254 lines
37 KiB
XML
254 lines
37 KiB
XML
<document id="108CDEF8C6FAC6EF0FED6725A5DDB64E" ID-DOI="10.11646/zootaxa.3999.1.7" ID-GBIF-Dataset="12d6ad4d-eb31-4cc5-811e-c9dd1879ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="243357" ID-ZooBank="60B5AB6B-C586-41E5-9370-80BF56025D1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461280129055" checkinUser="plazi" docAuthor="Zamotajlov, Alexandr S., Solodovnikov, Igor A. & Fominykh, Dmitriy D." docDate="2015" docId="03F28B45FFA5FFB1FF555959944EFECC" docLanguage="en" docName="zt03999p124.pdf" docOrigin="Zootaxa 3999 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Aphaonus) mariamae Zamotajlov, Solodovnikov & Fominykh, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="116" masterDocId="FFCBF33DFFA7FFB4FFC25E359157FF8C" masterDocTitle="Description of two new species related to Pterostichus (Aphaonus) arcanoides Lorenz, 1998 from the West Transcaucasia (Coleoptera: Carabidae: Pterostichini)" masterLastPageNumber="124" masterPageNumber="111" pageNumber="113" updateTime="1698613781523" updateUser="plazi">
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<mods:mods id="7B834F98E47545E064C98D8C05447DD5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8FA7648B5F545FFB3B2D4AE89F79D078">
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<mods:title id="DAF5CACDA4CDA8FE3EE53639560522B7">Description of two new species related to Pterostichus (Aphaonus) arcanoides Lorenz, 1998 from the West Transcaucasia (Coleoptera: Carabidae: Pterostichini)</mods:title>
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<mods:roleTerm id="F0E47D94EFF73AF53C2FB419B3FDF86B">Author</mods:roleTerm>
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<mods:namePart id="FA12E2B6E25D71FA641B474510A6BD55">Zamotajlov, Alexandr S.</mods:namePart>
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<mods:namePart id="B2B3E82177B9F5955CDE2DB407435031">Solodovnikov, Igor A.</mods:namePart>
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<mods:roleTerm id="31E56C2DB14AB3B64DB8EB5FD945DC25">Author</mods:roleTerm>
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<mods:namePart id="C28DE63C54AEFC04A40DA96A4B340F22">Fominykh, Dmitriy D.</mods:namePart>
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<mods:typeOfResource id="1E5F8A5CDA7C5015EC43B4699A67110B">text</mods:typeOfResource>
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<mods:title id="6AFE5F318AAB5A475C9534BB064D84D0">Zootaxa</mods:title>
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<mods:date id="C20ECB5D2F7700B74B482C3B5A7121FC">2015</mods:date>
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<mods:number id="2BFA1DC21ABB419B91010E2EEE77273E">3999</mods:number>
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<treatment id="03F28B45FFA5FFB1FF555959944EFECC" ID-DOI="http://doi.org/10.5281/zenodo.6114396" ID-GBIF-Taxon="119640445" ID-Zenodo-Dep="6114396" LSID="urn:lsid:plazi:treatment:03F28B45FFA5FFB1FF555959944EFECC" httpUri="http://treatment.plazi.org/id/03F28B45FFA5FFB1FF555959944EFECC" lastPageId="5" lastPageNumber="116" pageId="2" pageNumber="113">
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<subSubSection id="C34169D8FFA5FFB6FF55595990B5F824" pageId="2" pageNumber="113" type="nomenclature">
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<paragraph id="8BE43A53FFA5FFB6FF55595993FDF80B" blockId="2.[151,682,1900,1960]" box="[151,682,1900,1927]" pageId="2" pageNumber="113">
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<heading id="D0AC8D3FFFA5FFB6FF55595993FDF80B" bold="true" box="[151,682,1900,1927]" fontSize="11" level="1" pageId="2" pageNumber="113" reason="1">
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<emphasis id="B92FE641FFA5FFB6FF55595993FDF80B" bold="true" box="[151,682,1900,1927]" pageId="2" pageNumber="113">
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<taxonomicName id="4C5B41D0FFA5FFB6FF5559599312F80A" ID-CoL="8M7FW" box="[151,581,1900,1927]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov." subGenus="Aphaonus">
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<emphasis id="B92FE641FFA5FFB6FF555959907AF80A" bold="true" box="[151,301,1900,1926]" italics="true" pageId="2" pageNumber="113">Pterostichus</emphasis>
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(
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<emphasis id="B92FE641FFA5FFB6FEFF595990EEF80A" bold="true" box="[317,441,1900,1926]" italics="true" pageId="2" pageNumber="113">Aphaonus</emphasis>
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)
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<emphasis id="B92FE641FFA5FFB6FE0959599312F80A" bold="true" box="[459,581,1900,1926]" italics="true" pageId="2" pageNumber="113">mariamae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C5B3AFFA5FFB6FD8E595893FDF80B" box="[588,682,1901,1927]" pageId="2" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE43A53FFA5FFB6FF5559BA90B5F824" blockId="2.[151,682,1900,1960]" box="[151,482,1935,1960]" pageId="2" pageNumber="113">
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(
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<figureCitation id="136026D6FFA5FFB6FF5D59BA905CF824" box="[159,267,1935,1960]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="2" pageNumber="113">Figs. 3–5</figureCitation>
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,
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<figureCitation id="136026D6FFA5FFB6FED459A5900AF824" box="[278,349,1936,1960]" captionStart="FIGURES 10 – 14" captionStartId="7.[151,264,1460,1483]" captionTargetBox="[169,1412,218,1409]" captionTargetId="figure@7.[151,1436,193,1440]" captionTargetPageId="7" captionText="FIGURES 10 – 14. Pterostichus spp., mentum, dorsal view: 10 — Pterostichus mariamae sp. nov., holotype, male; 11 — Pterostichus mariamae sp. nov., paratype, female; 12 — Pterostichus arcanoides Lorenz, male, Chedym Mt. Range; 13 — Pterostichus arcanoides Lorenz, female, Chedym Mt. Range; 14 — Pterostichus svetlanae sp. nov., holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="2" pageNumber="113">10, 11</figureCitation>
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,
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<figureCitation id="136026D6FFA5FFB6FEAB59BA90E5F824" box="[361,434,1935,1960]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[191,1395,206,1907]" captionTargetId="figure@9.[169,1419,193,1932]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Pterostichus mariamae sp. nov., holotype, male genitalia: 15 – 16 — aedeagus: 15 – left lateral view; 16 — right lateral view; 17 – 18 — parameres: 17 — dorsal view; 18 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243365/files/figure.png" pageId="2" pageNumber="113">15–18</figureCitation>
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,
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<figureCitation id="136026D6FFA5FFB6FE7F59A5908DF824" box="[445,474,1936,1960]" captionStart="FIGURE 23" captionStartId="11.[151,250,938,960]" captionTargetBox="[282,1307,202,913]" captionTargetId="figure@11.[275,1312,193,916]" captionTargetPageId="11" captionText="FIGURE 23. Female head comparative proportions of Pterostichus- species: left— Pt. svetlanae sp. nov.; middle— Pt. mariamae sp. nov.; right— Pt. arcanoides Lorenz." httpUri="https://zenodo.org/record/243367/files/figure.png" pageId="2" pageNumber="113">23</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34169D8FFA5FFB7FF5559E295CDFF74" lastPageId="3" lastPageNumber="114" pageId="2" pageNumber="113" type="materials_examined">
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<paragraph id="8BE43A53FFA5FFB7FF5559E295CDFF74" blockId="2.[151,1436,2007,2032]" lastBlockId="3.[151,1437,151,1977]" lastPageId="3" lastPageNumber="114" pageId="2" pageNumber="113">
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<emphasis id="B92FE641FFA5FFB6FF5559E29013F87C" bold="true" box="[151,324,2007,2032]" pageId="2" pageNumber="113">
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<typeStatus id="54E084F1FFA5FFB6FF5559E29185F87C" box="[151,210,2007,2032]" pageId="2" pageNumber="113">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54E084F1FFA5FFB6FE8859E290E0F87C" box="[330,439,2007,2032]" pageId="2" pageNumber="113" type="holotype">Holotype</typeStatus>
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: male, labelled: Abkhazia, Gulripsh Distr., Tsebelda Plateau near village Ablukhvara, left bank of River Kelasur, slopes of Mt. Bashlug,
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||
<quantity id="4CA397B6FFA4FFB7FD015EA29246FF23" box="[707,785,151,176]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="114" unit="m" value="650.0">650 m</quantity>
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, 15.05.-
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<date id="FFE51C93FFA4FFB7FCAE5EA292A5FF3C" box="[876,1010,151,176]" pageId="3" pageNumber="114" value="2012-07-19">19.07.2012</date>
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||
, pitfall traps (D. Fominykh) (
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||
<collectionCode id="ED4AA296FFA4FFB7FA935EAD94C7FF3C" box="[1361,1424,152,176]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="114">ZISP</collectionCode>
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||
).
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||
<typeStatus id="54E084F1FFA4FFB7FF555E88905EFF59" box="[151,265,189,213]" pageId="3" pageNumber="114" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="9D5DF1DAFFA4FFB7FED05E8990D6FF59" box="[274,385,188,213]" pageId="3" pageNumber="114" type="female">2 females</specimenCount>
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||
, same locality,
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||
<quantity id="4CA397B6FFA4FFB7FDE95E8993FBFF58" box="[555,684,188,213]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="3" pageNumber="114" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500–650 m</quantity>
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||
, 15.05.–
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||
<date id="FFE51C93FFA4FFB7FCCA5E8992DBFF59" box="[776,908,188,213]" pageId="3" pageNumber="114" value="2012-07-19">19.07.2012</date>
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||
, pitfall traps (D. Fominykh) (
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||
<collectionCode id="ED4AA296FFA4FFB7FB165E89944CFF58" box="[1236,1307,188,212]" pageId="3" pageNumber="114">CBIA</collectionCode>
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||
,
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||
<collectionCode id="ED4AA296FFA4FFB7FAE45E89942BFF58" box="[1318,1404,188,212]" pageId="3" pageNumber="114">CZAM</collectionCode>
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||
);
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||
<specimenCount id="9D5DF1DAFFA4FFB7FA4C5E8991B1FF74" pageId="3" pageNumber="114" type="female">1 female</specimenCount>
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||
, same locality,
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||
<quantity id="4CA397B6FFA4FFB7FE505EEA9344FF7B" box="[402,531,223,248]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="114" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500–600 m</quantity>
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||
, 09.04.–
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||
<date id="FFE51C93FFA4FFB7FDB35EEA93A4FF74" box="[625,755,223,248]" pageId="3" pageNumber="114" value="2014-04-06">04.06.2014</date>
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||
, pitfall traps (D. Fominykh) (
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||
<collectionCode id="ED4AA296FFA4FFB7FBFF5ED595D9FF74" box="[1085,1166,224,248]" pageId="3" pageNumber="114">CFDD</collectionCode>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34169D8FFA4FFB7FF055F31936DF835" pageId="3" pageNumber="114" type="description">
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||
<paragraph id="8BE43A53FFA4FFB7FF055F319503FEE9" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
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<emphasis id="B92FE641FFA4FFB7FF055F31900EFE91" bold="true" box="[199,345,260,285]" pageId="3" pageNumber="114">Description.</emphasis>
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Habitus (
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<figureCitation id="136026D6FFA4FFB7FE0C5F319313FE90" box="[462,580,260,285]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="3" pageNumber="114">Figs. 3, 4</figureCitation>
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||
). Large and robust species, body length
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||
<quantity id="4CA397B6FFA4FFB7FBE45F319587FE90" box="[1062,1232,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.435" metricValueMax="2.51" metricValueMin="2.36" pageId="3" pageNumber="114" unit="mm" value="24.35" valueMax="25.1" valueMin="23.6">23.6–25.1 mm</quantity>
|
||
, males
|
||
<quantity id="4CA397B6FFA4FFB7FAEF5F3194CFFE90" box="[1325,1432,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="3" pageNumber="114" unit="mm" value="23.6">23.6 mm</quantity>
|
||
, females
|
||
<quantity id="4CA397B6FFA4FFB7FF375F1290CEFEB3" box="[245,409,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.485" metricValueMax="2.51" metricValueMin="2.46" pageId="3" pageNumber="114" unit="mm" value="24.85" valueMax="25.1" valueMin="24.6">24.6–25.1 mm</quantity>
|
||
long, width
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||
<quantity id="4CA397B6FFA4FFB7FDEA5F1293E6FEB3" box="[552,689,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.2" metricValueMin="5.7" pageId="3" pageNumber="114" unit="mm" value="5.95" valueMax="6.2" valueMin="5.7">5.7–6.2 mm</quantity>
|
||
, males
|
||
<quantity id="4CA397B6FFA4FFB7FCC45F129209FEB3" box="[774,862,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="114" unit="mm" value="5.7">5.7 mm</quantity>
|
||
, females
|
||
<quantity id="4CA397B6FFA4FFB7FC0B5F129506FEB3" box="[969,1105,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.2" metricValueMin="6.1" pageId="3" pageNumber="114" unit="mm" value="6.15" valueMax="6.2" valueMin="6.1">6.1–6.2 mm</quantity>
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||
wide. Dorsum dark brown to black, polished, underside dark brown to black, legs, antennae, and palpi dark brown.
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||
</paragraph>
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||
<paragraph id="8BE43A53FFA4FFB7FF055F5A9208FD41" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
|
||
Head broad, inflated, 0.99 times as long as wide and 0.94–1.01 times as wide as pronotum, nearly square (viewed dorsally), index 2 = 1.00: 0.94: 1.16, genae prominent, compound eyes small, rudimentary, occupying generally lateral position, placed much closer to anterior angle of genae than to temples; temples long, tumid, more or less evenly rounded, post temporal tubercles completely missing, neck broad; frontal furrows prominent, faintly divergent posteriorly and nearly reaching eye level, extending forwards up to clypeal setae, frons with coarsely rugose transversal impression, arcuately connecting frontal furrows; surface smooth, 2 setiferous pores present on each side, anterior one situated in supraorbital groove at eye level, and posterior—at hind extremity of supraorbital groove at mid-temple level, index 1 = 0.75: 1.00: 0.98; tooth of mentum (
|
||
<figureCitation id="136026D6FFA4FFB7FC3B5C5995D6FD08" box="[1017,1153,620,645]" captionStart="FIGURES 10 – 14" captionStartId="7.[151,264,1460,1483]" captionTargetBox="[169,1412,218,1409]" captionTargetId="figure@7.[151,1436,193,1440]" captionTargetPageId="7" captionText="FIGURES 10 – 14. Pterostichus spp., mentum, dorsal view: 10 — Pterostichus mariamae sp. nov., holotype, male; 11 — Pterostichus mariamae sp. nov., paratype, female; 12 — Pterostichus arcanoides Lorenz, male, Chedym Mt. Range; 13 — Pterostichus arcanoides Lorenz, female, Chedym Mt. Range; 14 — Pterostichus svetlanae sp. nov., holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="3" pageNumber="114">Figs. 10, 11</figureCitation>
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||
) rather short, bifid, with rather shallow apical hollow; mandibles rather long with fairly bent apices; antennomere 2 rather long, index 3 = 1.57 (2.26): 1.00 (1.84): 1.66 (2.88): 1.66 (3.15): 1.28 (2.68).
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</paragraph>
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<paragraph id="8BE43A53FFA4FFB7FF055CED945EFC29" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">Pronotum somewhat back-trapeziform, usually longitudinal, 0.99–1.10 times as long as wide, faintly convex, somewhat constricted posteriorly, widest at apex; apex fairly emarginate in the middle, lateral sides narrowly reflexed, inconspicuously rounded up to hind angles, basal margin deeply emarginate in the middle, hind angles prominent, somewhat obtuse, pointed; disk coarsely rugose, anterior transverse impression rather shallow, although prominent, median line deep, reaching both extremities, basal foveae longitudinal, shallow, transversally rugose; lateral margins with 7–10 marginal setae in anterior two-thirds and 1 seta in hind angles at both sides.</paragraph>
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<paragraph id="8BE43A53FFA4FFB7FF055D9A95CFFBD4" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
|
||
Elytra elongate, rather narrow, 1.85–2.08 times as long as wide and 1.13–1.19 times as wide as pronotum, sides subparallel, shoulders rounded but prominent, forming obtuse angles; striae prominent, scutellary striole distinct, intervals subconvex; interval 3 with 4–6 pores usually adjoining stria 3, umbilicate series subinterrupted or rarefied in the middle, forming vague humeral (6–7 pores), median (2 pores), and apical (7–8 pores) groups; microsculpture composed of fine transverse wrinkles and nearly isodiametric meshes.
|
||
<collectingCountry id="F34C7AC3FFA4FFB7FC6D5A0A92B0FBD4" box="[943,999,1087,1112]" name="India" pageId="3" pageNumber="114">Hind</collectingCountry>
|
||
wings reduced.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFA4FFB7FF055A509301FA81" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
|
||
Mesosternum coarsely punctate, proepisterna rather sparsely and smoothly punctate, mesepisterna, metepisterna, and lateral sides of metasternum smooth, lateral areas of sternites rather coarsely rugose. Metatrochantera almost reaching apices of metafemora and somewhat curved apically in males (
|
||
<figureCitation id="136026D6FFA4FFB7FB1B5A99944BFB49" box="[1241,1308,1196,1221]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="3" pageNumber="114">Fig. 5</figureCitation>
|
||
) and about as long as two-thirds of metafemora in females, their apices faintly bent posteriorly. Ventral side of tarsal segment 5 pubescent both dorsally and ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFA4FFB7FF055B229556FAF4" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
|
||
Aedeagus (
|
||
<figureCitation id="136026D6FFA4FFB7FE855B229085FABC" box="[327,466,1303,1328]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[191,1395,206,1907]" captionTargetId="figure@9.[169,1419,193,1932]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Pterostichus mariamae sp. nov., holotype, male genitalia: 15 – 16 — aedeagus: 15 – left lateral view; 16 — right lateral view; 17 – 18 — parameres: 17 — dorsal view; 18 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243365/files/figure.png" pageId="3" pageNumber="114">Figs. 15, 16</figureCitation>
|
||
) more or less distinctly bent at base, normal for the bulk of known species, without ventral swelling; endophallus with rather faintly sclerotized, folded structures. Left paramere wide, right one fairly slender, curving (
|
||
<figureCitation id="136026D6FFA4FFB7FE9B5B6A90B6FAF4" box="[345,481,1375,1400]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[191,1395,206,1907]" captionTargetId="figure@9.[169,1419,193,1932]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Pterostichus mariamae sp. nov., holotype, male genitalia: 15 – 16 — aedeagus: 15 – left lateral view; 16 — right lateral view; 17 – 18 — parameres: 17 — dorsal view; 18 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243365/files/figure.png" pageId="3" pageNumber="114">Figs. 17, 18</figureCitation>
|
||
), both normal in shape and size in the subgenus.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFA4FFB7FF055BB19012F889" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
|
||
<emphasis id="B92FE641FFA4FFB7FF055BB19085FA11" bold="true" box="[199,466,1412,1437]" pageId="3" pageNumber="114">Differential diagnosis.</emphasis>
|
||
The habitus of the new species resembles that of
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FBEF5BB0959EFA10" box="[1069,1225,1412,1436]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFA4FFB7FBEF5BB0959EFA10" box="[1069,1225,1412,1436]" italics="true" pageId="3" pageNumber="114">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
but differs in the following main features (
|
||
<figureCitation id="136026D6FFA4FFB7FE755B929372FA4C" box="[439,549,1447,1472]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="3" pageNumber="114">Figs. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFA4FFB7FDF15B9C9336FA4C" box="[563,609,1448,1472]" captionStart="FIGURES 7 – 9" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[327,1261,213,1908]" captionTargetId="figure@6.[307,1280,193,1932]" captionTargetPageId="6" captionText="FIGURES 7 – 9. Pterostichus arcanoides Lorenz: 7 – 8 — dorsal habitus: 7 — male, Chedym Mt. Range; 8 — female, Chedym Mt. Range; 9 – right metafemur and metatrochanter apices, male, Chedym Mt. Range. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243363/files/figure.png" pageId="3" pageNumber="114">7, 9</figureCitation>
|
||
): plumper species, body more robust, females generally larger and more robust; genae less developed, rudimentary eyes occupy somewhat lateral position (
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FBF35BF89590FA68" box="[1073,1223,1484,1508]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFA4FFB7FBF35BF89590FA68" box="[1073,1223,1484,1508]" italics="true" pageId="3" pageNumber="114">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
possesses stronger tumid and broader genae, eyes occupying dorso-lateral position), temples shorter, more or less gradually rounded (in
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FF7F58209003F9A0" box="[189,340,1556,1580]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFA4FFB7FF7F58209003F9A0" box="[189,340,1556,1580]" italics="true" pageId="3" pageNumber="114">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
temples longer and somewhat angulate, s.
|
||
<figureCitation id="136026D6FFA4FFB7FCF6582192D1F9A0" box="[820,902,1556,1581]" captionStart="FIGURE 23" captionStartId="11.[151,250,938,960]" captionTargetBox="[282,1307,202,913]" captionTargetId="figure@11.[275,1312,193,916]" captionTargetPageId="11" captionText="FIGURE 23. Female head comparative proportions of Pterostichus- species: left— Pt. svetlanae sp. nov.; middle— Pt. mariamae sp. nov.; right— Pt. arcanoides Lorenz." httpUri="https://zenodo.org/record/243367/files/figure.png" pageId="3" pageNumber="114">Fig. 23</figureCitation>
|
||
), tooth of mentum (
|
||
<figureCitation id="136026D6FFA4FFB7FBA15821946BF9A0" box="[1123,1340,1556,1581]" captionStart="FIGURES 10 – 14" captionStartId="7.[151,264,1460,1483]" captionTargetBox="[169,1412,218,1409]" captionTargetId="figure@7.[151,1436,193,1440]" captionTargetPageId="7" captionText="FIGURES 10 – 14. Pterostichus spp., mentum, dorsal view: 10 — Pterostichus mariamae sp. nov., holotype, male; 11 — Pterostichus mariamae sp. nov., paratype, female; 12 — Pterostichus arcanoides Lorenz, male, Chedym Mt. Range; 13 — Pterostichus arcanoides Lorenz, female, Chedym Mt. Range; 14 — Pterostichus svetlanae sp. nov., holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243364/files/figure.png" pageId="3" pageNumber="114">Figs. 10, 11, 12, 13</figureCitation>
|
||
) usually with more shallow apical hollow; lateral margin of pronotum stronger rounded, disk more smooth, coarse rugosity less prominent; metatrochantera (
|
||
<figureCitation id="136026D6FFA4FFB7FDD258699308F9F9" box="[528,607,1628,1653]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="3" pageNumber="114">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFA4FFB7FDAF5869932BF9F8" box="[621,636,1628,1652]" captionStart="FIGURES 7 – 9" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[327,1261,213,1908]" captionTargetId="figure@6.[307,1280,193,1932]" captionTargetPageId="6" captionText="FIGURES 7 – 9. Pterostichus arcanoides Lorenz: 7 – 8 — dorsal habitus: 7 — male, Chedym Mt. Range; 8 — female, Chedym Mt. Range; 9 – right metafemur and metatrochanter apices, male, Chedym Mt. Range. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243363/files/figure.png" pageId="3" pageNumber="114">9</figureCitation>
|
||
) somewhat shorter in both sexes; elytra stronger elongate, lateral sides more parallel, striae somewhat deeper, well developed along their full length (in
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FBFE58B49582F914" box="[1084,1237,1664,1688]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="arcanoides">
|
||
<emphasis id="B92FE641FFA4FFB7FBFE58B49582F914" box="[1084,1237,1664,1688]" italics="true" pageId="3" pageNumber="114">P. arcanoides</emphasis>
|
||
</taxonomicName>
|
||
striae obliterated apically). Aedeagus (
|
||
<figureCitation id="136026D6FFA4FFB7FE465891935CF931" box="[388,523,1700,1725]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[191,1395,206,1907]" captionTargetId="figure@9.[169,1419,193,1932]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Pterostichus mariamae sp. nov., holotype, male genitalia: 15 – 16 — aedeagus: 15 – left lateral view; 16 — right lateral view; 17 – 18 — parameres: 17 — dorsal view; 18 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243365/files/figure.png" pageId="3" pageNumber="114">Figs. 15, 16</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFA4FFB7FDD45891930AF930" box="[534,605,1700,1724]" captionStart="FIGURES 19 – 22" captionStartId="10.[151,264,1952,1975]" captionTargetBox="[225,1365,214,1907]" captionTargetId="figure@10.[201,1385,193,1932]" captionTargetPageId="10" captionText="FIGURES 19 – 22. Pterostichus arcanoides Lorenz, Chedym Mt. Range, male genitalia: 19 – 20 — aedeagus: 19 — left lateral view; 20 — right lateral view; 21 – 22 — parameres: 21 — dorsal view; 22 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243366/files/figure.png" pageId="3" pageNumber="114">19, 20</figureCitation>
|
||
) more distinctly bent basally, apical lamella somewhat longer and broader (viewed laterally); sclerotized elements of endophallus of different structure; parameres (
|
||
<figureCitation id="136026D6FFA4FFB7FB5358F2944BF96C" box="[1169,1308,1735,1760]" captionStart="FIGURES 15 – 18" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[191,1395,206,1907]" captionTargetId="figure@9.[169,1419,193,1932]" captionTargetPageId="9" captionText="FIGURES 15 – 18. Pterostichus mariamae sp. nov., holotype, male genitalia: 15 – 16 — aedeagus: 15 – left lateral view; 16 — right lateral view; 17 – 18 — parameres: 17 — dorsal view; 18 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243365/files/figure.png" pageId="3" pageNumber="114">Figs. 17, 18</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFA4FFB7FAE858FD9424F96C" box="[1322,1395,1736,1760]" captionStart="FIGURES 19 – 22" captionStartId="10.[151,264,1952,1975]" captionTargetBox="[225,1365,214,1907]" captionTargetId="figure@10.[201,1385,193,1932]" captionTargetPageId="10" captionText="FIGURES 19 – 22. Pterostichus arcanoides Lorenz, Chedym Mt. Range, male genitalia: 19 – 20 — aedeagus: 19 — left lateral view; 20 — right lateral view; 21 – 22 — parameres: 21 — dorsal view; 22 — ventral view. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243366/files/figure.png" pageId="3" pageNumber="114">21, 22</figureCitation>
|
||
) of different shape.
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFA4FFB7FF055924936DF835" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="114">
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FF0559249000F8AB" box="[199,343,1808,1832]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFA4FFB7FF0559249000F8AB" box="[199,343,1808,1832]" italics="true" pageId="3" pageNumber="114">P. mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFA4FFB7FEA2593A90EBF8A4" box="[352,444,1807,1832]" pageId="3" pageNumber="114" rank="species">
|
||
<emphasis id="B92FE641FFA4FFB7FEA2593A90EBF8A4" bold="true" box="[352,444,1807,1832]" pageId="3" pageNumber="114">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FD9A592493B7F8A4" box="[600,736,1808,1832]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="svetlanae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFA4FFB7FD9A592493B7F8A4" box="[600,736,1808,1832]" italics="true" pageId="3" pageNumber="114">P. svetlanae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFA4FFB7FD2B593A9212F8A4" box="[745,837,1807,1832]" pageId="3" pageNumber="114" rank="species">
|
||
<emphasis id="B92FE641FFA4FFB7FD2B593A9212F8A4" bold="true" box="[745,837,1807,1832]" pageId="3" pageNumber="114">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the following main features (
|
||
<figureCitation id="136026D6FFA4FFB7FB7E593A947AF8A4" box="[1212,1325,1807,1832]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1952,1975]" captionTargetBox="[332,1257,211,1906]" captionTargetId="figure@4.[311,1276,193,1932]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Pterostichus mariamae sp. nov.: 3 – 4 — dorsal habitus: 3 — holotype, male; 4 — paratype, female; 5 — right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm." httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="3" pageNumber="114">Figs. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="136026D6FFA4FFB7FAF9593A941BF8A4" box="[1339,1356,1807,1832]" captionStart="FIGURE 6" captionStartId="5.[151,250,2000,2022]" captionTargetBox="[513,1436,572,1970]" captionTargetId="figure@5.[502,1085,631,1979]" captionTargetPageId="5" captionText="FIGURE 6. Pterostichus svetlanae sp. nov., dorsal habitus, holotype, female. Scale bar 1.0 mm." httpUri="https://zenodo.org/record/243362/files/figure.png" pageId="3" pageNumber="114">6</figureCitation>
|
||
): body smaller and more slender, head somewhat trapeziform, with more developed and much shorter genae, rudimentary eyes occupying somewhat lateral position (in
|
||
<taxonomicName id="4C5B41D0FFA4FFB7FD08596C9200F8FC" box="[714,855,1880,1904]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="svetlanae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFA4FFB7FD08596C9200F8FC" box="[714,855,1880,1904]" italics="true" pageId="3" pageNumber="114">P. svetlanae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFA4FFB7FCA45962929FF8FC" box="[870,968,1879,1904]" pageId="3" pageNumber="114" rank="species">
|
||
<emphasis id="B92FE641FFA4FFB7FCA45962929FF8FC" bold="true" box="[870,968,1879,1904]" pageId="3" pageNumber="114">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
eyes occupy a dorso-lateral position), antennomere 2 much longer, body and antennomere proportions are different (see below), microreticulation of head and elytra different (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF246ADBFFA3FFB0FF55599592D2F85A" httpUri="https://zenodo.org/record/243361/files/figure.png" pageId="4" pageNumber="115" targetBox="[332,1257,211,1906]" targetPageId="4">
|
||
<paragraph id="8BE43A53FFA3FFB0FF55599592D2F85A" blockId="4.[151,1436,1952,2006]" pageId="4" pageNumber="115">
|
||
<emphasis id="B92FE641FFA3FFB0FF5559959068F83A" bold="true" box="[151,319,1952,1975]" pageId="4" pageNumber="115">FIGURES 3–5.</emphasis>
|
||
<taxonomicName id="4C5B41D0FFA3FFB0FE8B5994936EF83B" box="[329,569,1953,1975]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="mariamae" status="sp. nov.">
|
||
<emphasis id="B92FE641FFA3FFB0FE8B5994936EF83B" box="[329,569,1953,1975]" italics="true" pageId="4" pageNumber="115">Pterostichus mariamae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C5B3AFFA3FFB0FD86599493CFF83A" box="[580,664,1953,1974]" pageId="4" pageNumber="115" rank="species">
|
||
<emphasis id="B92FE641FFA3FFB0FD86599493CFF83A" bold="true" box="[580,664,1953,1974]" pageId="4" pageNumber="115">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 3–4—dorsal habitus: 3—holotype, male; 4—paratype, female; 5—right metafemur and metatrochanter apices, holotype, male. Scale bars 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34169D8FFA2FFB1FF055EA2944EFECC" pageId="5" pageNumber="116" type="distribution">
|
||
<paragraph id="8BE43A53FFA2FFB1FF055EA2959CFF74" blockId="5.[151,1437,151,320]" pageId="5" pageNumber="116">
|
||
<emphasis id="B92FE641FFA2FFB1FF055EA29037FF3C" bold="true" box="[199,352,151,176]" pageId="5" pageNumber="116">Distribution.</emphasis>
|
||
Transcaucasia (= South Caucasus), Abkhazia, Tsebelda Plateau at the left bank of River Kelasur.
|
||
<emphasis id="B92FE641FFA2FFB1FF055E89901DFF59" bold="true" box="[199,330,188,213]" pageId="5" pageNumber="116">Bionomics.</emphasis>
|
||
<typeStatus id="54E084F1FFA2FFB1FE935E8890DEFF59" box="[337,393,189,213]" pageId="5" pageNumber="116">Type</typeStatus>
|
||
specimens were collected in a wet valley forest site on highly pronounced karstic landforms, composed of boxwood bushes and multispecies beech stand at altitudes of
|
||
<quantity id="4CA397B6FFA2FFB1FC1E5EEA950CFF7B" box="[988,1115,223,248]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="5" pageNumber="116" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500–650 m</quantity>
|
||
(
|
||
<figureCitation id="136026D6FFA2FFB1FBA95EEA95E8FF74" box="[1131,1215,223,248]" captionStart="FIGURE 24" captionStartId="11.[151,250,1989,2011]" captionTargetBox="[170,1413,1040,1973]" captionTargetId="figure@11.[170,1414,1040,1973]" captionTargetPageId="11" captionText="FIGURE 24. Habitat of Pterostichus mariamae sp. nov., Mt. Bashlug, 600 m, boxwood bushes." httpUri="https://zenodo.org/record/243368/files/figure.png" pageId="5" pageNumber="116">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE43A53FFA2FFB1FF055F31944EFECC" blockId="5.[151,1437,151,320]" pageId="5" pageNumber="116">
|
||
<emphasis id="B92FE641FFA2FFB1FF055F3190D2FE91" bold="true" box="[199,389,260,285]" pageId="5" pageNumber="116">Specific epithet.</emphasis>
|
||
We are pleased to name this unexpected species after the wife of the third author, Mariam, as a token of gratitude for her patience and continuous support during his collecting trips in the Caucasus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |