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<document id="0919D6B59D3B617C994F973C422CF02E" ID-DOI="10.5281/zenodo.189074" ID-GBIF-Dataset="ab03e35b-2874-407a-9071-9d887e0322ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="189074" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461099212284" checkinUser="plazi" docAuthor="Tan, Heok Hui" docDate="2009" docId="5C0887D37F4FE25761C1F9945F0BF808" docLanguage="en" docName="zt02165p068.pdf" docOrigin="Zootaxa 2165" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Betta anabatoides Bleeker 1851" docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="A031FFAB7F4EE2526156FFB25B64FD1B" masterDocTitle="Redescription of Betta anabatoides Bleeker, and a new species of Betta from West Kalimantan, Borneo (Teleostei: Osphronemidae)" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698590524541" updateUser="plazi">
<mods:mods id="A9CDECB367139C751E3105E8FBED1D33" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="718BFBAD2643533C104373846D375C12">
<mods:title id="DB8EC5D5015C35A24BB2D279AAA3F860">Redescription of Betta anabatoides Bleeker, and a new species of Betta from West Kalimantan, Borneo (Teleostei: Osphronemidae)</mods:title>
</mods:titleInfo>
<mods:name id="A2741B81580ABAA6633E869249786CA0" type="personal">
<mods:role id="DDEA925B70FC5CEA82EC8E7AA7399245">
<mods:roleTerm id="2708CA932C25FBCC7954F3824292663D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12DD4E286FCCAF0F836CEBEF89DA5D24">Tan, Heok Hui</mods:namePart>
</mods:name>
<mods:typeOfResource id="15BEB8BE9C2667462E3CA2562C69D26B">text</mods:typeOfResource>
<mods:relatedItem id="CD56808C4E76097F00EEA2BF8D6A7604" type="host">
<mods:titleInfo id="C28F6B5721E850C8A912849A6FFEBC6B">
<mods:title id="EFD07491E9A6A498121886C455D83D15">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BB28AC0514E23F41284C791F528E9BA3">
<mods:date id="9F5D5EAA8AC973E106E77D7C475A105A">2009</mods:date>
<mods:detail id="720BAD89519B9F80F7E7DFE0940FE4C9" type="volume">
<mods:number id="102690CBC1695552FBD9845E4B694195">2165</mods:number>
</mods:detail>
<mods:extent id="3DAE2762D827A25286C12F0DE2037AD2" unit="page">
<mods:start id="B05B1DDD8217D898BEC138D820A0A075">59</mods:start>
<mods:end id="9085E95A4EC02F33D0321A33DB9378A4">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="36A08DA317EBA764DA5692E892643210">journal article</mods:classification>
<mods:identifier id="8F04A7F5956C14798ECC1C8EA40D0C8D" type="DOI">10.5281/zenodo.189074</mods:identifier>
<mods:identifier id="58E381F3B598D7B8DD71C3613569EE96" type="GBIF-Dataset">ab03e35b-2874-407a-9071-9d887e0322ba</mods:identifier>
<mods:identifier id="689502C3D7C1DE04ED59777F40AC5E7B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="30AE6E60B314B13B330366D186039E52" type="Zenodo-Dep">189074</mods:identifier>
</mods:mods>
<treatment id="5C0887D37F4FE25761C1F9945F0BF808" ID-DOI="http://doi.org/10.5281/zenodo.6220714" ID-GBIF-Taxon="119621261" ID-Zenodo-Dep="6220714" LSID="urn:lsid:plazi:treatment:5C0887D37F4FE25761C1F9945F0BF808" httpUri="http://treatment.plazi.org/id/5C0887D37F4FE25761C1F9945F0BF808" lastPageId="5" lastPageNumber="64" pageId="1" pageNumber="60">
<subSubSection id="9CBB654E7F4FE25361C1F9945A73FB78" pageId="1" pageNumber="60" type="nomenclature">
<paragraph id="D41E36C57F4FE25361C1F9945978FB5B" blockId="1.[151,540,1574,1635]" box="[151,540,1574,1600]" pageId="1" pageNumber="60">
<heading id="8F5681A97F4FE25361C1F9945978FB5B" bold="true" box="[151,540,1574,1600]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
<taxonomicName id="13A14D467F4FE25361C1F9945978FB5B" ID-CoL="LNYM" authority="Bleeker, 1851" authorityName="Bleeker" authorityYear="1851" box="[151,540,1574,1600]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4FE25361C1F9945978FB5B" bold="true" box="[151,540,1574,1600]" pageId="1" pageNumber="60">
<emphasis id="E6D5EAD77F4FE25361C1F9945A0AFB5B" bold="true" box="[151,366,1574,1600]" italics="true" pageId="1" pageNumber="60">Betta anabatoides</emphasis>
<bibRefCitation id="B0304B347F4FE2536023F9945978FB5B" author="Bleeker" box="[373,540,1574,1600]" pageId="1" pageNumber="68" refString="Bleeker, P. (1851) Nieuwe bijrage tot de kennis der ichthyologische fauna van Borneo. Nat. Tijdschr. Ned. Indie, 1, 259 - 275." type="journal article" year="1851">Bleeker, 1851</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D41E36C57F4FE25361C1F9FB5A73FB78" blockId="1.[151,540,1574,1635]" box="[151,279,1609,1635]" pageId="1" pageNumber="60">
(
<figureCitation id="4C9A2A407F4FE25361C9F9FB5A6AFB78" box="[159,270,1609,1635]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,255,1067,1091]" captionStartId-1="2.[151,255,1805,1829]" captionTargetBox-0="[151,1434,622,1040]" captionTargetBox-1="[151,1434,1144,1780]" captionTargetId-0="figure@2.[151,1436,622,1041]" captionTargetId-1="figure@2.[151,1436,1144,1781]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURE 1. Betta anabatoides, ZRC 40971, 47.2 mm SL, neotype, Banjarmasin (right side, laterally inverted)." captionText-1="FIGURE 2. Betta anabatoides, not preserved, ca. 35 mm SL, Central Kalimantan." httpUri-0="https://zenodo.org/record/189075/files/figure.png" httpUri-1="https://zenodo.org/record/189076/files/figure.png" pageId="1" pageNumber="60">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9CBB654E7F4FE25361C1F9255869FBEB" pageId="1" pageNumber="60" type="reference_group">
<paragraph id="D41E36C57F4FE25361C1F925590DFBD4" blockId="1.[151,1436,1686,1776]" pageId="1" pageNumber="60">
<treatmentCitationGroup id="F4B111EB7F4FE25361C1F925590DFBD4" pageId="1" pageNumber="60">
<taxonomicName id="13A14D467F4FE25361C1F9255954FBB6" ID-CoL="LNYM" authority="Bleeker, 1851: 269" authorityName="Bleeker" authorityPageNumber="269" authorityYear="1851" box="[151,560,1686,1710]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4FE25361C1F9255A3DFBB5" box="[151,345,1687,1710]" italics="true" pageId="1" pageNumber="60">Betta anabatoides</emphasis>
<treatmentCitation id="550010D47F4FE2536034F9245954FBB6" author="Bleeker" box="[354,560,1686,1709]" page="269" pageId="1" pageNumber="60" year="1851">
<bibRefCitation id="B0304B347F4FE2536034F9245A98FBB6" author="Bleeker" box="[354,508,1686,1709]" pageId="1" pageNumber="68" refString="Bleeker, P. (1851) Nieuwe bijrage tot de kennis der ichthyologische fauna van Borneo. Nat. Tijdschr. Ned. Indie, 1, 259 - 275." type="journal article" year="1851">Bleeker, 1851</bibRefCitation>
: 269
</treatmentCitation>
</taxonomicName>
— Weber &amp; de
<treatmentCitation id="550010D47F4FE25363B1F92458A5FBB6" author="Beaufort" box="[743,961,1686,1709]" page="357" pageId="1" pageNumber="60" year="1922">
<bibRefCitation id="B0304B347F4FE25363B1F92458E9FBB6" author="Beaufort" box="[743,909,1686,1709]" pageId="1" pageNumber="68" refString="Weber, M &amp; Beaufort, L. F. de (1922) The fishes of the Indo - Australian archipelago. 4. Brill, Leiden, xiii + 410 pp." type="book" year="1922">Beaufort, 1922</bibRefCitation>
: 357
</treatmentCitation>
(part);
<treatmentCitation id="550010D47F4FE2536542F9245FBFFBB6" author="Vierke" box="[1044,1243,1686,1709]" page="390" pageId="1" pageNumber="60" year="1979">
<bibRefCitation id="B0304B347F4FE2536542F9245FC0FBB6" author="Vierke" box="[1044,1188,1686,1709]" pageId="1" pageNumber="68" refString="Vierke, J. (1979) Betta anabatoides und Betta foerschi spec. nov., zwei Kampffische aus Borneo. Zeitschrift fur Aquarien-und Terrarienfreunde, 13, 386 - 390." type="journal article" year="1979">Vierke, 1979</bibRefCitation>
: 390
</treatmentCitation>
;
<treatmentCitation id="550010D47F4FE25365B1F9245EFCFBB6" author="Linke" box="[1255,1432,1686,1709]" page="26" pageId="1" pageNumber="60" year="1991">
<bibRefCitation id="B0304B347F4FE25365B1F9245E0AFBB6" author="Linke" box="[1255,1390,1686,1709]" pageId="1" pageNumber="68" refString="Linke, H. (1991) Labyrinth fish. The Bubble-nest-builders. Their identification, care and breeding. Tetra-Press, Melle, Germany, 174 pp." type="book" year="1991">Linke, 1991</bibRefCitation>
: 26
</treatmentCitation>
;
<treatmentCitation id="550010D47F4FE2536191F90A5AB2FBD4" author="Kottelat" box="[199,470,1719,1744]" page="222" pageId="1" pageNumber="60" year="1993">
<bibRefCitation id="B0304B347F4FE2536191F90A5AC0FBD4" author="Kottelat" box="[199,420,1719,1744]" pageId="1" pageNumber="68" refString="Kottelat, M., Whitten, A. J., Kartikasari, S. N. &amp; Wirjoatmodjo, S. (1993) Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, 259 pp., 84 pls." type="book" year="1993">
Kottelat
<emphasis id="E6D5EAD77F4FE2536075F9055A3AFBCB" box="[291,350,1719,1744]" italics="true" pageId="1" pageNumber="60">et al.</emphasis>
, 1993
</bibRefCitation>
: 222
</treatmentCitation>
(part), pl. 75.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D41E36C57F4FE25361C1F96B5869FBEB" blockId="1.[151,1436,1686,1776]" box="[151,781,1753,1776]" pageId="1" pageNumber="60">
<treatmentCitationGroup id="F4B111EB7F4FE25361C1F96B5869FBEB" box="[151,781,1753,1776]" pageId="1" pageNumber="60">
<taxonomicName id="13A14D467F4FE25361C1F96B5A6FFBEB" ID-CoL="LP2C" box="[151,267,1753,1776]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="picta">
<emphasis id="E6D5EAD77F4FE25361C1F96B5A6FFBEB" box="[151,267,1753,1776]" italics="true" pageId="1" pageNumber="60">Betta picta</emphasis>
</taxonomicName>
(non Valenciennes) —
<treatmentCitation id="550010D47F4FE2536351F96B59A0FBEB" author="Bleeker" box="[519,708,1753,1776]" page="26" pageId="1" pageNumber="60" year="1879">
<bibRefCitation id="B0304B347F4FE2536351F96B59FBFBEB" author="Bleeker" box="[519,671,1753,1776]" pageId="1" pageNumber="68" refString="Bleeker, P. (1879) Memoire sur les Poissons a Pharyngiens Labyrinthiformes de l'Inde Archipelagique. Natuurk. Verh. der Koninkl. Akademie, vol. 9, 1 - 56." type="journal article" year="1879">Bleeker, 1879</bibRefCitation>
: 26
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9CBB654E7F4FE25061C1F8AD58BEFF2F" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60" type="materials_examined">
<paragraph id="D41E36C57F4FE25361C1F8AD58D0FA7B" blockId="1.[151,1436,1823,2004]" pageId="1" pageNumber="60">
<emphasis id="E6D5EAD77F4FE25361C1F8AD5AE3FA22" bold="true" box="[151,391,1823,1849]" pageId="1" pageNumber="60">Material examined</emphasis>
.
<emphasis id="E6D5EAD77F4FE25360C0F8AD5961FA22" bold="true" box="[406,517,1823,1849]" pageId="1" pageNumber="60">
<typeStatus id="0B1A88677F4FE25360C0F8AD5964FA22" box="[406,512,1823,1849]" pageId="1" pageNumber="60" type="neotype">Neotype</typeStatus>
.
</emphasis>
<collectionCode id="B2B0AE007F4FE2536358F8925923FA21" box="[526,583,1824,1850]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="60">ZRC</collectionCode>
40971,
<quantity id="13599B207F4FE25363FFF8925877FA21" box="[681,787,1824,1850]" metricMagnitude="-2" metricUnit="m" metricValue="4.72" pageId="1" pageNumber="60" unit="mm" value="47.2">47.2 mm</quantity>
<collectionCode id="B2B0AE007F4FE253624DF8925825FA21" box="[795,833,1824,1850]" pageId="1" pageNumber="60">SL</collectionCode>
; South Kalimantan: Banjarmasin, stream at km 55 on Rantau–Martapura road; H. H. Ng &amp;
<collectingCountry id="ACB676557F4FE2536325F8F459EFFA7B" box="[627,651,1862,1888]" name="Iceland" pageId="1" pageNumber="60">O</collectingCountry>
. K. S. Chia,
<date id="A01F10057F4FE2536249F8F458D4FA7B" box="[799,944,1862,1888]" pageId="1" pageNumber="60" value="1996-06-06">6 June 1996</date>
.
</paragraph>
<paragraph id="D41E36C57F4FE2506190F8DF58BEFF2F" blockId="1.[151,1436,1823,2004]" lastBlockId="2.[151,1436,152,564]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
<emphasis id="E6D5EAD77F4FE2536190F8DF5AEEFA9C" bold="true" box="[198,394,1901,1927]" pageId="1" pageNumber="60">Other material.</emphasis>
South Kalimantan —
<collectionCode id="B2B0AE007F4FE25363C5F8DF59A8FA9C" box="[659,716,1901,1927]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="60">ZRC</collectionCode>
43710, 1 ex.,
<quantity id="13599B207F4FE2536223F8DF58BAFA9C" box="[885,990,1901,1927]" metricMagnitude="-2" metricUnit="m" metricValue="5.529999999999999" pageId="1" pageNumber="60" unit="mm" value="55.3">55.3 mm</quantity>
<collectionCode id="B2B0AE007F4FE25362B3F8DF5F6FFA9C" box="[997,1035,1901,1927]" pageId="1" pageNumber="60">SL</collectionCode>
; same locality data as
<typeStatus id="0B1A88677F4FE2536445F8DF5E17FA9C" box="[1299,1395,1901,1927]" pageId="1" pageNumber="60" type="neotype">neotype</typeStatus>
. — CMK 16740, 1 ex.,
<quantity id="13599B207F4FE25360D1F8265A96FAB5" box="[391,498,1940,1966]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="1" pageNumber="60" unit="mm" value="37.4">37.4 mm</quantity>
<collectionCode id="B2B0AE007F4FE25360ACF8265944FAB5" box="[506,544,1940,1966]" pageId="1" pageNumber="60">SL</collectionCode>
; Barito River basin, area of Tamiyang Layang (
<geoCoordinate id="B19550027F4FE2536509F8265FCAFAB5" box="[1119,1198,1940,1966]" direction="south" orientation="latitude" pageId="1" pageNumber="60" precision="925" value="-2.0166667">02°1'S</geoCoordinate>
<geoCoordinate id="B19550027F4FE25365E0F8265E7CFAB5" box="[1206,1304,1940,1966]" direction="east" orientation="longitude" pageId="1" pageNumber="60" precision="925" value="115.11667">115°7'E</geoCoordinate>
); T. Idei,
<date id="A01F10057F4FE25364DBF8265A6EFACF" pageId="1" pageNumber="60" value="2000-09-07">7 Sep 2000</date>
. —
<collectionCode id="B2B0AE007F4FE253606DF8085A10FACF" box="[315,372,1978,2004]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="60">ZRC</collectionCode>
51536, 9 ex., 33.0–
<quantity id="13599B207F4FE2536334F80859A8FACF" box="[610,716,1978,2004]" metricMagnitude="-2" metricUnit="m" metricValue="8.22" pageId="1" pageNumber="60" unit="mm" value="82.2">82.2 mm</quantity>
<collectionCode id="B2B0AE007F4FE2536382F808599EFACF" box="[724,762,1978,2004]" pageId="1" pageNumber="60">SL</collectionCode>
; Tamiyang Layang area; T. Idei, 2004. —
<collectionCode id="B2B0AE007F4FE25365A4F8085E4FFACF" box="[1266,1323,1978,2004]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="60">ZRC</collectionCode>
51533, 1 ex.,
<quantity id="13599B207F4CE250619CFF2A5A50FDA9" box="[202,308,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="4.89" pageId="2" pageNumber="61" unit="mm" value="48.9">48.9 mm</quantity>
<collectionCode id="B2B0AE007F4CE250606AFF2A5A06FDA9" box="[316,354,152,178]" pageId="2" pageNumber="61">SL</collectionCode>
; Relaihari, Gunung Raya, South of Banjarmasin; T. Idei, 2004. —
<collectionCode id="B2B0AE007F4CE25065D4FF2A5FD8FDA9" box="[1154,1212,152,178]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="61">ZRC</collectionCode>
51535, 1 ex.,
<quantity id="13599B207F4CE250643EFF2A5BA1FDC3" metricMagnitude="-2" metricUnit="m" metricValue="5.24" pageId="2" pageNumber="61" unit="mm" value="52.4">52.4 mm</quantity>
<collectionCode id="B2B0AE007F4CE250619BFF0C5B97FDC3" box="[205,243,190,216]" pageId="2" pageNumber="61">SL</collectionCode>
; Pelaihari, Gunung Raya (
<geoCoordinate id="B19550027F4CE2506364FF0C59DAFDC3" box="[562,702,190,216]" direction="south" orientation="latitude" pageId="2" pageNumber="61" precision="15" value="-3.6686113">03º40’07”S</geoCoordinate>
<geoCoordinate id="B19550027F4CE2506390FF0C5800FDC3" box="[710,868,190,216]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="15" value="114.676384">114º40’35”E</geoCoordinate>
); T. Idei, 2004. —
<collectionCode id="B2B0AE007F4CE250651CFF0C5FE0FDC3" box="[1098,1156,190,216]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="61">ZRC</collectionCode>
51534, 1 ex.,
<quantity id="13599B207F4CE2506467FF0C5EF8FDC3" box="[1329,1436,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="5.34" pageId="2" pageNumber="61" unit="mm" value="53.4">53.4 mm</quantity>
<collectionCode id="B2B0AE007F4CE25061C1FF575BDAFDE4" box="[151,190,229,255]" pageId="2" pageNumber="61">SL</collectionCode>
; Satai, East Point; T. Idei, 2004. Central Kalimantan — CMK 16707, 1 ex.,
<quantity id="13599B207F4CE2506508FF575FADFDE4" box="[1118,1225,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="2" pageNumber="61" unit="mm" value="33.3">33.3 mm</quantity>
<collectionCode id="B2B0AE007F4CE2506584FF575F9CFDE4" box="[1234,1272,229,255]" pageId="2" pageNumber="61">SL</collectionCode>
; Sampit river basin,
<quantity id="13599B207F4CE25061B3FEBE5A55FC3D" box="[229,305,268,294]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="61" unit="km" value="45.0">45 km</quantity>
west of Sampit (
<geoCoordinate id="B19550027F4CE25060A9FEBE593AFC3D" box="[511,606,268,294]" direction="south" orientation="latitude" pageId="2" pageNumber="61" precision="925" value="-2.5">02°30'S</geoCoordinate>
<geoCoordinate id="B19550027F4CE2506330FEBE59B3FC3D" box="[614,727,268,294]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="925" value="112.63333">112°38'E</geoCoordinate>
); T. Idei,
<date id="A01F10057F4CE250621DFEBE58BFFC3D" box="[843,987,268,294]" pageId="2" pageNumber="61" value="2000-07-20">20 Jul 2000</date>
. — CMK 16696, 1 ex.,
<quantity id="13599B207F4CE25065A8FEBE5E0DFC3D" box="[1278,1385,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="5.59" pageId="2" pageNumber="61" unit="mm" value="55.9">55.9 mm</quantity>
<collectionCode id="B2B0AE007F4CE2506427FEBE5EF3FC3D" box="[1393,1431,268,294]" pageId="2" pageNumber="61">SL</collectionCode>
; Sampit river basin,
<quantity id="13599B207F4CE250602CFE805AA0FC57" box="[378,452,306,332]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="61" unit="km" value="15.0">15 km</quantity>
north of Parenggean (
<geoCoordinate id="B19550027F4CE2506390FE805846FC57" box="[710,802,306,332]" direction="south" orientation="latitude" pageId="2" pageNumber="61" precision="925" value="-1.9333334">01°56'S</geoCoordinate>
<geoCoordinate id="B19550027F4CE250627FFE8058FCFC57" box="[809,920,306,332]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="925" value="112.71667">112°43'E</geoCoordinate>
); T. Idei,
<date id="A01F10057F4CE2506553FE805FF3FC57" box="[1029,1175,306,332]" pageId="2" pageNumber="61" value="2000-06-16">16 Jun 2000</date>
. —
<collectionCode id="B2B0AE007F4CE2506593FE805F9AFC57" box="[1221,1278,306,332]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="61">ZRC</collectionCode>
51537, 8 ex., 34.8–60.0 mm
<collectionCode id="B2B0AE007F4CE2506019FEEB5A12FC68" box="[335,374,345,371]" pageId="2" pageNumber="61">SL</collectionCode>
; Sampit river basin, southeast of Kuala Kuayang; T. Idei, 2004. —
<collectionCode id="B2B0AE007F4CE25065FAFEEB5F82FC68" box="[1196,1254,345,371]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="61">ZRC</collectionCode>
51538, 10 ex., 47.0–
<quantity id="13599B207F4CE250618AFE325A2DFC81" box="[220,329,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="7.02" pageId="2" pageNumber="61" unit="mm" value="70.2">70.2 mm</quantity>
<collectionCode id="B2B0AE007F4CE2506004FE325A1DFC81" box="[338,377,384,410]" pageId="2" pageNumber="61">SL</collectionCode>
; Plantalang Hulu (
<geoCoordinate id="B19550027F4CE2506337FE325994FC81" box="[609,752,384,410]" direction="south" orientation="latitude" pageId="2" pageNumber="61" precision="15" value="-2.0277777">02º01’40”S</geoCoordinate>
<geoCoordinate id="B19550027F4CE25063AFFE3258F8FC81" box="[761,924,384,410]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="15" value="112.996666">112º59’48”E</geoCoordinate>
); T. Idei, 2004. —
<collectionCode id="B2B0AE007F4CE25065DFFE325FA0FC81" box="[1161,1220,384,410]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="61">ZRC</collectionCode>
22842–43, 2 ex.,
<quantity id="13599B207F4CE25061C1FE145A23FCDB" box="[151,327,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" metricValueMax="5.59" metricValueMin="4.67" pageId="2" pageNumber="61" unit="mm" value="51.3" valueMax="55.9" valueMin="46.7">46.7–55.9 mm</quantity>
<collectionCode id="B2B0AE007F4CE2506019FE145A11FCDB" box="[335,373,422,448]" pageId="2" pageNumber="61">SL</collectionCode>
,
<collectionCode id="B2B0AE007F4CE25060D5FE145AD9FCDB" LSID="urn:lsid:biocol.org:col:34700" box="[387,445,422,448]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="2" pageNumber="61">CAS</collectionCode>
96958, 4 ex.,
<quantity id="13599B207F4CE2506338FE145879FCDB" box="[622,797,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="4.755" metricValueMax="5.9399999999999995" metricValueMin="3.57" pageId="2" pageNumber="61" unit="mm" value="47.55" valueMax="59.4" valueMin="35.7">35.7–59.4 mm</quantity>
<collectionCode id="B2B0AE007F4CE2506270FE145829FCDB" box="[806,845,422,448]" pageId="2" pageNumber="61">SL</collectionCode>
; Mentaya basin, Sungai Ramban,
<quantity id="13599B207F4CE25065B8FE145E5EFCDB" box="[1262,1338,422,448]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="61" unit="km" value="22.0">22 km</quantity>
west of Sampit on road to Pembuanghulu; T. R. Roberts,
<date id="A01F10057F4CE25063A5FE7F58EFFCFC" box="[755,907,461,487]" pageId="2" pageNumber="61" value="1992-06-11">11 Jun 1992</date>
. —
<collectionCode id="B2B0AE007F4CE25062E8FE7F589CFCFC" box="[958,1016,461,487]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="61">ZRC</collectionCode>
51539, 45 ex.,
<quantity id="13599B207F4CE25065EEFE7F5E0CFCFC" box="[1208,1384,461,487]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="6.56" metricValueMin="4.04" pageId="2" pageNumber="61" unit="mm" value="53.0" valueMax="65.6" valueMin="40.4">40.4–65.6 mm</quantity>
<collectionCode id="B2B0AE007F4CE2506426FE7F5EF3FCFC" box="[1392,1431,461,487]" pageId="2" pageNumber="61">SL</collectionCode>
; Katingan river basin, east of Kasongan (
<geoCoordinate id="B19550027F4CE2506321FE4659BCFF15" box="[631,728,500,526]" direction="south" orientation="latitude" pageId="2" pageNumber="61" precision="925" value="-1.9">01°54’S</geoCoordinate>
<geoCoordinate id="B19550027F4CE25063B7FE465831FF15" box="[737,853,500,526]" direction="east" orientation="longitude" pageId="2" pageNumber="61" precision="925" value="113.416664">113°25’E</geoCoordinate>
); T. Idei, 2004. — CMK 16764, 2 ex., 45.5–48.0 mm
<collectionCode id="B2B0AE007F4CE250619AFDA85B95FF2F" box="[204,241,538,564]" pageId="2" pageNumber="61">SL</collectionCode>
; Barito river basin:
<quantity id="13599B207F4CE250608CFDA85940FF2F" box="[474,548,538,564]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="61" unit="km" value="15.0">15 km</quantity>
east of Buntok; T. Idei,
<date id="A01F10057F4CE2506216FDA858B3FF2F" box="[832,983,538,564]" pageId="2" pageNumber="61" value="2000-09-13">13 Sep 2000</date>
.
</paragraph>
</subSubSection>
<caption id="80DE664D7F4CE25061C1FB995E5EF959" box="[151,1338,1067,1091]" httpUri="https://zenodo.org/record/189075/files/figure.png" pageId="2" pageNumber="61" targetBox="[151,1434,622,1040]" targetPageId="2">
<paragraph id="D41E36C57F4CE25061C1FB995E5EF959" blockId="2.[151,1338,1067,1091]" box="[151,1338,1067,1091]" pageId="2" pageNumber="61">
<emphasis id="E6D5EAD77F4CE25061C1FB995A7DF958" bold="true" box="[151,281,1067,1091]" pageId="2" pageNumber="61">FIGURE 1.</emphasis>
<taxonomicName id="13A14D467F4CE2506076FB995A85F959" box="[288,481,1067,1090]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4CE2506076FB995A85F959" box="[288,481,1067,1090]" italics="true" pageId="2" pageNumber="61">Betta anabatoides</emphasis>
</taxonomicName>
, ZRC 40971, 47.2 mm SL, neotype, Banjarmasin (right side, laterally inverted).
</paragraph>
</caption>
<caption id="80DE664D7F4CE25061C1F8BF5F6CFA3F" box="[151,1032,1805,1829]" httpUri="https://zenodo.org/record/189076/files/figure.png" pageId="2" pageNumber="61" targetBox="[151,1434,1144,1780]" targetPageId="2">
<paragraph id="D41E36C57F4CE25061C1F8BF5F6CFA3F" blockId="2.[151,1032,1805,1829]" box="[151,1032,1805,1829]" pageId="2" pageNumber="61">
<emphasis id="E6D5EAD77F4CE25061C1F8BF5B9BFA3E" bold="true" box="[151,255,1805,1829]" pageId="2" pageNumber="61">FIGURE</emphasis>
2.
<taxonomicName id="13A14D467F4CE2506076F8BC5A85FA3E" box="[288,481,1806,1829]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4CE2506076F8BC5A85FA3E" box="[288,481,1806,1829]" italics="true" pageId="2" pageNumber="61">Betta anabatoides</emphasis>
</taxonomicName>
, not preserved, ca. 35 mm SL, Central Kalimantan.
</paragraph>
</caption>
<subSubSection id="9CBB654E7F4CE2516190F8E15B87FDC3" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61" type="diagnosis">
<paragraph id="D41E36C57F4CE2516190F8E15B87FDC3" blockId="2.[151,1436,1875,2017]" lastBlockId="3.[151,1437,152,448]" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
<emphasis id="E6D5EAD77F4CE2506190F8E15A22FA76" bold="true" box="[198,326,1875,1901]" pageId="2" pageNumber="61">Diagnosis.</emphasis>
<taxonomicName id="13A14D467F4CE2506007F8E1594DFA77" box="[337,553,1875,1900]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4CE2506007F8E1594DFA77" box="[337,553,1875,1900]" italics="true" pageId="2" pageNumber="61">Betta anabatoides</emphasis>
</taxonomicName>
is unique in having the following combination of characters: a thick-set body (body depth 30.5–33.7 % SL) with the anterior part of the body reminiscent of the taxon
<taxonomicName id="13A14D467F4CE25065AAF8C85E30FA88" box="[1276,1364,1914,1939]" class="Actinopterygii" family="Anabantidae" genus="Anabas" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="E6D5EAD77F4CE25065AAF8C85E30FA88" box="[1276,1364,1914,1939]" italics="true" pageId="2" pageNumber="61">Anabas</emphasis>
</taxonomicName>
; head devoid of pattern except for a preorbital stripe that runs from the black lower lip to the eye, and continuous with a postorbital stripe to the posterior edge of the opercle; in living specimens, the opercle with faint gold iridescent scales; and some median caudal rays projecting beyond the fin margin (adapted from
<bibRefCitation id="B0304B347F4DE25165A3FF2A5BB2FDC3" author="Kottelat" pageId="3" pageNumber="68" refString="Kottelat, M., Whitten, A. J., Kartikasari, S. N. &amp; Wirjoatmodjo, S. (1993) Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, 259 pp., 84 pls." type="book" year="1993">
Kottelat
<emphasis id="E6D5EAD77F4DE251640CFF2A5EF1FDAA" box="[1370,1429,152,177]" italics="true" pageId="3" pageNumber="62">et al.</emphasis>
, 1993
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9CBB654E7F4DE2576190FF575A1EFC81" lastPageId="5" lastPageNumber="64" pageId="3" pageNumber="62" type="description">
<paragraph id="D41E36C57F4DE2516190FF575E1FFCDB" blockId="3.[151,1437,152,448]" pageId="3" pageNumber="62">
<emphasis id="E6D5EAD77F4DE2516190FF575A3BFDE4" bold="true" box="[198,351,229,255]" pageId="3" pageNumber="62">Description.</emphasis>
General appearance as illustrated in
<figureCitation id="4C9A2A407F4DE2516271FF5758FFFDE4" box="[807,923,229,255]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,255,1067,1091]" captionStartId-1="2.[151,255,1805,1829]" captionTargetBox-0="[151,1434,622,1040]" captionTargetBox-1="[151,1434,1144,1780]" captionTargetId-0="figure@2.[151,1436,622,1041]" captionTargetId-1="figure@2.[151,1436,1144,1781]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURE 1. Betta anabatoides, ZRC 40971, 47.2 mm SL, neotype, Banjarmasin (right side, laterally inverted)." captionText-1="FIGURE 2. Betta anabatoides, not preserved, ca. 35 mm SL, Central Kalimantan." httpUri-0="https://zenodo.org/record/189075/files/figure.png" httpUri-1="https://zenodo.org/record/189076/files/figure.png" pageId="3" pageNumber="62">Figs. 1–2</figureCitation>
. Meristic and morphometric data listed in Table 1. Head broadly pointed (anterior body reminiscent of
<taxonomicName id="13A14D467F4DE251620AFEBE58D0FC3E" box="[860,948,268,293]" class="Actinopterygii" family="Anabantidae" genus="Anabas" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="E6D5EAD77F4DE251620AFEBE58D0FC3E" box="[860,948,268,293]" italics="true" pageId="3" pageNumber="62">Anabas</emphasis>
</taxonomicName>
:
<figureCitation id="4C9A2A407F4DE2516295FEBE5F61FC3D" box="[963,1029,268,294]" captionStart="FIGURE 1" captionStartId="2.[151,255,1067,1091]" captionTargetBox="[151,1434,622,1040]" captionTargetId="figure@2.[151,1436,622,1041]" captionTargetPageId="2" captionText="FIGURE 1. Betta anabatoides, ZRC 40971, 47.2 mm SL, neotype, Banjarmasin (right side, laterally inverted)." httpUri="https://zenodo.org/record/189075/files/figure.png" pageId="3" pageNumber="62">Fig. 1</figureCitation>
), relatively long (HL 32.8–34.9 % SL); body thick-set (body depth 30.5–33.7 % SL); pectoral fin rounded; pelvic fin filamentous; dorsal fin situated far back (predorsal length 66.8–69.7 % SL); dorsal and anal fins pointed, with elongated posterior rays; caudal peduncle deep (caudal peduncle depth 19.8–22.1 % SL); caudal fin lanceolate, some median rays projecting beyond fin margin. Large male specimens with longer unpaired fins and darker body coloration.
</paragraph>
<paragraph id="D41E36C57F4DE25161C1FE5D5F0EFAA9" pageId="3" pageNumber="62">
<table id="A6A1C4657F4D1DAD61C1FE5D5E3DFAA9" box="[151,1369,495,1970]" gridcols="4" gridrows="34" pageId="3" pageNumber="62">
<tr id="6A9134877F4D1DAD61C1FE5D5E3DFF1C" box="[151,1369,495,519]" gridrow="0" pageId="3" pageNumber="62" rowspan-2="1" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FE5D5950FF1C" box="[151,564,495,519]" gridcol="0" gridrow="0" pageId="3" pageNumber="62">
<emphasis id="E6D5EAD77F4DE25161C1FE5D5A6CFF1C" bold="true" box="[151,264,495,519]" pageId="3" pageNumber="62">TABLE 1.</emphasis>
Meristic and morphometric
</th>
<th id="29405DFB7F4D1DAD636DFE5D5880FF1C" box="[571,996,495,519]" gridcol="1" gridrow="0" pageId="3" pageNumber="62">
data of
<taxonomicName id="13A14D467F4DE25163DDFE42582FFF1C" box="[651,843,496,519]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4DE25163DDFE42582FFF1C" box="[651,843,496,519]" italics="true" pageId="3" pageNumber="62">Betta anabatoides</emphasis>
</taxonomicName>
and
<taxonomicName id="13A14D467F4DE25162D6FE4258B9FF1C" box="[896,989,496,519]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="midas">
<emphasis id="E6D5EAD77F4DE25162D6FE4258B9FF1C" box="[896,989,496,519]" italics="true" pageId="3" pageNumber="62">B. midas</emphasis>
</taxonomicName>
.
</th>
</tr>
<tr id="6A9134877F4D1DAD61C1FD905E3DFF22" box="[151,1369,546,569]" gridrow="1" pageId="3" pageNumber="62" rowspan-0="1" rowspan-3="1">
<td id="29405DFB7F4D1DAD636DFD905880FF22" box="[571,996,546,569]" gridcol="1" gridrow="1" pageId="3" pageNumber="62">
<taxonomicName id="13A14D467F4DE251636AFD9059B8FF22" box="[572,732,546,569]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4DE251636AFD9059B8FF22" box="[572,732,546,569]" italics="true" pageId="3" pageNumber="62">B. anabatoides</emphasis>
</taxonomicName>
</td>
<td id="29405DFB7F4D1DAD6556FD905E4BFF22" box="[1024,1327,546,569]" gridcol="2" gridrow="1" pageId="3" pageNumber="62">
<taxonomicName id="13A14D467F4DE2516556FD905F39FF22" box="[1024,1117,546,569]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="midas">
<emphasis id="E6D5EAD77F4DE2516556FD905F39FF22" box="[1024,1117,546,569]" italics="true" pageId="3" pageNumber="62">B. midas</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FDFF5E3DFF7F" box="[151,1369,589,612]" gridrow="2" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FDFF5950FF7F" box="[151,564,589,612]" gridcol="0" gridrow="2" pageId="3" pageNumber="62">sample size</th>
<td id="29405DFB7F4D1DAD636DFDFF5880FF7F" box="[571,996,589,612]" gridcol="1" gridrow="2" pageId="3" pageNumber="62">7</td>
<td id="29405DFB7F4D1DAD6556FDFF5E4BFF7F" box="[1024,1327,589,612]" gridcol="2" gridrow="2" pageId="3" pageNumber="62">7</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FDCA5E3DFF94" box="[151,1369,632,655]" gridrow="3" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FDCA5950FF94" box="[151,564,632,655]" gridcol="0" gridrow="3" pageId="3" pageNumber="62">SL (mm)</th>
<td id="29405DFB7F4D1DAD636DFDCA5880FF94" box="[571,996,632,655]" gridcol="1" gridrow="3" pageId="3" pageNumber="62">46.7–59.4</td>
<td id="29405DFB7F4D1DAD6556FDCA5E4BFF94" box="[1024,1327,632,655]" gridcol="2" gridrow="3" pageId="3" pageNumber="62">41.6–58.0</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FD105E3DFFA2" box="[151,1369,674,697]" gridrow="4" pageId="3" pageNumber="62" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FD105950FFA2" box="[151,564,674,697]" gridcol="0" gridrow="4" pageId="3" pageNumber="62">MERISTICS (mode)</th>
</tr>
<tr id="6A9134877F4D1DAD61C1FD7F5E3DFE1C" box="[151,1369,717,775]" gridrow="5" pageId="3" pageNumber="62">
<th id="29405DFB7F4D1DAD61C1FD7F5950FE1C" box="[151,564,717,775]" gridcol="0" gridrow="5" pageId="3" pageNumber="62">Vertebrae</th>
<td id="29405DFB7F4D1DAD636DFD7F5880FE1C" box="[571,996,717,775]" gridcol="1" gridrow="5" pageId="3" pageNumber="62">10–11 + 19–20 (total 30–31, 30 or 31) (n=4)</td>
<td id="29405DFB7F4D1DAD6556FD7F5E4BFE1C" box="[1024,1327,717,775]" gridcol="2" gridrow="5" pageId="3" pageNumber="62">10–11 + 20–22 (total 31–32, (n=7)</td>
<td id="29405DFB7F4D1DAD6460FD7F5E3DFE1C" box="[1334,1369,717,775]" gridcol="3" gridrow="5" pageId="3" pageNumber="62">31)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FCA85E3DFE2A" box="[151,1369,794,817]" gridrow="6" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FCA85950FE2A" box="[151,564,794,817]" gridcol="0" gridrow="6" pageId="3" pageNumber="62">dorsal fin rays</th>
<td id="29405DFB7F4D1DAD636DFCA85880FE2A" box="[571,996,794,817]" gridcol="1" gridrow="6" pageId="3" pageNumber="62">0–I, 8–9 (total 8–10, 10)</td>
<td id="29405DFB7F4D1DAD6556FCA85E4BFE2A" box="[1024,1327,794,817]" gridcol="2" gridrow="6" pageId="3" pageNumber="62">I, 8–10 (total 9–11, 9)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FCF75E3DFE47" box="[151,1369,837,860]" gridrow="7" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FCF75950FE47" box="[151,564,837,860]" gridcol="0" gridrow="7" pageId="3" pageNumber="62">anal fin rays</th>
<td id="29405DFB7F4D1DAD636DFCF75880FE47" box="[571,996,837,860]" gridcol="1" gridrow="7" pageId="3" pageNumber="62">I–II, 27–29 (total 29–31, 30 or 31)</td>
<td id="29405DFB7F4D1DAD6556FCF75E4BFE47" box="[1024,1327,837,860]" gridcol="2" gridrow="7" pageId="3" pageNumber="62">I–II, 25–29 (total 27–31)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FCC25E3DFE9C" box="[151,1369,880,903]" gridrow="8" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FCC25950FE9C" box="[151,564,880,903]" gridcol="0" gridrow="8" pageId="3" pageNumber="62">caudal fin rays</th>
<td id="29405DFB7F4D1DAD636DFCC25880FE9C" box="[571,996,880,903]" gridcol="1" gridrow="8" pageId="3" pageNumber="62">ii, 5 + 7, i</td>
<td id="29405DFB7F4D1DAD6556FCC25E4BFE9C" box="[1024,1327,880,903]" gridcol="2" gridrow="8" pageId="3" pageNumber="62">ii, 5 + 7, i</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FC285E3DFEAA" box="[151,1369,922,945]" gridrow="9" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FC285950FEAA" box="[151,564,922,945]" gridcol="0" gridrow="9" pageId="3" pageNumber="62">pelvic fin rays</th>
<td id="29405DFB7F4D1DAD636DFC285880FEAA" box="[571,996,922,945]" gridcol="1" gridrow="9" pageId="3" pageNumber="62">I, 1, 4</td>
<td id="29405DFB7F4D1DAD6556FC285E4BFEAA" box="[1024,1327,922,945]" gridcol="2" gridrow="9" pageId="3" pageNumber="62">I, 1, 4</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FC775E3DFEC7" box="[151,1369,965,988]" gridrow="10" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FC775950FEC7" box="[151,564,965,988]" gridcol="0" gridrow="10" pageId="3" pageNumber="62">pectoral fin rays</th>
<td id="29405DFB7F4D1DAD636DFC775880FEC7" box="[571,996,965,988]" gridcol="1" gridrow="10" pageId="3" pageNumber="62">12–14 (14)</td>
<td id="29405DFB7F4D1DAD6556FC775E4BFEC7" box="[1024,1327,965,988]" gridcol="2" gridrow="10" pageId="3" pageNumber="62">13</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FC425E3DF913" box="[151,1369,1008,1032]" gridrow="11" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FC425950F913" box="[151,564,1008,1032]" gridcol="0" gridrow="11" pageId="3" pageNumber="62">subdorsal scales</th>
<td id="29405DFB7F4D1DAD636DFC425880F913" box="[571,996,1008,1032]" gridcol="1" gridrow="11" pageId="3" pageNumber="62">51/2–61/2 (61/2)</td>
<td id="29405DFB7F4D1DAD6556FC425E4BF913" box="[1024,1327,1008,1032]" gridcol="2" gridrow="11" pageId="3" pageNumber="62">61/2–71/2 (61/2, 7)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FBA95E3DF928" box="[151,1369,1051,1075]" gridrow="12" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FBA95950F928" box="[151,564,1051,1075]" gridcol="0" gridrow="12" pageId="3" pageNumber="62">transverse scales</th>
<td id="29405DFB7F4D1DAD636DFBA95880F928" box="[571,996,1051,1075]" gridcol="1" gridrow="12" pageId="3" pageNumber="62">91/2–101/2 (91/2)</td>
<td id="29405DFB7F4D1DAD6556FBA95E4BF928" box="[1024,1327,1051,1075]" gridcol="2" gridrow="12" pageId="3" pageNumber="62">10</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FBF45E3DF946" box="[151,1369,1094,1117]" gridrow="13" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FBF45950F946" box="[151,564,1094,1117]" gridcol="0" gridrow="13" pageId="3" pageNumber="62">lateral scales</th>
<td id="29405DFB7F4D1DAD636DFBF45880F946" box="[571,996,1094,1117]" gridcol="1" gridrow="13" pageId="3" pageNumber="62">30–32 (31, 32)</td>
<td id="29405DFB7F4D1DAD6556FBF45E4BF946" box="[1024,1327,1094,1117]" gridcol="2" gridrow="13" pageId="3" pageNumber="62">32–33 (32)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FBC35E3DF993" box="[151,1369,1137,1160]" gridrow="14" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FBC35950F993" box="[151,564,1137,1160]" gridcol="0" gridrow="14" pageId="3" pageNumber="62">predorsal scales</th>
<td id="29405DFB7F4D1DAD636DFBC35880F993" box="[571,996,1137,1160]" gridcol="1" gridrow="14" pageId="3" pageNumber="62">23–25 (23, 24)</td>
<td id="29405DFB7F4D1DAD6556FBC35E4BF993" box="[1024,1327,1137,1160]" gridcol="2" gridrow="14" pageId="3" pageNumber="62">22–24 (23)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FB295E3DF9A9" box="[151,1369,1179,1202]" gridrow="15" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FB295950F9A9" box="[151,564,1179,1202]" gridcol="0" gridrow="15" pageId="3" pageNumber="62">postdorsal scales</th>
<td id="29405DFB7F4D1DAD636DFB295880F9A9" box="[571,996,1179,1202]" gridcol="1" gridrow="15" pageId="3" pageNumber="62">10–11 (11)</td>
<td id="29405DFB7F4D1DAD6556FB295E4BF9A9" box="[1024,1327,1179,1202]" gridcol="2" gridrow="15" pageId="3" pageNumber="62">10–11 (11)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FB745E3DF9C6" box="[151,1369,1222,1245]" gridrow="16" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FB745950F9C6" box="[151,564,1222,1245]" gridcol="0" gridrow="16" pageId="3" pageNumber="62">lateral scales below dorsal fin origin</th>
<td id="29405DFB7F4D1DAD636DFB745880F9C6" box="[571,996,1222,1245]" gridcol="1" gridrow="16" pageId="3" pageNumber="62">16–17 (17)</td>
<td id="29405DFB7F4D1DAD6556FB745E4BF9C6" box="[1024,1327,1222,1245]" gridcol="2" gridrow="16" pageId="3" pageNumber="62">16–18 (17)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FB435E3DF813" box="[151,1369,1265,1288]" gridrow="17" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FB435950F813" box="[151,564,1265,1288]" gridcol="0" gridrow="17" pageId="3" pageNumber="62">lateral scales above anal fin origin</th>
<td id="29405DFB7F4D1DAD636DFB435880F813" box="[571,996,1265,1288]" gridcol="1" gridrow="17" pageId="3" pageNumber="62">5–6 (6)</td>
<td id="29405DFB7F4D1DAD6556FB435E4BF813" box="[1024,1327,1265,1288]" gridcol="2" gridrow="17" pageId="3" pageNumber="62">5–6½ (6)</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FAA95E3DF829" box="[151,1369,1307,1330]" gridrow="18" pageId="3" pageNumber="62" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FAA95950F829" box="[151,564,1307,1330]" gridcol="0" gridrow="18" pageId="3" pageNumber="62">MORPHOMETRICS - % SL</th>
</tr>
<tr id="6A9134877F4D1DAD61C1FAF45E3DF846" box="[151,1369,1350,1373]" gridrow="19" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FAF45950F846" box="[151,564,1350,1373]" gridcol="0" gridrow="19" pageId="3" pageNumber="62">total length</th>
<td id="29405DFB7F4D1DAD636DFAF45880F846" box="[571,996,1350,1373]" gridcol="1" gridrow="19" pageId="3" pageNumber="62">135.5–142.3</td>
<td id="29405DFB7F4D1DAD6556FAF45E4BF846" box="[1024,1327,1350,1373]" gridcol="2" gridrow="19" pageId="3" pageNumber="62">136.5–144.1</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FAC35E3DF893" box="[151,1369,1393,1416]" gridrow="20" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FAC35950F893" box="[151,564,1393,1416]" gridcol="0" gridrow="20" pageId="3" pageNumber="62">predorsal length</th>
<td id="29405DFB7F4D1DAD636DFAC35880F893" box="[571,996,1393,1416]" gridcol="1" gridrow="20" pageId="3" pageNumber="62">66.8–69.7</td>
<td id="29405DFB7F4D1DAD6556FAC35E4BF893" box="[1024,1327,1393,1416]" gridcol="2" gridrow="20" pageId="3" pageNumber="62">65.5–67.4</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FA295E3DF8A9" box="[151,1369,1435,1458]" gridrow="21" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FA295950F8A9" box="[151,564,1435,1458]" gridcol="0" gridrow="21" pageId="3" pageNumber="62">postdorsal length</th>
<td id="29405DFB7F4D1DAD636DFA295880F8A9" box="[571,996,1435,1458]" gridcol="1" gridrow="21" pageId="3" pageNumber="62">19.3–23.3</td>
<td id="29405DFB7F4D1DAD6556FA295E4BF8A9" box="[1024,1327,1435,1458]" gridcol="2" gridrow="21" pageId="3" pageNumber="62">19.9–24.3</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FA745E3DF8C6" box="[151,1369,1478,1501]" gridrow="22" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FA745950F8C6" box="[151,564,1478,1501]" gridcol="0" gridrow="22" pageId="3" pageNumber="62">body depth</th>
<td id="29405DFB7F4D1DAD636DFA745880F8C6" box="[571,996,1478,1501]" gridcol="1" gridrow="22" pageId="3" pageNumber="62">30.5–33.7</td>
<td id="29405DFB7F4D1DAD6556FA745E4BF8C6" box="[1024,1327,1478,1501]" gridcol="2" gridrow="22" pageId="3" pageNumber="62">24.3–29.0</td>
</tr>
<tr id="6A9134877F4D1DAD61C1FA435E3DFB13" box="[151,1369,1521,1544]" gridrow="23" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1FA435950FB13" box="[151,564,1521,1544]" gridcol="0" gridrow="23" pageId="3" pageNumber="62">caudal peduncle depth</th>
<td id="29405DFB7F4D1DAD636DFA435880FB13" box="[571,996,1521,1544]" gridcol="1" gridrow="23" pageId="3" pageNumber="62">19.8–22.1</td>
<td id="29405DFB7F4D1DAD6556FA435E4BFB13" box="[1024,1327,1521,1544]" gridcol="2" gridrow="23" pageId="3" pageNumber="62">16.8–19.0</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F9A95E3DFB29" box="[151,1369,1563,1586]" gridrow="24" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F9A95950FB29" box="[151,564,1563,1586]" gridcol="0" gridrow="24" pageId="3" pageNumber="62">preanal length</th>
<td id="29405DFB7F4D1DAD636DF9A95880FB29" box="[571,996,1563,1586]" gridcol="1" gridrow="24" pageId="3" pageNumber="62">44.9–46.7</td>
<td id="29405DFB7F4D1DAD6556F9A95E4BFB29" box="[1024,1327,1563,1586]" gridcol="2" gridrow="24" pageId="3" pageNumber="62">45.3–47.6</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F9F45E3DFB46" box="[151,1369,1606,1629]" gridrow="25" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F9F45950FB46" box="[151,564,1606,1629]" gridcol="0" gridrow="25" pageId="3" pageNumber="62">head length</th>
<td id="29405DFB7F4D1DAD636DF9F45880FB46" box="[571,996,1606,1629]" gridcol="1" gridrow="25" pageId="3" pageNumber="62">32.8–34.9</td>
<td id="29405DFB7F4D1DAD6556F9F45E4BFB46" box="[1024,1327,1606,1629]" gridcol="2" gridrow="25" pageId="3" pageNumber="62">31.9–34.4</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F9C35E3DFB93" box="[151,1369,1649,1672]" gridrow="26" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F9C35950FB93" box="[151,564,1649,1672]" gridcol="0" gridrow="26" pageId="3" pageNumber="62">pelvic fin length</th>
<td id="29405DFB7F4D1DAD636DF9C35880FB93" box="[571,996,1649,1672]" gridcol="1" gridrow="26" pageId="3" pageNumber="62">29.4–41.6</td>
<td id="29405DFB7F4D1DAD6556F9C35E4BFB93" box="[1024,1327,1649,1672]" gridcol="2" gridrow="26" pageId="3" pageNumber="62">29.0–37.7</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F9295E3DFBA9" box="[151,1369,1691,1714]" gridrow="27" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F9295950FBA9" box="[151,564,1691,1714]" gridcol="0" gridrow="27" pageId="3" pageNumber="62">anal fin base length</th>
<td id="29405DFB7F4D1DAD636DF9295880FBA9" box="[571,996,1691,1714]" gridcol="1" gridrow="27" pageId="3" pageNumber="62">55.2–61.1</td>
<td id="29405DFB7F4D1DAD6556F9295E4BFBA9" box="[1024,1327,1691,1714]" gridcol="2" gridrow="27" pageId="3" pageNumber="62">52.4–57.9</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F9745E3DFBC6" box="[151,1369,1734,1757]" gridrow="28" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F9745950FBC6" box="[151,564,1734,1757]" gridcol="0" gridrow="28" pageId="3" pageNumber="62">dorsal fin base length</th>
<td id="29405DFB7F4D1DAD636DF9745880FBC6" box="[571,996,1734,1757]" gridcol="1" gridrow="28" pageId="3" pageNumber="62">11.4–15.0</td>
<td id="29405DFB7F4D1DAD6556F9745E4BFBC6" box="[1024,1327,1734,1757]" gridcol="2" gridrow="28" pageId="3" pageNumber="62">12.3–14.3</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F9435E3DFA13" box="[151,1369,1777,1800]" gridrow="29" pageId="3" pageNumber="62" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F9435950FA13" box="[151,564,1777,1800]" gridcol="0" gridrow="29" pageId="3" pageNumber="62">% HL</th>
</tr>
<tr id="6A9134877F4D1DAD61C1F8A95E3DFA29" box="[151,1369,1819,1842]" gridrow="30" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F8A95950FA29" box="[151,564,1819,1842]" gridcol="0" gridrow="30" pageId="3" pageNumber="62">orbit diameter</th>
<td id="29405DFB7F4D1DAD636DF8A95880FA29" box="[571,996,1819,1842]" gridcol="1" gridrow="30" pageId="3" pageNumber="62">25.0–27.7</td>
<td id="29405DFB7F4D1DAD6556F8A95E4BFA29" box="[1024,1327,1819,1842]" gridcol="2" gridrow="30" pageId="3" pageNumber="62">25.9–28.3</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F8F45E3DFA46" box="[151,1369,1862,1885]" gridrow="31" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F8F45950FA46" box="[151,564,1862,1885]" gridcol="0" gridrow="31" pageId="3" pageNumber="62">postorbital length</th>
<td id="29405DFB7F4D1DAD636DF8F45880FA46" box="[571,996,1862,1885]" gridcol="1" gridrow="31" pageId="3" pageNumber="62">44.0–49.5</td>
<td id="29405DFB7F4D1DAD6556F8F45E4BFA46" box="[1024,1327,1862,1885]" gridcol="2" gridrow="31" pageId="3" pageNumber="62">43.8–51.0</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F8C35E3DFA93" box="[151,1369,1905,1928]" gridrow="32" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F8C35950FA93" box="[151,564,1905,1928]" gridcol="0" gridrow="32" pageId="3" pageNumber="62">interorbital width</th>
<td id="29405DFB7F4D1DAD636DF8C35880FA93" box="[571,996,1905,1928]" gridcol="1" gridrow="32" pageId="3" pageNumber="62">32.7–38.3</td>
<td id="29405DFB7F4D1DAD6556F8C35E4BFA93" box="[1024,1327,1905,1928]" gridcol="2" gridrow="32" pageId="3" pageNumber="62">33.0–37.2</td>
</tr>
<tr id="6A9134877F4D1DAD61C1F8295E3DFAA9" box="[151,1369,1947,1970]" gridrow="33" pageId="3" pageNumber="62" rowspan-3="1">
<th id="29405DFB7F4D1DAD61C1F8295950FAA9" box="[151,564,1947,1970]" gridcol="0" gridrow="33" pageId="3" pageNumber="62">snout length</th>
<td id="29405DFB7F4D1DAD636DF8295880FAA9" box="[571,996,1947,1970]" gridcol="1" gridrow="33" pageId="3" pageNumber="62">23.6–28.3</td>
<td id="29405DFB7F4D1DAD6556F8295E4BFAA9" box="[1024,1327,1947,1970]" gridcol="2" gridrow="33" pageId="3" pageNumber="62">23.1–25.8</td>
</tr>
</table>
</paragraph>
<caption id="80DE664D7F4AE25661C1F9425E48FA1C" box="[151,1324,1776,1800]" httpUri="https://zenodo.org/record/189077/files/figure.png" pageId="4" pageNumber="63" targetBox="[156,1436,197,1745]" targetPageId="4">
<paragraph id="D41E36C57F4AE25661C1F9425E48FA1C" blockId="4.[151,1324,1776,1800]" box="[151,1324,1776,1800]" pageId="4" pageNumber="63">
<emphasis id="E6D5EAD77F4AE25661C1F9425A7DFA13" bold="true" box="[151,281,1776,1800]" pageId="4" pageNumber="63">FIGURE 3.</emphasis>
Map showing known distribution of
<taxonomicName id="13A14D467F4AE25663F0F9425821FA1C" box="[678,837,1776,1799]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4AE25663F0F9425821FA1C" box="[678,837,1776,1799]" italics="true" pageId="4" pageNumber="63">B. anabatoides</emphasis>
</taxonomicName>
(circle) and
<taxonomicName id="13A14D467F4AE25662B2F9425F26FA1C" box="[996,1090,1776,1799]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="midas">
<emphasis id="E6D5EAD77F4AE25662B2F9425F26FA1C" box="[996,1090,1776,1799]" italics="true" pageId="4" pageNumber="63">B. midas</emphasis>
</taxonomicName>
(square) in Borneo.
</paragraph>
</caption>
<paragraph id="D41E36C57F4AE2576190F8845BD9FDC3" blockId="4.[151,1437,1846,2027]" lastBlockId="5.[151,1437,152,1299]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
<emphasis id="E6D5EAD77F4AE2566190F8845AC6FA4B" bold="true" box="[198,418,1846,1872]" pageId="4" pageNumber="63">Coloration in life.</emphasis>
See
<figureCitation id="4C9A2A407F4AE256608DF8845946FA4B" box="[475,546,1846,1872]" captionStart="FIGURE 2" captionStartId="2.[151,255,1805,1829]" captionTargetBox="[151,1434,1144,1780]" captionTargetId="figure@2.[151,1436,1144,1781]" captionTargetPageId="2" captionText="FIGURE 2. Betta anabatoides, not preserved, ca. 35 mm SL, Central Kalimantan." httpUri="https://zenodo.org/record/189076/files/figure.png" pageId="4" pageNumber="63">Fig. 2</figureCitation>
, and refer to
<bibRefCitation id="B0304B347F4AE25663EAF88458FAFA4B" author="Vierke" box="[700,926,1846,1872]" pageId="4" pageNumber="68" refString="Vierke, J. (1979) Betta anabatoides und Betta foerschi spec. nov., zwei Kampffische aus Borneo. Zeitschrift fur Aquarien-und Terrarienfreunde, 13, 386 - 390." type="journal article" year="1979">Vierke (1979: 389)</bibRefCitation>
,
<bibRefCitation id="B0304B347F4AE25662FFF8845F0BFA4B" author="Linke" box="[937,1135,1846,1872]" pageId="4" pageNumber="68" refString="Linke, H. (1991) Labyrinth fish. The Bubble-nest-builders. Their identification, care and breeding. Tetra-Press, Melle, Germany, 174 pp." type="book" year="1991">Linke (1991: 26)</bibRefCitation>
and
<bibRefCitation id="B0304B347F4AE25665FEF8845EFCFA4B" author="Kottelat" box="[1192,1432,1846,1872]" pageId="4" pageNumber="68" refString="Kottelat, M., Whitten, A. J., Kartikasari, S. N. &amp; Wirjoatmodjo, S. (1993) Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, 259 pp., 84 pls." type="book" year="1993">
Kottelat
<emphasis id="E6D5EAD77F4AE2566458F8855E2CFA4B" box="[1294,1352,1847,1872]" italics="true" pageId="4" pageNumber="63">et al.</emphasis>
(1993
</bibRefCitation>
: pl. 75) for colour illustrations. Dorsum of body brown, venter lighter brown. Dorsum of head brown with darker brown blotches behind eye; opercle with faint gold iridescence. Distinct pre- and postorbital black stripes on head; lower lip black; chin-bar absent. No iridescence on body. Sides of body plain brown, sometimes with 5–7 faint, irregularly-spaced vertical brown bars. Fins brownish, dorsal fin with 5–6 darkbrown transverse bars;caudal fin without pattern. Pectoral fin hyaline. Pelvic fin with a bright whitish distal tip.
</paragraph>
<paragraph id="D41E36C57F4BE2576190FF575A1EFC81" blockId="5.[151,1437,152,1299]" pageId="5" pageNumber="64">
<emphasis id="E6D5EAD77F4BE2576190FF57597FFDE4" bold="true" box="[198,539,229,255]" pageId="5" pageNumber="64">Coloration in preservative.</emphasis>
See
<figureCitation id="4C9A2A407F4BE257630CFF5759C7FDE4" box="[602,675,229,255]" captionStart="FIGURE 1" captionStartId="2.[151,255,1067,1091]" captionTargetBox="[151,1434,622,1040]" captionTargetId="figure@2.[151,1436,622,1041]" captionTargetPageId="2" captionText="FIGURE 1. Betta anabatoides, ZRC 40971, 47.2 mm SL, neotype, Banjarmasin (right side, laterally inverted)." httpUri="https://zenodo.org/record/189075/files/figure.png" pageId="5" pageNumber="64">Fig. 1</figureCitation>
. Dorsum of body dark brown, venter lighter brown to whitish. Dorsum of head dark brown, opercular area whitish. Distinct pre- and postorbital black stripes on head; chinbar absent. Body with 5–7 irregularly spaced vertical dark-brown bars. Fins brownish; dorsal fin with 5–6 transverse black bars, caudal fin without pattern. Pectoral fin hyaline. Pelvic fin light brown, its filamentous second ray hyaline.
</paragraph>
</subSubSection>
<subSubSection id="9CBB654E7F4BE2576190FE14589DFE58" pageId="5" pageNumber="64" type="distribution">
<paragraph id="D41E36C57F4BE2576190FE145A19FF15" blockId="5.[151,1437,152,1299]" pageId="5" pageNumber="64">
<emphasis id="E6D5EAD77F4BE2576190FE145A03FCDB" bold="true" box="[198,359,422,448]" pageId="5" pageNumber="64">Distribution.</emphasis>
<taxonomicName id="13A14D467F4BE2576025FE155928FCDB" box="[371,588,423,448]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4BE2576025FE155928FCDB" box="[371,588,423,448]" italics="true" pageId="5" pageNumber="64">Betta anabatoides</emphasis>
</taxonomicName>
is known from the lower Barito basin in South Kalimantan and the Kotawaringin, Katingan, Sampit, Mentaya and Kahayan basins in Central Kalimantan (
<figureCitation id="4C9A2A407F4BE25765F5FE7F5F88FCFC" box="[1187,1260,461,487]" captionStart="FIGURE 3" captionStartId="4.[151,255,1776,1800]" captionTargetBox="[156,1436,197,1745]" captionTargetId="figure@4.[151,1436,194,1750]" captionTargetPageId="4" captionText="FIGURE 3. Map showing known distribution of B. anabatoides (circle) and B. midas (square) in Borneo." httpUri="https://zenodo.org/record/189077/files/figure.png" pageId="5" pageNumber="64">Fig. 3</figureCitation>
; pers. obs.), in Indonesian Borneo.
</paragraph>
<paragraph id="D41E36C57F4BE2576190FDA85E49FFD4" blockId="5.[151,1437,152,1299]" pageId="5" pageNumber="64">
<emphasis id="E6D5EAD77F4BE2576190FDA85A3CFF2F" bold="true" box="[198,344,538,564]" pageId="5" pageNumber="64">Field notes.</emphasis>
The stream in which the
<typeStatus id="0B1A88677F4BE25763C4FDA8598BFF2F" box="[658,751,538,564]" pageId="5" pageNumber="64" type="neotype">neotype</typeStatus>
material of
<taxonomicName id="13A14D467F4BE25762D2FDA85F53FF28" box="[900,1079,538,563]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4BE25762D2FDA85F53FF28" box="[900,1079,538,563]" italics="true" pageId="5" pageNumber="64">B. anabatoides</emphasis>
</taxonomicName>
was obtained in Banjarmasin was murky, up to
<quantity id="13599B207F4BE257603CFDF35AC9FF40" box="[362,429,577,603]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="64" unit="m" value="1.2">1.2 m</quantity>
deep and
<collectingCountry id="ACB676557F4BE2576370FDF3592EFF40" box="[550,586,577,603]" name="Philippines" pageId="5" pageNumber="64">pH</collectingCountry>
7.5. The water was sluggish and drained from cultivated area upstream. Syntopic fishes included:
<taxonomicName id="13A14D467F4BE2576086FDDA59C5FF9A" box="[464,673,616,641]" class="Actinopterygii" family="Cyprinidae" genus="Boraras" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="brigittae">
<emphasis id="E6D5EAD77F4BE2576086FDDA59C5FF9A" box="[464,673,616,641]" italics="true" pageId="5" pageNumber="64">Boraras brigittae</emphasis>
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE25763E7FDDA58CAFF9A" box="[689,942,616,641]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="agyrotaenia">
<emphasis id="E6D5EAD77F4BE25763E7FDDA58CAFF9A" box="[689,942,616,641]" italics="true" pageId="5" pageNumber="64">Rasbora agyrotaenia</emphasis>
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE25762E8FDDA5F01FF9A" box="[958,1125,616,641]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="einthovenii">
<emphasis id="E6D5EAD77F4BE25762E8FDDA5F01FF9A" box="[958,1125,616,641]" italics="true" pageId="5" pageNumber="64">R. einthovenii</emphasis>
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE2576523FDDA5EF3FF99" box="[1141,1431,616,642]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="E6D5EAD77F4BE2576523FDDA5F97FF9A" box="[1141,1267,616,641]" italics="true" pageId="5" pageNumber="64">R. gracilis</emphasis>
(Cyprinidae)
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE25761C1FD3D59EEFFB3" box="[151,650,654,680]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalichthys" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="hasseltii">
<emphasis id="E6D5EAD77F4BE25761C1FD3D5A9CFFB3" box="[151,504,655,680]" italics="true" pageId="5" pageNumber="64">Lepidocephalichthys hasseltii</emphasis>
(Cobitidae)
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE25763C1FD3D5E68FFB3" box="[663,1292,654,680]" class="Actinopterygii" family="Zenarchopteridae" genus="Hemirhamphodon" kingdom="Animalia" order="Beloniformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="pogonognathus">
<emphasis id="E6D5EAD77F4BE25763C1FD3D5F4AFFB3" box="[663,1070,655,680]" italics="true" pageId="5" pageNumber="64">Hemirhamphodon pogonognathus</emphasis>
(Hemiramphidae)
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE257644EFD3D5AF2FFD4" family="Gobiidae" genus="Calamiana" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="paludosus">
<emphasis id="E6D5EAD77F4BE257644EFD3D5A6AFFD5" italics="true" pageId="5" pageNumber="64">Calamiana paludosus</emphasis>
(Gobiidae)
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE25760F5FD07595AFFD5" box="[419,574,693,718]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="edithae">
<emphasis id="E6D5EAD77F4BE25760F5FD07595AFFD5" box="[419,574,693,718]" italics="true" pageId="5" pageNumber="64">Betta edithae</emphasis>
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE257631AFD075811FFD5" box="[588,885,693,718]" class="Actinopterygii" family="Osphronemidae" genus="Trichopodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="trichopterus">
<emphasis id="E6D5EAD77F4BE257631AFD075811FFD5" box="[588,885,693,718]" italics="true" pageId="5" pageNumber="64">Trichopodus trichopterus</emphasis>
</taxonomicName>
,
<taxonomicName id="13A14D467F4BE25762D2FD075E4DFFD4" box="[900,1321,693,719]" class="Actinopterygii" family="Osphronemidae" genus="Trichopsis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="vittata">
<emphasis id="E6D5EAD77F4BE25762D2FD075F35FFD5" box="[900,1105,693,718]" italics="true" pageId="5" pageNumber="64">Trichopsis vittata</emphasis>
(Osphronemidae)
</taxonomicName>
.
</paragraph>
<paragraph id="D41E36C57F4BE2576190FD6E589DFE58" blockId="5.[151,1437,152,1299]" pageId="5" pageNumber="64">
In Central Kalimantan, the streams in which
<taxonomicName id="13A14D467F4BE25763B7FD6E58F6FFEE" box="[737,914,732,757]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4BE25763B7FD6E58F6FFEE" box="[737,914,732,757]" italics="true" pageId="5" pageNumber="64">B. anabatoides</emphasis>
</taxonomicName>
can be found lie within a mix of heath and peat swamp forest habitats. These habitats are usually typified by clear, highly tannified and very acidic waters (
<collectingCountry id="ACB676557F4BE25761A5FC9B5A7CFE58" box="[243,280,809,835]" name="Philippines" pageId="5" pageNumber="64">pH</collectingCountry>
as low as 3.5). The substrate is a mix of silicate sand and peat.
</paragraph>
</subSubSection>
<subSubSection id="9CBB654E7F4BE2576190FCFD5F0BF808" pageId="5" pageNumber="64" type="discussion">
<paragraph id="D41E36C57F4BE2576190FCFD59CDF930" blockId="5.[151,1437,152,1299]" pageId="5" pageNumber="64">
<emphasis id="E6D5EAD77F4BE2576190FCFD5A24FE72" bold="true" box="[198,320,847,873]" pageId="5" pageNumber="64">Remarks.</emphasis>
<taxonomicName id="13A14D467F4BE257601CFCE25945FE72" box="[330,545,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4BE257601CFCE25945FE72" box="[330,545,848,873]" italics="true" pageId="5" pageNumber="64">Betta anabatoides</emphasis>
</taxonomicName>
can be differentiated from its closest relative,
<taxonomicName id="13A14D467F4BE257650FFCE25FA7FE72" box="[1113,1219,848,873]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="midas">
<emphasis id="E6D5EAD77F4BE257650FFCE25FA7FE72" box="[1113,1219,848,873]" italics="true" pageId="5" pageNumber="64">B. midas</emphasis>
</taxonomicName>
, by the following characters: a more thick-set body (body depth 30.5–33.7 % SL, vs. 24.3–29.0); a deeper caudal peduncle (19.8–22.1 % SL, vs. 16.8–19.0); absence (vs. presence) of caudal transverse bars; faint (vs. distinct) iridescent gold opercle scales; absence (vs. presence) of gold body scales; pointed (vs. blunt) lateral head profile; a slightly greater predorsal length (66.8–69.7 % SL, vs. 65.5–67.4); and a slightly greater anal-fin base length (55.2–61.1 % SL, vs. 52.4–57.9).
</paragraph>
<paragraph id="D41E36C57F4BE2576190FB8A5F0BF808" blockId="5.[151,1437,152,1299]" pageId="5" pageNumber="64">
<taxonomicName id="13A14D467F4BE2576190FB8A5AFBF94A" box="[198,415,1080,1105]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="anabatoides">
<emphasis id="E6D5EAD77F4BE2576190FB8A5AFBF94A" box="[198,415,1080,1105]" italics="true" pageId="5" pageNumber="64">Betta anabatoides</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="13A14D467F4BE25763A5FB8A58E3F94A" box="[755,903,1080,1105]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="akarensis">
<emphasis id="E6D5EAD77F4BE25763A5FB8A58E3F94A" box="[755,903,1080,1105]" italics="true" pageId="5" pageNumber="64">B. akarensis</emphasis>
</taxonomicName>
and
<taxonomicName id="13A14D467F4BE257629FFB8A5F04F94A" box="[969,1120,1080,1105]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="ibanorum">
<emphasis id="E6D5EAD77F4BE257629FFB8A5F04F94A" box="[969,1120,1080,1105]" italics="true" pageId="5" pageNumber="64">B. ibanorum</emphasis>
</taxonomicName>
by the following suite of characters: absence (vs. presence) of chin-bar; caudal fin without pattern (vs. caudal transverse bars present); a greater body depth (30.5–33.7 % SL, vs. 23.3–30.9); a greater anal-fin base length (55.2–61.1 % SL, vs. 49.6–56.8); a shorter caudal fin length (total length 135.5–142.3 % SL, vs. 134.4–165.6); a greater number of anal fin rays than
<taxonomicName id="13A14D467F4BE257603AFB615964F9F7" box="[364,512,1235,1260]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="ibanorum">
<emphasis id="E6D5EAD77F4BE257603AFB615964F9F7" box="[364,512,1235,1260]" italics="true" pageId="5" pageNumber="64">B. ibanorum</emphasis>
</taxonomicName>
(29–31, vs. 27–29) and a smaller preanal length than
<taxonomicName id="13A14D467F4BE25765D7FB615E70F9F7" box="[1153,1300,1235,1260]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="ibanorum">
<emphasis id="E6D5EAD77F4BE25765D7FB615E70F9F7" box="[1153,1300,1235,1260]" italics="true" pageId="5" pageNumber="64">B. ibanorum</emphasis>
</taxonomicName>
(44.9–46.7 % SL, vs. 45.8–49.4) (data on
<taxonomicName id="13A14D467F4BE25760ABFB4B59E9F809" box="[509,653,1273,1298]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="akarensis">
<emphasis id="E6D5EAD77F4BE25760ABFB4B59E9F809" box="[509,653,1273,1298]" italics="true" pageId="5" pageNumber="64">B. akarensis</emphasis>
</taxonomicName>
and
<taxonomicName id="13A14D467F4BE2576393FB4B583CF809" box="[709,856,1273,1298]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="ibanorum">
<emphasis id="E6D5EAD77F4BE2576393FB4B583CF809" box="[709,856,1273,1298]" italics="true" pageId="5" pageNumber="64">B. ibanorum</emphasis>
</taxonomicName>
from
<bibRefCitation id="B0304B347F4BE25762F6FB4B5F06F808" author="Tan" box="[928,1122,1273,1299]" pageId="5" pageNumber="68" refString="Tan, H. H. &amp; Ng, P. K. L. (2004) Two new species of freshwater fish (Teleostei: Balitoridae, Osphronemidae) from southern Sarawak. In: Sarawak Bau Limestone Biodiversity (eds. H. S. Yong, F. S. P. Ng and E. E. L. Yen). Sarawak Museum Journal, Vol. LIX, No. 80 (New Series); Special Issue No. 8, 267 - 284." type="book chapter" year="2004">Tan &amp; Ng, 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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