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<document id="F757D2F8B1DF3C544B42F9C3D9A302B8" ID-CLB-Dataset="3282" ID-DOI="10.11646/zootaxa.5072.1.4" ID-GBIF-Dataset="465ef1aa-c9de-4ce2-9d75-2a178c9a3487" ID-ISSN="1175-5326" ID-Zenodo-Dep="5728961" ID-ZooBank="7FFA2C8D-6ED2-497E-A55C-05F50417016A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637913539004" checkinUser="plazi" docAuthor="Daniel, Gimo M., Strümpher, Werner P. &amp; Snäll, Stanislav" docDate="2021" docId="038487892514FFECFF493491FC24F9A3" docLanguage="en" docName="zootaxa.5072.1.4.pdf" docOrigin="Zootaxa 5072 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Haroldius lyleae Daniel, Strumpher &amp; Snall 2021, new species" docType="treatment" docVersion="7" lastPageNumber="35" masterDocId="FFBDFFF12515FFEDFFDE305DFFCAFF88" masterDocTitle="A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa" masterLastPageNumber="42" masterPageNumber="34" pageNumber="35" updateTime="1699470339287" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8E7360EBCA459E663C77C67310EB97FC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9A0A83487215DBCFF81B3B7770617226">
<mods:title id="BF5ED6E83B35E2B109C79C5484F66178">A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa</mods:title>
</mods:titleInfo>
<mods:name id="42FAF1423B434788CAB208008B430D55" type="personal">
<mods:role id="226DA96E08AE0324AC9C20F5840C53FF">
<mods:roleTerm id="356E06F3EC1BA53D4E4075B413517EA7">Author</mods:roleTerm>
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<mods:namePart id="432EF0E8F5B8E8C3102493DB00B59F86">Daniel, Gimo M.</mods:namePart>
<mods:affiliation id="0ED6B145464952661AF5FD6194119F0E">Department of Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, South Africa. &amp; Department of Biological &amp; Environmental Sciences, Walter Sisulu University, Mthatha, South Africa.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="D1F10F2FA7B56C99C21222BF250A4A4A">Strümpher, Werner P.</mods:namePart>
<mods:affiliation id="674F6211BCE727ACF3288CE637133806">Ento Studio &amp; Research Collection, Kärrhöksvägen 6, SE- 147 34 Tumba, Sweden.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="1532ADA2E746E9C7D8BBF300DB9B4D6D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB8E161E953BFC34A77721BCD7501A8B">Snäll, Stanislav</mods:namePart>
<mods:affiliation id="450FC6F156F4C897F39B04C9ECF4F08E">Ento Studio &amp; Research Collection, Kärrhöksvägen 6, SE- 147 34 Tumba, Sweden.</mods:affiliation>
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<mods:title id="87DF7937C0D9D266D58E40A766841C2B">Zootaxa</mods:title>
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<mods:part id="A7BDBC6B25688D4CA726138656D5EEC0">
<mods:date id="DDBF6E7F340856911629CF7B4D734B8F">2021</mods:date>
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<mods:number id="28A144D750A20D84391993A7C59B2ED0">2021-11-26</mods:number>
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<mods:identifier id="99E8481EB8930475D136E1568DF815ED" type="DOI">10.11646/zootaxa.5072.1.4</mods:identifier>
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<treatment id="038487892514FFECFF493491FC24F9A3" ID-DOI="http://doi.org/10.5281/zenodo.5729826" ID-GBIF-Taxon="190868760" ID-Zenodo-Dep="5729826" LSID="urn:lsid:plazi:treatment:038487892514FFECFF493491FC24F9A3" httpUri="http://treatment.plazi.org/id/038487892514FFECFF493491FC24F9A3" lastPageNumber="35" pageId="1" pageNumber="35">
<subSubSection id="C33765142514FFECFF493491FC9FFB6E" box="[151,853,1228,1255]" pageId="1" pageNumber="35" type="nomenclature">
<paragraph id="8B92369F2514FFECFF493491FC9FFB6E" blockId="1.[151,853,1228,1291]" box="[151,853,1228,1255]" pageId="1" pageNumber="35">
<heading id="D0DA81F32514FFECFF493491FC9FFB6E" bold="true" box="[151,853,1228,1255]" fontSize="11" level="1" pageId="1" pageNumber="35" reason="1">
<emphasis id="B959EA8D2514FFECFF493491FC9FFB6E" bold="true" box="[151,853,1228,1255]" pageId="1" pageNumber="35">
<taxonomicName id="4C2D4D1C2514FFECFF493491FD72FB6E" ID-CoL="8SSQZ" authority="Daniel, Strumpher &amp; Snall" authorityName="Daniel, Strumpher &amp; Snall" authorityYear="2021" box="[151,696,1228,1255]" class="Insecta" family="Scarabaeidae" genus="Haroldius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="35" phylum="Arthropoda" rank="species" species="lyleae" status="sp. nov.">
<emphasis id="B959EA8D2514FFECFF493491FE95FB6F" bold="true" box="[151,351,1228,1255]" italics="true" pageId="1" pageNumber="35">Haroldius lyleae</emphasis>
Daniel, Strümpher &amp; Snäll
</taxonomicName>
,
<taxonomicNameLabel id="A26A57F62514FFECFD183490FC9FFB6E" box="[710,853,1228,1255]" pageId="1" pageNumber="35" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33765142514FFECFF4934ADFE89FA83" box="[151,323,1264,1291]" pageId="1" pageNumber="35" type="description">
<paragraph id="8B92369F2514FFECFF4934ADFE89FA83" blockId="1.[151,853,1228,1291]" box="[151,323,1264,1291]" pageId="1" pageNumber="35">
(
<figureCitation id="13162A1A2514FFECFF4134ADFED8FA83" box="[159,274,1264,1291]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–9" captionStart-2="FIGURES 10–14" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1943,1967]" captionStartId-2="4.[151,264,1944,1968]" captionTargetBox-0="[184,1403,192,1902]" captionTargetBox-1="[319,1262,198,1908]" captionTargetBox-2="[229,1353,203,1910]" captionTargetId-0="figure-18@2.[171,1417,179,1918]" captionTargetId-1="figure-26@3.[309,1277,181,1920]" captionTargetId-2="figure-18@4.[222,1365,181,1921]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–4. Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species, holotype (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, habitus, lateral view; 4, holotype labels." captionText-1="FIGURES 5–9. Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species, holotype (TMSA). 5, prothorax; 6, pygidium; 7, aedeagus, ventral view; 8, aedeagus, lateral view; 9, aedeagus, dorsal view." captionText-2="FIGURES 10–14. Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species, paratype (ESRC). 10, habitus, dorsal view; 11, paratype labels; 12, habitus, ventral view; 13, clypeus; 14, host ant: Monomorium albopilosum, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.5728963" figureDoi-1="http://doi.org/10.5281/zenodo.5728967" figureDoi-2="http://doi.org/10.5281/zenodo.5728971" httpUri-0="https://zenodo.org/record/5728963/files/figure.png" httpUri-1="https://zenodo.org/record/5728967/files/figure.png" httpUri-2="https://zenodo.org/record/5728971/files/figure.png" pageId="1" pageNumber="35">Figs 1–13</figureCitation>
,
<figureCitation id="13162A1A2514FFECFEC034ADFEF1FA83" box="[286,315,1264,1291]" captionStart="FIGURE 15" captionStartId="5.[151,250,1120,1144]" captionTargetBox="[157,1426,186,1085]" captionTargetId="figure-26@5.[151,1436,181,1096]" captionTargetPageId="5" captionText="FIGURE 15. Distribution of Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species from South Africa (Yellow circles; circle with black point indicate type locality – “Soetvlakte Farm”)." figureDoi="http://doi.org/10.5281/zenodo.5728973" httpUri="https://zenodo.org/record/5728973/files/figure.png" pageId="1" pageNumber="35">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33765142514FFECFF493564FC24F9A3" pageId="1" pageNumber="35" type="materials_examined">
<paragraph id="8B92369F2514FFECFF493564FB3FFADB" blockId="1.[151,1437,1336,1579]" box="[151,1269,1336,1365]" pageId="1" pageNumber="35">
<materialsCitation id="3B453CC22514FFECFF493564FB38FADB" ID-GBIF-Occurrence="3412195305" box="[151,1266,1336,1365]" country="South Africa" latitude="-26.75" location="Northern Cape Province" longLatPrecision="1240" longitude="22.833334" pageId="1" pageNumber="35" specimenCount="1" stateProvince="Northern Cape" typeStatus="holotype">
<emphasis id="B959EA8D2514FFECFF493564FEFAFADA" bold="true" box="[151,304,1336,1363]" pageId="1" pageNumber="35">
<typeStatus id="5496883D2514FFECFF493564FF18FADB" box="[151,210,1337,1363]" pageId="1" pageNumber="35">Type</typeStatus>
locality
</emphasis>
: Soetvlakte Farm [
<geoCoordinate id="EE1950582514FFECFDDB3567FDABFADD" box="[517,609,1338,1365]" degrees="26" direction="south" minutes="45" orientation="latitude" pageId="1" pageNumber="35" precision="925" value="-26.75">26°45′S</geoCoordinate>
<geoCoordinate id="EE1950582514FFECFDB93567FD0DFADD" box="[615,711,1338,1365]" degrees="22" direction="east" minutes="50" orientation="longitude" pageId="1" pageNumber="35" precision="925" value="22.833334">22°50′E</geoCoordinate>
],
<collectingRegion id="49E9F87D2514FFECFD073565FC26FADA" box="[729,1004,1336,1363]" country="South Africa" name="Northern Cape" pageId="1" pageNumber="35">Northern Cape Province</collectingRegion>
,
<collectingCountry id="F33A760F2514FFECFC293565FB4CFADA" box="[1015,1158,1336,1362]" name="South Africa" pageId="1" pageNumber="35">South Africa</collectingCountry>
(
<figureCitation id="13162A1A2514FFECFB4B3565FB22FADB" box="[1173,1256,1336,1363]" captionStart="FIGURE 15" captionStartId="5.[151,250,1120,1144]" captionTargetBox="[157,1426,186,1085]" captionTargetId="figure-26@5.[151,1436,181,1096]" captionTargetPageId="5" captionText="FIGURE 15. Distribution of Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species from South Africa (Yellow circles; circle with black point indicate type locality – “Soetvlakte Farm”)." figureDoi="http://doi.org/10.5281/zenodo.5728973" httpUri="https://zenodo.org/record/5728973/files/figure.png" pageId="1" pageNumber="35">Fig. 15</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B92369F2514FFECFF193500FC24F9A3" blockId="1.[151,1437,1336,1579]" pageId="1" pageNumber="35">
<emphasis id="B959EA8D2514FFECFF193500FEB6FAFE" bold="true" box="[199,380,1372,1399]" pageId="1" pageNumber="35">Type material.</emphasis>
<materialsCitation id="3B453CC22514FFECFE573500FE1CFA6B" ID-GBIF-Occurrence="3412195302" collectionCode="TMSA" collectorName="S. Af. &amp; Robin Lyle" country="South Africa" elevation="1027" location="Soetvlakte Farm" municipality="Province" pageId="1" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="holotype">
<emphasis id="B959EA8D2514FFECFE573500FE35FAFC" bold="true" box="[393,511,1373,1399]" pageId="1" pageNumber="35">
<typeStatus id="5496883D2514FFECFE573500FE35FAFC" box="[393,511,1373,1399]" pageId="1" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
(
<specimenCount id="9D2BFD162514FFECFDC83500FDE7FAFF" box="[534,557,1373,1399]" pageId="1" pageNumber="35" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE5A2514FFECFDE13500FD5AFAFF" box="[575,656,1373,1399]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="1" pageNumber="35">TMSA</collectionCode>
, aedeagus extracted) (
<figureCitation id="13162A1A2514FFECFC423501FBC3FAFF" box="[924,1033,1372,1399]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–9" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1943,1967]" captionTargetBox-0="[184,1403,192,1902]" captionTargetBox-1="[319,1262,198,1908]" captionTargetId-0="figure-18@2.[171,1417,179,1918]" captionTargetId-1="figure-26@3.[309,1277,181,1920]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–4. Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species, holotype (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, habitus, lateral view; 4, holotype labels." captionText-1="FIGURES 5–9. Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species, holotype (TMSA). 5, prothorax; 6, pygidium; 7, aedeagus, ventral view; 8, aedeagus, lateral view; 9, aedeagus, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.5728963" figureDoi-1="http://doi.org/10.5281/zenodo.5728967" httpUri-0="https://zenodo.org/record/5728963/files/figure.png" httpUri-1="https://zenodo.org/record/5728967/files/figure.png" pageId="1" pageNumber="35">Figs 1–9</figureCitation>
): “
<collectorName id="26D853492514FFECFBEE3501FBA6FAFE" box="[1072,1132,1372,1398]" pageId="1" pageNumber="35">S.Af.</collectorName>
[
<collectingCountry id="F33A760F2514FFECFB5E3501FAD5FAFF" box="[1152,1311,1372,1399]" name="South Africa" pageId="1" pageNumber="35">South Africa</collectingCountry>
];
<collectingRegion id="49E9F87D2514FFECFAE93501FF1BFA13" country="South Africa" name="Northern Cape" pageId="1" pageNumber="35">Northern Cape</collectingRegion>
[
<collectingMunicipality id="6BF6ACE52514FFECFF3F35DDFE80FA13" box="[225,330,1408,1435]" pageId="1" pageNumber="35">Province</collectingMunicipality>
] |
<location id="8EF260442514FFECFEB535DDFDE0FA13" LSID="urn:lsid:plazi:treatment:038487892514FFECFF493491FC24F9A3:8EF260442514FFECFEB535DDFDE0FA13" box="[363,554,1408,1435]" country="South Africa" municipality="Province" name="Soetvlakte Farm" pageId="1" pageNumber="35" stateProvince="Northern Cape">Soetvlakte Farm</location>
,
<quantity id="4CD59B7A2514FFECFDE635DCFD4CFA13" box="[568,646,1409,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.027" pageId="1" pageNumber="35" unit="m" value="1027.0">
<elevation id="0000D1AC2514FFECFDE635DCFD4CFA13" box="[568,646,1409,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.027" pageId="1" pageNumber="35" unit="m" value="1027.0">1027m</elevation>
</quantity>
| 26[°].45[̓′]S – 22[°].50[′]E” || “17-19.2[February].2010; E-Y: 3873 |
<collectingMethod id="526C4E882514FFECFF7935F9FF21FA36" box="[167,235,1444,1470]" pageId="1" pageNumber="35">pitfall</collectingMethod>
;Vaalbos in grassl [grassland] | leg.
<collectorName id="26D853492514FFECFD5935F9FCC0FA36" box="[647,778,1444,1470]" pageId="1" pageNumber="35">Robin Lyle</collectorName>
” || “
<typeStatus id="5496883D2514FFECFCE135F8FC66FA35" box="[831,940,1445,1471]" pageId="1" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C2D4D1C2514FFECFC1B35F9FB49FA36" authorityName="Daniel, Strumpher &amp; Snall" authorityYear="2021" box="[965,1155,1444,1470]" class="Insecta" family="Scarabaeidae" genus="Haroldius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="35" phylum="Arthropoda" rank="species" species="lyleae">
<emphasis id="B959EA8D2514FFECFC1B35F9FB49FA36" box="[965,1155,1444,1470]" italics="true" pageId="1" pageNumber="35">Haroldius lyleae</emphasis>
</taxonomicName>
| Daniel, Strümpher &amp; | Snäll 2021 (red cardstock)”
</materialsCitation>
.
<materialsCitation id="3B453CC22514FFECFE3C3594FC21F9A3" ID-GBIF-Occurrence="3412195301" collectingDate="2006-09-02" collectionCode="ESRC, PARA-, TYPE" collectorName="Stanislav Snall" country="South Africa" latitude="-26.241667" location="Springs" longLatPrecision="20" longitude="28.506111" municipality="Province" pageId="1" pageNumber="35" specimenCount="1" stateProvince="Gauteng" typeStatus="paratype">
<emphasis id="B959EA8D2514FFECFE3C3594FD98FA69" bold="true" box="[482,594,1481,1507]" pageId="1" pageNumber="35">
<typeStatus id="5496883D2514FFECFE3C3594FD98FA69" box="[482,594,1481,1507]" pageId="1" pageNumber="35" type="paratype">PARATYPE</typeStatus>
</emphasis>
(female,
<collectionCode id="ED3CAE5A2514FFECFD1F3594FCC1FA6B" box="[705,779,1481,1507]" country="Canada" httpUri="http://grbio.org/cool/etmf-fiec" name="Nova Scotia Department Natural Resources" pageId="1" pageNumber="35">ESRC</collectionCode>
; https://www.inaturalist.org/observations/20160703) (
<figureCitation id="13162A1A2514FFECFAB43595FF28F98F" captionStart="FIGURES 10–14" captionStartId="4.[151,264,1944,1968]" captionTargetBox="[229,1353,203,1910]" captionTargetId="figure-18@4.[222,1365,181,1921]" captionTargetPageId="4" captionText="FIGURES 10–14. Haroldius lyleae Daniel, Strümpher &amp; Snäll, new species, paratype (ESRC). 10, habitus, dorsal view; 11, paratype labels; 12, habitus, ventral view; 13, clypeus; 14, host ant: Monomorium albopilosum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5728971" httpUri="https://zenodo.org/record/5728971/files/figure.png" pageId="1" pageNumber="35">Figs 10–13</figureCitation>
): “
<collectingCountry id="F33A760F2514FFECFF2135B0FEF6F98F" box="[255,316,1517,1543]" name="South Africa" pageId="1" pageNumber="35">RSA</collectingCountry>
:
<collectingRegion id="49E9F87D2514FFECFE9935B0FE6CF98F" box="[327,422,1517,1543]" country="South Africa" name="Gauteng" pageId="1" pageNumber="35">Gauteng</collectingRegion>
pr [
<collectingMunicipality id="6BF6ACE52514FFECFE0F35B1FDF0F98F" box="[465,570,1516,1543]" pageId="1" pageNumber="35">Province</collectingMunicipality>
] |
<location id="8EF260442514FFECFD8A35B1FD60F98E" LSID="urn:lsid:plazi:treatment:038487892514FFECFF493491FC24F9A3:8EF260442514FFECFD8A35B1FD60F98E" box="[596,682,1516,1542]" country="South Africa" latitude="-26.241667" longLatPrecision="20" longitude="28.506111" municipality="Province" name="Springs" pageId="1" pageNumber="35" stateProvince="Gauteng">Springs</location>
<date id="FF93105F2514FFECFD6E35B1FCE8F98E" box="[688,802,1516,1542]" pageId="1" pageNumber="35" value="2006-09-02">
<collectingDate id="EFD7E9B72514FFECFD6E35B1FCE8F98E" box="[688,802,1516,1542]" pageId="1" pageNumber="35" value="2006-09-02">2.IX.2006</collectingDate>
</date>
|
<geoCoordinate id="EE1950582514FFECFCEB35B3FC7EF981" box="[821,948,1518,1545]" degrees="26" direction="south" minutes="14" orientation="latitude" pageId="1" pageNumber="35" precision="15" seconds="30" value="-26.241667">26°14′30″S</geoCoordinate>
,
<geoCoordinate id="EE1950582514FFECFC6735B3FBF4F981" box="[953,1086,1518,1545]" degrees="28" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="35" precision="15" seconds="22" value="28.506111">28°30′22″E</geoCoordinate>
|
<collectorName id="26D853492514FFECFB8F35B1FB33F98E" box="[1105,1273,1516,1542]" pageId="1" pageNumber="35">Stanislav Snäll</collectorName>
leg.” || “
<collectionCode id="ED3CAE5A2514FFECFA8435B0FF02F9A0" pageId="1" pageNumber="35">
<subScript id="17A934DA2514FFECFA8435B0FF02F9A0" attach="left" fontSize="7" pageId="1" pageNumber="35">PARATYPE</subScript>
</collectionCode>
|
<emphasis id="B959EA8D2514FFECFF02364DFE6FF9A3" box="[220,421,1552,1579]" italics="true" pageId="1" pageNumber="35">
<taxonomicName id="4C2D4D1C2514FFECFF02364DFE5CF9A2" authorityName="Daniel, Strumpher &amp; Snall" authorityYear="2021" box="[220,406,1552,1578]" class="Insecta" family="Scarabaeidae" genus="Haroldius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="35" phylum="Arthropoda" rank="species" species="lyleae">Haroldius lyleae</taxonomicName>
|
</emphasis>
Daniel, Strümpher &amp; | Snäll, 2021 (red cardstock)”
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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