180 lines
20 KiB
XML
180 lines
20 KiB
XML
<document id="AB8F41105334269A397F532111892EC5" ID-CLB-Dataset="27906" ID-DOI="10.11646/zootaxa.4525.1.1" ID-GBIF-Dataset="aab5d525-1d70-4a48-9469-43d4e446685d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610805" ID-ZooBank="9D378750-982F-4061-A419-B28E8DDFF825" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553626294178" checkinUser="plazi" docAuthor="E. B, Neo, Adrain, Jonathan M. & Karim, Talia S." docDate="2018" docId="0384871CFF945062FF77FB2EFC1EFE1D" docLanguage="en" docName="zootaxa.4525.1.1.pdf" docOrigin="Zootaxa 4525 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Millardaspis knoxi E.B & Adrain & Karim 2018, n. sp." docType="treatment" docVersion="10" lastPageNumber="41" masterDocId="FFBDFF64FFB3504AFFE0FF82FF8BFFE8" masterDocTitle="The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA" masterLastPageNumber="152" masterPageNumber="1" pageNumber="40" updateTime="1698714183241" updateUser="ExternalLinkService">
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<mods:titleInfo id="F6C68B1DEF816D40C2695348CDE64945">
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<mods:title id="1129065414D4A4711DEA75837604BEC5">The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA</mods:title>
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</mods:titleInfo>
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<mods:name id="159B5B0CEF8B3F8220AD150758813FCB" type="personal">
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<mods:roleTerm id="FEE4677630C0DA542A12BC516BD5DC35">Author</mods:roleTerm>
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<mods:namePart id="2DBDB28FF86E9563580A8BCE1C02A216">E. B, Neo</mods:namePart>
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<mods:namePart id="71CCF9E54C5B1492467E3A93958BE3A6">Adrain, Jonathan M.</mods:namePart>
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<mods:name id="EEF75F0B1F3644BF2878C8E2879E1CC6" type="personal">
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<mods:roleTerm id="B66EDE2D9074DD8B8524C6FE14684249">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="50947A6E90BD704C8BE45E5E62F91FDE">Karim, Talia S.</mods:namePart>
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<mods:typeOfResource id="0F16636DEEA00ABB9AA842C92A4BC173">text</mods:typeOfResource>
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<mods:title id="FDC6CD79F688A7E4086828C13FCA8848">Zootaxa</mods:title>
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<mods:part id="FA63E9AE9AA5073C77A816CC19DB55F0">
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<mods:date id="586B6980DA2AB64226B975EE2E0EE840">2018</mods:date>
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<mods:number id="5027C5EFFB3BD5E5FD09A2D37FFA7EDC">2018-11-27</mods:number>
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<mods:number id="5A613D98A09E32953066EA82D2045CD2">4525</mods:number>
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<treatment id="0384871CFF945062FF77FB2EFC1EFE1D" ID-DOI="http://doi.org/10.5281/zenodo.5969688" ID-GBIF-Taxon="154499796" ID-Zenodo-Dep="5969688" LSID="urn:lsid:plazi:treatment:0384871CFF945062FF77FB2EFC1EFE1D" httpUri="http://treatment.plazi.org/id/0384871CFF945062FF77FB2EFC1EFE1D" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<subSubSection id="C3376581FF94506DFF77FB2EFE4FFB2E" box="[151,452,1196,1222]" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="8B92360AFF94506DFF77FB2EFE4FFB2E" blockId="39.[151,452,1196,1255]" box="[151,452,1196,1222]" pageId="39" pageNumber="40">
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<heading id="D0DA8166FF94506DFF77FB2EFE4FFB2E" bold="true" box="[151,452,1196,1222]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
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<emphasis id="B959EA18FF94506DFF77FB2EFE4FFB2E" bold="true" box="[151,452,1196,1222]" pageId="39" pageNumber="40">
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<taxonomicName id="4C2D4D89FF94506DFF77FB2EFEF0FB2E" authority="E.B & Adrain & Karim, 2018" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[151,379,1196,1222]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="knoxi" status="sp. nov.">
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<emphasis id="B959EA18FF94506DFF77FB2EFEF0FB2E" bold="true" box="[151,379,1196,1222]" italics="true" pageId="39" pageNumber="40">Millardaspis knoxi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26A5763FF94506DFE61FB2EFE4FFB2E" box="[385,452,1196,1222]" pageId="39" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3376581FF94506DFF77FB4CFF79FB0F" box="[151,242,1230,1255]" pageId="39" pageNumber="40" type="description">
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<paragraph id="8B92360AFF94506DFF77FB4CFF79FB0F" blockId="39.[151,452,1196,1255]" box="[151,242,1230,1255]" pageId="39" pageNumber="40">
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<figureCitation id="13162A8FFF94506DFF77FB4CFF79FB0F" box="[151,242,1230,1255]" captionStart="PLATE 55" captionStartId="151.[151,234,1658,1680]" captionText="PLATE 55. Millardaspis knoxi n. sp., from Section H 163.3 m, Fillmore Formation (Floian; Tulean; Panisaspis sevierensis Zone), southern Confusion Range, Ibex area, Millard County, western Utah, USA." pageId="39" pageNumber="40">Plate 55</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF94506DFF77FA94FCF2FA9F" pageId="39" pageNumber="40" type="materials_examined">
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<paragraph id="8B92360AFF94506DFF77FA94FCF2FA9F" blockId="39.[151,1436,1302,2011]" pageId="39" pageNumber="40">
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<emphasis id="B959EA18FF94506DFF77FA94FE8DFAC7" bold="true" box="[151,262,1302,1327]" pageId="39" pageNumber="40">Material.</emphasis>
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<materialsCitation id="3B453C57FF94506DFEEDFA94FCF2FA9F" ID-GBIF-Occurrence="2028522474" collectionCode="SUI" collectorName="Section H & Fillmore Formation" country="United States of America" county="Millard County" location="southern Confusion Range" municipality="Ibex area" pageId="39" pageNumber="40" specimenCode="SUI 129548, SUI 129544-129547, 129549-129552" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
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<typeStatus id="549688A8FF94506DFEEDFA94FEF3FAC7" box="[269,376,1302,1327]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
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, pygidium,
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<specimenCode id="DB8B9E71FF94506DFE1DFA95FD0EFAC7" box="[509,645,1302,1327]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="39" pageNumber="40" type="University or college">SUI 129548</specimenCode>
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(Pl. 55, figs 12, 17, 18) and assigned specimens
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<specimenCode id="DB8B9E71FF94506DFB4BFA95FA13FAC7" box="[1195,1432,1302,1327]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="39" pageNumber="40" type="University or college">SUI 129544–129547</specimenCode>
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,
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<specimenCode id="DB8B9E71FF94506DFF77FAB8FEC6FABB" box="[151,333,1338,1363]" collectionCode="SUI" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="39" pageNumber="40" type="University or college">129549–129552</specimenCode>
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from
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<collectorName id="26D853DCFF94506DFE79FAB8FD87FABA" box="[409,524,1338,1363]" pageId="39" pageNumber="40">Section H</collectorName>
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163.3 m,
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<collectorName id="26D853DCFF94506DFD69FAB8FCE1FABB" box="[649,874,1338,1363]" pageId="39" pageNumber="40">Fillmore Formation</collectorName>
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(Tulean;
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<emphasis id="B959EA18FF94506DFC3DFABEFB57FABB" box="[989,1244,1340,1363]" italics="true" pageId="39" pageNumber="40">Panisaspis sevierensis</emphasis>
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Zone),
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<location id="8EF260D1FF94506DFADBFAB8FED6FA9F" LSID="urn:lsid:plazi:treatment:0384871CFF945062FF77FB2EFC1EFE1D:8EF260D1FF94506DFADBFAB8FED6FA9F" country="United States of America" county="Millard County" municipality="Ibex area" name="southern Confusion Range" pageId="39" pageNumber="40" stateProvince="Utah">southern Confusion Range</location>
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,
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<collectingMunicipality id="6BF6AC70FF94506DFE88FADCFE58FA9E" box="[360,467,1374,1399]" pageId="39" pageNumber="40">Ibex area</collectingMunicipality>
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,
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<collectingCounty id="62F34E86FF94506DFE3EFADCFD07FA9F" box="[478,652,1374,1399]" pageId="39" pageNumber="40">Millard County</collectingCounty>
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, western
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<collectingRegion id="49E9F8E8FF94506DFD15FADCFCA5FA9F" box="[757,814,1374,1399]" country="United States of America" name="Utah" pageId="39" pageNumber="40">Utah</collectingRegion>
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,
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<collectingCountry id="F33A769AFF94506DFCD9FADDFCFEFA9F" box="[825,885,1375,1399]" name="United States of America" pageId="39" pageNumber="40">USA</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF94506DFF27FA00FDDBFA72" box="[199,592,1410,1435]" pageId="39" pageNumber="40" type="etymology">
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<paragraph id="8B92360AFF94506DFF27FA00FDDBFA72" blockId="39.[151,1436,1302,2011]" box="[199,592,1410,1435]" pageId="39" pageNumber="40">
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<emphasis id="B959EA18FF94506DFF27FA00FEC5FA73" bold="true" box="[199,334,1410,1435]" pageId="39" pageNumber="40">Etymology.</emphasis>
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After Benjamin Knox.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF94506DFF27FA24FD6CF9EF" pageId="39" pageNumber="40" type="diagnosis">
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<paragraph id="8B92360AFF94506DFF27FA24FD6CF9EF" blockId="39.[151,1436,1302,2011]" pageId="39" pageNumber="40">
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<emphasis id="B959EA18FF94506DFF27FA24FECAFA57" bold="true" box="[199,321,1446,1471]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
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Librigena with short posterior branch of facial suture; pygidium with very coarse tuberculate sculpture and line of large tubercles running down the middle of each pygidial pleura and onto the spine; shallow and wide pygidial axial furrow; lacks terminal piece.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376581FF945062FF27F990FC1EFE1D" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="discussion">
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<paragraph id="8B92360AFF94506DFF27F990FC7CF937" blockId="39.[151,1436,1302,2011]" pageId="39" pageNumber="40">
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<emphasis id="B959EA18FF94506DFF27F990FEC7F9C3" bold="true" box="[199,332,1554,1579]" pageId="39" pageNumber="40">Discussion.</emphasis>
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<taxonomicName id="4C2D4D89FF94506DFEB3F990FDAFF9C3" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[339,548,1554,1579]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="knoxi">
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<emphasis id="B959EA18FF94506DFEB3F990FDAFF9C3" box="[339,548,1554,1579]" italics="true" pageId="39" pageNumber="40">Millardaspis knoxi</emphasis>
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</taxonomicName>
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is compared to
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<taxonomicName id="4C2D4D89FF94506DFD37F990FC59F9C3" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[727,978,1554,1579]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="milsteadi">
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<emphasis id="B959EA18FF94506DFD37F990FC59F9C3" box="[727,978,1554,1579]" italics="true" pageId="39" pageNumber="40">Millardaspis milsteadi</emphasis>
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</taxonomicName>
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, noting all morphological differences, in the discussion of that species. Compared to the stratigraphically nearest species of the
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<taxonomicName id="4C2D4D89FF94506DFB39F9B4FAC8F9A7" box="[1241,1347,1590,1615]" class="Trilobita" family="Pliomeridae" genus="Ibexaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA18FF94506DFB39F9B4FAC8F9A7" box="[1241,1347,1590,1615]" italics="true" pageId="39" pageNumber="40">Ibexaspis</emphasis>
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</taxonomicName>
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group,
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<taxonomicName id="4C2D4D89FF94506DFF77F9D8FE3AF99B" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[151,433,1626,1651]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="heimbergi">
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<emphasis id="B959EA18FF94506DFF77F9D8FE3AF99B" box="[151,433,1626,1651]" italics="true" pageId="39" pageNumber="40">Deltapliomera heimbergi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2D4D89FF94506DFE5FF9D8FD1AF99B" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[447,657,1626,1651]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="knoxi">
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<emphasis id="B959EA18FF94506DFE5FF9D8FD1AF99B" box="[447,657,1626,1651]" italics="true" pageId="39" pageNumber="40">Millardaspis knoxi</emphasis>
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</taxonomicName>
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(cf. Pl. 24, fig. 1, Pl. 55, fig. 2) has a narrower, more strongly vaulted (sag., tr.) cranidium; a more anteriorly bowed anterior border; a shorter, slightly more inflated glabella with slightly longer and shallower lateral furrows; a longer, more laterally tapered LO; more laterally directed palpebral lobes; shorter and narrower posterior fixigenae; and more densely spaced sculpture.
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</paragraph>
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<paragraph id="8B92360AFF94506DFF27F968FEB2F8A3" blockId="39.[151,1436,1302,2011]" pageId="39" pageNumber="40">
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Librigenae (cf. Pl. 24, figs 16, 17, Pl. 55, fig. 3) of
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<taxonomicName id="4C2D4D89FF94506DFCE7F968FC52F8EB" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[775,985,1770,1795]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="knoxi">
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<emphasis id="B959EA18FF94506DFCE7F968FC52F8EB" box="[775,985,1770,1795]" italics="true" pageId="39" pageNumber="40">Millardaspis knoxi</emphasis>
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</taxonomicName>
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possesses a shorter librigenal field with more tubercles, and a lateral border with a relatively longer anterior projection and coarser sculpture near the border furrow.
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</paragraph>
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<paragraph id="8B92360AFF945062FF27F8D4FCECFF3D" blockId="39.[151,1436,1302,2011]" lastBlockId="40.[151,1436,151,501]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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Thoracic segments (cf. Pl. 25, fig. 27, Pl. 55, fig. 11) mainly differ in that the specimen assigned to
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<taxonomicName id="4C2D4D89FF94506DFF77F8F8FEE2F87B" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[151,361,1914,1939]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="knoxi">
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<emphasis id="B959EA18FF94506DFF77F8F8FEE2F87B" box="[151,361,1914,1939]" italics="true" pageId="39" pageNumber="40">Millardaspis knoxi</emphasis>
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</taxonomicName>
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has a more inflated axial ring and pleurae than that of
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<taxonomicName id="4C2D4D89FF94506DFC2DF8FEFBEBF87B" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[973,1120,1914,1939]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="heimbergi">
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<emphasis id="B959EA18FF94506DFC2DF8FEFBEBF87B" box="[973,1120,1914,1939]" italics="true" pageId="39" pageNumber="40">D. heimbergi</emphasis>
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</taxonomicName>
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; the axial ring is longer; the axial furrows are shallower; there is a transverse line of granules on the posterior margin of the articulating half ring; and the sculpture is also more densely spaced, particularly on the pleural spine. The longer pleural spines of
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<taxonomicName id="4C2D4D89FF9B5062FF77FF15FEE2FF58" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[151,361,151,176]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="knoxi">
|
||
<emphasis id="B959EA18FF9B5062FF77FF15FEE2FF58" box="[151,361,151,176]" italics="true" pageId="40" pageNumber="41">Millardaspis knoxi</emphasis>
|
||
</taxonomicName>
|
||
may be attributable to a different position in the thorax, or may be another difference, but more thoracic segments from both species are needed for comparison.
|
||
</paragraph>
|
||
<paragraph id="8B92360AFF9B5062FF27FF5DFD8DFE45" blockId="40.[151,1436,151,501]" pageId="40" pageNumber="41">
|
||
The pygidia (cf. Pl. 26, fig. 2, Pl. 27, fig. 13, pygidia of Pl. 55) are slightly narrower and much more strongly vaulted (sag., tr.) in both the axis and the pleurae; the pleurae are more strongly backswept, much more strongly downturned, and less splayed; the axial rings are more strongly independently inflated, as are the pleural ribs; the axial furrows are wider, especially across the first and second pleurae; the sculpture is composed of larger, more closely spaced tubercles; and the terminal piece is apparently never expressed, as opposed to its presence in about half of the
|
||
<taxonomicName id="4C2D4D89FF9B5062FEF1FE17FE28FE45" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[273,419,404,429]" class="Trilobita" family="Pliomeridae" genus="Deltapliomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="heimbergi">
|
||
<emphasis id="B959EA18FF9B5062FEF1FE17FE28FE45" box="[273,419,404,429]" italics="true" pageId="40" pageNumber="41">D. heimbergi</emphasis>
|
||
</taxonomicName>
|
||
pygidia.
|
||
</paragraph>
|
||
<paragraph id="8B92360AFF9B5062FF27FE35FC1EFE1D" blockId="40.[151,1436,151,501]" pageId="40" pageNumber="41">
|
||
<taxonomicName id="4C2D4D89FF9B5062FF27FE35FE10FE38" authorityName="E.B & Adrain & Karim" authorityYear="2018" box="[199,411,439,464]" class="Trilobita" family="Pliomeridae" genus="Millardaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="knoxi">
|
||
<emphasis id="B959EA18FF9B5062FF27FE35FE10FE38" box="[199,411,439,464]" italics="true" pageId="40" pageNumber="41">Millardaspis knoxi</emphasis>
|
||
</taxonomicName>
|
||
uniquely (among all members of the
|
||
<taxonomicName id="4C2D4D89FF9B5062FCB0FE35FC31FE38" box="[848,954,439,464]" class="Trilobita" family="Pliomeridae" genus="Ibexaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B959EA18FF9B5062FCB0FE35FC31FE38" box="[848,954,439,464]" italics="true" pageId="40" pageNumber="41">Ibexaspis</emphasis>
|
||
</taxonomicName>
|
||
group) possesses a line of large tubercles running down the middle of each pygidial pleura and onto the spine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |