236 lines
32 KiB
XML
236 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1108.85156" ID-GBIF-Dataset="82ad6418-e2ba-4e94-b288-54cacb69506a" ID-PMC="PMC9848670" ID-Pensoft-Pub="1313-2970-1108-89" ID-Pensoft-UUID="4C588406675B5E2F8199E4C9DF13E485" ID-PubMed="36760703" ID-ZooBank="DB9FDD0F5FDF4B028E0B003EAE84EC91" ModsDocID="1313-2970-1108-89" checkinTime="1656059516731" checkinUser="pensoft" docAuthor="Zhao, Yi, Guo, Wan-Ru, Golovatch, Sergei I. & Liu, Wei-Xin" docDate="2022" docId="2F6C4A2FA26D5C6AB2EFC35CFD97C050" docLanguage="en" docName="ZooKeys 1108: 89-118" docOrigin="ZooKeys 1108" docPubDate="2022-06-23" docSource="http://dx.doi.org/10.3897/zookeys.1108.85156" docTitle="Glyphiulus portaliformis Zhao & Liu 2022, sp. nov." docType="treatment" docUuid="5879B2B8-A5DE-46BE-8710-07E3E41BAE46" docUuidSource="ZooBank" docVersion="4" id="4C588406675B5E2F8199E4C9DF13E485" lastPageNumber="89" masterDocId="4C588406675B5E2F8199E4C9DF13E485" masterDocTitle="Revision of the javanicus species group of the millipede genus Glyphiulus Gervais, 1847, with descriptions of five new species from China (Diplopoda, Spirostreptida, Cambalopsidae)" masterLastPageNumber="118" masterPageNumber="89" pageNumber="89" updateTime="1732480599592" updateUser="ExternalLinkService">
|
|
<mods:mods id="E4F50B6052C7B5CB87886CB8BE26AE05" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="DBBE0B68648EF115B99D41A7EEFE1845">
|
|
<mods:title id="F65CB75764CE7F50133B32EFC9297B6B">Revision of the javanicus species group of the millipede genus Glyphiulus Gervais, 1847, with descriptions of five new species from China (Diplopoda, Spirostreptida, Cambalopsidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="53FA078E00AC2FD83AF3274BB9770A3B" type="personal">
|
|
<mods:role id="8A585CF82F96956DE09A9269E83BCEE0">
|
|
<mods:roleTerm id="62283BB98A187E9EF8A56397DE1CCD27">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="D7797C1674C9DABCCBFF09DAFECAE9EF">Zhao, Yi</mods:namePart>
|
|
<mods:affiliation id="840FB7EED67290DF4D4410A1A0CBBACD">College of Plant Protection, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="C1B55E8512CA43A590A06F366FF1EA46" type="personal">
|
|
<mods:role id="C2DCA37A0D72D9AFF7B3E4A236364E41">
|
|
<mods:roleTerm id="981B71100B0AE81B9F3EA0A61CF1277F">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="995EA147608BC964E07A09ECF6CB4239">Guo, Wan-Ru</mods:namePart>
|
|
<mods:affiliation id="F1D6045C033C40724459A8B728C4D940">College of Plant Protection, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="1E06269F35F304D11B1C1C8F573C2E33" type="personal">
|
|
<mods:role id="036A5ACEC2737A4006ED8AC111DBC2AC">
|
|
<mods:roleTerm id="E9A8495F6815231A2C7822B757AFFF77">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="053928586B1F125868AD2C75F2FEE02F">Golovatch, Sergei I.</mods:namePart>
|
|
<mods:affiliation id="7401CAAA7A5DBF13DF2294BE5A8CEAE4">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
|
|
<mods:nameIdentifier id="844AE93B72F020DA70C511B0FC525493" type="email">sgolovatch@yandex.ru</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="DF0462FEE46CE93C2ECDC9EC93F7B497" type="personal">
|
|
<mods:role id="B659B4C899A00455AA23DF74757D126F">
|
|
<mods:roleTerm id="69E694D2D4BA474A13AD5EF482EA6A62">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="2A2671085F9B58F87E39DB1AD9F5FEF0">Liu, Wei-Xin</mods:namePart>
|
|
<mods:affiliation id="04F18A960B876E4636E56B1E24F0EE1A">College of Plant Protection, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
|
|
<mods:nameIdentifier id="6D098C5CBD8D68E01552DABBE0718AE8" type="email">da2000wei@163.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource id="BF1EC530AF464446D8F89927D589ECB8">text</mods:typeOfResource>
|
|
<mods:relatedItem id="7ECBA2191DDA827725615BCB7E79FF3F" type="host">
|
|
<mods:titleInfo id="06689FEE3B5C76AD733878D70A311F4C">
|
|
<mods:title id="2CC0AF09D8F0E1E15855B4E43921F9AE">ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="F3F4EEE6EA4C520FDD449E475C09F2CA">
|
|
<mods:date id="AD2FBAEFB5861A8532F497C953E6D6D3">2022</mods:date>
|
|
<mods:detail id="1CED3CCE3C43889BFF7E63DE9024F3A1" type="pubDate">
|
|
<mods:number id="13EE1BF52DB72DA8A53A589EAEAB0698">2022-06-23</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="A12A6D497B09B4205B8E64058F7B9A2F" type="volume">
|
|
<mods:number id="794FB9127A0ABB18AD35AF5263E187EF">1108</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="70A0C8A55C57A0A8F3E2FF847F825508" unit="page">
|
|
<mods:start id="EC4CF064A353F8EEB574E83D77C908DD">89</mods:start>
|
|
<mods:end id="0086B60E238EAC9E5B3A85A26C63EE0B">118</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="88F7067B1C855F6697E6B8F129E767E1">
|
|
<mods:url id="E952A936E3D7CB94F531D2EB35D9A74F">http://dx.doi.org/10.3897/zookeys.1108.85156</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="4C2F439EA55A35C76E512E42B98E93AA">journal article</mods:classification>
|
|
<mods:identifier id="26680872C3A2F893B0CB7BB4AFB7ACE7" type="DOI">http://dx.doi.org/10.3897/zookeys.1108.85156</mods:identifier>
|
|
<mods:identifier id="7D9A23FD76ED6D62993B6880AE94061E" type="Pensoft-Pub">1313-2970-1108-89</mods:identifier>
|
|
<mods:identifier id="F0971043628B2503C60ED751270BB874" type="ZooBank">DB9FDD0F5FDF4B028E0B003EAE84EC91</mods:identifier>
|
|
<mods:identifier id="C8DC3FDCFB2B77A3A6671AE4F72D57BF" type="Pensoft-UUID">4C588406675B5E2F8199E4C9DF13E485</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="2F6C4A2FA26D5C6AB2EFC35CFD97C050" ID-GBIF-Taxon="196167196" LSID="urn:lsid:zoobank.org:act:5879B2B8-A5DE-46BE-8710-07E3E41BAE46" httpUri="http://treatment.plazi.org/id/2F6C4A2FA26D5C6AB2EFC35CFD97C050" lastPageNumber="89" pageId="0" pageNumber="89" scope_class="Diplopoda" scope_family="Cambalopsidae" scope_order="Spirostreptida">
|
|
<subSubSection id="174B46D434719D92DF9E4E4937BB61CA" pageId="0" pageNumber="89" type="nomenclature">
|
|
<paragraph id="2DBB1EB743CE5403F2485A630E20690B" pageId="0" pageNumber="89">
|
|
<taxonomicName id="1300F97437BFD5B6C4C9756C3A33F545" ID-CoL="9FW7Q" LSID="http://zoobank.org/5879B2B8-A5DE-46BE-8710-07E3E41BAE46" authority="Zhao & Liu" authorityName="Zhao & Liu" authorityYear="2022" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus portaliformis" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="portaliformis" status="sp. nov.">Glyphiulus portaliformis Zhao & Liu</taxonomicName>
|
|
<taxonomicNameLabel id="E0BEFEF2DA914BF918CFA8A670B1E56A" pageId="0" pageNumber="89">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6F53551EB260E89CA5815FAFA7B18812" pageId="0" pageNumber="89" type="description">
|
|
<paragraph id="C111C40BF93CD9E477B1F79D8BD1BEAC" pageId="0" pageNumber="89">
|
|
<figureCitation id="E362A74A428F46FB166349B6BB4CED7E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pictures of live animals A G. sinuatoprocessus Zhao & Liu, sp. nov. from Cave Niubi Yan B G. scutatus Zhao & Liu, sp. nov. from Cave Bianfu Dong C G. portaliformis Zhao & Liu, sp. nov. from Cave Baiyan Dong D G. xiniudong Zhao & Liu, sp. nov. from Cave Xiniu Dong E G. conuliformis Zhao & Liu, sp. nov. from Cave Yanzi Dong." figureDoi="10.3897/zookeys.1108.85156.figure5" httpUri="https://binary.pensoft.net/fig/705255" pageId="0" pageNumber="89">Figs 5C</figureCitation>
|
|
<figureCitation id="F3EAFFE22005579813B814A7ED3CB211" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">, 14</figureCitation>
|
|
<figureCitation id="8F07FBE25EB8BD0ECAE7DED051A7C645" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">, 15</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B0901983982775679B6ECD4292D8A710" pageId="0" pageNumber="89" type="type material">
|
|
<paragraph id="DCF5D1E49BA64CBC6FD3862E1D3A2E1B" pageId="0" pageNumber="89">Type material.</paragraph>
|
|
<paragraph id="816729DF39530102EF9884EE3D58543B" pageId="0" pageNumber="89">
|
|
<emphasis id="F446F3833C876DF595705F0361F863C8" bold="true" italics="true" pageId="0" pageNumber="89">Holotype</emphasis>
|
|
♂ (SCAU WL40), China, Guangxi Zhuang Autonomous Region, Hechi City, Bama County, Cave Baiyan Dong,
|
|
<geoCoordinate id="79C056A644E7E41C97C4F2BEC8ADBE12" degrees="24" direction="north" minutes="03" orientation="latitude" precision="15" seconds="40" value="24.06111">24°03'40"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="E554585DDF0F8B12436FC3F58CA7C595" degrees="107" direction="east" minutes="08" orientation="longitude" precision="15" seconds="16" value="107.13778">107°08'16"E</geoCoordinate>
|
|
, 400 m alt., 2015-VII-31, leg. Chen Jujian, Wang Xinhui & Tang Mingruo.
|
|
<emphasis id="467F284759AEBF7C3DC1602C35008DEC" bold="true" italics="true" pageId="0" pageNumber="89">Paratypes</emphasis>
|
|
: 7 ♂, 18 ♀ (SCAU WL40), same data as the holotype.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="938F3541B5A46848D52FA0287C4F442F" pageId="0" pageNumber="89" type="etymology">
|
|
<paragraph id="2B907965BB8034C1DD908937BD949817" pageId="0" pageNumber="89">Etymology.</paragraph>
|
|
<paragraph id="B50573CBA35DD88E258348DC5BD13C58" pageId="0" pageNumber="89">To emphasise the coxosternum of the anterior gonopods being portal-shaped.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="195399ED5A029F319797A508C15EBB07" pageId="0" pageNumber="89" type="diagnosis">
|
|
<paragraph id="95448BD900A3BAB48FF2C7BB37DD9FA6" pageId="0" pageNumber="89">Diagnosis.</paragraph>
|
|
<paragraph id="9839707281C147521F386E3984430AFA" pageId="0" pageNumber="89">
|
|
Differs from congeners of the
|
|
<taxonomicName id="E9EB595DF611888C8B406772882FF7EB" baseAuthorityName="Pocock" baseAuthorityYear="1895" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus formosus" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="formosus">
|
|
<emphasis id="C2A115210474733FEA04C9AF9E45E39E" italics="true" pageId="0" pageNumber="89">Glyphiulus formosus</emphasis>
|
|
</taxonomicName>
|
|
-group by the epiproct showing a small caudal protrusion and the anterior gonopods being portal-shaped, combined with a foliate flagellum of the posterior gonopod. Based on molecular evidence,
|
|
<taxonomicName id="62D4C34083F279569774EECC5F713298" lsidName="G. portaliformis" pageId="0" pageNumber="89" rank="species" species="portaliformis">
|
|
<emphasis id="1AEF89841AC31220DC88C79B3EC47F6F" italics="true" pageId="0" pageNumber="89">G. portaliformis</emphasis>
|
|
</taxonomicName>
|
|
Zhao & Liu, sp. nov. Differs from all other
|
|
<taxonomicName id="71A29E4C3C2B4E624E2A42CFFFD28BC6" authorityName="Gervais" authorityYear="1847" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="0AA26C28F8D1DA5197AE0901B7BAF0BC" italics="true" pageId="0" pageNumber="89">Glyphiulus</emphasis>
|
|
</taxonomicName>
|
|
species analysed in a> 14.4% uncorrected p-distance of the COI barcoding gene.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D30C73082DCD30F27016619ACC864EE0" pageId="0" pageNumber="89" type="description">
|
|
<paragraph id="A3A564B94BC3278F03B551BC0EA58859" pageId="0" pageNumber="89">Description.</paragraph>
|
|
<paragraph id="52101F1281870F2EE670EA01427CAE45" pageId="0" pageNumber="89">
|
|
Length of both sexes ca. 41.0-56.0 mm, mid-body rings round in cross-section, their width and height similar, 2.2-3.0 mm. Body with 55-67 podous + 1 apodous ring + telson. Colouration brownish, legs almost transparent (Fig.
|
|
<figureCitation id="2FBC2D591DBF568A888D6636F2D4781D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pictures of live animals A G. sinuatoprocessus Zhao & Liu, sp. nov. from Cave Niubi Yan B G. scutatus Zhao & Liu, sp. nov. from Cave Bianfu Dong C G. portaliformis Zhao & Liu, sp. nov. from Cave Baiyan Dong D G. xiniudong Zhao & Liu, sp. nov. from Cave Xiniu Dong E G. conuliformis Zhao & Liu, sp. nov. from Cave Yanzi Dong." figureDoi="10.3897/zookeys.1108.85156.figure5" httpUri="https://binary.pensoft.net/fig/705255" pageId="0" pageNumber="89">5C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="6A2427768F550F098778E5D95CA58A5F" pageId="0" pageNumber="89">
|
|
<emphasis id="9A3EDAD26256ED3EA3BB944318292EAF" bold="true" italics="true" pageId="0" pageNumber="89">Head</emphasis>
|
|
surface smooth. Labrum with 4 teeth anteromedially (Fig.
|
|
<figureCitation id="D45423FAC84E1276491D711805EA8C2E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15A</figureCitation>
|
|
). Ocellaria blackish, with 14-17 ommatidia arranged in 2-3 irregular linear rows (Figs
|
|
<figureCitation id="8E8254ACF73DA081F76D6D76645B89E7" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">14B</figureCitation>
|
|
,
|
|
<figureCitation id="5B2FEE7894CF6470AC04B7C02D21A860" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15B</figureCitation>
|
|
). Antennae short, slightly clavate, reaching back to ring 4; in length, antennomeres 5> 3> 2> 4> 6> 1> 7. Antennomeres 5-7 each with a distodorsal field or corolla of bacilliform sensilla (sensory bacilli). Antennomere 7 with four sensory cones (Fig.
|
|
<figureCitation id="12DD62BDC66A85EE6F68D93200CD9B77" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15A</figureCitation>
|
|
). Gnathochilarium with a separate promentum, polytrichous on promentum and mentum, lamellae linguales each with 6-7 setae (Fig.
|
|
<figureCitation id="6ED56D5534F8E18279E8346DDB72BD66" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15A</figureCitation>
|
|
). Mandible not dissected.
|
|
</paragraph>
|
|
<caption id="DAD935938DEA86F461377328D0B377F3" doi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89" start="Figure 14" startId="F14">
|
|
<paragraph id="CE46B7FE372D7329DDC99E36C329DCB7" pageId="0" pageNumber="89">
|
|
<emphasis id="C8F4DB9D30B1BBB4A43D7E6CBD2DE985" bold="true" pageId="0" pageNumber="89">Figure 14.</emphasis>
|
|
<taxonomicName id="30F04D0B2AA6E95B451B28C26292213E" authorityName="Zhao & Liu" authorityYear="2022" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus portaliformis" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="portaliformis">
|
|
<emphasis id="91EDF5E4B4619B0D3550AECCB04DF1EC" italics="true" pageId="0" pageNumber="89">Glyphiulus portaliformis</emphasis>
|
|
</taxonomicName>
|
|
Zhao & Liu, sp. nov., ♂ paratype
|
|
<emphasis id="217A059954F3E81C95E0F6A0E0A09EB1" bold="true" pageId="0" pageNumber="89">A, B</emphasis>
|
|
anterior body rings, dorsal and lateral views
|
|
<emphasis id="BE43C7F1DDB29FC3F7B36C89D5E8BD7F" bold="true" pageId="0" pageNumber="89">C, D</emphasis>
|
|
mid-body rings, dorsal and lateral views, respectively
|
|
<emphasis id="77C5D71BF803F5FF717952341F116D57" bold="true" pageId="0" pageNumber="89">E, F</emphasis>
|
|
posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="A02FD9DC0E4C2EEBFC0C39C21867A190" doi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89" start="Figure 15" startId="F15">
|
|
<paragraph id="2CF14290B977CB37109FBAC57A90573E" pageId="0" pageNumber="89">
|
|
<emphasis id="1E26B48BF496802BFBA510954B22147E" bold="true" pageId="0" pageNumber="89">Figure 15.</emphasis>
|
|
<taxonomicName id="36DC3F6558AC5E7B1B91697015F37F8D" authorityName="Zhao & Liu" authorityYear="2022" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus portaliformis" order="Spirostreptida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="portaliformis">
|
|
<emphasis id="8A141A16395D831DFF22CCB0A83AD2AB" italics="true" pageId="0" pageNumber="89">Glyphiulus portaliformis</emphasis>
|
|
</taxonomicName>
|
|
Zhao & Liu, sp. nov., ♂ paratype
|
|
<emphasis id="F27C3F989AF1CDCC5C69D5FC5FD71B3D" bold="true" pageId="0" pageNumber="89">A</emphasis>
|
|
gnathochilarium and left antenna
|
|
<emphasis id="5B3A691122D8DA5D2CB032FF5ABDF278" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
head and collum
|
|
<emphasis id="30C84FD1238BD4E51A262F1657D955B3" bold="true" pageId="0" pageNumber="89">C</emphasis>
|
|
penes
|
|
<emphasis id="843787470D451373C1B9C3D2D4311B4B" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
mid-leg and claw
|
|
<emphasis id="B97534216D0AD010AFCEE9974A8A2F04" bold="true" pageId="0" pageNumber="89">E</emphasis>
|
|
leg 3, caudal view
|
|
<emphasis id="E4D74130540D49F4560D2D1015A2D886" bold="true" pageId="0" pageNumber="89">F</emphasis>
|
|
leg 1, frontal view
|
|
<emphasis id="BAE5E76551C1B05DE260548AA9CE90D1" bold="true" pageId="0" pageNumber="89">G</emphasis>
|
|
♀ paratype, vulvae
|
|
<emphasis id="F7DADAD184023A9454DF4CD02E8538DF" bold="true" pageId="0" pageNumber="89">H</emphasis>
|
|
anterior gonopods, caudal view
|
|
<emphasis id="76E1F3F47D15F7E568BD2E6740ADDEDA" bold="true" pageId="0" pageNumber="89">I, J</emphasis>
|
|
posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (
|
|
<emphasis id="8B1E19C64367B199A264AD352EB9C291" bold="true" pageId="0" pageNumber="89">A, B, D, E</emphasis>
|
|
), 0.2 mm (
|
|
<emphasis id="8EE1863F7B40B0411B50AE034FEDF512" bold="true" pageId="0" pageNumber="89">C, F-J</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="07B83C51DEDA54CC726A0E633059C293" pageId="0" pageNumber="89">
|
|
<emphasis id="3032AB8B85993D449DD9E4D62B5E1123" bold="true" italics="true" pageId="0" pageNumber="89">Collum</emphasis>
|
|
: crests complete and evident; carinotaxy formula I-III+P+M (Figs
|
|
<figureCitation id="E10DE9B6346480DE545C778910F16C6F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">14A, B</figureCitation>
|
|
,
|
|
<figureCitation id="45ADD1AFB1A78E4EAA58896B3C2144A6" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15B</figureCitation>
|
|
). Following metaterga strongly crested; carinotaxy formula 2/2+I/i+3/3 (Fig.
|
|
<figureCitation id="EF859D177F240534B438E9EB32968B78" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">14</figureCitation>
|
|
). Ozoporiferous tubercles very large, coniform. Prozonae delicately alveolate-areolate, fine longitudinal striations in front of stricture. Metazonae with an obvious, corrugate, carved texture (Fig.
|
|
<figureCitation id="859D4DBAAA561569EE4596FA05AB3C25" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">14</figureCitation>
|
|
). Rings 2 and 3 with long pleural flaps.
|
|
</paragraph>
|
|
<paragraph id="C35DC971B46D21F2425CFB292BD687CC" pageId="0" pageNumber="89">
|
|
<emphasis id="B000DCECEBA702278D72347F98E0C9D3" bold="true" italics="true" pageId="0" pageNumber="89">Epiproct</emphasis>
|
|
simple, caudal edge with a central conical protrusion and dorsally with a strong central rounded tubercle (Fig.
|
|
<figureCitation id="E8227BA77A141120B42706C1697FBF3D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">14E</figureCitation>
|
|
). Paraprocts regularly convex, each with several irregular rows of setae. Hypoproct transversely bean-shaped, no setae visible (Fig.
|
|
<figureCitation id="42059460E7EE2D34C462E398DF6E6420" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A, B anterior body rings, dorsal and lateral views C, D mid-body rings, dorsal and lateral views, respectively E, F posterior body rings, dorsal and ventral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1108.85156.figure14" httpUri="https://binary.pensoft.net/fig/705264" pageId="0" pageNumber="89">14F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="004F5E5A06328807C856C7C999F87BBA" pageId="0" pageNumber="89">
|
|
<emphasis id="59DF67286A69BF438BEB08D21C2CCFB3" bold="true" italics="true" pageId="0" pageNumber="89">Legs</emphasis>
|
|
slender, about 1.1 times as long as mid-body height; claw with a small accessory spine at base, about 1/5 as long as claw (Fig.
|
|
<figureCitation id="791C32BBA43E29DC7179E3C622EF6CA4" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15D</figureCitation>
|
|
). ♂ legs 1 very strongly reduced, represented by a sternum showing a pair of small, fused, paramedian, subunciform prongs directed forward, with about ten long setae at base; flanked by strongly separated, rudimentary, 2-segmented leg vestiges, second segment very large, stout, rod-shaped (Fig.
|
|
<figureCitation id="13685BD5746A2F0BE7707DE117FFC04E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15F</figureCitation>
|
|
). ♂ legs 2 slightly hypertrophied, coxae large; penes small, much shorter than coxae, oblong-subtrapeziform, each with two or three strong setae distolaterally (Fig.
|
|
<figureCitation id="9EBF07D9A9F1CDF87F07ADAD555E2DCF" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15C</figureCitation>
|
|
). ♂ legs 3 modified through coxae being especially slender and elongate (Fig.
|
|
<figureCitation id="2267A38156654EAB0E949C3E8B482E73" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15E</figureCitation>
|
|
). Both ♂ femora 6 and 7 normal, without modifications.
|
|
</paragraph>
|
|
<paragraph id="233F4CF87BF06D312F68337E0F8A79F3" pageId="0" pageNumber="89">
|
|
<emphasis id="EA334D00D3EEE58A4404C9A5BA90543D" bold="true" italics="true" pageId="0" pageNumber="89">Anterior gonopods</emphasis>
|
|
(Fig.
|
|
<figureCitation id="C06C7700E8A2985C19255865CD292FC6" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15H</figureCitation>
|
|
) very tall, with a portal-shaped coxosternum with about 16-18 microsetae near distal margin; apicomesal process of coxite subtriangular, tip small, slightly curved inwards. Telopodite very large and clavate, 1-segmented, lateral in position, slightly higher than lateral corner of coxite, with a field of nine microspinules at base and six strong setae apically.
|
|
</paragraph>
|
|
<paragraph id="213397C4B43F0811527460FB7AFACE40" pageId="0" pageNumber="89">
|
|
<emphasis id="C38D2C7B5E8DCDD89A2A0112931EE257" bold="true" italics="true" pageId="0" pageNumber="89">Posterior gonopods</emphasis>
|
|
(Figs
|
|
<figureCitation id="4E19AB42CE403DD56CFDB043817772FA" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15I, J</figureCitation>
|
|
) compact. Coxite nearly parallelogram-shaped, with an elongated field of about 14 basolateral microspinules in frontal view and with a field of 16 median microspinules in caudal view; apical and mesal parts of coxite with dense, strong and curved setae. Lamelliform lobe membranous, with a relatively long, broad, distally foliate flagellum.
|
|
</paragraph>
|
|
<paragraph id="7A87A500DB844179C622F78443156E76" pageId="0" pageNumber="89">
|
|
<emphasis id="CF2EAE231CF4C019C7D47D2288C397FE" bold="true" italics="true" pageId="0" pageNumber="89">Vulvae</emphasis>
|
|
very simple, bare, faintly emarginate medially (Fig.
|
|
<figureCitation id="2978C7A201AFAFDC7068A5D1DEE12C70" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Glyphiulus portaliformis Zhao & Liu, sp. nov., ♂ paratype A gnathochilarium and left antenna B head and collum C penes D mid-leg and claw E leg 3, caudal view F leg 1, frontal view G ♀ paratype, vulvae H anterior gonopods, caudal view I, J posterior gonopod, caudal and frontal views, respectively. Scale bars: 0.5 mm (A, B, D, E), 0.2 mm (C, F-J)." figureDoi="10.3897/zookeys.1108.85156.figure15" httpUri="https://binary.pensoft.net/fig/705265" pageId="0" pageNumber="89">15G</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5736B9BB80A31E13BEFA7B09CA62586A" pageId="0" pageNumber="89" type="remarks">
|
|
<paragraph id="F98B1A445A67790334CCAEA21515368A" pageId="0" pageNumber="89">Remark.</paragraph>
|
|
<paragraph id="779541D3521A043ED06C0BC8309251EA" pageId="0" pageNumber="89">In the absence of direct troglomorphic traits, this species can only be considered as troglophilic at most.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |