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<document id="2E7420F63E11579359840D31F32E1C81" ID-DOI="10.11646/zootaxa.5555.1.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="14508653" ID-ZooBank="B9A7D42F-3518-40CF-BD83-4DAD5088CCBB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1734446921180" checkinUser="plazi" docAuthor="Schneider, Scott A. &amp; Lapolla, John S." docDate="2024" docId="1A3487C0FFCCFF8EA0E1FBD6FA7D5545" docLanguage="en" docName="zootaxa.5555.1.9.pdf" docOrigin="Zootaxa 5555 (1)" docSource="https://doi.org/10.11646/zootaxa.5555.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coccidella advena Schneider &amp; LaPolla 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="128" masterDocId="E60DFFB8FFCDFF8DA076FFCDFFE1504D" masterDocTitle="New species of Coccidella Hambleton and Rhizoecus Kunckel dHerculais from South America (Hemiptera: Rhizoecidae)" masterLastPageNumber="133" masterPageNumber="125" pageNumber="126" updateTime="1734955945360" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="4E20F97AA15378CBEE5E2196DDB36106">New species of Coccidella Hambleton and Rhizoecus Kunckel dHerculais from South America (Hemiptera: Rhizoecidae)</mods:title>
</mods:titleInfo>
<mods:name id="0A91C6D36828368A1F0058EE5A71401A" type="personal">
<mods:role id="75960711685621736E8BD2F1745C6E10">
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<mods:namePart id="45A2322ED8B633392D0150AFF8DB6183">Schneider, Scott A.</mods:namePart>
<mods:affiliation id="717D0AA8A58C1CC290F108075724917C">United States Department of Agriculture, Agricultural Research Service, Henry A. Wallace Beltsville Agricultural Research Center, Systematic Entomology Laboratory, 10300 Baltimore Ave, Beltsville, MD 20705, USA</mods:affiliation>
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<mods:namePart id="11E87B13903339FDF3AC7BC2AA18EA1F">Lapolla, John S.</mods:namePart>
<mods:affiliation id="CBDC4EF71222216A9A14AC7255C3BAAF">Department of Biological Sciences, Towson University, 8000 York Rd, Towson, MD 21252, USA</mods:affiliation>
</mods:name>
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<mods:title id="DA8B738AA5D3D5796555747DA594E5A9">Zootaxa</mods:title>
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<mods:date id="D69B55D7C360200723B2B07648EF49A8">2024</mods:date>
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<mods:number id="3C8E9D3CD99249702307F9D937DA4191">2024-12-17</mods:number>
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<mods:number id="D8D3710ED6B5ECBE1FD8E7DCF9A8B3A2">5555</mods:number>
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<mods:url id="CC16BA3E4DCFC04ACC197BF3CEE4288E">https://doi.org/10.11646/zootaxa.5555.1.9</mods:url>
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<mods:classification id="83C99F812F92BC41A370E0F6E04B8EC8">journal article</mods:classification>
<mods:identifier id="4FCF03A3D6F3D1A9F2BC51EC0C62092C" type="DOI">10.11646/zootaxa.5555.1.9</mods:identifier>
<mods:identifier id="1B548AA4A851CCA64B868DB1EED55472" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="1A3487C0FFCCFF8EA0E1FBD6FA7D5545" LSID="urn:lsid:plazi:treatment:1A3487C0FFCCFF8EA0E1FBD6FA7D5545" httpUri="http://treatment.plazi.org/id/1A3487C0FFCCFF8EA0E1FBD6FA7D5545" lastPageId="3" lastPageNumber="128" pageId="1" pageNumber="126">
<subSubSection id="DA87655DFFCCFF8CA0E1FBD6FD0C5478" box="[151,749,1051,1078]" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="922236D6FFCCFF8CA0E1FBD6FD0C5478" blockId="1.[151,749,1051,1113]" box="[151,749,1051,1078]" pageId="1" pageNumber="126">
<heading id="C96A81BAFFCCFF8CA0E1FBD6FD0C5478" bold="true" box="[151,749,1051,1078]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
<emphasis id="A0E9EAC4FFCCFF8CA0E1FBD6FD0C5478" bold="true" box="[151,749,1051,1078]" pageId="1" pageNumber="126">
<taxonomicName id="559D4D55FFCCFF8CA0E1FBD6FD625478" authority="Schneider &amp; LaPolla" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[151,643,1051,1078]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="advena" status="sp. nov.">
<emphasis id="A0E9EAC4FFCCFF8CA0E1FBD6FE95547B" bold="true" box="[151,372,1051,1078]" italics="true" pageId="1" pageNumber="126">Coccidella advena</emphasis>
Schneider &amp; LaPolla
</taxonomicName>
,
<taxonomicNameLabel id="BBDA57BFFFCCFF8CA2E6FBD6FD0C5478" box="[656,749,1051,1077]" pageId="1" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA87655DFFCCFF8CA0E1FBF2FF385414" box="[151,217,1087,1113]" pageId="1" pageNumber="126" type="description">
<paragraph id="922236D6FFCCFF8CA0E1FBF2FF385414" blockId="1.[151,749,1051,1113]" box="[151,217,1087,1113]" pageId="1" pageNumber="126">
<figureCitation id="0AA62A53FFCCFF8CA0E1FBF2FF385414" box="[151,217,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[199,1389,197,1899]" captionTargetId="figure-22@2.[181,1406,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Coccidella advena sp. nov. Adult female, full body view, illustrated from the holotype and paratypes. Illustration by SAS." figureDoi="http://doi.org/10.5281/zenodo.14508655" httpUri="https://zenodo.org/record/14508655/files/figure.png" pageId="1" pageNumber="126">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA87655DFFCCFF8CA0E1FB4AFB9E557C" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="922236D6FFCCFF8CA0E1FB4AFC9954A4" blockId="1.[151,1437,1159,1906]" pageId="1" pageNumber="126">
<emphasis id="A0E9EAC4FFCCFF8CA0E1FB4AFE9954EC" bold="true" box="[151,376,1159,1185]" pageId="1" pageNumber="126">Material examined</emphasis>
.
<materialsCitation id="22F53C8BFFCCFF8CA1F2FB4AFC9954A4" collectingDate="2019-05-31" collectionCode="UNMSM" collectorName="J. S. LaPolla &amp; S. A. Schneider" country="Peru" latitude="-13.0548" location="Manu Paradise Lodge" longLatPrecision="7" longitude="-71.544" municipality="Las Cruces" pageId="1" pageNumber="126" specimenCode="PER01-02" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="A0E9EAC4FFCCFF8CA1F2FB4AFE0D54EF" bold="true" box="[388,492,1159,1186]" italics="true" pageId="1" pageNumber="126">
<typeStatus id="4D268874FFCCFF8CA1F2FB4AFE0D54EF" box="[388,492,1159,1186]" pageId="1" pageNumber="126" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCountry id="EA8A7646FFCCFF8CA18FFB45FDA454EF" box="[505,581,1160,1186]" name="Peru" pageId="1" pageNumber="126">PERU</collectingCountry>
:
<specimenCount id="849BFD5FFFCCFF8CA23BFB4AFD5254EF" box="[589,691,1159,1186]" pageId="1" pageNumber="126">1 adult ♀</specimenCount>
,
<collectingRegion id="5059F834FFCCFF8CA2C9FB4AFC8454EC" box="[703,869,1159,1185]" country="Peru" name="Madre de Dios" pageId="1" pageNumber="126">Madre de Dios</collectingRegion>
,
<collectingMunicipality id="7246ACACFFCCFF8CA318FB45FC0C54EC" box="[878,1005,1159,1186]" pageId="1" pageNumber="126">Las Cruces</collectingMunicipality>
,
<location id="9742600DFFCCFF8CA380FB45FB0A54EC" LSID="urn:lsid:plazi:treatment:1A3487C0FFCCFF8EA0E1FBD6FA7D5545:9742600DFFCCFF8CA380FB45FB0A54EC" box="[1014,1259,1159,1186]" country="Peru" latitude="-13.0548" longLatPrecision="7" longitude="-71.544" municipality="Las Cruces" name="Manu Paradise Lodge" pageId="1" pageNumber="126" stateProvince="Madre de Dios">Manu Paradise Lodge</location>
, from
<taxonomicName id="559D4D55FFCCFF8CA546FB4AFEE65488" authorityName="LaPolla &amp; Schneider" authorityYear="2023" class="Insecta" family="Formicidae" genus="Acropyga" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="manuense">
<emphasis id="A0E9EAC4FFCCFF8CA546FB4AFEE65488" italics="true" pageId="1" pageNumber="126">Acropyga manuense</emphasis>
</taxonomicName>
nest behind lodge, in soil around small rotting branches,
<geoCoordinate id="F7A95011FFCCFF8CA3F1FB61FC0F548B" box="[903,1006,1195,1222]" degrees="13.0548" direction="south" orientation="latitude" pageId="1" pageNumber="126" precision="5" value="-13.0548">-13.0548</geoCoordinate>
,
<geoCoordinate id="F7A95011FFCCFF8CA38FFB61FB81548B" box="[1017,1120,1195,1222]" degrees="71.5440" direction="west" orientation="longitude" pageId="1" pageNumber="126" precision="5" value="-71.544">-71.5440</geoCoordinate>
,
<date id="E6231016FFCCFF8CA41DFB66FB3D5488" box="[1131,1244,1195,1221]" pageId="1" pageNumber="126" value="2019-05-31">
<collectingDate id="F667E9FEFFCCFF8CA41DFB66FB3D5488" box="[1131,1244,1195,1221]" pageId="1" pageNumber="126" value="2019-05-31">31.v.2019</collectingDate>
</date>
,
<collectorName id="3F685300FFCCFF8CA491FB66FA8C5488" box="[1255,1389,1195,1221]" pageId="1" pageNumber="126">J.S. LaPolla</collectorName>
and
<collectorName id="3F685300FFCCFF8CA0E1FB02FEA154A4" box="[151,320,1231,1257]" pageId="1" pageNumber="126">S.A. Schneider</collectorName>
leg.,
<collectionCode id="F48CAE13FFCCFF8CA108FB02FE0954A4" box="[382,488,1231,1257]" pageId="1" pageNumber="126">UNMSM</collectionCode>
(nest ID
<specimenCode id="C23B9EADFFCCFF8CA227FB02FD2854A4" box="[593,713,1231,1257]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="1" pageNumber="126" type="Herbarium">PER01-02</specimenCode>
; prep S0401B).
</materialsCitation>
</paragraph>
<paragraph id="922236D6FFCCFF8CA0B1FB3EFB9E557C" blockId="1.[151,1437,1159,1906]" pageId="1" pageNumber="126">
<materialsCitation id="22F53C8BFFCCFF8CA0B1FB3EFAA45540" box="[199,1349,1267,1294]" collectingDate="2019-05-31" collectionCode="USNM" collectorName="J. S. LaPolla &amp; S. A. Schneider" country="Peru" latitude="-13.0548" location="PERU" longLatPrecision="7" longitude="-71.544" municipality="Las Cruces" pageId="1" pageNumber="126" specimenCode="PER01-02" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="paratype">
<emphasis id="A0E9EAC4FFCCFF8CA0B1FB3EFED85540" bold="true" box="[199,313,1267,1293]" italics="true" pageId="1" pageNumber="126">
<typeStatus id="4D268874FFCCFF8CA0B1FB3EFED85540" box="[199,313,1267,1293]" pageId="1" pageNumber="126" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="EA8A7646FFCCFF8CA130FB39FE715543" box="[326,400,1268,1294]" name="Peru" pageId="1" pageNumber="126">PERU</collectingCountry>
, all with same data as holotype:
<specimenCount id="849BFD5FFFCCFF8CA281FB3EFCBF5543" box="[759,862,1267,1294]" pageId="1" pageNumber="126">1 adult ♀</specimenCount>
,
<collectionCode id="F48CAE13FFCCFF8CA31DFB3EFC5D5540" box="[875,956,1267,1293]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="126" type="Museum">USNM</collectionCode>
(nest ID
<specimenCode id="C23B9EADFFCCFF8CA454FB3EFB7B5540" box="[1058,1178,1267,1293]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="1" pageNumber="126" type="Herbarium">PER01-02</specimenCode>
; prep S0401D)
</materialsCitation>
;
<materialsCitation id="22F53C8BFFCCFF8CA526FB3EFE17557C" collectingDate="2019-05-31" collectionCode="USNM" collectorName="J. S. LaPolla &amp; S. A. Schneider" country="Peru" isTailEnumeration="true" latitude="-13.0548" location="PERU" longLatPrecision="7" longitude="-71.544" municipality="Las Cruces" pageId="1" pageNumber="126" specimenCode="PER01-02" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="849BFD5FFFCCFF8CA526FB3EFF4D557F" pageId="1" pageNumber="126">1 adult ♀</specimenCount>
,
<collectionCode id="F48CAE13FFCCFF8CA0CCFADAFEEA557C" box="[186,267,1303,1329]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="126" type="Museum">USNM</collectionCode>
(nest ID
<specimenCode id="C23B9EADFFCCFF8CA102FADAFE0F557C" box="[372,494,1303,1329]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="1" pageNumber="126" type="Herbarium">PER01-02</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="22F53C8BFFCCFF8CA277FADAFC70557C" box="[513,913,1303,1330]" collectingDate="2019-05-31" collectionCode="USNM" collectorName="J. S. LaPolla &amp; S. A. Schneider" country="Peru" isTailEnumeration="true" latitude="-13.0548" location="PERU" longLatPrecision="7" longitude="-71.544" municipality="Las Cruces" pageId="1" pageNumber="126" specimenCode="PER01-02" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="849BFD5FFFCCFF8CA277FADAFD8B557F" box="[513,618,1303,1330]" pageId="1" pageNumber="126">1 adult ♀</specimenCount>
in poor condition,
<collectionCode id="F48CAE13FFCCFF8CA336FADAFC70557C" box="[832,913,1303,1329]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="126" type="Museum">USNM</collectionCode>
</materialsCitation>
<materialsCitation id="22F53C8BFFCCFF8CA3EEFADAFB9A557C" box="[920,1147,1303,1330]" collectingDate="2019-05-31" collectionCode="PER" collectorName="J. S. LaPolla &amp; S. A. Schneider" country="Peru" isTailEnumeration="true" latitude="-13.0548" location="PERU" longLatPrecision="7" longitude="-71.544" municipality="Las Cruces" pageId="1" pageNumber="126" specimenCode="PER01-02" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="paratype">
(nest ID
<specimenCode id="C23B9EADFFCCFF8CA38CFADAFB92557C" box="[1018,1139,1303,1329]" collectionCode="PER" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="1" pageNumber="126" type="Herbarium">PER01-02</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA87655DFFCCFF8EA0B1FAF6FB915270" lastPageId="3" lastPageNumber="128" pageId="1" pageNumber="126" type="description">
<paragraph id="922236D6FFCCFF8CA0B1FAF6FDA55518" blockId="1.[151,1437,1159,1906]" box="[199,580,1339,1365]" pageId="1" pageNumber="126">
<emphasis id="A0E9EAC4FFCCFF8CA0B1FAF6FEB85518" bold="true" box="[199,345,1339,1365]" pageId="1" pageNumber="126">Description.</emphasis>
Adult female (N=4).
</paragraph>
<paragraph id="922236D6FFCCFF8CA0B1FA92FBAA5537" blockId="1.[151,1437,1159,1906]" box="[199,1099,1375,1402]" pageId="1" pageNumber="126">
<emphasis id="A0E9EAC4FFCCFF8CA0B1FA92FE775534" box="[199,406,1375,1401]" italics="true" pageId="1" pageNumber="126">Appearance in life</emphasis>
. Body white to cream-colored, dusted in powdery white wax.
</paragraph>
<paragraph id="922236D6FFCCFF8CA0B1FA49FBBE573C" blockId="1.[151,1437,1159,1906]" pageId="1" pageNumber="126">
<emphasis id="A0E9EAC4FFCCFF8CA0B1FA49FD4655D0" box="[199,679,1411,1437]" italics="true" pageId="1" pageNumber="126">Description of slide-mounted adult female</emphasis>
. Body elongate oval and membranous,
<quantity id="55659B33FFCCFF8CA419FA4EFAF255D3" box="[1135,1299,1411,1438]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.28" metricValueMin="0.92" pageId="1" pageNumber="126" unit="mm" value="1.1" valueMax="1.28" valueMin="0.92">0.921.28 mm</quantity>
long,
<quantity id="55659B33FFCCFF8CA52BFA4EFF1A558F" metricMagnitude="-4" metricUnit="m" metricValue="6.05" metricValueMax="7.5" metricValueMin="4.6" pageId="1" pageNumber="126" unit="mm" value="0.605" valueMax="0.75" valueMin="0.46">0.46 0.75 mm</quantity>
wide at widest point near abdominal segment I. Eyes present on head margin. Antennae geniculate, each 5 segmented, situated ventrally on submargin of head; overall length 200202 µm; segment lengths (in µm): segment I, 57; II, 1721; III, 3033; IV, 2126; and V, 70. Antennal setae mostly flagellate, each 1542 µm long; longest setae situated on apical segment (V), together with 4 falcate sensory setae. Cephalic plate present and sclerotized. Labium 3 segmented, 107 µm long, 50 µm wide. Anterior and posterior pairs of spiracles each about 16 µm in diameter. Legs well developed, measurements in µm: hind leg 433446, coxa 5967, trochanter + femur 171177, tibia + tarsus 170171, claw 32; ratio of trochanter + femur / tibia + tarsus 1.01.04. Leg setae mostly flagellate, each 1538 µm long, tarsal digitules spine-like, claw digitules simple and shorter than claw (about 6 µm long). Both pairs of ostioles present and heavily sclerotized, each about 50 µm in diameter across orifice; lips of ostioles bearing multiple setae and trilocular pores. Anal lobes indicated by slight projections at posterior end on either side of anal opening, each lobe bearing 3 distinctly enlarged setae, each about 50 µm long when intact but several broken or missing in
<typeStatus id="4D268874FFCCFF8CA165F8F9FEA25703" box="[275,323,1844,1870]" pageId="1" pageNumber="126">type</typeStatus>
series. Anal ring about 48 µm in diameter, with 2 rows of cells, those in outer ring each containing a spicule; ring bearing 6 setae, each 5164 µm long. Internal genital organ sclerotized.
</paragraph>
<caption id="C6E2665EFFCFFF8FA0E1F85AFF0D579E" ID-DOI="http://doi.org/10.5281/zenodo.14508655" ID-Zenodo-Dep="14508655" httpUri="https://zenodo.org/record/14508655/files/figure.png" pageId="2" pageNumber="127" startId="2.[151,250,1943,1967]" targetBox="[199,1389,197,1899]" targetPageId="2" targetType="figure">
<paragraph id="922236D6FFCFFF8FA0E1F85AFF0D579E" blockId="2.[151,1436,1943,2003]" pageId="2" pageNumber="127">
<emphasis id="A0E9EAC4FFCFFF8FA0E1F85AFEF557E2" bold="true" box="[151,276,1943,1967]" pageId="2" pageNumber="127">FIGURE 1.</emphasis>
<taxonomicName id="559D4D55FFCFFF8FA16DF85AFE3957E2" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[283,472,1943,1967]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="advena" status="sp. nov.">
<emphasis id="A0E9EAC4FFCFFF8FA16DF85AFE3957E2" box="[283,472,1943,1967]" italics="true" pageId="2" pageNumber="127">Coccidella advena</emphasis>
</taxonomicName>
<emphasis id="A0E9EAC4FFCFFF8FA1A9F85AFDD157E2" bold="true" box="[479,560,1943,1967]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="BBDA57BFFFCFFF8FA1A9F85AFDD157E2" box="[479,560,1943,1967]" pageId="2" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Adult female, full body view, illustrated from the holotype and paratypes. Illustration by SAS.
</paragraph>
</caption>
<paragraph id="922236D6FFCEFF8EA0B1FF5AFD6051C4" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="128">
<emphasis id="A0E9EAC4FFCEFF8EA0B1FF5AFEEE50FD" box="[199,271,151,176]" italics="true" pageId="3" pageNumber="128">Venter</emphasis>
. Body setae flagellate, 1228 µm long on head, 816 µm on thoracic segments, 1019 µm on abdominal segments. Multilocular disc-pores present near vulva, 21 pores situated primarily on segments VII and VIII, in
<specimenCount id="849BFD5FFFCEFF8EA502FF76FEE050B4" count="1" pageId="3" pageNumber="128" type="generic">one specimen</specimenCount>
2 are present on VI; most pores each apparently with 9 loculi. Tritubular cerores each 89 µm in diameter, situated on submargins of abdominal segments VI and VII and with 1 submedial pair on segment VI. Trilocular pores numerous, distributed among body setae, each about 3.5 µm wide. Enlarged trilocular pores, each about 5 µm wide, in medial clusters, with 1923 pores on abdominal segment IV, 2830 pores on V, and 1319 pores on VI. Oral collar tubular ducts absent. Circulus absent.
</paragraph>
<paragraph id="922236D6FFCEFF8EA0B1FE5EFB915270" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="128">
<emphasis id="A0E9EAC4FFCEFF8EA0B1FE5EFEC051E1" box="[199,289,403,428]" italics="true" pageId="3" pageNumber="128">Dorsum</emphasis>
. Body setae flagellate, 914 µm long on head, 1014 µm on thoracic segments, 1019 on abdominal segments. Multilocular disc-pores absent. Tritubular cerores slightly larger than on venter, each 1213 µm in diameter, situated on submargins of head, prothorax, mesothorax, metathorax, and abdominal segments III, V, VII; also medial cerores present on prothorax, metathorax, and abdominal segments III, V. Trilocular pores numerous, distributed among body setae, each about 3.5 µm wide. Oral collar tubular ducts absent.
</paragraph>
</subSubSection>
<subSubSection id="DA87655DFFCEFF8EA0B1FD8AFC1C52C8" pageId="3" pageNumber="128" type="etymology">
<paragraph id="922236D6FFCEFF8EA0B1FD8AFC1C52C8" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="128">
<emphasis id="A0E9EAC4FFCEFF8EA0B1FD8AFEA9522C" bold="true" box="[199,328,583,609]" pageId="3" pageNumber="128">Etymology</emphasis>
. The epithet
<taxonomicName id="559D4D55FFCEFF8EA19EFD8AFDD8522C" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[488,569,583,609]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="advena">
<emphasis id="A0E9EAC4FFCEFF8EA19EFD8AFDD8522C" box="[488,569,583,609]" italics="true" pageId="3" pageNumber="128">advena</emphasis>
</taxonomicName>
is a Latin noun meaning “visitor” or “stranger”. It signifies that the
<typeStatus id="4D268874FFCEFF8EA51AFD8AFA7D522C" box="[1388,1436,583,609]" pageId="3" pageNumber="128">type</typeStatus>
collection was collected with an ant colony but was unaffiliated with the ants.
</paragraph>
</subSubSection>
<subSubSection id="DA87655DFFCEFF8EA0B1FD43FA7D5545" pageId="3" pageNumber="128" type="discussion">
<paragraph id="922236D6FFCEFF8EA0B1FD43FE015384" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="128">
<emphasis id="A0E9EAC4FFCEFF8EA0B1FD43FEA952E5" bold="true" box="[199,328,654,680]" pageId="3" pageNumber="128">Comments</emphasis>
. The new species is placed in the genus
<taxonomicName id="559D4D55FFCEFF8EA36DFD42FC7552E4" authorityName="Hambleton" authorityYear="1946" box="[795,916,655,681]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="A0E9EAC4FFCEFF8EA36DFD42FC7552E4" box="[795,916,655,681]" italics="true" pageId="3" pageNumber="128">Coccidella</emphasis>
</taxonomicName>
because it possesses characteristic clusters of specialized trilocular pores on the ventral surface of the abdominal segments (
<bibRefCitation id="F60C4B27FFCEFF8EA473FD7EFA705280" author="Kozar, F. &amp; Konczne Benedicty, Z." box="[1029,1425,691,717]" pageId="3" pageNumber="128" refId="ref5767" refString="Kozar, F. &amp; Konczne Benedicty, Z. (2007) Rhizoecinae of the world. Plant Protection Institute, Hungarian Academy of Sciences, Budapest, 617 pp." type="book" year="2007">Kozár &amp; Konczné Benedicty 2007</bibRefCitation>
).
<taxonomicName id="559D4D55FFCEFF8EA0E1FD1AFE8D52BC" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[151,364,727,753]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="advena">
<emphasis id="A0E9EAC4FFCEFF8EA0E1FD1AFE8D52BC" box="[151,364,727,753]" italics="true" pageId="3" pageNumber="128">Coccidella advena</emphasis>
</taxonomicName>
is similar in appearance to
<taxonomicName id="559D4D55FFCEFF8EA2C2FD1AFB7B52BC" authority="Konczne Benedicty &amp; Kozar" authorityName="Konczne Benedicty &amp; Kozar" box="[692,1178,727,753]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="A0E9EAC4FFCEFF8EA2C2FD1AFCA352BC" box="[692,834,727,753]" italics="true" pageId="3" pageNumber="128">C. boliviana</emphasis>
Konczné Benedicty &amp; Kozár
</taxonomicName>
but differs from it by having (character states of
<taxonomicName id="559D4D55FFCEFF8EA1B6FD36FDAB5358" box="[448,586,763,789]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="A0E9EAC4FFCEFF8EA1B6FD36FDAB5358" box="[448,586,763,789]" italics="true" pageId="3" pageNumber="128">C. boliviana</emphasis>
</taxonomicName>
are given in parentheses): no multilocular disc-pores on the thorax (present); enlarged trilocular pores absent on the head (present); and with a ventral submedial pair of tritubular cerores on VI (absent). In a prior publication,
<taxonomicName id="559D4D55FFCEFF8EA18FFC8EFD8B5310" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[505,618,835,861]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="advena">
<emphasis id="A0E9EAC4FFCEFF8EA18FFC8EFD8B5310" box="[505,618,835,861]" italics="true" pageId="3" pageNumber="128">C. advena</emphasis>
</taxonomicName>
was referred to as
<taxonomicName id="559D4D55FFCEFF8EA34BFC8EFC575310" authorityName="Hambleton" authorityYear="1946" box="[829,950,835,861]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="A0E9EAC4FFCEFF8EA34BFC8EFC575310" box="[829,950,835,861]" italics="true" pageId="3" pageNumber="128">Coccidella</emphasis>
</taxonomicName>
near
<taxonomicName id="559D4D55FFCEFF8EA385FC8EFA865310" authority="(Schneider et al. 2022)" baseAuthorityName="Schneider" baseAuthorityYear="2022" box="[1011,1383,835,861]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="A0E9EAC4FFCEFF8EA385FC8EFBBC5310" box="[1011,1117,835,861]" italics="true" pageId="3" pageNumber="128">boliviana</emphasis>
(Schneider
<emphasis id="A0E9EAC4FFCEFF8EA492FC8EFAFD5310" box="[1252,1308,835,861]" italics="true" pageId="3" pageNumber="128">et al.</emphasis>
2022)
</taxonomicName>
. The host plant identity is unknown for this species. The
<typeStatus id="4D268874FFCEFF8EA290FCAAFCF753CC" box="[742,790,871,897]" pageId="3" pageNumber="128">type</typeStatus>
series was collected with a nest of
<taxonomicName id="559D4D55FFCEFF8EA4DAFCAAFADD53CD" authorityName="LaPolla &amp; Schneider" authorityYear="2023" box="[1196,1340,871,897]" class="Insecta" family="Formicidae" genus="Acropyga" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="manuense">
<emphasis id="A0E9EAC4FFCEFF8EA4DAFCAAFADD53CD" box="[1196,1340,871,897]" italics="true" pageId="3" pageNumber="128">A. manuense</emphasis>
</taxonomicName>
but was ignored by the worker ants during a period of observation so the species is presumed to have been living freely in the vicinity of the ant colony.
</paragraph>
<paragraph id="922236D6FFCEFF8EA0B1FC1EFD355478" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="128">
In the key to adult females of
<taxonomicName id="559D4D55FFCEFF8EA26EFC1EFD7053A0" authorityName="Hambleton" authorityYear="1946" box="[536,657,979,1005]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="A0E9EAC4FFCEFF8EA26EFC1EFD7053A0" box="[536,657,979,1005]" italics="true" pageId="3" pageNumber="128">Coccidella</emphasis>
</taxonomicName>
by
<bibRefCitation id="F60C4B27FFCEFF8EA2CDFC1EFC4753A0" author="Kaydan, M. &amp; Konczne Benedicty, Z. &amp; Kondo, D. &amp; Ramos-Portilla, A. A. &amp; Szita, E." box="[699,934,979,1005]" pageId="3" pageNumber="128" pagination="369 - 378" refId="ref5619" refString="Kaydan, M., Konczne Benedicty, Z., Kondo, D., Ramos-Portilla, A. A. &amp; Szita, E. (2018) The genus Coccidella Hambleton (Hemiptera: Rhizoecidae) with description of two new species. Neotropical Entomology, 47, 369 - 378. https: // doi. org / 10.1007 / s 13744 - 017 - 0566 - 3" type="journal article" year="2018">
Kaydan
<emphasis id="A0E9EAC4FFCEFF8EA36FFC1EFCB353A0" box="[793,850,979,1005]" italics="true" pageId="3" pageNumber="128">et al.</emphasis>
(2018)
</bibRefCitation>
,
<taxonomicName id="559D4D55FFCEFF8EA3C4FC1EFBC553A0" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[946,1060,979,1005]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="advena">
<emphasis id="A0E9EAC4FFCEFF8EA3C4FC1EFBC553A0" box="[946,1060,979,1005]" italics="true" pageId="3" pageNumber="128">C. advena</emphasis>
</taxonomicName>
departs from the key at couplet 5 because it has the combination of eyes present and ostiole lips sclerotized. Their key can be modified at this point to accommodate
<taxonomicName id="559D4D55FFCEFF8EA120FBD6FE295478" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[342,456,1051,1077]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="advena">
<emphasis id="A0E9EAC4FFCEFF8EA120FBD6FE295478" box="[342,456,1051,1077]" italics="true" pageId="3" pageNumber="128">C. advena</emphasis>
</taxonomicName>
by using the following:
</paragraph>
<paragraph id="922236D6FFCEFF8EA0E1FB92FA7C5438" blockId="3.[151,1437,1119,1288]" box="[151,1437,1119,1142]" pageId="3" pageNumber="128">
5 Eyes absent........................................................
<taxonomicName id="559D4D55FFCEFF8EA38AFBADFA7C5438" authority="Konczne Benedicty &amp; Kozar" authorityName="Konczne Benedicty &amp; Kozar" box="[1020,1437,1120,1142]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="kissbalazsi">
<emphasis id="A0E9EAC4FFCEFF8EA38AFBADFB615438" box="[1020,1152,1120,1141]" italics="true" pageId="3" pageNumber="128">C. kissbalazsi</emphasis>
Konczné Benedicty &amp; Kozár
</taxonomicName>
</paragraph>
<paragraph id="922236D6FFCEFF8EA0E1FBB0FA7D54DE" blockId="3.[151,1437,1119,1288]" box="[151,1436,1149,1171]" pageId="3" pageNumber="128">- Eyes present......................................................................................... 6</paragraph>
<paragraph id="922236D6FFCEFF8EA0E1FB57FA7D54E2" blockId="3.[151,1437,1119,1288]" box="[151,1436,1178,1200]" pageId="3" pageNumber="128">
6 Ostiole lips sclerotized..................................................
<taxonomicName id="559D4D55FFCEFF8EA46FFB56FAAB54FD" authority="Schneider &amp; LaPolla" authorityName="Schneider &amp; LaPolla" authorityYear="2024" box="[1049,1354,1178,1200]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="advena" status="sp. nov.">
<emphasis id="A0E9EAC4FFCEFF8EA46FFB56FB9B54E2" box="[1049,1146,1178,1200]" italics="true" pageId="3" pageNumber="128">C. advena</emphasis>
Schneider &amp; LaPolla
</taxonomicName>
<emphasis id="A0E9EAC4FFCEFF8EA526FB57FA7D54E2" bold="true" box="[1360,1436,1178,1199]" pageId="3" pageNumber="128">
<taxonomicNameLabel id="BBDA57BFFFCEFF8EA526FB57FA7D54E2" box="[1360,1436,1178,1199]" pageId="3" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="922236D6FFCEFF8EA0E1FB75FA7D5483" blockId="3.[151,1437,1119,1288]" box="[151,1436,1208,1230]" pageId="3" pageNumber="128">- Ostiole lips membranous.............................................................................. 6a</paragraph>
<paragraph id="922236D6FFCEFF8EA0E1FB18FA7D54A6" blockId="3.[151,1437,1119,1288]" box="[151,1436,1237,1259]" pageId="3" pageNumber="128">
6a Antennae 6 segmented; dorsal setae hair-like...........................................
<taxonomicName id="559D4D55FFCEFF8EA4D6FB18FA7D54A6" authority="(Hambleton)" baseAuthorityName="Hambleton" box="[1184,1436,1237,1259]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="globocula">
<emphasis id="A0E9EAC4FFCEFF8EA4D6FB18FAFD54A7" box="[1184,1308,1237,1258]" italics="true" pageId="3" pageNumber="128">C. globocula</emphasis>
(Hambleton)
</taxonomicName>
</paragraph>
<paragraph id="922236D6FFCEFF8EA0E1FB3EFA7D5545" blockId="3.[151,1437,1119,1288]" box="[151,1436,1266,1288]" pageId="3" pageNumber="128">
- Antennae 5 segmented; dorsal setae spine-like...........................................
<taxonomicName id="559D4D55FFCEFF8EA4D3FB3EFA7D5545" authority="Kaydan &amp; Szita" authorityName="Kaydan &amp; Szita" authorityYear="2018" box="[1189,1436,1266,1288]" class="Insecta" family="Pseudococcidae" genus="Coccidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="kozari">
<emphasis id="A0E9EAC4FFCEFF8EA4D3FB3EFB1C554A" box="[1189,1277,1266,1288]" italics="true" pageId="3" pageNumber="128">C. kozari</emphasis>
Kaydan &amp; Szita
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>