232 lines
18 KiB
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232 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.418.7629" ID-PMC="PMC4104466" ID-Pensoft-Pub="1313-2970-418-1" ID-Pensoft-UUID="F034FFA3FFAFFFBEB578C42DD65AFFA2" ID-PubMed="25061358" ID-Zenodo-Dep="578347" ID-ZooBank="D71C8A3DA6CA40A5B3A034A1FD1C16A0" ModsDocID="1313-2970-418-1" checkinTime="1451245725588" checkinUser="pensoft" docAuthor="Skvarla, Michael J., Fisher, J. Ray & Dowling, Ashley P. G." docDate="2014" docId="00DAB24A69BFED12759928F466AC6204" docLanguage="en" docName="ZooKeys 418: 1-103" docOrigin="ZooKeys 418" docPubDate="2014-06-20" docSource="http://dx.doi.org/10.3897/zookeys.418.7629" docTitle="Denheyernaxoides Smiley 1992" docType="treatment" docVersion="5" id="F034FFA3FFAFFFBEB578C42DD65AFFA2" lastPageNumber="23" masterDocId="F034FFA3FFAFFFBEB578C42DD65AFFA2" masterDocTitle="A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records" masterLastPageNumber="103" masterPageNumber="1" pageNumber="21" updateTime="1668158708043" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skvarla, Michael J.</mods:namePart>
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<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, J. Ray</mods:namePart>
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<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
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<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-06-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>418</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>103</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.418.7629</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.418.7629</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-418-1</mods:identifier>
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<mods:identifier type="ZooBank">D71C8A3DA6CA40A5B3A034A1FD1C16A0</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F034FFA3FFAFFFBEB578C42DD65AFFA2</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578347</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152053736" LSID="urn:lsid:plazi:treatment:00DAB24A69BFED12759928F466AC6204" httpUri="http://treatment.plazi.org/id/00DAB24A69BFED12759928F466AC6204" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<subSubSection pageId="20" pageNumber="21" type="nomenclature">
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<paragraph pageId="20" pageNumber="21">
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<taxonomicName LSID="00DAB24A-69BF-ED12-7599-28F466AC6204" authority="Smiley, 1992" authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Denheyernaxoides" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Denheyernaxoides Smiley, 1992</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="historical review">
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<paragraph pageId="20" pageNumber="21">Historical review.</paragraph>
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<paragraph pageId="20" pageNumber="21">
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Canestrini (1885) described
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<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brevirostris">
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<emphasis italics="true" pageId="20" pageNumber="21">Eupalus brevirostris</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Berlese, A" journalOrPublisher="Salmin" pageId="83" pageNumber="84" refId="B36" refString="Berlese, A, 1894. Acari, Myriapoda et (Pseudo) Scorpiones hucusque in Italia Tratr. Salmin 71, 5 pp." title="Acari, Myriapoda et (Pseudo) Scorpiones hucusque in Italia Tratr." volume="71" year="1894">Berlese (1894</bibRefCitation>
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,
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<bibRefCitation author="Berlese, A" journalOrPublisher="Salmin" pageId="83" pageNumber="84" pagination="138 - 139" refId="B37" refString="Berlese, A, 1897. Gill Acari Agrarii. In: Revista patologia vegetale.: 138 - 139" title="Gill Acari Agrarii." volumeTitle="Revista patologia vegetale." year="1897">1897</bibRefCitation>
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) redescribed
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<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brevirostris">
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<emphasis italics="true" pageId="20" pageNumber="21">Eupalus brevirostris</emphasis>
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</taxonomicName>
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and provided illustrations of the dorsal idiosoma, chelicera, and palp.
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<bibRefCitation author="Baker, EW" journalOrPublisher="Anales de la Escuela Nacional de Ciencias Biologicas Mexico" pageId="82" pageNumber="83" pagination="229 - 273" refId="B11" refString="Baker, EW, Hoffmann, A, 1948. Acaros de la familia Cunaxidae. Anales de la Escuela Nacional de Ciencias Biologicas Mexico 5 (3-4): 229 - 273" title="Acaros de la familia Cunaxidae." volume="5" year="1948">Baker and Hoffmann (1948)</bibRefCitation>
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proposed
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<taxonomicName authorityName="Baker & Hoffmann" authorityYear="1948" class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="20" pageNumber="21">Cunaxoides</emphasis>
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</taxonomicName>
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as
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<emphasis italics="true" pageId="20" pageNumber="21">nomen novum</emphasis>
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as
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<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="20" pageNumber="21">Eupalus</emphasis>
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</taxonomicName>
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was preoccupied.
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<bibRefCitation author="Smiley, RL" journalOrPublisher="Indira Publishing House, West Bloomington, Michigan" pageId="93" pageNumber="94" refId="B239" refString="Smiley, RL, 1992. The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomington, Michigan" title="The predatory mite family Cunaxidae (Acari) of the world with a new classification." year="1992">Smiley (1992)</bibRefCitation>
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erected
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<taxonomicName lsidName="" pageId="20" pageNumber="21" rank="subFamily" subFamily="Denheyernaxoidinae">Denheyernaxoidinae</taxonomicName>
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and
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<taxonomicName authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="20" pageNumber="21">Denheyernaxoides</emphasis>
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</taxonomicName>
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for
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<taxonomicName authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="martini">
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<emphasis italics="true" pageId="20" pageNumber="21">Denheyernaxoides martini</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Lin, J" journalOrPublisher="Wuyi Science Journal" pageId="90" pageNumber="91" pagination="1 - 5" refId="B174" refString="Lin, J, 2001. New record and new combination of the genus Denheyernaxoides from Fujian, China (Acari: Cunaxidae). Wuyi Science Journal 17: 1 - 5" title="New record and new combination of the genus Denheyernaxoides from Fujian, China (Acari: Cunaxidae)." volume="17" year="2001">Lin (2001)</bibRefCitation>
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moved transferred
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<taxonomicName class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brevirostris">
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<emphasis italics="true" pageId="20" pageNumber="21">Cunaxoides brevirostris</emphasis>
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</taxonomicName>
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to
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<taxonomicName authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="20" pageNumber="21">Denheyernaxoides</emphasis>
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</taxonomicName>
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and redescribed the species based on specimens from China.
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<bibRefCitation author="Den Heyer, J" journalOrPublisher="Arthropod fauna of the UAE" pageId="86" pageNumber="87" pagination="17 - 25" refId="B92" refString="Den Heyer, J, 2009. . Arthropod fauna of the UAE Vol. 2: 17 - 25" volume="Vol. 2" year="2009">Den Heyer (2009)</bibRefCitation>
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considered
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<taxonomicName lsidName="" pageId="20" pageNumber="21" rank="subFamily" subFamily="Denheyernaxoidinae">Denheyernaxoidinae</taxonomicName>
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as a junior synonym of
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<taxonomicName authorityName="Den Heyer" authorityYear="1978" lsidName="" pageId="20" pageNumber="21" rank="subFamily" subFamily="Cunaxoidinae">Cunaxoidinae</taxonomicName>
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.
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<bibRefCitation author="Den Heyer, J" journalOrPublisher="Zootaxa" pageId="86" pageNumber="87" pagination="1 - 15" refId="B100" refString="Den Heyer, J, Castro, TMMG, 2009. Four new cunaxoidine genera (Acari: Prostigmata: Cunaxidae) and the description of two new Neotropical species. Zootaxa 2140: 1 - 15" title="Four new cunaxoidine genera (Acari: Prostigmata: Cunaxidae) and the description of two new Neotropical species." volume="2140" year="2009">Den Heyer and Castro (2009)</bibRefCitation>
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considered
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<taxonomicName authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="20" pageNumber="21">Denheyernaxoides</emphasis>
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</taxonomicName>
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to belong to
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<taxonomicName lsidName="" pageId="20" pageNumber="21" rank="tribe" tribe="Cunaxoidini">Cunaxoidini</taxonomicName>
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. Sergeyenko (2011) reported
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<taxonomicName baseAuthorityName="Canestrini" baseAuthorityYear="1885" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brevirostris">
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<emphasis italics="true" pageId="20" pageNumber="21">Denheyernaxoides brevirostris</emphasis>
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</taxonomicName>
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from Ukraine and erected
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<taxonomicName lsidName="" pageId="20" pageNumber="21" rank="tribe" tribe="Denheyernaxoidini">Denheyernaxoidini</taxonomicName>
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for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
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<paragraph pageId="21" pageNumber="22">
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<pageBreakToken pageId="21" pageNumber="22" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<emphasis italics="true" pageId="21" pageNumber="22">Gnathosoma</emphasis>
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.
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<emphasis bold="true" pageId="21" pageNumber="22">Pedipalps</emphasis>
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3-segmented. Femurogenua at least twice as long as wide, complemented with 5 setae. Tibiotarsi at least twice as long as wide, usually complemented with 6 setae. A small apophysis occurs basally and a pointed process occurs near the terminal tip; a ridge runs between the apophysis and pointed process.
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<emphasis bold="true" pageId="21" pageNumber="22">Subcapitulum</emphasis>
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with 4 pairs of setae (
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<emphasis italics="true" pageId="21" pageNumber="22">hg1</emphasis>
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-
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<emphasis italics="true" pageId="21" pageNumber="22">4</emphasis>
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); setae
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<emphasis italics="true" pageId="21" pageNumber="22">hg4</emphasis>
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often the longest. Adoral setae absent.
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<emphasis bold="true" pageId="21" pageNumber="22">Chelicera</emphasis>
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without seta.
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</paragraph>
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<paragraph pageId="22" pageNumber="23">
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<emphasis italics="true" pageId="22" pageNumber="23">
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<pageBreakToken pageId="22" pageNumber="23" start="start">Idiosoma</pageBreakToken>
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, dorsal
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</emphasis>
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. Proterosoma lacks a shield, complemented with 2 pairs of setae (
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<emphasis italics="true" pageId="22" pageNumber="23">lps</emphasis>
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and
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<emphasis italics="true" pageId="22" pageNumber="23">mps</emphasis>
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) and 2 pairs of setose sensillae (
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<emphasis italics="true" pageId="22" pageNumber="23">at</emphasis>
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and
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<emphasis italics="true" pageId="22" pageNumber="23">pt</emphasis>
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). Dorsal hysterosoma lacks a plate. Setae
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<emphasis italics="true" pageId="22" pageNumber="23">c1</emphasis>
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-
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<emphasis italics="true" pageId="22" pageNumber="23">h1</emphasis>
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,
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<emphasis italics="true" pageId="22" pageNumber="23">c2</emphasis>
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, and
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<emphasis italics="true" pageId="22" pageNumber="23">f2</emphasis>
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,
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<emphasis italics="true" pageId="22" pageNumber="23">h2</emphasis>
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present. Cupule
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<emphasis italics="true" pageId="22" pageNumber="23">im</emphasis>
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present laterad and posterior of
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<emphasis italics="true" pageId="22" pageNumber="23">e1</emphasis>
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. Integument not covered in shields or plates is striated.
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</paragraph>
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<paragraph pageId="22" pageNumber="23">
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<emphasis italics="true" pageId="22" pageNumber="23">Idiosoma, ventral</emphasis>
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.
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<emphasis bold="true" pageId="22" pageNumber="23">Coxae</emphasis>
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I-II connected by small apodemes. Coxae III-IV fused. Each coxa complemented with 2-4 setae. Genital plates each bear 4 setae (
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<emphasis italics="true" pageId="22" pageNumber="23">g1</emphasis>
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-
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<emphasis italics="true" pageId="22" pageNumber="23">4</emphasis>
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); 2 pairs of genital papillae visible underneath the plates. Anal plates bear 1 pair of setae; 1 pair of setae present ventrally on the integument near the anal plates. 5 pairs of setae present on the integument between the coxal and genital plates. Cupule
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<emphasis italics="true" pageId="22" pageNumber="23">ih</emphasis>
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present ventrally laterad the integumental setae associated with the anal plates. Integument not covered in shields or plates is striated.
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<emphasis bold="true" pageId="22" pageNumber="23">Legs.</emphasis>
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Femora I and II not divided. Trichobothrium on tibia IV absent. Tarsi never constricted apically so as to end in lobes. Ambulacral claws on either side of a 4-rayed empodium present.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="key to adult female denheyernaxoides">
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<paragraph pageId="22" pageNumber="23">
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Key to adult female
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<taxonomicName authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="22" pageNumber="23">Denheyernaxoides</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph pageId="22" pageNumber="23">
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<table inLine="true" pageId="22" pageNumber="23">
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<tr pageId="22" pageNumber="23">
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<td colspan="1" pageId="22" pageNumber="23" rowspan="1">1</td>
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<td colspan="1" pageId="22" pageNumber="23" rowspan="1">Coxa I with 1 sts; trochanters I-IV setal count 1-1-1-1; femora I-II setal count 2-2; gnathosoma with deep indention posterioventrally</td>
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<td colspan="1" pageId="22" pageNumber="23" rowspan="1">
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<taxonomicName authorityName="Smiley" authorityYear="1992" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="martini">
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<emphasis italics="true" pageId="22" pageNumber="23">Denheyernaxoides martini</emphasis>
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</taxonomicName>
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Smiley, 1992
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</td>
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</tr>
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<tr pageId="22" pageNumber="23">
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<td colspan="1" pageId="22" pageNumber="23" rowspan="1">-</td>
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<td colspan="1" pageId="22" pageNumber="23" rowspan="1">Coxa I with 3 sts; trochanters I-IV setal count 0-0-1-0; femora I-II setal count 4-5; gnathosoma with slight indention posterioventrally</td>
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<td colspan="1" pageId="22" pageNumber="23" rowspan="1">
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<taxonomicName baseAuthorityName="Canestrini" baseAuthorityYear="1885" class="Arachnida" family="Cunaxidae" genus="Denheyernaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brevirostris">
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<emphasis italics="true" pageId="22" pageNumber="23">Denheyernaxoides brevirostris</emphasis>
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</taxonomicName>
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(Canestrini 1885)
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |