plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/F3/92/03F39207FFB3FFD07EFCF8A2F1F4DC98.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

298 lines
37 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="94217DBCC120D85EB5DC206768E8D718" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFB3FFD07EFCF8A2F1F4DC98" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopus igaranus Needham &amp; Murphy" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="11" updateTime="1698238640945" updateUser="plazi">
<mods:mods id="8BDC78F0ADC6586CF435557C817B8702" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AEA1A2031C3C459D1886D5F6910F6B7A">
<mods:title id="7C7C251A6467B27566D59C562B935A61">A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
</mods:titleInfo>
<mods:name id="38C268D09D97F954D4B567841735033D" type="personal">
<mods:role id="F192A6245F23BD76F4BA5718E43E7421">
<mods:roleTerm id="45CC8AC4E0F36080D7E3B3ADDC712823">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2BCA021E36F7BE30B2ED32AD8974A929">Molineri, Carlos</mods:namePart>
</mods:name>
<mods:typeOfResource id="2C7BB9C11598D2271ACAC158B910850B">text</mods:typeOfResource>
<mods:relatedItem id="9FBA35D568A8C5D418C2E6E03559944D" type="host">
<mods:titleInfo id="77FE52D7096FC6FF2BE119E25DF356F5">
<mods:title id="14B1E1C60BB6F17B8AC5F9A26F1A7A41">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="63D7A4786D1288F89DF337C18A207031">
<mods:date id="54D84FCE6CBFBA03B83923588277603E">2010</mods:date>
<mods:detail id="3F8361E875C91BAA87CC32B6098C9EAC" type="volume">
<mods:number id="6BAC8AA3505E3552982B9D2713014641">2481</mods:number>
</mods:detail>
<mods:extent id="56FB1CF842825EAD81F2127D320F8667" unit="page">
<mods:start id="AFC5800BE37B69F7131DE24C4012B5F2">1</mods:start>
<mods:end id="3D0BDF00A8B9AAC9CF7A7104D3994110">36</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="14F866E09E0A1639A63DAD6EEE73172C">journal article</mods:classification>
<mods:identifier id="94793A264E576B179EEE5521D09053B8" type="DOI">10.5281/zenodo.195431</mods:identifier>
<mods:identifier id="83C9E1CC9D498B7D8D383AFE0CB97B2E" type="GBIF-Dataset">2df8bd5a-e125-42aa-aa6f-ae3e78b83929</mods:identifier>
<mods:identifier id="B5046BD9358272FC90446ED3D369CBBB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="332F97D53A627F1E3FA01997E757CF6A" type="Zenodo-Dep">195431</mods:identifier>
</mods:mods>
<treatment id="03F39207FFB3FFD07EFCF8A2F1F4DC98" ID-DOI="http://doi.org/10.5281/zenodo.5690565" ID-GBIF-Taxon="119386657" ID-Zenodo-Dep="5690565" LSID="urn:lsid:plazi:treatment:03F39207FFB3FFD07EFCF8A2F1F4DC98" httpUri="http://treatment.plazi.org/id/03F39207FFB3FFD07EFCF8A2F1F4DC98" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11">
<subSubSection id="C340709AFFB3FFD47EFCF8A2F78ADE3B" box="[151,641,1824,1850]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BE52311FFB3FFD47EFCF8A2F78ADE3B" blockId="10.[151,641,1824,1850]" box="[151,641,1824,1850]" pageId="10" pageNumber="11">
<heading id="D0AD947DFFB3FFD47EFCF8A2F78ADE3B" bold="true" box="[151,641,1824,1850]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C5A5892FFB3FFD47EFCF8A2F78ADE3B" ID-CoL="57H2R" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy" box="[151,641,1824,1850]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB3FFD47EFCF8A2F78ADE3B" bold="true" box="[151,641,1824,1850]" pageId="10" pageNumber="11">
<emphasis id="B92EFF03FFB3FFD47EFCF8A2F47FDE3B" bold="true" box="[151,372,1824,1850]" italics="true" pageId="10" pageNumber="11">Tortopus igaranus</emphasis>
Needham &amp; Murphy
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFB3FFD47EFCF8EAF645DE7E" box="[151,846,1896,1919]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BE52311FFB3FFD47EFCF8EAF645DE7E" blockId="10.[151,846,1896,1919]" box="[151,846,1896,1919]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4A043FFFB3FFD47EFCF8EAF645DE7E" box="[151,846,1896,1919]" pageId="10" pageNumber="11">
<taxonomicName id="4C5A5892FFB3FFD47EFCF8EAF7AADE7E" ID-CoL="57H2R" authority="Needham &amp; Murphy, 1924:24" authorityName="Needham &amp; Murphy" authorityPageNumber="24" authorityYear="1924" box="[151,673,1896,1919]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB3FFD47EFCF8EAF451DE7E" box="[151,346,1896,1919]" italics="true" pageId="10" pageNumber="11">Tortopus igaranus</emphasis>
<treatmentCitation id="0AFB0500FFB3FFD47F0AF8EAF7AADE7E" author="Needham" box="[353,673,1896,1919]" page="24" pageId="10" pageNumber="11" year="1924">
<bibRefCitation id="EFCB5EE0FFB3FFD47F0AF8EAF789DE7E" author="Needham" box="[353,642,1896,1919]" pageId="10" pageNumber="35" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, 24, Entomological Series, 4, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy, 1924</bibRefCitation>
:24
</treatmentCitation>
</taxonomicName>
(female) in part
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFB3FFD07EFCF82CF1F4DC98" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BE52311FFB3FFD57EFCF82CF734D893" blockId="10.[151,1436,1966,2029]" lastBlockId="11.[151,1437,152,2010]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<typeStatus id="54E19DB3FFB3FFD47EFCF82CF5D9DEC9" box="[151,210,1966,1992]" pageId="10" pageNumber="11">Type</typeStatus>
material (
<collectionCode id="ED4BBBD4FFB3FFD47F27F82CF492DEC9" box="[332,409,1966,1992]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="10" pageNumber="11">CUIC</collectionCode>
).
<typeStatus id="54E19DB3FFB3FFD47FC5F82CF4FFDEC9" box="[430,500,1966,1992]" pageId="10" pageNumber="11">Types</typeStatus>
in alcohol:
<typeStatus id="54E19DB3FFB3FFD47CEFF82CF7E7DEC9" box="[644,748,1966,1992]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(female abdomen) and
<specimenCount id="9D5CE898FFB3FFD47A62F82CF17ADEC9" box="[1033,1137,1966,1992]" pageId="10" pageNumber="11" type="female">4 female</specimenCount>
<typeStatus id="54E19DB3FFB3FFD47A12F82CF1E0DEC9" box="[1145,1259,1966,1992]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="F34D6381FFB3FFD47B5EF82CF064DEC9" box="[1333,1391,1966,1992]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
, río Putumayo between Puerto Alfonso and the mouth of the Igará-Paraná,
<date id="FFE405D1FFB3FFD47D82F851F186DEEC" box="[1001,1165,2003,2029]" pageId="10" pageNumber="11" value="1920-08-14">14-VIII-1920</date>
. One female specimen from
<collectingCountry id="F34D6381FFB2FFD57EBCFF1AF41BD9B3" box="[215,272,152,178]" name="Peru" pageId="11" pageNumber="12">Perú</collectingCountry>
, río Igará-Paraná,
<date id="FFE405D1FFB2FFD57F82FF1AF7AFD9B3" box="[489,676,152,178]" pageId="11" pageNumber="12" value="1920-07-15" valueMax="1920-07-17" valueMin="1920-07-15">15-17-VII-1920</date>
(1 pair of wings dissected,
<collectionCode id="ED4BBBD4FFB2FFD57D82FF1AF124D9B3" box="[1001,1071,152,178]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="11" pageNumber="12">CUIC</collectionCode>
slide No. 629.3); and
<specimenCount id="9D5CE898FFB2FFD57B5DFF1AF097D9B3" box="[1334,1436,152,178]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
(body without wings, very damaged) from
<collectingCountry id="F34D6381FFB2FFD57CC4FF3FF7E0D9D6" box="[687,747,189,215]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
, Puerto Bermudez (no more data).
<typeStatus id="54E19DB3FFB2FFD57AC9FF3FF1D4D9D6" box="[1186,1247,189,215]" pageId="11" pageNumber="12">Type</typeStatus>
slides: the two
<typeStatus id="54E19DB3FFB2FFD57EFCFF60F40BD9FD" box="[151,256,226,252]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
slides (No. 629.1) were not studied,
<collectionCode id="ED4BBBD4FFB2FFD57CA9FF60F602D9FD" box="[706,777,226,252]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="11" pageNumber="12">CUIC</collectionCode>
reported them as completely broken and with missing parts.
<typeStatus id="54E19DB3FFB2FFD57E8BFE8AF44CD823" box="[224,327,264,290]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
slides: No. 629.3 correspond to the female from Igará-Paraná mentioned above, but the slide (1 wing pair) presents the following data (presumably a mistake): blue carton “
<typeStatus id="54E19DB3FFB2FFD57A06FEAFF000D846" box="[1133,1291,301,327]" pageId="11" pageNumber="12" type="paratype">PARATYPE</typeStatus>
/Cornell U./ No.629.3”, white carton “Cornell University/N0 654/
<collectionCode id="ED4BBBD4FFB2FFD57D5DFED0F678D86D" box="[822,883,338,364]" httpUri="http://grbio.org/cool/iz3p-2r7s" name="Universitat Bonn" pageId="11" pageNumber="12">SUB</collectionCode>
. 14 SL.Ƥ/
<taxonomicName id="4C5A5892FFB2FFD57D98FED0F16BD86D" box="[1011,1120,338,364]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Tortopus</taxonomicName>
/
<taxonomicName id="4C5A5892FFB2FFD57A01FED0F1FED86D" box="[1130,1269,338,364]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Campsurus</taxonomicName>
/
<taxonomicName id="4C5A5892FFB2FFD57A95FED0F06DD86D" box="[1278,1382,338,364]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="igaranus">igaranus</taxonomicName>
/Rio Putumayo/
<collectionCode id="ED4BBBD4FFB2FFD57F7FFEFAF46AD893" box="[276,353,376,402]" pageId="11" pageNumber="12">DATE</collectionCode>
<date id="FFE405D1FFB2FFD57F03FEFAF4CCD893" box="[360,455,376,402]" pageId="11" pageNumber="12" value="1914-08">14 Aug.</date>
´20
<collectingCountry id="F34D6381FFB2FFD57F9EFEFAF73AD893" box="[501,561,376,402]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
”.
</paragraph>
<paragraph id="8BE52311FFB2FFD57EADFE1FF6B3DBBD" blockId="11.[151,1437,152,2010]" pageId="11" pageNumber="12">
The following material is removed from the
<typeStatus id="54E19DB3FFB2FFD57CB7FE1FF604D8B6" box="[732,783,413,439]" pageId="11" pageNumber="12">type</typeStatus>
series since they represent female adults of a different species: six females originally designated as
<typeStatus id="54E19DB3FFB2FFD57CC5FE40F614D8DD" box="[686,799,450,476]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
(
<bibRefCitation id="EFCB5EE0FFB2FFD57D44FE40F164D8DD" author="Needham" box="[815,1135,450,476]" pageId="11" pageNumber="35" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, 24, Entomological Series, 4, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924</bibRefCitation>
), were removed from the vial containing the
<typeStatus id="54E19DB3FFB2FFD57F11FE6AF4EBDB03" box="[378,480,488,514]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(
<collectingCountry id="F34D6381FFB2FFD57F9BFE6AF721DB03" box="[496,554,488,514]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
, río Putumayo between Puerto Alfonso and the mouth of the Igará-Paraná,
<date id="FFE405D1FFB2FFD57EFCFD8FF435DB26" box="[151,318,525,551]" pageId="11" pageNumber="12" value="1920-08-14">14-VIII-1920</date>
). The slides No. 629.2, 629.4 and 629.5 are not conspecific with the
<typeStatus id="54E19DB3FFB2FFD57AEDFD8FF1F8DB26" box="[1158,1267,525,551]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
, the bodies in alcohol that correspond to the slides 629.2 and 629.4 are among the
<specimenCount id="9D5CE898FFB2FFD57DBCFDB0F147DB4D" box="[983,1100,562,588]" pageId="11" pageNumber="12" type="female">6 females</specimenCount>
mentioned above (the slide 629.5 does not have an associated specimen in alcohol). Additional material (
<collectionCode id="ED4BBBD4FFB2FFD57A58FDDAF164DB73" box="[1075,1135,600,626]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="11" pageNumber="12">IML</collectionCode>
). Five male and
<specimenCount id="9D5CE898FFB2FFD57B5EFDDAF097DB73" box="[1333,1436,600,626]" pageId="11" pageNumber="12" type="female">4 female</specimenCount>
imagos from
<collectingCountry id="F34D6381FFB2FFD57F5DFDFFF4A4DB96" box="[310,431,637,663]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
: Depto. Amazonas, Leticia,
<collectingCountry id="F34D6381FFB2FFD57D6BFDFFF621DB96" box="[768,810,637,663]" name="Iceland" pageId="11" pageNumber="12">Isla</collectingCountry>
Beatriz,
<quantity id="4CA28EF4FFB2FFD57DF3FDFFF6D3DB96" box="[920,984,637,663]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="11" pageNumber="12" unit="m" value="93.0">93 m</quantity>
,
<date id="FFE405D1FFB2FFD57D8EFDFFF152DB96" box="[997,1113,637,663]" pageId="11" pageNumber="12" value="1999-02-08">8-II-1999</date>
,
<geoCoordinate id="EE6E45D6FFB2FFD57A0DFDFFF1EFDB96" box="[1126,1252,637,663]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="-4.0730553">S 4º 4' 23&quot;</geoCoordinate>
<geoCoordinate id="EE6E45D6FFB2FFD57A86FDFFF09DDB96" box="[1261,1430,637,663]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-69.986664">W 69º 59' 12&quot;</geoCoordinate>
, light trap 18–20 h, M. C. Zúñiga, E. Domínguez &amp; C. Molineri cols.
</paragraph>
<paragraph id="8BE52311FFB2FFD57EADFD4AF4EDDA76" blockId="11.[151,1437,152,2010]" pageId="11" pageNumber="12">
<typeStatus id="54E19DB3FFB2FFD57EADFD4AF438DBE3" box="[198,307,712,738]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
(only female abdominal segments VII–X). Eggs orangeish. Abdominal tergum VII damaged, incomplete; terga VIII–X shaded slightly with gray, lighter in X. Abdominal sterna without gray shading, parastyli receptors on sternum VIII relatively small and near the medial line, with paired shallow longitudinal furrows directed anteriorly from each socket (Fig. 7). Cerci translucent yellowish white, vestigial terminal filament translucent whitish.
</paragraph>
<paragraph id="8BE52311FFB2FFD57EADFC00F134DCF5" blockId="11.[151,1437,152,2010]" pageId="11" pageNumber="12">
<typeStatus id="54E19DB3FFB2FFD57EADFC00F426DA9D" box="[198,301,898,924]" pageId="11" pageNumber="12" type="paratype">Paratype</typeStatus>
female adults. One of the four specimens from Putumayo river is much damaged presenting only the thorax and the wings (fore wings,
<quantity id="4CA28EF4FFB2FFD57C30FC2AF7C3DAC3" box="[603,712,936,962]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="12" unit="mm" value="12.5">12.5 mm</quantity>
; hind wings, 5.0). The other 3 are in better condition (one of them with a pair of wings dissected, without associated slide), body length: 7.0 (shrunken specimen)-
<quantity id="4CA28EF4FFB2FFD57B50FC4FF09CDAE6" box="[1339,1431,973,999]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="12" unit="mm" value="8.5">8.5 mm</quantity>
; fore wings, 9.8–10.0 mm; hind wings,
<quantity id="4CA28EF4FFB2FFD57CE8FC70F614DD0D" box="[643,799,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="11" pageNumber="12" unit="mm" value="3.7" valueMax="3.9" valueMin="3.5">3.5–3.9 mm</quantity>
. The
<typeStatus id="54E19DB3FFB2FFD57D0FFC70F6DBDD0D" box="[868,976,1010,1036]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
specimen from Igaraná-Paraná (and associated slide) is in a rather good condition, measures: body,
<quantity id="4CA28EF4FFB2FFD57DEFFB9AF6D5DD33" box="[900,990,1048,1074]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="12" unit="mm" value="6.5">6.5 mm</quantity>
(shrunken), fore wing
<quantity id="4CA28EF4FFB2FFD57A84FB9AF050DD33" box="[1263,1371,1048,1074]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="11" pageNumber="12" unit="mm" value="10.2">10.2 mm</quantity>
, hind wing,
<quantity id="4CA28EF4FFB2FFD57E8FFBBFF44DDD56" box="[228,326,1085,1111]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="mm" value="3.9">3.9 mm</quantity>
. General coloration yellowish white, eggs orangeish. Head with a whitish triangular mark behind median ocellus, occiput with a pair of grayish oblique bands extending from each lateral ocellus to the medial line near hind margin (Fig. 5). Antennae: scape and pedicel yellowish tinted with gray, flagellum hyaline. Thorax. Pronotum markedly convex dorsally, with well developed anterior hump; shaded with gray except laterally, medial line whitish, anterior margin shaded with gray sublaterally. Mesonotum whitish yellow, except medioposterior zone whitish, anterior phragma orangeish. Metanotum whitish medially, rest yellowish. Thoracic sterna pale, furcasternal protuberances with divergent medial margins (Fig. 4). Legs whitish except coxae yellowish. Wings. Membrane hyaline to whitish translucent, longitudinal veins yellowish brown, cross veins whitish; R3 and short intercalary vein before it present in fore wings (Fig. 8); reticulate pattern of cross veins in anal region of hind wings slightly marked (Fig. 9). Abdomen. Terga translucent whitish slightly and widely shaded with gray, darker on terga VIII–IX, tergum X whitish. Abdominal sterna pale, parastyli receptors (Fig. 6) on sternum VIII as
<typeStatus id="54E19DB3FFB2FFD57DBBFA58F130DCF5" box="[976,1083,1498,1524]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BE52311FFB2FFD57EADF982F694DF3E" blockId="11.[151,1437,152,2010]" pageId="11" pageNumber="12">
Length of voucher female imagos (collected with the males described below): body, 9.0–
<quantity id="4CA28EF4FFB2FFD57A95F982F055DF1B" box="[1278,1374,1536,1562]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="11" pageNumber="12" unit="mm" value="9.2">9.2 mm</quantity>
; fore wings, 8.5–10.0 mm; hind wings,
<quantity id="4CA28EF4FFB2FFD57C4DF9A7F7BCDF3E" box="[550,695,1573,1599]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="11" pageNumber="12" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
; cerci,
<quantity id="4CA28EF4FFB2FFD57D61F9A7F691DF3E" box="[778,922,1573,1599]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="11" pageNumber="12" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2–2.7 mm</quantity>
.
</paragraph>
<paragraph id="8BE52311FFB2FFD57EADF9C8F62EDFD5" blockId="11.[151,1437,152,2010]" pageId="11" pageNumber="12">
Eggs (
<figureCitation id="13613F94FFB2FFD57F79F9C8F4A8DF65" box="[274,419,1610,1636]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="11" pageNumber="12">Figs. 67–69</figureCitation>
). Subcircular outline. Length, 325 µm; width, 300 µm. No attachment device present. The convex area of the egg is sculptured with pentagonal and hexagonal cells (
<figureCitation id="13613F94FFB2FFD57A3CF9F2F1B9DF8B" box="[1111,1202,1648,1674]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="11" pageNumber="12">Fig. 68</figureCitation>
). The concave area shows a regular arrangement of shallow circular grooves (
<figureCitation id="13613F94FFB2FFD57D07F917F6C1DFAE" box="[876,970,1685,1711]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="11" pageNumber="12">Fig. 69</figureCitation>
). The entire zone forming the margin between the above mentioned areas is smooth (
<figureCitation id="13613F94FFB2FFD57CABF938F613DFD5" box="[704,792,1722,1748]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="11" pageNumber="12">Fig. 67</figureCitation>
).
</paragraph>
<paragraph id="8BE52311FFB2FFD27EADF962F71ADB03" blockId="11.[151,1437,152,2010]" lastBlockId="12.[151,1437,152,514]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Male imago. Length (mm): body, 7.5–9.8; fore wing, 8.0–8.9; hind wing,
<quantity id="4CA28EF4FFB2FFD57A48F962F1B9DFFB" box="[1059,1202,1760,1786]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="11" pageNumber="12" unit="mm" value="3.65" valueMax="3.9" valueMin="3.4">3.4–3.9 mm</quantity>
; fore leg (from base of coxae to apex of claw), 3.7–4.0; cerci, 20.0–22.0. General coloration whitish yellow. Head whitish shaded gray on pair of oblique bands on occiput, each band extending from lateral ocellus posteriorly towards medial line. Antennae: scape and pedicel whitish shaded light gray, flagellum hyaline. Thorax. Pronotum dorsally convex, anterior ring hyaline translucent, projecting anteriorly (&quot;anterior hump&quot;), a blackish marks is present laterally on fore margin; posterior ring of pronotum whitish translucent shaded with gray medially; propleura and presternum hyaline to whitish translucent. Mesonotum whitish yellow with darker margins and carinae, shaded slightly with gray along medioparapsidal sutures and inner margins of posterior scutal protuberances. Metanotum whitish translucent shaded gray anteromedially. Meso- and metaplaurae and sterna pale. Legs yellowish white, shaded with light gray on fore leg, more markedly on fore tibiae and tarsi, except intertarsal joinings. Wings. Membrane hyaline except apical third of C and Sc area whitish translucent; veins whitish translucent shaded with gray on veins Sc and RA. Abdomen whitish translucent shaded gray dorsally, very slightly on anterior segments but darkening posteriorly; terga III–VII with a pair of submedian bands, each band turns laterally towards hind margin; terga VIII–X shaded more widely, stronger on hind margin of tergum X; medial line of all terga pale. Abdominal sterna pale, with blackish spot in the middle of sternum IX. Genitalia whitish except parastyli and outer margin of penes yellowish. Caudal filaments whitish translucent except basal segment yellowish.
</paragraph>
<caption id="DF257399FFB5FFD27EFCF814F669DECF" httpUri="https://zenodo.org/record/195434/files/figure.png" pageId="12" pageNumber="13" targetBox="[187,1400,564,1941]" targetPageId="12">
<paragraph id="8BE52311FFB5FFD27EFCF814F669DECF" blockId="12.[151,1436,1942,1998]" pageId="12" pageNumber="13">
<emphasis id="B92EFF03FFB5FFD27EFCF814F46CDEAF" bold="true" box="[151,359,1942,1966]" pageId="12" pageNumber="13">FIGURES 14–18.</emphasis>
<taxonomicName id="4C5A5892FFB5FFD27F04F815F755DEAF" box="[367,606,1943,1966]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="circumfluus">
<emphasis id="B92EFF03FFB5FFD27F04F815F755DEAF" box="[367,606,1943,1966]" italics="true" pageId="12" pageNumber="13">Tortopus circumfluus</emphasis>
</taxonomicName>
, female holotype. 14, fore wing; 15, hind wing (scheme); 16, hind wing, enlarged; 17, abdominal sternum VIII; 18, mesofurcasternal plates.
</paragraph>
</caption>
<caption id="DF257399FFB4FFD37EFCFB9FF05FDD55" httpUri="https://zenodo.org/record/195435/files/figure.png" pageId="13" pageNumber="14" targetBox="[255,1332,207,1019]" targetPageId="13">
<paragraph id="8BE52311FFB4FFD37EFCFB9FF05FDD55" blockId="13.[151,1436,1053,1108]" pageId="13" pageNumber="14">
<emphasis id="B92EFF03FFB4FFD37EFCFB9FF455DD34" bold="true" box="[151,350,1053,1077]" pageId="13" pageNumber="14">FIGURES 19–23.</emphasis>
<taxonomicName id="4C5A5892FFB4FFD37F0FFB9FF719DD35" box="[356,530,1053,1076]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFB4FFD37F0FFB9FF719DD35" box="[356,530,1053,1076]" italics="true" pageId="13" pageNumber="14">Tortopus harrisi</emphasis>
</taxonomicName>
, male holotype: 19, genitalia, v.v. (pa = penial arm); 20–21, details of apex of penes; 22, detail of pedestal and base of forceps, v.v. (k = knob, pd = pedestal, ps = parastylus); 23, female sternum VIII.
</paragraph>
</caption>
<paragraph id="8BE52311FFB4FFD37EADFB00F125DCCB" blockId="13.[151,1437,1154,2011]" pageId="13" pageNumber="14">
Discussion and diagnosis. From the study of the
<typeStatus id="54E19DB3FFB4FFD37D48FB00F65DDD9D" box="[803,854,1154,1180]" pageId="13" pageNumber="14">type</typeStatus>
series, two groups of females representing two different genera (
<taxonomicName id="4C5A5892FFB4FFD37F0CFB2AF4DBDDC0" box="[359,464,1192,1217]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB4FFD37F0CFB2AF4DBDDC0" box="[359,464,1192,1217]" italics="true" pageId="13" pageNumber="14">Tortopus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A5892FFB4FFD37C60FB2AF772DDC0" box="[523,633,1192,1217]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB4FFD37C60FB2AF772DDC0" box="[523,633,1192,1217]" italics="true" pageId="13" pageNumber="14">Tortopsis</emphasis>
</taxonomicName>
) were distinguished. Fortunately, the
<typeStatus id="54E19DB3FFB4FFD37A54FB2AF1ACDDC3" box="[1087,1191,1192,1218]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
abdomen in alcohol was preserved enough to show important characters for species identification (
<typeStatus id="54E19DB3FFB4FFD37A57FB4FF1AEDDE6" box="[1084,1189,1229,1255]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
slides were reported as broken and with lost parts). The
<typeStatus id="54E19DB3FFB4FFD37C22FB70F7B8DC0D" box="[585,691,1266,1292]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and conespecific
<typeStatus id="54E19DB3FFB4FFD37DFBFB70F10FDC0D" box="[912,1028,1266,1292]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C5A5892FFB4FFD37A44FB71F1B2DC0D" box="[1071,1209,1267,1292]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB4FFD37A44FB71F1B2DC0D" box="[1071,1209,1267,1292]" italics="true" pageId="13" pageNumber="14">T. igaranus</emphasis>
</taxonomicName>
were compared to newly collected material from sites near to the
<typeStatus id="54E19DB3FFB4FFD37CBDFA9AF601DC33" box="[726,778,1304,1330]" pageId="13" pageNumber="14">type</typeStatus>
localities. Descriptions given above are based on this morphotype. The abdomens of the above described
<typeStatus id="54E19DB3FFB4FFD37D75FABFF69EDC56" box="[798,917,1341,1367]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
coincide completely with the
<typeStatus id="54E19DB3FFB4FFD37B64FABFF070DC56" box="[1295,1403,1341,1367]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
in morphology and coloration, including the shape of the parastyli receptors. Veins R3 and the short intercalary before it on fore wings are hard to distinguish in the relatively damaged wings of the
<typeStatus id="54E19DB3FFB4FFD37ADEFA09F026DCA4" box="[1205,1325,1419,1445]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
, they are present in one slide (
<collectionCode id="ED4BBBD4FFB4FFD37FE2FA32F4DADCCB" box="[393,465,1456,1482]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="13" pageNumber="14">CUIC</collectionCode>
No. 629.3) and on one of the specimens in alcohol.
</paragraph>
<paragraph id="8BE52311FFB4FFD37EADFA57F68DDF89" blockId="13.[151,1437,1154,2011]" pageId="13" pageNumber="14">
The
<typeStatus id="54E19DB3FFB4FFD37F6AFA57F46CDCEE" box="[257,359,1493,1519]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and the 4 conspecific
<typeStatus id="54E19DB3FFB4FFD37CE7FA57F7F7DCEE" box="[652,764,1493,1519]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C5A5892FFB4FFD37D45FA54F6B2DCEE" box="[814,953,1494,1519]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB4FFD37D45FA54F6B2DCEE" box="[814,953,1494,1519]" italics="true" pageId="13" pageNumber="14">T. igaranus</emphasis>
</taxonomicName>
present the following characters: head coloration formed mainly by a pair of oblique bands running from lateral ocelli to medial posterior line; mesonotum without a gray marking before mesoscutellum; furcasternal protuberances diverging from its anterior portion; fore wing with R3 and short intercalary vein before it; cross veins in fore and hind wings whitish; parastyli receptors on sterna VIII small and submedian.
</paragraph>
<paragraph id="8BE52311FFB4FFD37EADF911F7BADEDA" blockId="13.[151,1437,1154,2011]" pageId="13" pageNumber="14">
The six females from río Putumayo here excluded from the
<typeStatus id="54E19DB3FFB4FFD37D14F911F6B9DFAC" box="[895,946,1683,1709]" pageId="13" pageNumber="14">type</typeStatus>
series represents a different species in the genus
<taxonomicName id="4C5A5892FFB4FFD37E89F93BF445DFD3" box="[226,334,1721,1746]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB4FFD37E89F93BF445DFD3" box="[226,334,1721,1746]" italics="true" pageId="13" pageNumber="14">Tortopsis</emphasis>
</taxonomicName>
. These specimens can be distinguished from the
<typeStatus id="54E19DB3FFB4FFD37DE7F93AF6F9DFD3" box="[908,1010,1720,1746]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E19DB3FFB4FFD37A40F93AF190DFD3" box="[1067,1179,1720,1746]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
referred above by the following characters: head coloration more extended in occiput; pronotum in dorsal view rectangular, with well separated anterior and posterior rings; mesonotum with a gray V-shaped marking before mesoscutellum; furcasternal protuberances with parallel sides on anterior half, diverging only on the posterior portion; veins R3 and short intercalary before it absent in fore wing; cross veins in fore and hind wings as dark as longitudinal veins; parastyli receptors on sterna VIII C-shaped and relatively large, more lateral in position. These specimens could be the female of
<taxonomicName id="4C5A5892FFB4FFD37C05F819F6D6DEB4" authority="Ulmer" authorityName="Ulmer" box="[622,989,1947,1973]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B92EFF03FFB4FFD37C05F819F672DEB5" box="[622,889,1947,1972]" italics="true" pageId="13" pageNumber="14">Tortopsis unguiculatus</emphasis>
(Ulmer)
</taxonomicName>
or a new species, but their overall bad condition necessitates leaving them unnamed.
</paragraph>
<caption id="DF257399FFB7FFD07EFCFBC6F7CDDDBA" httpUri="https://zenodo.org/record/195436/files/figure.png" pageId="14" pageNumber="15" targetBox="[243,1335,197,1049]" targetPageId="14">
<paragraph id="8BE52311FFB7FFD07EFCFBC6F7CDDDBA" blockId="14.[151,1436,1092,1211]" pageId="14" pageNumber="15">
<emphasis id="B92EFF03FFB7FFD07EFCFBC6F456DD5D" bold="true" box="[151,349,1092,1116]" pageId="14" pageNumber="15">FIGURES 24–30.</emphasis>
<taxonomicName id="4C5A5892FFB7FFD07F08FBC7F709DD5D" box="[355,514,1093,1116]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zottai">
<emphasis id="B92EFF03FFB7FFD07F08FBC7F709DD5D" box="[355,514,1093,1116]" italics="true" pageId="14" pageNumber="15">Tortopus zottai</emphasis>
</taxonomicName>
, male paratype. 24, genitalia, v.v. (f1 and f2 = forceps segment 1 and 2, k = knob, mf = median furrow, pe = penes, sIX = ninth abdominal sternum); 25, mesofurcasternal plates (fsi = furcasternal impression, fsp = furcasternal plates); 26–27, details of apex of penes; 28, detail of pedestal and base of forceps, v.v.; 29, pedestal; 30, pedestal and forceps (pd = pedestal, ps = parastilus).
</paragraph>
</caption>
<paragraph id="8BE52311FFB7FFD07EADFB68F1F4DC98" blockId="14.[151,1436,1258,1433]" pageId="14" pageNumber="15">
<taxonomicName id="4C5A5892FFB7FFD07EADFB68F496DC02" box="[198,413,1258,1283]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB7FFD07EADFB68F496DC02" box="[198,413,1258,1283]" italics="true" pageId="14" pageNumber="15">Tortopus igaranus</emphasis>
</taxonomicName>
can be distinguished from all other species in the genus by the following combination of characters: 1) fore wing length 8.0–
<quantity id="4CA28EF4FFB7FFD07C36FA8DF7BCDC28" box="[605,695,1295,1321]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="14" pageNumber="15" unit="mm" value="8.9">8.9 mm</quantity>
(female
<quantity id="4CA28EF4FFB7FFD07D4BFA8DF6C1DC28" box="[800,970,1295,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.25" metricValueMin="0.85" pageId="14" pageNumber="15" unit="mm" value="10.5" valueMax="12.5" valueMin="8.5">8.5–12.5 mm</quantity>
); 2) parastyli relatively long (1.5 times length of pedestal) and thin from its base (Figs. 2–3); 3) ventral knob relatively small (Fig. 2); 4) penes not strongly expanded (Fig. 2); 5) female with sockets on s. VIII with oblique opening as in Figs. 6–7; and 6) a very pale species, head shaded gray in a pair of oblique bands from lateral ocelli to hind margin.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink