270 lines
30 KiB
XML
270 lines
30 KiB
XML
<document id="1FBB879628BDCB51EED37926CA348693" ID-DOI="10.11646/zootaxa.4755.1.4" ID-GBIF-Dataset="70aae18d-e6ec-4bb1-abe0-27d1f24fbab8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724261" ID-ZooBank="53E41147-52AA-4A16-BB69-3A9279A303AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968507449" checkinUser="plazi" docAuthor="Barr, Cheryl B. & Shepard, William D." docDate="2020" docId="039CB74BFFD13A40F9DFFA49FE4494E4" docLanguage="en" docName="zootaxa.4755.1.4.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parygrus guarani Barr and Shepard 2020, new species" docType="treatment" docUuid="ADB49206-58A4-48C0-8578-8BC67B175FDB" docUuidSource="ZooBank" docVersion="6" lastPageNumber="117" masterDocId="FFA5CF33FFDE3A52F948FFB8FF87956E" masterDocTitle="Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)" masterLastPageNumber="128" masterPageNumber="99" pageNumber="114" updateTime="1698730760387" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6CC4F16324C977A2D82539EBE106A032">Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)</mods:title>
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<mods:namePart id="B5239DAC8C77E47B40BF00759DA975A8">Barr, Cheryl B.</mods:namePart>
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<mods:roleTerm id="989ECD1A18181403C7CF299D56143127">Author</mods:roleTerm>
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<mods:namePart id="DA0980C4A7334287C104565C34CE5EF9">Shepard, William D.</mods:namePart>
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<mods:title id="62522654C06FB22A4CCCF94F89A95C70">Zootaxa</mods:title>
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<mods:date id="EAC3F3BAAE9648374D5BC673C94B7BD3">2020</mods:date>
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<mods:number id="96AC6B3B58DE5539A834110FF8D3516B">2020-03-23</mods:number>
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<treatment id="039CB74BFFD13A40F9DFFA49FE4494E4" ID-DOI="http://doi.org/10.5281/zenodo.3730215" ID-GBIF-Taxon="162696915" ID-Zenodo-Dep="3730215" LSID="urn:lsid:zoobank.org:act:ADB49206-58A4-48C0-8578-8BC67B175FDB" httpUri="http://treatment.plazi.org/id/039CB74BFFD13A40F9DFFA49FE4494E4" lastPageId="18" lastPageNumber="117" pageId="15" pageNumber="114">
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<subSubSection id="C32F55D6FFD13A5DF9DFFA49FEC49341" pageId="15" pageNumber="114" type="nomenclature">
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<paragraph id="8B8A065DFFD13A5DF9DFFA49FD709365" blockId="15.[151,904,1521,1655]" box="[151,759,1521,1547]" pageId="15" pageNumber="114">
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<emphasis id="B941DA4FFFD13A5DF9DFFA49FD709365" bold="true" box="[151,759,1521,1547]" pageId="15" pageNumber="114">
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<taxonomicName id="4C357DDEFFD13A5DF9DFFA49FDDC9365" authority="Barr and Shepard" authorityName="Barr and Shepard" authorityYear="2020" box="[151,603,1521,1547]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="114" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
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<emphasis id="B941DA4FFFD13A5DF9DFFA49FEEA9365" bold="true" box="[151,365,1521,1547]" italics="true" pageId="15" pageNumber="114">Parygrus guarani</emphasis>
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Barr and Shepard
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2726734FFD13A5DFB2FFA49FD709365" box="[615,759,1521,1547]" pageId="15" pageNumber="114" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8A065DFFD13A5DF9DFF9ADFEC49341" blockId="15.[151,904,1521,1655]" box="[151,323,1556,1583]" pageId="15" pageNumber="114">(Figs 3, 14, 15)</paragraph>
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</subSubSection>
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<subSubSection id="C32F55D6FFD13A5DF9DFF9E4FC0F9319" box="[151,904,1628,1655]" pageId="15" pageNumber="114" type="description">
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<paragraph id="8B8A065DFFD13A5DF9DFF9E4FC0F9319" blockId="15.[151,904,1521,1655]" box="[151,904,1628,1655]" pageId="15" pageNumber="114">
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http://zoobank.org/
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</paragraph>
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</subSubSection>
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<subSubSection id="C32F55D6FFD13A5DF9DFF91DFE2092F9" pageId="15" pageNumber="114" type="materials_examined">
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<paragraph id="8B8A065DFFD13A5DF9DFF91DFF719245" blockId="15.[151,1437,1700,2016]" pageId="15" pageNumber="114">
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<emphasis id="B941DA4FFFD13A5DF9DFF91DFEC393D1" bold="true" box="[151,324,1700,1727]" pageId="15" pageNumber="114">Type material.</emphasis>
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||
<materialsCitation id="3B5D0C00FFD13A5DF802F91DFA1E938D" ID-GBIF-Occurrence="2579576905" collectingDate="1983-02-02" collectingDateMax="1983-02-04" collectingDateMin="1983-02-02" collectionCode="EMEC" collectorName="Coll. E. G. Riley" country="Paraguay" location="Hohenau" pageId="15" pageNumber="114" specimenCount="2" stateProvince="Asuncion" typeStatus="holotype">
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<emphasis id="B941DA4FFFD13A5DF802F91DFD2493D1" bold="true" box="[330,675,1700,1727]" pageId="15" pageNumber="114">
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<typeStatus id="548EB8FFFFD13A5DF802F91DFE3093D1" box="[330,439,1701,1727]" pageId="15" pageNumber="114" type="holotype">Holotype</typeStatus>
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||
male.
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<collectingCountry id="F32246CDFFD13A5DFB4BF91CFD1993D1" box="[515,670,1700,1727]" name="Paraguay" pageId="15" pageNumber="114">PARAGUAY</collectingCountry>
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.
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</emphasis>
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“
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<collectingRegion id="49F1C8BFFFD13A5DFBF3F91DFCD393D1" box="[699,852,1701,1727]" country="Paraguay" name="Asuncion" pageId="15" pageNumber="114">PARAGUAY</collectingRegion>
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:
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<collectingRegion id="49F1C8BFFFD13A5DFA14F91DFC3893D1" box="[860,959,1701,1727]" country="Paraguay" name="Itapua" pageId="15" pageNumber="114">ITAPUA</collectingRegion>
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/
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<quantity id="4CCDABB8FFD13A5DFA9AF91DFB9F93D1" box="[978,1048,1701,1727]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="114" unit="km" value="17.0">17 km</quantity>
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||
N
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||
<location id="8EEA5086FFD13A5DFD70F91DFB1A93D1" LSID="urn:lsid:plazi:treatment:039CB74BFFD13A40F9DFFA49FE4494E4:8EEA5086FFD13A5DFD70F91DFB1A93D1" box="[1080,1181,1701,1727]" country="Paraguay" name="Hohenau" pageId="15" pageNumber="114" stateProvince="Asuncion">Hohenau</location>
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/
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<date id="FF8B209DFFD13A5DFDF8F91DFAD693D1" box="[1200,1361,1701,1727]" pageId="15" pageNumber="114" value="1983-02-02" valueMax="1983-02-04" valueMin="1983-02-02">
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<collectingDate id="EFCFD975FFD13A5DFDF8F91DFAD693D1" box="[1200,1361,1701,1727]" pageId="15" pageNumber="114" value="1983-02-02" valueMax="1983-02-04" valueMin="1983-02-02">Feb. 2-4, 1983</collectingDate>
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</date>
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||
/
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||
<collectorName id="26C0638BFFD13A5DFC2DF91DFE8F938D" pageId="15" pageNumber="114">Coll. E.G.Riley</collectorName>
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||
//
|
||
<typeStatus id="548EB8FFFFD13A5DF86FF971FE3C938D" box="[295,443,1737,1763]" pageId="15" pageNumber="114" type="holotype">HOLOTYPE</typeStatus>
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/
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||
<taxonomicName id="4C357DDEFFD13A5DF89AF971FD16938D" authorityName="Barr and Shepard" authorityYear="2020" box="[466,657,1737,1763]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="114" phylum="Arthropoda" rank="species" species="guarani">Parygrus guarani</taxonomicName>
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||
/ Barr & Shepard [red label, handwritten] // EMEC49556” (
|
||
<collectionCode id="ED249E98FFD13A5DFC75F971FA16938D" box="[1341,1425,1737,1763]" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="15" pageNumber="114" type="University or college">EMEC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<emphasis id="B941DA4FFFD13A5DF9DFF955FEE39269" bold="true" box="[151,356,1772,1799]" pageId="15" pageNumber="114">
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||
<typeStatus id="548EB8FFFFD13A5DF9DFF955FE889269" box="[151,271,1773,1799]" pageId="15" pageNumber="114" type="paratype">Paratypes</typeStatus>
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||
(1 M).
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||
</emphasis>
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||
[as above] //
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||
<materialsCitation id="3B5D0C00FFD13A5DFB4FF955FF759245" ID-GBIF-Occurrence="2579576884" collectionCode="EMEC" collectorName="Barr & Shepard" country="Paraguay" pageId="15" pageNumber="114" specimenCount="1" stateProvince="Asuncion" typeStatus="paratype">
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||
<typeStatus id="548EB8FFFFD13A5DFB4FF955FD159269" box="[519,658,1773,1799]" pageId="15" pageNumber="114" type="paratype">PARATYPE</typeStatus>
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||
/
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||
<taxonomicName id="4C357DDEFFD13A5DFBE5F956FCFF9269" authorityName="Barr and Shepard" authorityYear="2020" box="[685,888,1774,1799]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="114" phylum="Arthropoda" rank="species" species="guarani">
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||
<emphasis id="B941DA4FFFD13A5DFBE5F956FCFF9269" box="[685,888,1774,1799]" italics="true" pageId="15" pageNumber="114">Parygrus guarani</emphasis>
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||
</taxonomicName>
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||
/
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||
<collectorName id="26C0638BFFD13A5DFADBF955FC429269" box="[915,965,1773,1799]" pageId="15" pageNumber="114">Barr</collectorName>
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&
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||
<collectorName id="26C0638BFFD13A5DFAA6F955FBCD9269" box="[1006,1098,1773,1799]" pageId="15" pageNumber="114">Shepard</collectorName>
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||
[yellow label, printed] (1 M,
|
||
<collectionCode id="ED249E98FFD13A5DF9DFF8A9FF6D9245" box="[151,234,1809,1835]" country="USA" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="15" pageNumber="114" type="University or college">EMEC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B8A065DFFD13A5DF98FF88CFE2092F9" blockId="15.[151,1437,1700,2016]" pageId="15" pageNumber="114">
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||
<emphasis id="B941DA4FFFD13A5DF98FF88CFD059221" bold="true" box="[199,642,1844,1871]" pageId="15" pageNumber="114">Additional material examined (2 F).</emphasis>
|
||
[
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<materialsCitation id="3B5D0C00FFD13A5DFBDCF88DFE2492F9" ID-GBIF-Occurrence="2579576892" collectingDate="2016-11-11" collectingDateMax="2016-11-14" collectingDateMin="2016-11-11" collectionCode="LSAM, FSCA" collectorName="J. E. Eger" country="Paraguay" county="Caacupe" elevation="247" location="Morment" municipality="Campamento Jack" pageId="15" pageNumber="114" specimenCount="1" stateProvince="Cordillera">
|
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as above] (1 F,
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||
<collectionCode id="ED249E98FFD13A5DFA07F88DFC259221" box="[847,930,1845,1871]" country="USA" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="15" pageNumber="114">LSAM</collectionCode>
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);
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<collectingCountry id="F32246CDFFD13A5DFAF0F88DFBD69221" box="[952,1105,1845,1871]" name="Paraguay" pageId="15" pageNumber="114">PARAGUAY</collectingCountry>
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:
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<collectingRegion id="49F1C8BFFFD13A5DFD17F88DFB569221" box="[1119,1233,1845,1871]" country="Paraguay" name="Cordillera" pageId="15" pageNumber="114">Cordillera</collectingRegion>
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Dept. /
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||
<collectingCounty id="62EB7ED1FFD13A5DFC7DF88DFA1F9221" box="[1333,1432,1845,1871]" pageId="15" pageNumber="114">Caacupé</collectingCounty>
|
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,
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||
<collectingMunicipality id="6BEE9C27FFD13A5DF9DFF8E1FEE2921D" box="[151,357,1881,1907]" pageId="15" pageNumber="114">Campamento Jack</collectingMunicipality>
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/
|
||
<location id="8EEA5086FFD13A5DF830F8E1FE63921D" LSID="urn:lsid:plazi:treatment:039CB74BFFD13A40F9DFFA49FE4494E4:8EEA5086FFD13A5DF830F8E1FE63921D" box="[376,484,1881,1907]" country="Paraguay" county="Caacupe" municipality="Campamento Jack" name="Morment" pageId="15" pageNumber="114" stateProvince="Cordillera">Morment</location>
|
||
;
|
||
<collectingMethod id="52747E4AFFD13A5DF8A6F8E1FD05921D" box="[494,642,1881,1907]" pageId="15" pageNumber="114">UV light trap</collectingMethod>
|
||
/ S25°22.116′, W057°07.917′/
|
||
<date id="FF8B209DFFD13A5DFA93F8E1FB01921D" box="[987,1158,1881,1907]" pageId="15" pageNumber="114" value="2016-11-11" valueMax="2016-11-14" valueMin="2016-11-11">
|
||
<collectingDate id="EFCFD975FFD13A5DFA93F8E1FB01921D" box="[987,1158,1881,1907]" pageId="15" pageNumber="114" value="2016-11-11" valueMax="2016-11-14" valueMin="2016-11-11">11-14-XI-2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C0638BFFD13A5DFDD9F8E1FB77921D" box="[1169,1264,1881,1907]" pageId="15" pageNumber="114">J.E.Eger</collectorName>
|
||
/
|
||
<quantity id="4CCDABB8FFD13A5DFC4CF8E1FACE921D" box="[1284,1353,1881,1907]" metricMagnitude="2" metricUnit="m" metricValue="2.474976" pageId="15" pageNumber="114" unit="ft" value="812.0">
|
||
<elevation id="0018E16EFFD13A5DFC4CF8E1FACE921D" box="[1284,1353,1881,1907]" metricMagnitude="2" metricUnit="m" metricValue="2.474976" pageId="15" pageNumber="114" unit="ft" value="812.0">812ft.</elevation>
|
||
</quantity>
|
||
, permit #271/2016 (1 F,
|
||
<collectionCode id="ED249E98FFD13A5DF818F8C5FE1C92F9" box="[336,411,1917,1943]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="15" pageNumber="114" type="Museum">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFD13A42F98FF818FEAE94E5" lastPageId="16" lastPageNumber="115" pageId="15" pageNumber="114" type="diagnosis">
|
||
<paragraph id="8B8A065DFFD13A5DF98FF818FA1B92B1" blockId="15.[151,1437,1700,2016]" pageId="15" pageNumber="114">
|
||
<emphasis id="B941DA4FFFD13A5DF98FF818FEC592D5" bold="true" box="[199,322,1952,1979]" pageId="15" pageNumber="114">Diagnosis.</emphasis>
|
||
The male genitalia of
|
||
<taxonomicName id="4C357DDEFFD13A5DFB06F81AFC9E92D5" authorityName="Barr and Shepard" authorityYear="2020" box="[590,793,1954,1979]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="114" phylum="Arthropoda" rank="species" species="guarani">
|
||
<emphasis id="B941DA4FFFD13A5DFB06F81AFC9E92D5" box="[590,793,1954,1979]" italics="true" pageId="15" pageNumber="114">Parygrus guarani</emphasis>
|
||
</taxonomicName>
|
||
are distinctive (Fig. 15); none of the other species (for which the genitalia are known) have a phallobase more than twice as long as the parameres in addition to parameres
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F9DFFF21FEAE94E5" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
with inner teeth. In both the males and females, the body is slender, parallel-sided, and narrowly tapered posteriorly (Fig. 14); nearly all of the other species are slightly narrowed anterior to the middle and widened at the posterior 1/3, and are broadly rounded at the elytral apices.
|
||
<taxonomicName id="4C357DDEFFCE3A42FB85FF5AFC319595" authorityName="Grouvelle" authorityYear="1896" box="[717,950,226,251]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="115" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFCE3A42FB85FF5AFC319595" box="[717,950,226,251]" italics="true" pageId="16" pageNumber="115">Parygrus angustatus</emphasis>
|
||
</taxonomicName>
|
||
is similarly parallel-sided (
|
||
<figureCitation id="130E1AD8FFCE3A42FDA4FF59FAB79595" box="[1260,1328,225,251]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="16" pageNumber="115">Fig. 8</figureCitation>
|
||
), but it is much longer (
|
||
<quantity id="4CCDABB8FFCE3A42F879FEBDFE089471" box="[305,399,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="16" pageNumber="115" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
|
||
) than
|
||
<taxonomicName id="4C357DDEFFCE3A42F89BFEBEFDC09471" authorityName="Barr and Shepard" authorityYear="2020" box="[467,583,262,287]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="115" phylum="Arthropoda" rank="species" species="guarani">
|
||
<emphasis id="B941DA4FFFCE3A42F89BFEBEFDC09471" box="[467,583,262,287]" italics="true" pageId="16" pageNumber="115">P. guarani</emphasis>
|
||
</taxonomicName>
|
||
(less than
|
||
<quantity id="4CCDABB8FFCE3A42FBF4FEBCFC929471" box="[700,789,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="16" pageNumber="115" unit="mm" value="5.6">5.6 mm</quantity>
|
||
). Because of the very small number of specimens on hand, it is possible that the above external characters may not be reliable for diagnosing the species. The two females were assigned to this species due to their association with a male and/or their similar morphology; they are not designated as
|
||
<typeStatus id="548EB8FFFFCE3A42F9FEFEC9FEA294E5" box="[182,293,369,395]" pageId="16" pageNumber="115" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFCE3A40F98FFE2CFEA8942D" lastPageId="18" lastPageNumber="117" pageId="16" pageNumber="115" type="description">
|
||
<paragraph id="8B8A065DFFCE3A42F98FFE2CFE3B9775" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FFE2CFD9D94C1" bold="true" box="[199,538,404,431]" pageId="16" pageNumber="115">
|
||
Description.
|
||
<typeStatus id="548EB8FFFFCE3A42F82BFE2DFE5794C1" box="[355,464,405,431]" pageId="16" pageNumber="115" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
Cuticle dark brown, legs, antennae and mouthparts lighter; slender, elongate, subparallel, elytra dorsally flattened along midline; length
|
||
<quantity id="4CCDABB8FFCE3A42FA75FE00FC2394BD" box="[829,932,440,467]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="115" unit="mm" value="5.5">5.50 mm</quantity>
|
||
(pronotum + elytra), width
|
||
<quantity id="4CCDABB8FFCE3A42FDA3FE01FAD394BD" box="[1259,1364,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="16" pageNumber="115" unit="mm" value="1.9">1.90 mm</quantity>
|
||
, more than 2x as long as wide; body covered with erect and semi-erect, moderately long, golden setae and much shorter, recumbent setae (Fig. 14).
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F98FFD9DFBB29655" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FFD9DFE839751" box="[199,260,549,575]" italics="true" pageId="16" pageNumber="115">Head</emphasis>
|
||
dark brown; punctures separated by 1x puncture diameter; setae moderately long; vertex flat; frons protruding slightly forward between antennal bases, anterior margin barely arcuate, distance between antennal bases less than length of antennomere 1. Antenna with antennomeres 1 and 2 dark brown, setose; antennomeres 3–11 yellow-brown, more densely setose; antennomere 1 slightly shorter than antennomere 2, antennomere 2 covering antennomeres 3–6. Clypeus densely setose, deeply punctate, margin nearly straight. Maxillary palpus with terminal palpomere slightly clavate, ventral face flat. Labrum emarginate, densely setose, bordered with dense fringe of short setae. Labial palpus with terminal palpomere dorsoventrally flattened, vaguely kidney-shaped with one side straight, one side curved; length shorter than that of terminal maxillary palpomere.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F98FFCFEFD20917D" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FFCFEFEB19631" box="[199,310,838,863]" italics="true" pageId="16" pageNumber="115">Pronotum</emphasis>
|
||
dark brown; convex, disc weakly flattened at center; length
|
||
<quantity id="4CCDABB8FFCE3A42FA9BFCFDFBBF9631" box="[979,1080,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="115" unit="mm" value="1.2">1.20 mm</quantity>
|
||
, width
|
||
<quantity id="4CCDABB8FFCE3A42FDC1FCFCFB699631" box="[1161,1262,836,863]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="16" pageNumber="115" unit="mm" value="1.55">1.55 mm</quantity>
|
||
, widest at basal 1/3 to 1/2; anterior margin straight except at strongly acute, depressed, anterolateral angles; lateral margins weakly arcuate, sinuate at base, narrowly margined, fringed with erect setae; posterior border trisinuate; posterolateral angles acute, moderately explanate. Disc evenly punctate, punctures separated by about 1x puncture diameter; setae moderately long.
|
||
<taxonomicName id="4C357DDEFFCE3A42F813FC6DFE4E9681" box="[347,457,981,1007]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="16" pageNumber="115" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFCE3A42F813FC6DFE4E9681" box="[347,457,981,1007]" italics="true" pageId="16" pageNumber="115">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
dark brown, subcordate, wider than long; anterior margin strongly arcuate between two anterolateral notches; disc flat; punctation fine.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F98FFBA5FC6B9185" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FFBA5FE9D9159" box="[199,282,1053,1079]" italics="true" pageId="16" pageNumber="115">Elytron</emphasis>
|
||
dark brown; length
|
||
<quantity id="4CCDABB8FFCE3A42FB4CFBA5FDEC9159" box="[516,619,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="115" unit="mm" value="4.3">4.30 mm</quantity>
|
||
, width
|
||
<quantity id="4CCDABB8FFCE3A42FB89FBA4FCAE9159" box="[705,809,1052,1079]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="16" pageNumber="115" unit="mm" value="0.95">0.95 mm</quantity>
|
||
; flattened; little variation in width except at apical 1/3. Humeral angle rounded; lateral margin widened and shallowly sulcate at basal 1/3, entire length narrowly margined; apex acute. Disc punctate and shallowly striate; central disc with moderately deep, closely spaced, punctures; base and apex with punctures shallow, striae effaced; stria 9 at basal 1/3 with larger, deeper punctures; disc at anterior 3/4 flattened at midline between suture and interval 2, other intervals weakly convex; setae shorter than on pronotum, semi-erect and recumbent, uniformly distributed.
|
||
<emphasis id="B941DA4FFFCE3A42FB8BFB69FCC59185" box="[707,834,1233,1259]" italics="true" pageId="16" pageNumber="115">Hind wing:</emphasis>
|
||
Macropterous.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F98FFB4EFA0B9341" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FFB4EFE839061" box="[199,260,1270,1295]" italics="true" pageId="16" pageNumber="115">Legs.</emphasis>
|
||
Profemur dark red-brown, covered with fine, evenly spaced punctures and long, recumbent golden setae; protibia red-brown, mostly bare and shiny, dorsal surface with sparse short setae and a few very long setae, ventral surface with dense row of short, stiff setae, arcuate with narrow spines at ventral apex; protarsus red-brown, shiny, dorsal surface with a few sparse, very long setae, ventral surface with dense, shorter setae, protarsomeres 4 and 5 each with a few much-longer dorsal setae. Mesofemur similar to profemur; mesotibia red-brown, mostly bare and shiny, dorsal surface with a few long setae, ventral surface with a patch of dense, short setae at apical 1/2, weakly arcuate with spinose ventral apex; mesotarsus similar to protarsus. Metafemur similar to pro- and mesofemur; metatibia weakly arcuate, setation like that of pro- and mesotibia; both metatarsi missing, but in the male
|
||
<typeStatus id="548EB8FFFFCE3A42FC74FA49FA1B9365" box="[1340,1436,1521,1547]" pageId="16" pageNumber="115" type="paratype">paratype</typeStatus>
|
||
the metatarsi are similar to the pro- and mesotarsi. [Note: left metatarsus was lost after being imaged for Fig. 14.]
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F98FF982FAC19221" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FF982FE88933D" box="[199,271,1594,1619]" italics="true" pageId="16" pageNumber="115">Venter</emphasis>
|
||
dark brown; heavily setose. Prosternum with anterior border narrowly margined; prosternal process wide between procoxae, widest at midlength, lateral margins weakly arcuate, thickly margined; process with median longitudinal carina, bordered laterally by sulci, terminating in a low, elongate protuberance near apex. Metaventrite with intercoxal process margined and depressed; posterior disc weakly depressed at junction of sulcate metakatepisternal suture and metathoracic discrimen. Abdomen with some setae longer than on rest of body except legs; ventrite 1 with triangular intercoxal process weakly depressed between metacoxae, thickly margined laterally; ventrites 2–4 of equal length, finely and evenly punctate at center of disc, coarser laterally, punctures aligned with faint, transverse strigae; ventrite 5 longest, disc coarsely punctate, covered with long setae prominent at margins.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCE3A42F98FF8E1FA1B92B1" blockId="16.[151,1437,153,2015]" pageId="16" pageNumber="115">
|
||
<emphasis id="B941DA4FFFCE3A42F98FF8E1FEBE921D" box="[199,313,1881,1907]" italics="true" pageId="16" pageNumber="115">Genitalia.</emphasis>
|
||
Aedeagus elongate (Fig. 15). Phallobase about 3x longer than parameres and wider than paramere bases together. Parameres each blade-like, inner surface concave; in dorsal view each with an inward-facing tooth near midlength; each paramere laterally sinuate, widest at base, narrowest near midlength, apex narrowly rounded; in lateral view aedeagus curved ventrally, each paramere wide, ventral surface nearly straight, dorsal surface weakly
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A40F98FFF20FC659594" blockId="18.[151,1436,152,394]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFF20FE5E95DD" bold="true" box="[199,473,152,179]" pageId="18" pageNumber="117">Intraspecific variation.</emphasis>
|
||
No significant non-sexual variation was noted among the small number of specimens, except that in the females the elytra are more convex in the posterior 1/2 than in the males. The two males, 5.30 and
|
||
<quantity id="4CCDABB8FFCC3A40F9DFFF58FF7C9595" box="[151,251,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="117" unit="mm" value="5.5">5.50 mm</quantity>
|
||
long, are close in size to the two females, 5.45 and
|
||
<quantity id="4CCDABB8FFCC3A40FA76FF58FC259595" box="[830,930,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" pageId="18" pageNumber="117" unit="mm" value="5.55">5.55 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFCC3A40F98FFEBCFEA8942D" blockId="18.[151,1436,152,394]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFEBCFE779471" bold="true" box="[199,496,260,287]" pageId="18" pageNumber="117">Geographic distribution.</emphasis>
|
||
Known from only two localities in the departments of
|
||
<collectingRegion id="49F1C8BFFFCC3A40FD2AFEBCFB539470" box="[1122,1236,260,286]" country="Paraguay" name="Cordillera" pageId="18" pageNumber="117">Cordillera</collectingRegion>
|
||
and
|
||
<collectingRegion id="49F1C8BFFFCC3A40FC44FEBDFAD29471" box="[1292,1365,261,287]" country="Paraguay" name="Itapua" pageId="18" pageNumber="117">Itapúa</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32246CDFFCC3A40FC29FEBDFF4A942D" name="Paraguay" pageId="18" pageNumber="117">Paraguay</collectingCountry>
|
||
(Fig. 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFCC3A40F98FFEF5FE4494E4" pageId="18" pageNumber="117" type="etymology">
|
||
<paragraph id="8B8A065DFFCC3A40F98FFEF5FE4494E4" blockId="18.[151,1436,152,394]" pageId="18" pageNumber="117">
|
||
<emphasis id="B941DA4FFFCC3A40F98FFEF5FEC99409" bold="true" box="[199,334,333,359]" pageId="18" pageNumber="117">Etymology.</emphasis>
|
||
This new species is named in honor of the
|
||
<emphasis id="B941DA4FFFCC3A40FA77FEF5FC199409" box="[831,926,333,359]" italics="true" pageId="18" pageNumber="117">Guaraní</emphasis>
|
||
, an indigenous people who inhabit
|
||
<collectingCountry id="F32246CDFFCC3A40FC7CFEF5FA1B9409" box="[1332,1436,333,359]" name="Paraguay" pageId="18" pageNumber="117">Paraguay</collectingCountry>
|
||
and neighboring countries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |