146 lines
17 KiB
XML
146 lines
17 KiB
XML
<document id="C49CC5B693D34F96931CF7DC3E2E7EB2" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFAFFFC97EFCFA5FF183D846" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis unguiculatus Ulmer, New combination" docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="23" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="5BB0AD2ABF987BB9D52A9F5E581833CA">A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:namePart id="7D4E3382CAC89223C5E9CBF1A4DB2B8A">Molineri, Carlos</mods:namePart>
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<mods:title id="AA4F1177117BCF2D6FDC5D09E09C7A11">Zootaxa</mods:title>
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<mods:date id="67808A09341BCB8C5E36BC9710B9FF05">2010</mods:date>
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<mods:number id="DA7905C34AF65AF181E0EA885C458825">2481</mods:number>
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<treatment id="03F39207FFAFFFC97EFCFA5FF183D846" ID-DOI="http://doi.org/10.5281/zenodo.5690579" ID-GBIF-Taxon="119386660" ID-Zenodo-Dep="5690579" LSID="urn:lsid:plazi:treatment:03F39207FFAFFFC97EFCFA5FF183D846" httpUri="http://treatment.plazi.org/id/03F39207FFAFFFC97EFCFA5FF183D846" lastPageId="23" lastPageNumber="23" pageId="22" pageNumber="23">
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<subSubSection id="C340709AFFAFFFC87EFCFA5FF7F3DCF6" box="[151,760,1501,1527]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BE52311FFAFFFC87EFCFA5FF7F3DCF6" blockId="22.[151,760,1501,1527]" box="[151,760,1501,1527]" pageId="22" pageNumber="23">
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<heading id="D0AD947DFFAFFFC87EFCFA5FF7F3DCF6" bold="true" box="[151,760,1501,1527]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="B92EFF03FFAFFFC87EFCFA5FF7F3DCF6" bold="true" box="[151,760,1501,1527]" pageId="22" pageNumber="23">
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<taxonomicName id="4C5A5892FFAFFFC87EFCFA5FF71DDCF6" ID-CoL="57H2L" authority="Ulmer" authorityName="Ulmer" box="[151,534,1501,1527]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="unguiculatus" status="comb. nov.">
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<emphasis id="B92EFF03FFAFFFC87EFCFA5FF4A0DCF6" bold="true" box="[151,427,1501,1527]" italics="true" pageId="22" pageNumber="23">Tortopsis unguiculatus</emphasis>
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(Ulmer)
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFAFFFC87C76FA5FF7F3DCF6" box="[541,760,1501,1527]" pageId="22" pageNumber="23">New combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFAFFFC87EFCF9A4F61DDF5D" pageId="22" pageNumber="23" type="reference_group">
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<paragraph id="8BE52311FFAFFFC87EFCF9A4F741DF3D" blockId="22.[151,790,1573,1629]" box="[151,586,1573,1597]" pageId="22" pageNumber="23">
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<treatmentCitationGroup id="AB4A043FFFAFFFC87EFCF9A4F741DF3D" box="[151,586,1573,1597]" pageId="22" pageNumber="23">
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<taxonomicName id="4C5A5892FFAFFFC87EFCF9A4F74EDF3D" authority="Ulmer 1920: 4" authorityName="Ulmer" authorityPageNumber="4" authorityYear="1920" box="[151,581,1573,1597]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="unguiculatus">
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<emphasis id="B92EFF03FFAFFFC87EFCF9A4F4AADF3C" box="[151,417,1574,1597]" italics="true" pageId="22" pageNumber="23">Campsurus unguiculatus</emphasis>
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<treatmentCitation id="0AFB0500FFAFFFC87FCCF9A7F74EDF3D" author="Ulmer" box="[423,581,1573,1596]" page="4" pageId="22" pageNumber="23" year="1920">
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<bibRefCitation id="EFCB5EE0FFAFFFC87FCCF9A7F720DF3D" author="Ulmer" box="[423,555,1573,1596]" pageId="22" pageNumber="35" refString="Ulmer, G. (1920) Neue Ephemeropteren. Archiv fur Naturgeschichte, 85 (A), 1 - 80." type="journal article" year="1920">Ulmer 1920</bibRefCitation>
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: 4
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE52311FFAFFFC87EFCF9C4F61DDF5D" blockId="22.[151,790,1573,1629]" box="[151,790,1605,1629]" pageId="22" pageNumber="23">
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<treatmentCitationGroup id="AB4A043FFFAFFFC87EFCF9C4F61DDF5D" box="[151,790,1605,1629]" pageId="22" pageNumber="23">
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<taxonomicName id="4C5A5892FFAFFFC87EFCF9C4F48EDF5C" box="[151,389,1606,1629]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="unguiculatus">
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<emphasis id="B92EFF03FFAFFFC87EFCF9C4F48EDF5C" box="[151,389,1606,1629]" italics="true" pageId="22" pageNumber="23">Tortopus unguiculatus</emphasis>
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</taxonomicName>
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;
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<treatmentCitation id="0AFB0500FFAFFFC87FF8F9C7F747DF5D" author="Ulmer" box="[403,588,1605,1628]" page="109" pageId="22" pageNumber="23" year="1942">
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<bibRefCitation id="EFCB5EE0FFAFFFC87FF8F9C7F71CDF5D" author="Ulmer" box="[403,535,1605,1628]" pageId="22" pageNumber="35" refString="Ulmer, G. (1942) Alte und neue Eintagsfliegen (Ephemeropteren) aus Sud- und Mittelamerika. Stettiner Entomologische Zeitung, 103, 98 - 128." type="journal article" year="1942">Ulmer 1942</bibRefCitation>
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: 109
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</treatmentCitation>
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;
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<treatmentCitation id="0AFB0500FFAFFFC87C3CF9C7F619DF5D" author="Traver" box="[599,786,1605,1628]" page="600" pageId="22" pageNumber="23" year="1950">
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<bibRefCitation id="EFCB5EE0FFAFFFC87C3CF9C7F7D6DF5D" author="Traver" box="[599,733,1605,1628]" pageId="22" pageNumber="35" refString="Traver, J. R. (1950) Notes on Neotropical mayflies. Part. IV. Family Ephemeridae (continued). Revista de Entomologia, 21, 593 - 614." type="journal article" year="1950">Traver 1950</bibRefCitation>
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: 600
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C340709AFFAFFFC97EFCF909F183D846" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="8BE52311FFAFFFC87EFCF909F5EDDFCB" blockId="22.[151,1436,1675,2037]" pageId="22" pageNumber="23">
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Material (
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<collectionCode id="ED4BBBD4FFAFFFC87F65F909F453DFA4" box="[270,344,1675,1701]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="22" pageNumber="23">ZMH</collectionCode>
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):
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<specimenCount id="9D5CE898FFAFFFC87F06F909F4CBDFA4" box="[365,448,1675,1701]" pageId="22" pageNumber="23" type="male">3 male</specimenCount>
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labeled “
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<typeStatus id="54E19DB3FFAFFFC87C58F909F77CDFA4" box="[563,631,1675,1701]" pageId="22" pageNumber="23">Type</typeStatus>
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”, “Columbien/Behn vend.
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<date id="FFE405D1FFAFFFC87DD5F909F140DFA4" box="[958,1099,1675,1701]" pageId="22" pageNumber="23" value="1900-12-03">3.XII.1900</date>
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.”, “G. Ulmer determ./
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<date id="FFE405D1FFAFFFC87B3AF909F5DCDFCB" pageId="22" pageNumber="23" value="1916" valueMax="1923">1916– 1923</date>
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”.
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</paragraph>
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<paragraph id="8BE52311FFAFFFC87EADF954F728DE84" blockId="22.[151,1436,1675,2037]" pageId="22" pageNumber="23">
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Male imago. General coloration faint in the old pinned
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<typeStatus id="54E19DB3FFAFFFC87DE8F954F6CEDFF1" box="[899,965,1750,1776]" pageId="22" pageNumber="23">types</typeStatus>
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. Genitalia: long and cylindrical penes completely separated from their bases (
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<figureCitation id="13613F94FFAFFFC87C01F979F7C8DE14" box="[618,707,1787,1813]" captionStart="FIGURES 41 – 43" captionStartId="16.[151,273,1978,2002]" captionTargetBox="[221,1357,197,1936]" captionTargetId="figure@16.[221,1366,194,1957]" captionTargetPageId="16" captionText="FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes)." httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="22" pageNumber="23">Fig. 41</figureCitation>
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); sclerotized spine on inner apical margin with wide rounded base and slightly hooked apex (
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<figureCitation id="13613F94FFAFFFC87C67F8A2F76FDE3B" box="[524,612,1824,1850]" captionStart="FIGURES 41 – 43" captionStartId="16.[151,273,1978,2002]" captionTargetBox="[221,1357,197,1936]" captionTargetId="figure@16.[221,1366,194,1957]" captionTargetPageId="16" captionText="FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes)." httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="22" pageNumber="23">Fig. 43</figureCitation>
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); parastyli long and acute, markedly curved dorsally, the main axis of the basal third, form a 90° angle with the axis of the distal third (
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<figureCitation id="13613F94FFAFFFC87DC6F8C4F103DE61" box="[941,1032,1862,1888]" captionStart="FIGURES 41 – 43" captionStartId="16.[151,273,1978,2002]" captionTargetBox="[221,1357,197,1936]" captionTargetId="figure@16.[221,1366,194,1957]" captionTargetPageId="16" captionText="FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes)." httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="22" pageNumber="23">Fig. 42</figureCitation>
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); parastyli conical and entire, not furrowed longitudinally (
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<figureCitation id="13613F94FFAFFFC87FD5F8E9F71DDE84" box="[446,534,1899,1925]" captionStart="FIGURES 41 – 43" captionStartId="16.[151,273,1978,2002]" captionTargetBox="[221,1357,197,1936]" captionTargetId="figure@16.[221,1366,194,1957]" captionTargetPageId="16" captionText="FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes)." httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="22" pageNumber="23">Fig. 41</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE52311FFAFFFC97EADF812F183D846" blockId="22.[151,1436,1675,2037]" lastBlockId="23.[151,1437,152,327]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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Discussion and diagnosis.
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<bibRefCitation id="EFCB5EE0FFAFFFC87C6AF812F7A9DEAB" author="Ulmer" box="[513,674,1936,1962]" pageId="22" pageNumber="35" refString="Ulmer, G. (1920) Neue Ephemeropteren. Archiv fur Naturgeschichte, 85 (A), 1 - 80." type="journal article" year="1920">Ulmer (1920)</bibRefCitation>
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described this species from
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<specimenCount id="9D5CE898FFAFFFC87D9DF812F14DDEAB" box="[1014,1094,1936,1962]" pageId="22" pageNumber="23" type="male">3 male</specimenCount>
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imagos from
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<collectingCountry id="F34D6381FFAFFFC87A85F812F06EDEAB" box="[1262,1381,1936,1962]" name="Colombia" pageId="22" pageNumber="23">Colombia</collectingCountry>
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, and later (
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<bibRefCitation id="EFCB5EE0FFAFFFC87EB6F834F473DED1" author="Ulmer" box="[221,376,1974,2000]" pageId="22" pageNumber="35" refString="Ulmer, G. (1942) Alte und neue Eintagsfliegen (Ephemeropteren) aus Sud- und Mittelamerika. Stettiner Entomologische Zeitung, 103, 98 - 128." type="journal article" year="1942">Ulmer, 1942</bibRefCitation>
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) provided a description of a female imago from
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<collectingCountry id="F34D6381FFAFFFC87DACF834F136DED1" box="[967,1085,1974,2000]" name="Colombia" pageId="22" pageNumber="23">Colombia</collectingCountry>
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and cited some male imagos from Reventazón (
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<collectingCountry id="F34D6381FFAFFFC87F1EF859F4F4DEF4" box="[373,511,2011,2037]" name="Costa Rica" pageId="22" pageNumber="23">Costa Rica</collectingCountry>
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). Additional records exist for this species in
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<collectingCountry id="F34D6381FFAFFFC87A7CF859F17EDEF4" box="[1047,1141,2011,2037]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
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,
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<collectingCountry id="F34D6381FFAFFFC87AE9F859F1F7DEF4" box="[1154,1276,2011,2037]" name="Nicaragua" pageId="22" pageNumber="23">Nicaragua</collectingCountry>
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(Zelaya) and
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<collectingCountry id="F34D6381FFAEFFC97EFCFF1AF410D9B3" box="[151,283,152,178]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
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(
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<bibRefCitation id="EFCB5EE0FFAEFFC97F40FF1AF4B8D9B3" author="Maes" box="[299,435,152,178]" pageId="23" pageNumber="34" refString="Maes, J. M. (1988) Catalogo de los Ephemeroptera y Plecoptera de Nicaragua. Revista Nicaraguense de Entomologia, 2, 49 - 50." type="journal article" year="1988">Maés 1988</bibRefCitation>
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) and
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<collectingCountry id="F34D6381FFAEFFC97F93FF1AF771D9B3" box="[504,634,152,178]" name="Guatemala" pageId="23" pageNumber="24">Guatemala</collectingCountry>
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and
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<collectingCountry id="F34D6381FFAEFFC97CDDFF1AF620D9B3" box="[694,811,152,178]" name="Honduras" pageId="23" pageNumber="24">Honduras</collectingCountry>
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(
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<bibRefCitation id="EFCB5EE0FFAEFFC97D57FF1AF1CAD9B3" author="Lugo-Ortiz" box="[828,1217,152,178]" pageId="23" pageNumber="34" refString="Lugo-Ortiz, C. R. & McCafferty W. P. (1996) Central American Tortopus (Ephemeroptera: Polymitarcyidae): a unique new species and new country records. Entomological News, 107, 23 - 27." type="journal article" year="1996">Lugo-Ortiz & McCafferty 1996</bibRefCitation>
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), but this material should be revised with the new data offered here to confirm its specific identity.
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<taxonomicName id="4C5A5892FFAEFFC97A54FF3FF041D9D7" box="[1087,1354,189,214]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="unguiculatus">
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<emphasis id="B92EFF03FFAEFFC97A54FF3FF041D9D7" box="[1087,1354,189,214]" italics="true" pageId="23" pageNumber="24">Tortopsis unguiculatus</emphasis>
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</taxonomicName>
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can be distinguished from all other species of the genus by: 1) fore wing length 9.0–11.0 mm (male), 11.0–12.0 mm (female); 2) wings hyaline translucent, veins whitish translucent; 3) parastyli long and very strongly curved (
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||
<figureCitation id="13613F94FFAEFFC97EF4FEAFF5FCD846" box="[159,247,301,327]" captionStart="FIGURES 41 – 43" captionStartId="16.[151,273,1978,2002]" captionTargetBox="[221,1357,197,1936]" captionTargetId="figure@16.[221,1366,194,1957]" captionTargetPageId="16" captionText="FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes)." httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="23" pageNumber="24">Fig. 42</figureCitation>
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); 4) penes cylindrical, apical spine with wide base and a small hook (
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<figureCitation id="13613F94FFAEFFC97A48FEAFF170D846" box="[1059,1147,301,327]" captionStart="FIGURES 41 – 43" captionStartId="16.[151,273,1978,2002]" captionTargetBox="[221,1357,197,1936]" captionTargetId="figure@16.[221,1366,194,1957]" captionTargetPageId="16" captionText="FIGURES 41 – 43. Tortopsis unguiculatus, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes)." httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="23" pageNumber="24">Fig. 43</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |