia (not collected). Photographs by P. Pawangkhanant (A-C) and Azrul Azrizal Azmy (D-E)." figureDoi="10.3897/vz.73.e109854.figure7" httpUri="https://binary.pensoft.net/fig/908089" pageId="0" pageNumber="697">7A</figureCitation>
); one specimen was found after heavy rain around 18h00 while perching vertically on a limestone rock surface ca. 5 m above the ground.
</paragraph>
<paragraph id="2F8FAF0851EEDB025669930E8A7F969C" pageId="0" pageNumber="697">
In captivity, due to its small size, this species of pitviper mostly feeds on small geckos and microhylid frogs [
<taxonomicName id="761DD7A476FA964C266D5909ED62CAB3" class="Reptilia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microhyla butleri" order="Anura" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="butleri">
<emphasis id="0E93721408E0DB06C79DAB172EDB3DE7" italics="true" pageId="0" pageNumber="697">Microhyla butleri</emphasis>
</taxonomicName>
Boulenger,
<taxonomicName id="E7E4C4CA42E10CD9E7CBEE0C7A4A4D67" lsidName="M. heymonsi" pageId="0" pageNumber="697" rank="species" species="heymonsi">M. cf. heymonsi</taxonomicName>
Vogt, and
<taxonomicName id="2256AA477F0C8C21829DAF6670BF02A3" authorityName="Taylor" authorityYear="1962" class="Amphibia" family="Microhylidae" genus="Micryletta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Micryletta lineata" order="Anura" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="lineata">Micryletta cf. lineata</taxonomicName>
(Taylor)]. Nothing is known about the diet of the new species in the wild, but in one specimen from Mueang District, Satun Province, caudal luring behavior was observed: the male specimen was wiggling its tail tip when an adult
<taxonomicName id="0079EE7D7D986F620C50F016050A4910" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gekko gecko" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="gecko">Gekko gecko</taxonomicName>
<taxonomicName id="0D1045801E5EC64ECC9B504E08B078E0" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cnemaspis biocellata" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="biocellata">
<emphasis id="B04D2B4AC9B5EAF3E18313A2EA3FFE14" italics="true" pageId="0" pageNumber="697">Cnemaspis biocellata</emphasis>
</taxonomicName>
Grismer, Chan, Nasir &amp; Sumontha approached its shelter. Other amphibian and reptile species recorded in syntopy with the new species include:
<taxonomicName id="9DD97785E9B6BA9E8414B9A7286972F5" authorityName="Boulenger" authorityYear="1887" baseAuthorityName="Boulenger" baseAuthorityYear="1887" class="Amphibia" family="Bufonidae" genus="Ingerophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ingerophrynus parvus" order="Anura" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="parvus">
<emphasis id="DDF8C559313EF970D6DE3FD5395739C9" italics="true" pageId="0" pageNumber="697">Ingerophrynus parvus</emphasis>
</taxonomicName>
(Boulenger),
<taxonomicName id="2F7F5C44482DAC8C5A903DB6A49481D7" authorityName="Taylor" authorityYear="1962" class="Amphibia" family="Microhylidae" genus="Micryletta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Micryletta lineata" order="Anura" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="lineata">Micryletta cf. lineata</taxonomicName>
,
<taxonomicName id="65B3992353FAA03B82C848DE84C8438D" class="Reptilia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microhyla butleri" order="Anura" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="butleri">
<emphasis id="61DD1DE66259DBCECAA9075156C59E02" italics="true" pageId="0" pageNumber="697">Microhyla butleri</emphasis>
</taxonomicName>
,
<taxonomicName id="4E5047E683ADFC674E4ECA140921D0C1" authorityName="Chan, Grismer &amp; Brown" authorityYear="2014" class="Amphibia" family="Microhylidae" genus="Kaloula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kaloula latidisca" order="Anura" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="latidisca">
<emphasis id="71BE41A75CB569A85A87CEDC710184C0" italics="true" pageId="0" pageNumber="697">Kaloula latidisca</emphasis>
</taxonomicName>
(Chan, Grismer &amp; Brown),
<taxonomicName id="DECE4701C42F4456CF6AEE6114E77481" authorityName="Grismer, Sumontha, Cota, Grismer, Wood, Pauwels &amp; Kunya" authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cnemaspis niyomwanae" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="niyomwanae">
<emphasis id="2B0A2240E0EA680FD0FCB29AADE4F6E9" italics="true" pageId="0" pageNumber="697">Cnemaspis niyomwanae</emphasis>
</taxonomicName>
Grismer, Sumontha, Cota, Grismer, Wood, Pauwels &amp; Kunya,
<taxonomicName id="B4B572756C0C231DEB4DD151D6F1A148" lsidName="C. biocellata" pageId="0" pageNumber="697" rank="species" species="biocellata">
<emphasis id="456C60827DF0EE04A37B09BA29C6DC6B" italics="true" pageId="0" pageNumber="697">C. biocellata</emphasis>
</taxonomicName>
,
<taxonomicName id="7A499F6FC866C7BD218381A3ED23EEEF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gekko gecko" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="gecko">
<emphasis id="8DE08F0F15AC1E2DEE44CA3CCFF8E1F8" italics="true" pageId="0" pageNumber="697">Gekko gecko</emphasis>
</taxonomicName>
(Linnaeus),
<taxonomicName id="244E674E58D63CFC9C5F3A6643EADD98" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus astrum" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="astrum">
<emphasis id="9D3AE7241F84DE64C98687D4ECFAD7BD" italics="true" pageId="0" pageNumber="697">Cyrtodactylus astrum</emphasis>
</taxonomicName>
Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels,
<taxonomicName id="DCD99BA3003C122FBE65D5203AB6F9A9" lsidName="C. quadrivirgatus" pageId="0" pageNumber="697" rank="species" species="quadrivirgatus">
<emphasis id="9BB186563FFDC0497B928BC865B95112" italics="true" pageId="0" pageNumber="697">C. quadrivirgatus</emphasis>
</taxonomicName>
Taylor,
<taxonomicName id="1B14E332AAA4D513ACB5F4EB8D6FE0B6" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyrtodactylus" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="genus">
<emphasis id="9169B559B5549AD4080085D413C70529" italics="true" pageId="0" pageNumber="697">Cyrtodactylus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="C3FEB7BB191A54EA9BDA560EC69670C7" baseAuthorityName="Butler" baseAuthorityYear="1899" class="Reptilia" family="Colubridae" genus="Elaphe" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Elaphe taeniura subsp. ridleyi" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="subSpecies" species="taeniura" subSpecies="ridleyi">
<emphasis id="A276A8CD68C152F184FB72390BFEE596" italics="true" pageId="0" pageNumber="697">Elaphe taeniura ridleyi</emphasis>
</taxonomicName>
(Cope), and
<taxonomicName id="50773E27FC78E4C0792D51E5E09C20B9" class="Reptilia" family="Viperidae" genus="Tropidolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tropidolaemus wagleri" order="Squamata" pageId="0" pageNumber="697" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="A10AA4ACF6CA60EA56C76FC569AF38EE" italics="true" pageId="0" pageNumber="697">Tropidolaemus wagleri</emphasis>
</taxonomicName>
(Boie).
</paragraph>
</subSubSection>
<subSubSection id="B5B6C8BEB7656943F1345563CFF2CE95" pageId="0" pageNumber="697" type="etymology">
<paragraph id="03848C30E462FB5B2251676E229A8552" pageId="0" pageNumber="697">Etymology.</paragraph>
<paragraph id="C08D7700C326533DE5B21FBD4074C30C" pageId="0" pageNumber="697">
The species name &quot;
<taxonomicName id="333C10A7620DAE03BF7EAD53A3C8D356" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="765C101C49215728C1EE252302578B8D" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
&quot; is a Latin adjective in the nominative singular, masculine gender, derived from Latin word &quot;
<emphasis id="B32776CA630C54E99E3F369CA08A5129" italics="true" pageId="0" pageNumber="697">cilium</emphasis>
&quot; meaning &quot;an eyelash&quot;, and is given in reference to the characteristic small and distinct supraocular scales in the new species, which resemble eyelashes or eyebrows in lateral view. We suggest the following common names for the new species: &quot;Ngu Hang Mhai Khao Hin Poon&quot; (งูหางไหม้เขาหินปูน) (in Thai), &quot;Limestone Eyelash Pitviper&quot; (in English), and &quot;
<emphasis id="8078FD01B19D936C921B6FDDFC65BF24" italics="true" pageId="0" pageNumber="697">Resni-tchataya ukrashennaya kufiya</emphasis>
&quot; (in Russian).
</paragraph>
</subSubSection>
<subSubSection id="263F6BB48CD7E884403AC8AB0CF288AA" pageId="0" pageNumber="697" type="comparisons">
<paragraph id="9258622E2ED4388E3E32ADD452FFBF87" pageId="0" pageNumber="697">Comparisons.</paragraph>
<paragraph id="71E2ED278E29A05FE48E1D69A0A3B179" pageId="0" pageNumber="697">
The new species is morphologically and phylogenetically placed within the subgenus
<taxonomicName id="2BAC1B4273C53567C1CAD7D8029BCCB9" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="(Trimeresurus)" order="Squamata" pageId="0" pageNumber="697" rank="subGenus" subGenus="Trimeresurus">
<emphasis id="586D9F10FF46E184F2C7AE0D17FD46F8" italics="true" pageId="0" pageNumber="697">Trimeresurus</emphasis>
</taxonomicName>
(
<bibRefCitation id="357C796DF3ED4345E89011E66EFFC06D" DOI="https://doi.org/10.1016/j.ympev.2004.02.008" author="Malhotra, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="697" pagination="83 - 100" refId="B35" refString="Malhotra, A, Thorpe, RS, 2004a. A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution 32: 83 - 100, DOI: https://doi.org/10.1016/j.ympev.2004.02.008" title="A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis)." url="https://doi.org/10.1016/j.ympev.2004.02.008" volume="32" year="2004 a">Malhotra and Thorpe 2004a</bibRefCitation>
;
<bibRefCitation id="F5D31E81822634A83EAC7194BAFF0489" author="David, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="697" pagination="1 - 51" refId="B14" refString="David, P, Vogel, G, Dubois, A, 2011. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae). Zootaxa 2992: 1 - 51" title="On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae)." volume="2992" year="2011">David et al. 2011</bibRefCitation>
) and morphologically is overall most similar to other limestone-dwelling pitviper species, including
<taxonomicName id="D4094027AC96CC5C33DCD9A8B3569C0B" lsidName="T. kanburiensis" pageId="0" pageNumber="697" rank="species" species="kanburiensis">
<emphasis id="1FCEE5D237B78E11512DD77ACB2E3DA2" italics="true" pageId="0" pageNumber="697">T. kanburiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2E3C0180AB9565C8A4495843EF4A58C8" lsidName="T. kuiburi" pageId="0" pageNumber="697" rank="species" species="kuiburi">
<emphasis id="52F4681438918CEF3741802B339B2C8C" italics="true" pageId="0" pageNumber="697">T. kuiburi</emphasis>
</taxonomicName>
, and
<taxonomicName id="65DE56FA7E68F084793A4A8B8C4D66BE" lsidName="T. venustus" pageId="0" pageNumber="697" rank="species" species="venustus">
<emphasis id="A19D4E00C50B50345636DD4BBB993552" italics="true" pageId="0" pageNumber="697">T. venustus</emphasis>
</taxonomicName>
s. str., so the comparisons with these three species appear to be the most pertinent.
<taxonomicName id="7E2708FCE3FA7D0B0F95B772CFB51639" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="CE4409D3D2869DAA15FC3227B11F8D72" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="882B3418A9E7578ACDAB181790DEB405" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
can be easily distinguished from other congeners by having: 17 midbody scale rows, three or four small supraocular scales, and a small scale between the nasal and the scale formed by the fused second supralabial and loreal. The main diagnostic characters separating
<taxonomicName id="675B5C99DC4B526EA6CC92C0B56F448E" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="66CF77A21B3BD11DE44B4B66137AFB42" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="1F1881FAA635D5D49E0431B760F0EACF" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
from these three species are summarized in Table
<tableCitation id="7F909C6FB060C45FCF7DE4E5CF98AC03" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Comparison of morphological characters of Trimeresurus ciliaris sp. nov. with T. kanburiensis, T. kuiburi, and T. venustus s. str. Diagnostic differences from the new species are marked in bold." httpUri="http://table.plazi.org/id/4AFC56C5F0F756D648F29C541F308407" pageId="0" pageNumber="697" tableUuid="4AFC56C5F0F756D648F29C541F308407">5</tableCitation>
. The comparison of body coloration and head scalation of these species is presented in Figures
<figureCitation id="DC4BE3263C2685F45A58F67E3556732F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5</figureCitation>
and
<figureCitation id="E451D49D3BA71B661071EEA211439F7B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6</figureCitation>
, respectively. From the superficially similar
<taxonomicName id="54231616C693BF9F5F5CD7897D5FC104" authorityName="Orlov, Ryabov, Thanh &amp; H Cuc" authorityYear="2004" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus truongsonensis" order="Squamata" pageId="0" pageNumber="697" rank="species" species="truongsonensis">
<emphasis id="25D7364BA22D76B533C346F3F07C78F7" italics="true" pageId="0" pageNumber="697">Trimeresurus truongsonensis</emphasis>
</taxonomicName>
Orlov, Ryabov, Vu &amp; Ho, a member of the subgenus
<taxonomicName id="7F2D5F1D7D7F5642E463806CC9EE593A" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus (Viridovipera)" order="Squamata" pageId="0" pageNumber="697" rank="subGenus" subGenus="Viridovipera">
<emphasis id="B681CB5434B771C4B099244E0E547821" italics="true" pageId="0" pageNumber="697">Trimeresurus Viridovipera</emphasis>
</taxonomicName>
, the new species can be easily distinguished by having a long papillose and deeply forked hemipenis (vs. short spinose hemipenis) and partially fused first supralabial and nasal scales (vs. separate).
</paragraph>
<caption id="4AFC56C5F0F756D648F29C541F308407" ID-Table-UUID="4AFC56C5F0F756D648F29C541F308407" httpUri="http://table.plazi.org/id/4AFC56C5F0F756D648F29C541F308407" pageId="0" pageNumber="697" start="Table 5" startId="T5">
<paragraph id="5330628E8DD19892144883FEE434F939" pageId="0" pageNumber="697">
<emphasis id="D7AB7259725F97CA98AA11472FA38295" bold="true" pageId="0" pageNumber="697">Table 5.</emphasis>
Comparison of morphological characters of
<taxonomicName id="1BB21370FF010255D0ED7EA3A81C4CDF" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="65ABB6388E48F73BA4702B4991DE3827" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="3DFDF7AFA0373CDA8FAECF7ACA1AD079" bold="true" pageId="0" pageNumber="697">sp. nov</emphasis>
. with
<taxonomicName id="E1D5D1862619DD699A9D8A36320DD136" lsidName="T. kanburiensis" pageId="0" pageNumber="697" rank="species" species="kanburiensis">
<emphasis id="7191BFB1A76AAEF465B121A4B009859E" italics="true" pageId="0" pageNumber="697">T. kanburiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B40F8E357DB5B8D56BAEFA8BA201B8F8" lsidName="T. kuiburi" pageId="0" pageNumber="697" rank="species" species="kuiburi">
<emphasis id="691B308734E81B1C9669DE95EAADFB17" italics="true" pageId="0" pageNumber="697">T. kuiburi</emphasis>
</taxonomicName>
, and
<taxonomicName id="B303DCB10775C2AF270506DF304E4DF5" lsidName="T. venustus" pageId="0" pageNumber="697" rank="species" species="venustus">
<emphasis id="8772FCEB3A2B4552C04B53973D294E3B" italics="true" pageId="0" pageNumber="697">T. venustus</emphasis>
</taxonomicName>
s. str. Diagnostic differences from the new species are marked in bold.
</paragraph>
</caption>
<paragraph id="4D8CB8DB99CFA404CB8CECB745CC6E74" pageId="0" pageNumber="697">
<table id="1451675C1A95CEC28EB45592383FF5D7" pageId="0" pageNumber="697">
<tr id="6BB581BF0ED6DA554562673DC41FB0DE" pageId="0" pageNumber="697">
<td id="1049ED9E86798C9B3A15DE5803C51FDF" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="36857255A8A3BA3F89AFF8D786B8FFAC" bold="true" pageId="0" pageNumber="697">Character</emphasis>
</td>
<td id="FFF3D985B6D9F072DEEA80FF6CC01368" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<taxonomicName id="9D735B0197EE8B8FD327F88DE3E6827A" lsidName="T. ciliaris" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="D57E604483E5429AC65DA9FAB16B857C" italics="true" pageId="0" pageNumber="697">T. ciliaris</emphasis>
</taxonomicName>
<emphasis id="3FF21E30BEFD5A0E9686AB26B44AEACC" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
</td>
<td id="2DCE019D752C2D776C04D37EB2551A57" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<taxonomicName id="BE6A8FB6C09A2B873A157F0FC47E6F8F" lsidName="T. kanburiensis" pageId="0" pageNumber="697" rank="species" species="kanburiensis">
<emphasis id="DB3F70756A90962F9439CBC2E3BD8A08" italics="true" pageId="0" pageNumber="697">T. kanburiensis</emphasis>
</taxonomicName>
</td>
<td id="67229D2BDF648FF0C953229856E433A2" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<taxonomicName id="C00117B87837949F9BB639FDE725D086" lsidName="T. kuiburi" pageId="0" pageNumber="697" rank="species" species="kuiburi">
<emphasis id="C9EF5ABF3FCE9FD3A75CA69CC468969B" italics="true" pageId="0" pageNumber="697">T. kuiburi</emphasis>
</taxonomicName>
</td>
<td id="E739AC7E7F37C5C12AF094817A592C3A" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<taxonomicName id="B4FAF2127E3C3F19FE26F39B2378EF6C" lsidName="T. venustus" pageId="0" pageNumber="697" rank="species" species="venustus">
<emphasis id="F608D498A1C3B8DCB4B44DAB5EC720F3" italics="true" pageId="0" pageNumber="697">T. venustus</emphasis>
</taxonomicName>
<emphasis id="6E20AA66ECE1472CDAA24F0E9742C0FB" bold="true" pageId="0" pageNumber="697">s. str.</emphasis>
</td>
</tr>
<tr id="D473F26E1ACE229FCA54EE233C05AD62" pageId="0" pageNumber="697">
<td id="6DF0D38C3AA4B991457836074F440433" colspan="1" pageId="0" pageNumber="697" rowspan="1">Max SVL (males, mm)</td>
<td id="23B826D1BF66296089C4301C8BE7A1C6" colspan="1" pageId="0" pageNumber="697" rowspan="1">360</td>
<td id="F4624ECE598EEF63EBA8D1AA7D86E364" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="7CD357EBAA2ECD58F00EC2D8804476F1" bold="true" pageId="0" pageNumber="697">412</emphasis>
</td>
<td id="DCEB8B81C6D497BFBC2CF2719674B59C" colspan="1" pageId="0" pageNumber="697" rowspan="1">376</td>
<td id="33513050C38D815B4142970C0501C005" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="4ADF40EC09A0CF849D87F06F0CD22E8D" bold="true" pageId="0" pageNumber="697">490</emphasis>
</td>
</tr>
<tr id="1B7D774BBAE00314669DD669E4AF0CBB" pageId="0" pageNumber="697">
<td id="6F742A99C2A869F2F535E46A4C4E02F0" colspan="1" pageId="0" pageNumber="697" rowspan="1">Max SVL (females, mm)</td>
<td id="558C22E17EE0BDAE54DBDB8BAE9105DE" colspan="1" pageId="0" pageNumber="697" rowspan="1">333</td>
<td id="076CB4491CA1EB952E0BA6E3CE2F9F63" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="931429EEDD5A633CDBF2484D6532E8D3" bold="true" pageId="0" pageNumber="697">572</emphasis>
</td>
<td id="E57DC6887A84506FE298D5FBF51A9ACF" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="CB9F6776C745F5EA25479B84428F2B20" bold="true" pageId="0" pageNumber="697">451</emphasis>
</td>
<td id="1E6A24E480A6F66C906D63409CE4CD52" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="9584200BF61DB9ED6554E71D1EDE4EF3" bold="true" pageId="0" pageNumber="697">456</emphasis>
</td>
</tr>
<tr id="ABCB0ED3D5A54514F9616A568B8034AE" pageId="0" pageNumber="697">
<td id="97EBFA5EEF9EC93985C4879A46D831B8" colspan="1" pageId="0" pageNumber="697" rowspan="1">IOS</td>
<td id="9A92142E382AEB8597C351942FF80F38" colspan="1" pageId="0" pageNumber="697" rowspan="1">11-14</td>
<td id="CB087BF6A4CE4201CD04FDFACEA85DA7" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="1A3D4E59C67CCDAFCA8975735C81D64F" bold="true" pageId="0" pageNumber="697">7-9</emphasis>
</td>
<td id="7B8A5B05295C48F0C8434D04AE662760" colspan="1" pageId="0" pageNumber="697" rowspan="1">9-11</td>
<td id="01BF6F0E94EAF9D49E67DFEAE8A30155" colspan="1" pageId="0" pageNumber="697" rowspan="1">8-12</td>
</tr>
<tr id="463B36654052582BFCED7AF03522A085" pageId="0" pageNumber="697">
<td id="9F1C55CCF0083F4E58E5CCB36870BA2E" colspan="1" pageId="0" pageNumber="697" rowspan="1">Internasals in contact</td>
<td id="80D784B8BB1C941D269118789F8FA856" colspan="1" pageId="0" pageNumber="697" rowspan="1">0%</td>
<td id="E6B91CE6E7BE3F962A61FB6565D2EC3F" colspan="1" pageId="0" pageNumber="697" rowspan="1">0%</td>
<td id="C51ACBACE7E61EFD0648EE52DF34A35B" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="575FCF3B002F1291AC0C1084C8ED56F7" bold="true" pageId="0" pageNumber="697">100</emphasis>
%
</td>
<td id="92F870648C44A42008813204DF3CA307" colspan="1" pageId="0" pageNumber="697" rowspan="1">45%</td>
</tr>
<tr id="27F937912571D7210ED386645C988CD4" pageId="0" pageNumber="697">
<td id="3F14B203293B9407464DE323D3D40665" colspan="1" pageId="0" pageNumber="697" rowspan="1">ASR</td>
<td id="93F97A8F3F18C8FA2044EF009BE007F5" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="A2669F5B8597CA39A71731F66121DCC8" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="5B7C6C65009CB4BA8A6187A735ADBD2A" bold="true" pageId="0" pageNumber="697">23 (21 or 22)</emphasis>
</td>
<td id="401218C6BB9095DD27D61076A6C6937C" colspan="1" pageId="0" pageNumber="697" rowspan="1">21 (23)</td>
<td id="CC1FD30DD388E344F36A5F94F36FD687" colspan="1" pageId="0" pageNumber="697" rowspan="1">23 (21 or 25)</td>
</tr>
<tr id="D67B2F1646D35E198CB04EBC7CFDB782" pageId="0" pageNumber="697">
<td id="E7860638F884D564E29B926CEFE77E82" colspan="1" pageId="0" pageNumber="697" rowspan="1">MSR</td>
<td id="908070B9CFCA5B33020EE3CC704FD447" colspan="1" pageId="0" pageNumber="697" rowspan="1">17</td>
<td id="7C989DDA3C3568B08EDD579B4AFEE48E" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="4BF9F5F3B14A1D77722D43C748CEB868" bold="true" pageId="0" pageNumber="697">19</emphasis>
</td>
<td id="486F2060DCED3EB4FC69E796C7B5E43D" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="AA593D48758DDAE69419E47D29F4A972" bold="true" pageId="0" pageNumber="697">19</emphasis>
</td>
<td id="7400E17F7E2A7F5A0E6678050124F7E4" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="E09528FEC77EBCA0F920516A980D4FA9" bold="true" pageId="0" pageNumber="697">21 (19)</emphasis>
</td>
</tr>
<tr id="036D82B646F94FA17173CFCB23955FC7" pageId="0" pageNumber="697">
<td id="7B2FAECB955DA5189DBD7012F03362D9" colspan="1" pageId="0" pageNumber="697" rowspan="1">PSR</td>
<td id="0C17C6668819958A842833F856A7A6C8" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
<td id="8B820AAA61B7280AC2F0D5C11FE9611F" colspan="1" pageId="0" pageNumber="697" rowspan="1">15 (16)</td>
<td id="C65CA6968701E257A28CAE994593ED92" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
<td id="F33785C6FFAEAD37150F4FAFA93CD8C0" colspan="1" pageId="0" pageNumber="697" rowspan="1">15</td>
</tr>
<tr id="D00BE05D4AB59C057981F0717277A556" pageId="0" pageNumber="697">
<td id="C2F8229B379EF6E36508FE8D40810E03" colspan="1" pageId="0" pageNumber="697" rowspan="1">VEN males</td>
<td id="348C64E4876EBF0B471C4E9541C1A2FF" colspan="1" pageId="0" pageNumber="697" rowspan="1">172-175</td>
<td id="E151BF2C11078518791AA66E90A90604" colspan="1" pageId="0" pageNumber="697" rowspan="1">172</td>
<td id="0FBE611D0614D1612CFB9CE8EAA6B3F4" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="6EC37E66BEDB8EAAB9FF141BF31D3F9F" bold="true" pageId="0" pageNumber="697">164-166</emphasis>
</td>
<td id="E085CD0C2494DFC8F988E680B9AF1D57" colspan="1" pageId="0" pageNumber="697" rowspan="1">166-177</td>
</tr>
<tr id="9C1389A26B168905C18BB8C6283DA349" pageId="0" pageNumber="697">
<td id="E013B3C54C009E560D3444EE5261AEC9" colspan="1" pageId="0" pageNumber="697" rowspan="1">VEN females</td>
<td id="E7C67B4F5EAB77B80FC0DD255D677300" colspan="1" pageId="0" pageNumber="697" rowspan="1">171</td>
<td id="F2D8A502798A054640A15767DCE951B9" colspan="1" pageId="0" pageNumber="697" rowspan="1">170-178</td>
<td id="F7D303BD42CB7CCE6B580CEB0D9A4331" colspan="1" pageId="0" pageNumber="697" rowspan="1">164-171</td>
<td id="083801623754030FF0C61AD44CEF3E2A" colspan="1" pageId="0" pageNumber="697" rowspan="1">172-181</td>
</tr>
<tr id="848B990E787A62DBB2BA9BB089A845A8" pageId="0" pageNumber="697">
<td id="FDE3124AA2091CB9960623C62E6D3E0D" colspan="1" pageId="0" pageNumber="697" rowspan="1">SC males</td>
<td id="09B2434AC37719DE2270643034FE6F89" colspan="1" pageId="0" pageNumber="697" rowspan="1">59-63</td>
<td id="5BEF1E6D5859157E8B357B4AA5A6A80F" colspan="1" pageId="0" pageNumber="697" rowspan="1">59</td>
<td id="32CDD594C57F5FDE560422F5C4004340" colspan="1" pageId="0" pageNumber="697" rowspan="1">63-65</td>
<td id="7E9EEBF5713FF6A6E9979FA98F28FAF8" colspan="1" pageId="0" pageNumber="697" rowspan="1">63-72</td>
</tr>
<tr id="37A4258B06DD0E4B885D0D6808F94781" pageId="0" pageNumber="697">
<td id="02DA614FD61D10AD9ED46A02ECE84285" colspan="1" pageId="0" pageNumber="697" rowspan="1">SC females</td>
<td id="9DFC11C78955C526D889863D1C9F7448" colspan="1" pageId="0" pageNumber="697" rowspan="1">52</td>
<td id="8A6535A39DAECEAF21DAD9D00D6FAECB" colspan="1" pageId="0" pageNumber="697" rowspan="1">41-51</td>
<td id="157B687C2256ECE997F7104AEF2B1ED9" colspan="1" pageId="0" pageNumber="697" rowspan="1">51-53</td>
<td id="2CC0E1E244CC75106BB507501A1B071E" colspan="1" pageId="0" pageNumber="697" rowspan="1">50-66</td>
</tr>
<tr id="62CDFF471995483BE732E0C48F022DCC" pageId="0" pageNumber="697">
<td id="29C14388E205246E1963BDCDF976E176" colspan="1" pageId="0" pageNumber="697" rowspan="1">Small scale between nasal and the scale formed by the fused second supralabial and loreal</td>
<td id="8EF18D14C8E38E5B2604B1D3B1FB9E15" colspan="1" pageId="0" pageNumber="697" rowspan="1">present</td>
<td id="B2DFA1F4B10CB9D7DB8D7ECFB2A2C764" colspan="1" pageId="0" pageNumber="697" rowspan="1">present/absent</td>
<td id="CFC317323A70C75FFC12B69E4E2BA215" colspan="1" pageId="0" pageNumber="697" rowspan="1">absent</td>
<td id="E033076C92F9653F9942EF2DBED2B17B" colspan="1" pageId="0" pageNumber="697" rowspan="1">absent</td>
</tr>
<tr id="DEB20286024B6326FEEBD1A74F54E3AE" pageId="0" pageNumber="697">
<td id="262D1EC6605A0F3FEAA1EF03F5DD824D" colspan="1" pageId="0" pageNumber="697" rowspan="1">Dorsum color in life</td>
<td id="B2B519E2D049441E3A8C6E97B5DBB25B" colspan="1" pageId="0" pageNumber="697" rowspan="1">Reddish-brown bands on emerald-green background</td>
<td id="E0FD06AE2A20E714D06FDD9102E969DF" colspan="1" pageId="0" pageNumber="697" rowspan="1">Dark olive-brown bands on olive-grayish background</td>
<td id="D2661A83773336C12158A070AC5C2647" colspan="1" pageId="0" pageNumber="697" rowspan="1">Red/purple bands on bottle green background</td>
<td id="CAAACB94C65BB1E6CDB0C9F68E1C5AAB" colspan="1" pageId="0" pageNumber="697" rowspan="1">Red/purple bands on dark/bottle green background</td>
</tr>
<tr id="B35E97B166508EE0A882651660CB8175" pageId="0" pageNumber="697">
<td id="70EB794345CAB44CC82B91D78873F91C" colspan="1" pageId="0" pageNumber="697" rowspan="1">White vertebral dots (in males)</td>
<td id="DBBB4680D23BE3AA95EA2F48AE3057DB" colspan="1" pageId="0" pageNumber="697" rowspan="1">Present, every 2-3 scales</td>
<td id="FFA5F5112F404F1C54DF2EE1469CAE9C" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="BAE302BB0A19237D447817FBE3EB478F" bold="true" pageId="0" pageNumber="697">Present, every 3-5 scales</emphasis>
</td>
<td id="F52311BE9DDCA87C5574B43A57F2C914" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="6E6F1DF2B616787763DB5813379C49F1" bold="true" pageId="0" pageNumber="697">Present, every 5-6 scales</emphasis>
</td>
<td id="95068DED9F19FA350B9DBCC94681D001" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="E85CC9A2FC3BB6CD78C7EB87AE94E6D9" bold="true" pageId="0" pageNumber="697">Absent</emphasis>
</td>
</tr>
<tr id="2EE4EE4B6DAA5E4B076A4B72AD2B7801" pageId="0" pageNumber="697">
<td id="2B5AAD93C8A14DE622A87EE48BD9D2CD" colspan="1" pageId="0" pageNumber="697" rowspan="1">White vertebral dots (in females)</td>
<td id="8D9C6453663232DB95935E2350D1DDEC" colspan="1" pageId="0" pageNumber="697" rowspan="1">Present, every 2-3 scales</td>
<td id="9263FB4BB5E5F53A6BBF083A8FC9C138" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="245D32B7D2CF6E3B92B7B4D929A1F26A" bold="true" pageId="0" pageNumber="697">Absent</emphasis>
</td>
<td id="58778FA753048DF1A8BFAC62964273A5" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="CE63406FEA6F7C5FC88B8D76896B3F13" bold="true" pageId="0" pageNumber="697">Absent</emphasis>
</td>
<td id="D598F3A3C37AF251ACD5D1B6606D4EFD" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="366BCB6DA10A7E8239FF558BCC106E11" bold="true" pageId="0" pageNumber="697">Absent</emphasis>
</td>
</tr>
<tr id="25F14D38F6C5ABE1572881239711FFCE" pageId="0" pageNumber="697">
<td id="3AD06BF025A1060F035A92FC3F043D43" colspan="1" pageId="0" pageNumber="697" rowspan="1">Belly background color in life</td>
<td id="30329164F42CC95ABECC76A35280F691" colspan="1" pageId="0" pageNumber="697" rowspan="1">creamy white</td>
<td id="E36BD97D2B17A3D6B346848A2CE4C5E9" colspan="1" pageId="0" pageNumber="697" rowspan="1">creamy white</td>
<td id="4028885AF7E74D5B822C8216B519C4F4" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="BE1A228458E2E48041CD3B453A2041D1" bold="true" pageId="0" pageNumber="697">pale green</emphasis>
</td>
<td id="AA8074D05E95EEE1FEE0198C6557A52B" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="B703479A808893450A582DDC7FB6B707" bold="true" pageId="0" pageNumber="697">pale green</emphasis>
</td>
</tr>
<tr id="2EC16F719C03247A77023DDFF47DB445" pageId="0" pageNumber="697">
<td id="C1856B6374F35BEA6D8F86D8110A330C" colspan="1" pageId="0" pageNumber="697" rowspan="1">Lateral dark stripe on ventrals</td>
<td id="0C7BC779BA50B34E68EF12C67ED3A881" colspan="1" pageId="0" pageNumber="697" rowspan="1">Absent</td>
<td id="E6E7081DA09E2502003AAEB2D9C57E91" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="00ABF96E1E5D3AD99E7BCB3C12700CAB" bold="true" pageId="0" pageNumber="697">Always present, discontinuous, olive-brown</emphasis>
</td>
<td id="8FABDBCAB78EBB736C13D2D700D372B0" colspan="1" pageId="0" pageNumber="697" rowspan="1">Absent</td>
<td id="A04D0D018E957677B8A2CB51A9A44979" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="1CA87C072226A83B678C6A873D6E58B3" bold="true" pageId="0" pageNumber="697">Always present, continuous or discontinuous, red</emphasis>
</td>
</tr>
<tr id="C52C6B13B9072DAA3CA42064C5E905F5" pageId="0" pageNumber="697">
<td id="24C7C27FFDC949826984FA9ECB45FD56" colspan="1" pageId="0" pageNumber="697" rowspan="1">Supraocular scale</td>
<td id="01DA7CDCB5D30C2E6799B0B7F878A7DD" colspan="1" pageId="0" pageNumber="697" rowspan="1">Three or four, small</td>
<td id="EC7E8306D05387E6FDF99941EAAB6091" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="51E9A53642F2DAC74D79C36FE55FFD12" bold="true" pageId="0" pageNumber="697">One, large</emphasis>
</td>
<td id="EBFD3C554B68C113E0A7F26919E52AE8" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="7B4B6510B647D1D80895C1F28E3D5709" bold="true" pageId="0" pageNumber="697">One, large</emphasis>
</td>
<td id="790EB471BC94C098429ECE5774875139" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<emphasis id="D5F1BC00E30FFEC1FE8A53B1F184A13C" bold="true" pageId="0" pageNumber="697">One, large</emphasis>
</td>
</tr>
<tr id="E9A7B1AC073EE008DFCA5AF0E5E59013" pageId="0" pageNumber="697">
<td id="6E9815FB6F9AF4FB7FA1354B4B593BA3" colspan="1" pageId="0" pageNumber="697" rowspan="1">Iris color</td>
<td id="C43741504572A756C974B8F11B9CA791" colspan="1" pageId="0" pageNumber="697" rowspan="1">Olive-green with faded brown horizontal stripe</td>
<td id="B9C95733586443F945A76C2084FFA22F" colspan="1" pageId="0" pageNumber="697" rowspan="1">brown, slightly golden</td>
<td id="D29E3CF16C80867C0AAE8E11652580B7" colspan="1" pageId="0" pageNumber="697" rowspan="1">copper</td>
<td id="4C12D7129D776AD1973BB73E9FE12A48" colspan="1" pageId="0" pageNumber="697" rowspan="1">yellowish-brown/gold</td>
</tr>
<tr id="2CDDFF5A926881648BBC9D06ED8BA550" pageId="0" pageNumber="697">
<td id="4C715B67572942A814885EB629255F56" colspan="1" pageId="0" pageNumber="697" rowspan="1">Tail color</td>
<td id="90817A26B125E713D96DA46AFC27B35B" colspan="1" pageId="0" pageNumber="697" rowspan="1">dark brown mottled with rusty</td>
<td id="95EE64A4AF2918DC7280C25902D8C741" colspan="1" pageId="0" pageNumber="697" rowspan="1">brownish-gray with olive-brown blotches</td>
<td id="99B11CF904BE6D1AB56705852978B4EB" colspan="1" pageId="0" pageNumber="697" rowspan="1">red with some thin lighter bands</td>
<td id="BD735C527AE2F1FBA97948D10FB21802" colspan="1" pageId="0" pageNumber="697" rowspan="1">brown with dark purplish-brown crossbars</td>
</tr>
<tr id="B3033F5F613C484B9468F400B50E859D" pageId="0" pageNumber="697">
<td id="70CD769C7432AE775C5490248E642732" colspan="1" pageId="0" pageNumber="697" rowspan="1">Distribution</td>
<td id="D0F9FE4B1D631BB46F802F736983AE87" colspan="1" pageId="0" pageNumber="697" rowspan="1">Thailand (Trang, Satun), Malaysia (Perlis)</td>
<td id="E3EA58064D760A1F7D03009731861C9A" colspan="1" pageId="0" pageNumber="697" rowspan="1">Thailand (Kanchanaburi)</td>
<td id="4E34BE446F4AD6D58D3187E0BB9461A9" colspan="1" pageId="0" pageNumber="697" rowspan="1">Thailand (Prachuap Khiri Khan)</td>
<td id="4146D0431DE7AF64739022FB410CB0CF" colspan="1" pageId="0" pageNumber="697" rowspan="1">Thailand (Surat Thani, Phang-Nga, Trang, Nakhon Si Thammarat, Surat Thani, Krabi, Chumphon?), Malaysia (Kedah?)</td>
</tr>
<tr id="7D5FC32557C5DC272797AE0D11417DED" pageId="0" pageNumber="697">
<td id="528E3B33E34DD7544C965B56525C80CA" colspan="1" pageId="0" pageNumber="697" rowspan="1">Sources</td>
<td id="C5CA8E9579A008DD147210C15E08C4A2" colspan="1" pageId="0" pageNumber="697" rowspan="1">our data</td>
<td id="D673590116082F65B427E8F763EEBF8C" colspan="1" pageId="0" pageNumber="697" rowspan="1">
Smith (1943);
<bibRefCitation id="5FB8F61479AF76EFD42DC7EB8136A97C" author="David, P" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="697" pagination="81 - 90" refId="B15" refString="David, P, Vogel, G, Sumontha, M, Pauwels, OSG, Chanhome, L, 2004. Expanded description of the poorly known pitviper Trimeresurus kanburiensis Smith, 1943, with confirmation of the validity of Trimeresurus venustus Vogel, 1991 (Reptilia: Serpentes: Crotalidae). Russian Journal of Herpetology 11: 81 - 90" title="Expanded description of the poorly known pitviper Trimeresurus kanburiensis Smith, 1943, with confirmation of the validity of Trimeresurus venustus Vogel, 1991 (Reptilia: Serpentes: Crotalidae)." volume="11" year="2004">David et al. (2004)</bibRefCitation>
; our data
</td>
<td id="38BAAE6A8C68FE00C2334CE96986CEBC" colspan="1" pageId="0" pageNumber="697" rowspan="1">
<bibRefCitation id="056A21AEC4B51264ACD9CEA41344661D" DOI="https://doi.org/10.11646/zootaxa.4974.2.4" author="Sumontha, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="697" pagination="307 - 332" refId="B54" refString="Sumontha, M, Suntrarachun, S, Pauwels, OSG, Pawangkhanant, P, Chomngam, N, Iamwiriyakul, P, Chanhome, L, 2021. A new karst-dwelling, colorful pitviper (Viperidae: Trimeresurus) from northern Peninsular Thailand. Zootaxa 4974: 307 - 332, DOI: https://doi.org/10.11646/zootaxa.4974.2.4" title="A new karst-dwelling, colorful pitviper (Viperidae: Trimeresurus) from northern Peninsular Thailand." url="https://doi.org/10.11646/zootaxa.4974.2.4" volume="4974" year="2021">Sumontha et al. (2021)</bibRefCitation>
; our data
</td>
<td id="A177BE212A5CBECADD5377CD95FE8751" colspan="1" pageId="0" pageNumber="697" rowspan="1">
Vogel (1991);
<bibRefCitation id="2DCFE45ED81D5AE4B6190D218F513A1A" author="David, P" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="697" pagination="81 - 90" refId="B15" refString="David, P, Vogel, G, Sumontha, M, Pauwels, OSG, Chanhome, L, 2004. Expanded description of the poorly known pitviper Trimeresurus kanburiensis Smith, 1943, with confirmation of the validity of Trimeresurus venustus Vogel, 1991 (Reptilia: Serpentes: Crotalidae). Russian Journal of Herpetology 11: 81 - 90" title="Expanded description of the poorly known pitviper Trimeresurus kanburiensis Smith, 1943, with confirmation of the validity of Trimeresurus venustus Vogel, 1991 (Reptilia: Serpentes: Crotalidae)." volume="11" year="2004">David et al. (2004)</bibRefCitation>
;
<bibRefCitation id="907EAC5F1A78AFC039176016EFA83B41" author="Malhotra, A" journalOrPublisher="Herpetological Journal" pageId="0" pageNumber="697" pagination="21 - 33" refId="B36" refString="Malhotra, A, Thorpe, RS, 2004b. Reassessment of the validity and diagnosis of the pitviper Trimeresurus venustus Vogel, 1991. Herpetological Journal 14: 21 - 33" title="Reassessment of the validity and diagnosis of the pitviper Trimeresurus venustus Vogel, 1991." volume="14" year="2004 b">Malhotra and Thorpe (2004b)</bibRefCitation>
; our data
</td>
</tr>
</table>
</paragraph>
<paragraph id="A5F61F036A85EA8DFB03F37A05A9008E" pageId="0" pageNumber="697">
In particular,
<taxonomicName id="9E2C1F08111E9B0B16F5A663C4658426" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="7A0B1D928D490803B4BEE1ADFC3DB92A" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="700839A72B709E3126AB232C16F168F9" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
differs from
<taxonomicName id="A1A8D2926C65927474CB4A49C74658DC" lsidName="T. kanburiensis" pageId="0" pageNumber="697" rank="species" species="kanburiensis">
<emphasis id="20E3A4D4BAA63D688489DD24DEA35169" italics="true" pageId="0" pageNumber="697">T. kanburiensis</emphasis>
</taxonomicName>
by having: smaller maximal SVL (360 mm in male, 333 mm in female vs. 412 mm in male, 572 mm in female); lower anterior number of dorsal scale rows (17 vs. 23, rarely 21 or 22); lower number of midbody scale rows (17 vs. 19); absence of dark lateral stripe on ventrals, Fig.
<figureCitation id="C55B3545866D5AD009325CBB1913EC0C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5A</figureCitation>
(vs. dark lateral stripe on ventrals always present, discontinuous, olive-brown, Fig.
<figureCitation id="18F4D138F912F45BE63542F1D223ACB5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5B</figureCitation>
); three or four small supraocular scales, Fig.
<figureCitation id="E7635D45B110AD1AED423F396B0B26AB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6B</figureCitation>
(vs. single large supraocular scale, Fig.
<figureCitation id="506FDC9AC3AFB7E2181CF57AB59AA964" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6D</figureCitation>
); body color in life (reddish-brown bands on an emerald-green background [Fig.
<figureCitation id="CD174C9F5AC78FBBE150AC83B80E8A86" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5A</figureCitation>
] vs. dark olive-brown bands on olive-greyish background [Fig.
<figureCitation id="F4B2B2F29B8AC45F0671003A9D2B540D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5B</figureCitation>
]); white vertebral spots present in both sexes separated by 2-3 dorsal scales vs. white vertebral spots present only in males and are separated by 3-5 dorsal scales; iris color (olive-green with faded brown horizontal stripe [Fig.
<figureCitation id="7E12660A84FC183E4C1CCDE9189CC698" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6A</figureCitation>
] vs. brown, slightly golden [Fig.
<figureCitation id="86FE6DF1E293DE4868A64E40878DF13A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6C</figureCitation>
]); and tail color (dark brown mottled with rusty spots vs. brownish-gray with olive-brown blotches).
</paragraph>
<paragraph id="6B8348EBE0813E152DAB62FE5ABC9C8E" pageId="0" pageNumber="697">
<taxonomicName id="7F9BA5B6AF6AF50C0FAF7C4CBA529C7A" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="5DF656022885854123D9384C04798B01" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="ADBEED56D40BCA653AFE578700D20684" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
further differs from
<taxonomicName id="45F33499DC44ABE4681B4273FB3457CE" lsidName="T. kuiburi" pageId="0" pageNumber="697" rank="species" species="kuiburi">
<emphasis id="7B4F970EC5C2F53AAE5BB72E2E3B1D60" italics="true" pageId="0" pageNumber="697">T. kuiburi</emphasis>
</taxonomicName>
by having: smaller maximal SVL in female (333 mm vs. 451 mm); internasals separated by two or three scales (vs. always in contact); lower anterior number of dorsal scale rows (17 vs. 21 rarely 23); lower number midbody scale rows (17 vs. 19); higher number of ventrals in females (172-175 vs. 164-166); ventral surfaces of body creamy-white (vs. pale-green); three or four small supraocular scales, Fig.
<figureCitation id="9EC0EE7D4A5E8EE0FBD5CBB45EBED858" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6A</figureCitation>
(vs. one large supraocular scale, Fig.
<figureCitation id="6DC5F93CD177F092AD173FED2CB04596" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6F</figureCitation>
); small scale between nasal and second supralabial present vs. absent; body color in life (reddish-brown bands on emerald-green background [Fig.
<figureCitation id="A53B078905A9C918C4FE20BD1D5E2BD9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5A</figureCitation>
] vs. red to purple bands on bottle-green background [Fig.
<figureCitation id="57AEB483DF00BBA62ADA515E747BDF6A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5C</figureCitation>
]); white vertebral spots present in both sexes separated by 2-3 dorsal scales vs. white vertebral dots present only in males and are separated by 5-6 dorsal scales; iris color (olive-green with faded brown horizontal stripe [Fig.
<figureCitation id="430A3E9E83F089CA010E1205F59D46E1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6A</figureCitation>
] vs. copper [Fig.
<figureCitation id="B091239FA92DF59513182FB2E16CBC6C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6E</figureCitation>
]); and tail color (dark brown mottled with rusty spots vs. red with some thin lighter bands).
</paragraph>
<paragraph id="974DEE9B16DD554CBF02F7BF59E99F7B" pageId="0" pageNumber="697">
<taxonomicName id="E71E05B409D09DCA111F10E413E8BE5A" authorityName="Idiiatullina &amp; Pawangkhanant &amp; Tawan &amp; Worranuch &amp; Dechochai &amp; Suwannapoom &amp; Nguyen &amp; Chanhome &amp; Poyarkov" authorityYear="2023" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus ciliaris" order="Squamata" pageId="0" pageNumber="697" rank="species" species="ciliaris">
<emphasis id="E5CA1DF60DF4435F81C1F34B7C3E9CAF" italics="true" pageId="0" pageNumber="697">Trimeresurus ciliaris</emphasis>
</taxonomicName>
<emphasis id="D565066A95B1D3B87F1D96C3D46A59A8" bold="true" pageId="0" pageNumber="697">sp. nov.</emphasis>
can be further differentiated from
<taxonomicName id="66EB480CAE25FE6CDF357FDFEA9F66E4" lsidName="T. venustus" pageId="0" pageNumber="697" rank="species" species="venustus">
<emphasis id="170BA4378560A20C9B331A588EFA027A" italics="true" pageId="0" pageNumber="697">T. venustus</emphasis>
</taxonomicName>
s. str. by having: smaller maximal SVL (360 mm in male, 333 mm in female vs. 490 mm in male, 456 mm in female); lower anterior number of dorsal scale rows (17 vs. 23, rarely 21 or 25); lower number midbody scale rows (17 vs. 21, rarely 19); white vertebral spots present in both sexes separated by 2-3 dorsal scales (vs. absent); ventral surfaces creamy-white (vs. pale-green); lateral dark stripe on ventrals absent (vs. lateral dark stripe on ventrals always present, discontinuous, red); three or four small supraocular scales, Fig.
<figureCitation id="C049BC4C285DCD623972E1493276F8BA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6A</figureCitation>
(vs. one large supraocular scale, Fig.
<figureCitation id="8B5E3B16CC21CFCBE8B3C3D6759892C4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6H</figureCitation>
); small scale between nasal and second supralabial present vs. absent; body color in life (reddish-brown bands on emerald-green background [Fig.
<figureCitation id="15630D5022418B7A43CC6F87FEC15704" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5A</figureCitation>
] vs. red to purple bands on dark bottle-green background [Fig.
<figureCitation id="9304246E94D844E976D734C7FE29B220" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body coloration between males of A Trimeresurus ciliaris sp. nov.; B T. kanburiensis; C T. kuiburi; and D T. venustus s. str. Photographs by P. Pawangkhanant (A, B, D) and T. Woranuch (C)." figureDoi="10.3897/vz.73.e109854.figure5" httpUri="https://binary.pensoft.net/fig/908087" pageId="0" pageNumber="697">5D</figureCitation>
]); white vertebral spots present in both sexes separated by 2-3 dorsal scales vs. white vertebral dots absent in both sexes; iris color (olive-green with faded brown horizontal stripe [Fig.
<figureCitation id="73027A4D5843879E7CE688288739FCCF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6A</figureCitation>
] vs. yellowish-brown to gold [Fig.
<figureCitation id="60A5AA66D2441EBD3B4F29FCA449D893" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of head coloration (left profile and dorsal view of the head) between males of A-B Trimeresurus ciliaris sp. nov.; C - D T. kanburiensis; E - F T. kuiburi; and G - H T. venustus s. str. Photographs by P. Pawangkhanant (A-D), A. Kaosung (E-F, H) and M. Naiduangchan (G)." figureDoi="10.3897/vz.73.e109854.figure6" httpUri="https://binary.pensoft.net/fig/908088" pageId="0" pageNumber="697">6G</figureCitation>
]); and tail coloration (dark brown mottled with rusty spots vs. brown with dark purplish-brown crossbars).
</paragraph>
<paragraph id="7B4ECB24EA17CA31950DE1C82F22D9EE" pageId="0" pageNumber="697">
Among the other species in the subgenus
<taxonomicName id="5BEB1161D04F5F4A91B0B5BDD0A40A0E" authority="Trimeresurus" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" lsidName="Trimeresurus (Trimeresurus)" order="Squamata" pageId="0" pageNumber="697" rank="subGenus" subGenus="Trimeresurus">Trimeresurus Trimeresurus</taxonomicName>
, the new species can be readily distinguished from
<taxonomicName id="F72B4B9D2FDE4A802C4898D3B7C9DDF3" lsidName="T. albolabris" pageId="0" pageNumber="697" rank="species" species="albolabris">
<emphasis id="EC920D0EA9C9B49859B4DB00E002F552" italics="true" pageId="0" pageNumber="697">T. albolabris</emphasis>
</taxonomicName>
,
<taxonomicName id="5BD1F4D3C2E886DA3C0416A5771E8222" lsidName="T. andersonii" pageId="0" pageNumber="697" rank="species" species="andersonii">
<emphasis id="700AB231289BA34BDF7A018189B1B973" italics="true" pageId="0" pageNumber="697">T. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="1439ACF5A4217AFD1A24CCDED71DC0BB" lsidName="T. cantori" pageId="0" pageNumber="697" rank="species" species="cantori">
<emphasis id="D628A48B32A36A643D14EF3B498CC03A" italics="true" pageId="0" pageNumber="697">T. cantori</emphasis>
</taxonomicName>
,
<taxonomicName id="091A017AA4ABA9F9B56D9F94A7E61EDB" lsidName="T. cardamomensis" pageId="0" pageNumber="697" rank="species" species="cardamomensis">
<emphasis id="94BEE9F5A2A76746E09610DA6F73062B" italics="true" pageId="0" pageNumber="697">T. cardamomensis</emphasis>
</taxonomicName>
,
<taxonomicName id="998FA5A18C2E5A57E0D3F3B3A900A2D1" lsidName="T. caudornatus" pageId="0" pageNumber="697" rank="species" species="caudornatus">
<emphasis id="4FC45E2960F7DF779AE3BDED50442455" italics="true" pageId="0" pageNumber="697">T. caudornatus</emphasis>
</taxonomicName>
,
<taxonomicName id="801EF3B16DF42E313F24C255D60F9B4A" lsidName="T. davidi" pageId="0" pageNumber="697" rank="species" species="davidi">
<emphasis id="8F2A95BFD742306BA8150A05DD546B44" italics="true" pageId="0" pageNumber="697">T. davidi</emphasis>
</taxonomicName>
,
<taxonomicName id="04C8373C93032C3E4BB0052E9891E789" lsidName="T. erythrurus" pageId="0" pageNumber="697" rank="species" species="erythrurus">
<emphasis id="219A6A325978505A34E5CED680EA5EF1" italics="true" pageId="0" pageNumber="697">T. erythrurus</emphasis>
</taxonomicName>
,
<taxonomicName id="FC1040607933B9E7FFF5FBE75E944097" lsidName="T. fasciatus" pageId="0" pageNumber="697" rank="species" species="fasciatus">
<emphasis id="AC48AEE2039B2013E1186F5EBFBA13BB" italics="true" pageId="0" pageNumber="697">T. fasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="1304C00D674F06599648CA8016740374" lsidName="T. guoi" pageId="0" pageNumber="697" rank="species" species="guoi">
<emphasis id="E7B68F148CA20DC8BEE7D4001F5DEC50" italics="true" pageId="0" pageNumber="697">T. guoi</emphasis>
</taxonomicName>
,
<taxonomicName id="E848ED4016286696E97FF0B90F93FC4A" lsidName="T. honsonensis" pageId="0" pageNumber="697" rank="species" species="honsonensis">
<emphasis id="098614F88E4B88E74F6B1FABD1CB38A5" italics="true" pageId="0" pageNumber="697">T. honsonensis</emphasis>
</taxonomicName>
,
<taxonomicName id="721FF34BEEB42289F19C1A6AC013B29E" lsidName="T. insularis" pageId="0" pageNumber="697" rank="species" species="insularis">
<emphasis id="DB3B59A5C1FAC6E9FC039DFDA5D735E6" italics="true" pageId="0" pageNumber="697">T. insularis</emphasis>
</taxonomicName>
,
<taxonomicName id="3177A9155D253313DD76C8EEEF91B685" lsidName="T. labialis" pageId="0" pageNumber="697" rank="species" species="labialis">
<emphasis id="FE0CDE21B29359228CC46A5EBCA9EF69" italics="true" pageId="0" pageNumber="697">T. labialis</emphasis>
</taxonomicName>
,
<taxonomicName id="B1F4115BB90471264962557FC6DE6ED2" lsidName="T. macrops" pageId="0" pageNumber="697" rank="species" species="macrops">
<emphasis id="DBA48D60635013971BDD502C8433809F" italics="true" pageId="0" pageNumber="697">T. macrops</emphasis>
</taxonomicName>
,
<taxonomicName id="D6F323F4424CB5CB12DFCC9C44B8E849" lsidName="T. mutabilis" pageId="0" pageNumber="697" rank="species" species="mutabilis">
<emphasis id="C3F7E7456B9FC4CD060663D276B33804" italics="true" pageId="0" pageNumber="697">T. mutabilis</emphasis>
</taxonomicName>
,
<taxonomicName id="759E6010B3836B4049402266978725D3" lsidName="T. purpureomaculatus" pageId="0" pageNumber="697" rank="species" species="purpureomaculatus">
<emphasis id="B802D4A607FA3E9D57E5D4F41F261B17" italics="true" pageId="0" pageNumber="697">T. purpureomaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="BBEEA77EE8D30F02BB544008DC1671AF" lsidName="T. rubeus" pageId="0" pageNumber="697" rank="species" species="rubeus">
<emphasis id="619DA44F9FC33003387E798BE2C64B46" italics="true" pageId="0" pageNumber="697">T. rubeus</emphasis>
</taxonomicName>
,
<taxonomicName id="E4B9E2EA1C2524E2A2116A16C1492FA4" lsidName="T. salazar" pageId="0" pageNumber="697" rank="species" species="salazar">
<emphasis id="B83FB100401B4FF4D09692090F82CBE3" italics="true" pageId="0" pageNumber="697">T. salazar</emphasis>
</taxonomicName>
, and
<taxonomicName id="B8BAEF24363D1B16E8417FED90DFB031" lsidName="T. septentrionalis" pageId="0" pageNumber="697" rank="species" species="septentrionalis">
<emphasis id="D2CB30738BBD592E2F24467AAF7B2914" italics="true" pageId="0" pageNumber="697">T. septentrionalis</emphasis>
</taxonomicName>
by body coloration and pattern (reddish-brown bands on emerald-green background in the new species vs. uniform green or green coloration with no pattern, or pattern consisting of small brownish spots or speckles in other species, or straw-yellow background with irregular, dark-brown transverse body bands in
<taxonomicName id="5C9D535AB4D09F4C8F25941A1D2384C2" lsidName="T. honsonensis" pageId="0" pageNumber="697" rank="species" species="honsonensis">
<emphasis id="665AAB5C9FDC5709AAB75CA8A17A7FC8" italics="true" pageId="0" pageNumber="697">T. honsonensis</emphasis>
</taxonomicName>
). Moreover, by having 17 MSR, the new species can be further separated from
<taxonomicName id="EC084064A5A2E4BBB0265DB613ACD4E2" lsidName="T. albolabris" pageId="0" pageNumber="697" rank="species" species="albolabris">
<emphasis id="72049E6A6DF5C3493FDEA739766B66A7" italics="true" pageId="0" pageNumber="697">T. albolabris</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="61A458FE369C5B379B7C02CF35173468" lsidName="T. andersonii" pageId="0" pageNumber="697" rank="species" species="andersonii">
<emphasis id="97ABC44D0AB1F4F46C98ADD693DA3803" italics="true" pageId="0" pageNumber="697">T. andersonii</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="4C28C30C5AE9573F93D7BBC2F1BD7AB1" lsidName="T. cantori" pageId="0" pageNumber="697" rank="species" species="cantori">
<emphasis id="A1C388B471B9601F08B90E4A1B6A3CCB" italics="true" pageId="0" pageNumber="697">T. cantori</emphasis>
</taxonomicName>
(27, 29 or 31 MSR),
<taxonomicName id="D06501E005102B15A70BCFA3EB711ACC" lsidName="T. cardamomensis" pageId="0" pageNumber="697" rank="species" species="cardamomensis">
<emphasis id="B62024503E9626C29B5716A5174E3881" italics="true" pageId="0" pageNumber="697">T. cardamomensis</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="784B1976760C30C923B9699FDC367CAC" lsidName="T. caudornatus" pageId="0" pageNumber="697" rank="species" species="caudornatus">
<emphasis id="B5BBE1D7DA857A8BAD31185D6BC8D623" italics="true" pageId="0" pageNumber="697">T. caudornatus</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="4EF87ABB216845502BF9C75961FDD760" lsidName="T. davidi" pageId="0" pageNumber="697" rank="species" species="davidi">
<emphasis id="49800C049F534FAF981CF4710D9E7627" italics="true" pageId="0" pageNumber="697">T. davidi</emphasis>
</taxonomicName>
(21 or 23 MSR),
<taxonomicName id="C9C038CCA6563495E50640512F5E0C3C" lsidName="T. erythrurus" pageId="0" pageNumber="697" rank="species" species="erythrurus">
<emphasis id="1C16049E6F39E0BA536A608AFD95DD64" italics="true" pageId="0" pageNumber="697">T. erythrurus</emphasis>
</taxonomicName>
(23 rarely 21, 25 MSR),
<taxonomicName id="FE3407385DCD968AC3E3A64321500313" lsidName="T. fasciatus" pageId="0" pageNumber="697" rank="species" species="fasciatus">
<emphasis id="C0A8047241B68365BD59651F88EF450C" italics="true" pageId="0" pageNumber="697">T. fasciatus</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="A5348145FDA19D3B770B7E8BB9E50765" lsidName="T. guoi" pageId="0" pageNumber="697" rank="species" species="guoi">
<emphasis id="C6B4045B25634E65E17794954EDE4B97" italics="true" pageId="0" pageNumber="697">T. guoi</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="6AEF3A2DD09C53DEED1794878002A690" lsidName="T. honsonensis" pageId="0" pageNumber="697" rank="species" species="honsonensis">
<emphasis id="7ACD4F4536D71040BC0E7B608B0446BC" italics="true" pageId="0" pageNumber="697">T. honsonensis</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="0BF51667039ACEE9F86864F6C41D8F08" lsidName="T. insularis" pageId="0" pageNumber="697" rank="species" species="insularis">
<emphasis id="D46D3D290C78171BFA92C92924EB765C" italics="true" pageId="0" pageNumber="697">T. insularis</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="BF3C6DE087A3CD1F0FD5E25D49F2F07F" lsidName="T. labialis" pageId="0" pageNumber="697" rank="species" species="labialis">
<emphasis id="10B99713175A498D3905B2ED047E9878" italics="true" pageId="0" pageNumber="697">T. labialis</emphasis>
</taxonomicName>
(21 or 23 MSR),
<taxonomicName id="6D2F7746090149A5AC5112695741DD22" lsidName="T. macrops" pageId="0" pageNumber="697" rank="species" species="macrops">
<emphasis id="7A502424791610763C640E1F2B845E24" italics="true" pageId="0" pageNumber="697">T. macrops</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="33B27C437B5F7F16C3940450A7842BC3" lsidName="T. mutabilis" pageId="0" pageNumber="697" rank="species" species="mutabilis">
<emphasis id="DE11DF94CEA891C9AA11A4CB14194175" italics="true" pageId="0" pageNumber="697">T. mutabilis</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="B73056C5BAE3DB90DFCBA031C4AD5B0F" lsidName="T. purpureomaculatus" pageId="0" pageNumber="697" rank="species" species="purpureomaculatus">
<emphasis id="2CE557B37044BC63BC3E42B7D42B3C76" italics="true" pageId="0" pageNumber="697">T. purpureomaculatus</emphasis>
</taxonomicName>
(25 rarely 27, 29 MSR),
<taxonomicName id="C0A3B739C24265C8543BCF7B44DFF5BD" lsidName="T. rubeus" pageId="0" pageNumber="697" rank="species" species="rubeus">
<emphasis id="CA8DB5EAA636A4DF7355FB8885F1E11A" italics="true" pageId="0" pageNumber="697">T. rubeus</emphasis>
</taxonomicName>
(21 MSR),
<taxonomicName id="A8A2B3D3F5E2131F0EF17E4B6D26F7C3" lsidName="T. salazar" pageId="0" pageNumber="697" rank="species" species="salazar">
<emphasis id="0969EFC052B4B360C46029DF7564522D" italics="true" pageId="0" pageNumber="697">T. salazar</emphasis>
</taxonomicName>
(21 MSR), and
<taxonomicName id="0F6320D003ED8EB08767F5CEB933B487" lsidName="T. septentrionalis" pageId="0" pageNumber="697" rank="species" species="septentrionalis">
<emphasis id="5340A0DA90C2226C447AA77E92BC9D45" italics="true" pageId="0" pageNumber="697">T. septentrionalis</emphasis>
</taxonomicName>
(21 MSR) (see
<bibRefCitation id="7F100DFDC8014EB9D132BE6BA9EAF5B8" author="Gumprecht, A" journalOrPublisher="Hamadryad" pageId="0" pageNumber="697" refId="B25" refString="Gumprecht, A, Tillack, F, Orlov, NL, Captain, A, Ryabov, S, 2004. Asian Pit Vipers. Geitje Books Berlin, 368 pp." title="Asian Pit Vipers. Geitje Books Berlin, 368 pp." year="2004">Gumprecht et al. 2004</bibRefCitation>
;
<bibRefCitation id="61CF31B1D970F1634A300FA6C814BE10" DOI="https://doi.org/10.11646/­zootaxa.1715.1.4" author="Grismer, LL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="697" pagination="57 - 68" refId="B23" refString="Grismer, LL, Van Tri, N, Grismer, JL, 2008. A new species of insular pitviper of the genus Cryptelytrops (Squamata: Viperidae) from southern Vietnam. Zootaxa 68: 57 - 68, DOI: https://doi.org/10.11646/-zootaxa.1715.1.4" title="A new species of insular pitviper of the genus Cryptelytrops (Squamata: Viperidae) from southern Vietnam." url="https://doi.org/10.11646/­zootaxa.1715.1.4" volume="68" year="2008">Grismer et al. 2008</bibRefCitation>
;
<bibRefCitation id="B314445C8B421184F9AC1B95B429DF23" author="Malhotra, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="697" pagination="1 - 23" refId="B39" refString="Malhotra, A, Thorpe, RS, Mrinalini, , Stuart, BL, 2011. Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia. Zootaxa 2757: 1 - 23" title="Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia." volume="2757" year="2011">Malhotra et al. 2011</bibRefCitation>
;
<bibRefCitation id="9C705A7674610633F101175D860FC211" author="Chandramouli, SR" journalOrPublisher="Amphibian and Reptile Conservation" pageId="0" pageNumber="697" pagination="169 - 176" refId="B7" refString="Chandramouli, SR, Campbell, PD, Vogel, G, 2020. A new species of green pit viper of the genus Trimeresurus Lacepede, 1804 (Reptilia: Serpentes: Viperidae) from the Nicobar Archipelago, Indian Ocean. Amphibian and Reptile Conservation 14: 169 - 176" title="A new species of green pit viper of the genus Trimeresurus Lacepede, 1804 (Reptilia: Serpentes: Viperidae) from the Nicobar Archipelago, Indian Ocean." volume="14" year="2020">Chandramouli et al. 2020</bibRefCitation>
;
<bibRefCitation id="D3D3D111B59257FFB5F9530FDF105088" DOI="https://doi.org/10.11646/zootaxa.4768.1.7" author="Chen, Z" journalOrPublisher="Zootaxa" pageId="0" pageNumber="697" pagination="112 - 128" refId="B10" refString="Chen, Z, Yu, J, Vogel, G, Shi, S, Song, Z, Tang, Y, Yang, J, Ding, L, Chen, C, 2020. A new pit viper of the genus Trimeresurus (Lacepede, 1804) (Squamata: Viperidae) from Southwest China. Zootaxa 4768: 112 - 128, DOI: https://doi.org/10.11646/zootaxa.4768.1.7" title="A new pit viper of the genus Trimeresurus (Lacepede, 1804) (Squamata: Viperidae) from Southwest China." url="https://doi.org/10.11646/zootaxa.4768.1.7" volume="4768" year="2020">Chen et al. 2020</bibRefCitation>
,
<bibRefCitation id="AB83E3862C71EB01A1A4C24A2DF14D3A" DOI="https://doi.org/10.16373/j.cnki.ahr.200084" author="Chen, Z" journalOrPublisher="Asian Herpetological Research" pageId="0" pageNumber="697" pagination="167 - 177" refId="B9" refString="Chen, Z, Shi, S, Gao, J, Vogel, G, Song, Z, Ding, L, Dai, R, 2021. A new species of Trimeresurus Lacepede, 1804 (Squamata: Viperidae) from Southwestern China, Vietnam, Thailand and Myanmar. Asian Herpetological Research 12: 167 - 177, DOI: https://doi.org/10.16373/j.cnki.ahr.200084" title="A new species of Trimeresurus Lacepede, 1804 (Squamata: Viperidae) from Southwestern China, Vietnam, Thailand and Myanmar." url="https://doi.org/10.16373/j.cnki.ahr.200084" volume="12" year="2021">2021</bibRefCitation>
;
<bibRefCitation id="5DA3453A69B61FE5B878A0696ACBFDED" DOI="https://doi.org/10.3897/zse.96.48431" author="Mirza, ZA" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="697" pagination="123 - 138" refId="B41" refString="Mirza, ZA, Bhosale, HS, Phansalkar, PU, Sawant, M, Gowande, GG, Patel, H, 2020. A new species of green pit vipers of the genus Trimeresurus Lacepede, 1804 (Reptilia, Serpentes, Viperidae) from western Arunachal Pradesh, India. Zoosystematics and Evolution 96: 123 - 138, DOI: https://doi.org/10.3897/zse.96.48431" title="A new species of green pit vipers of the genus Trimeresurus Lacepede, 1804 (Reptilia, Serpentes, Viperidae) from western Arunachal Pradesh, India." url="https://doi.org/10.3897/zse.96.48431" volume="96" year="2020">Mirza et al. 2020</bibRefCitation>
; our data).
</paragraph>
</subSubSection>
</treatment>
</document>
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