185 lines
29 KiB
XML
185 lines
29 KiB
XML
<document id="A26510D8C95C60AA4D652074FAAEE7E7" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFADFFCB7EFCF99AF6B1DF65" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopus arenales Molineri, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="21" updateTime="1698238640945" updateUser="plazi">
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<mods:mods id="10B8FF610BC88EC55BF2C2564462CCAA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="32783CFF30AEE59B342340FDC5046FE9">
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<mods:title id="8E965B75CA9DADFEC240E7B40A3E24CB">A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3637408527DCD1EFAD36F6B9811228FA" type="personal">
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<mods:role id="5C2E9255B2783EBA0E2120EEB57BDB65">
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<mods:roleTerm id="E25FCF890BA24A0796A6ECB6D55FC3E1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5819D91264220B4AF8C6CD302912D127">Molineri, Carlos</mods:namePart>
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<mods:typeOfResource id="6DC6A40B5EB0A52840387D3B72F1A0FC">text</mods:typeOfResource>
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<mods:title id="2B901E6DEDB7B5D8E5FC111C20DB72FB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="03A2827B1C11FBE1BF90BD90345F0AA8">
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<mods:date id="909C85A696D0A9E76496D716C9A558B7">2010</mods:date>
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<mods:detail id="61AF1B6388690B033A21A8CE3A9668DC" type="volume">
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<mods:number id="DAA16138FB432086C6655B359372670A">2481</mods:number>
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<mods:extent id="54E5A284B278EC021B1CFFAC58B7FE0B" unit="page">
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<mods:start id="8572107EC30177434D62CA01184B3B45">1</mods:start>
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<mods:end id="60343F34C2B8B45CA7A2840447DF51FF">36</mods:end>
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<mods:classification id="3334009AE5D1B21F9733134A69DCDE9B">journal article</mods:classification>
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<mods:identifier id="1CCF87030EC30BAE14ACFD99916D1892" type="DOI">10.5281/zenodo.195431</mods:identifier>
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<mods:identifier id="5195D08AA7BEEBFE6B2734AD2560B82C" type="GBIF-Dataset">2df8bd5a-e125-42aa-aa6f-ae3e78b83929</mods:identifier>
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<mods:identifier id="DFAE3D4E181B7874CFEB164430F33333" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BED8DA7CCB78F7F7313C79A7A8D15A37" type="Zenodo-Dep">195431</mods:identifier>
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</mods:mods>
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<treatment id="03F39207FFADFFCB7EFCF99AF6B1DF65" ID-DOI="http://doi.org/10.5281/zenodo.5690575" ID-GBIF-Taxon="119386663" ID-Zenodo-Dep="5690575" LSID="urn:lsid:plazi:treatment:03F39207FFADFFCB7EFCF99AF6B1DF65" httpUri="http://treatment.plazi.org/id/03F39207FFADFFCB7EFCF99AF6B1DF65" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="C340709AFFADFFCA7EFCF99AF4DFDF33" box="[151,468,1560,1586]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BE52311FFADFFCA7EFCF99AF4DFDF33" blockId="20.[151,468,1560,1586]" box="[151,468,1560,1586]" pageId="20" pageNumber="21">
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<heading id="D0AD947DFFADFFCA7EFCF99AF4DFDF33" bold="true" box="[151,468,1560,1586]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="B92EFF03FFADFFCA7EFCF99AF4DFDF33" bold="true" box="[151,468,1560,1586]" pageId="20" pageNumber="21">
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<taxonomicName id="4C5A5892FFADFFCA7EFCF99AF464DF33" ID-CoL="57H2M" box="[151,367,1560,1586]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arenales" status="sp. nov.">
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<emphasis id="B92EFF03FFADFFCA7EFCF99AF464DF33" bold="true" box="[151,367,1560,1586]" italics="true" pageId="20" pageNumber="21">Tortopus arenales</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFADFFCA7F1DF99AF4DFDF33" box="[374,468,1560,1586]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFADFFCB7EFCF9E2F6B1DF65" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8BE52311FFADFFCA7EFCF9E2F744DE7E" blockId="20.[151,1436,1632,2031]" pageId="20" pageNumber="21">
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Material:
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<typeStatus id="54E19DB3FFADFFCA7F64F9E2F471DF7B" box="[271,378,1632,1658]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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male imago from
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<collectingCountry id="F34D6381FFADFFCA7C34F9E2F7C2DF7B" box="[607,713,1632,1658]" name="Ecuador" pageId="20" pageNumber="21">Ecuador</collectingCountry>
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: Prov. Esmeralda, Canton Eloy Alfaro, Parroquia Telembí, Estero
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<taxonomicName id="4C5A5892FFADFFCA7E81F907F45EDF9E" box="[234,341,1669,1695]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="29" phylum="Arthropoda" rank="species" species="arenales">Arenales</taxonomicName>
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,
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<quantity id="4CA28EF4FFADFFCA7F0AF907F4ABDF9E" box="[353,416,1669,1695]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="21" unit="m" value="80.0">80 m</quantity>
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,
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||
<date id="FFE405D1FFADFFCA7FC6F907F73CDF9E" box="[429,567,1669,1695]" pageId="20" pageNumber="21" value="2002-04-15">15-IV-2002</date>
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||
,
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||
<geoCoordinate id="EE6E45D6FFADFFCA7C2FF907F7CADF9E" box="[580,705,1669,1695]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-0.66833335">S 0º 40' 6"</geoCoordinate>
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||
<geoCoordinate id="EE6E45D6FFADFFCA7CA2F907F67ADF9E" box="[713,881,1669,1695]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-78.99027">W 78º 59' 25"</geoCoordinate>
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, E. Domínguez col.
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<typeStatus id="54E19DB3FFADFFCA7A0FF907F1D1DF9E" box="[1124,1242,1669,1695]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<specimenCount id="9D5CE898FFADFFCA7A8DF907F04FDF9E" box="[1254,1348,1669,1695]" pageId="20" pageNumber="21" type="male">13 male</specimenCount>
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||
and
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<specimenCount id="9D5CE898FFADFFCA7B14F907F5ECDFC5" pageId="20" pageNumber="21" type="female">21 female</specimenCount>
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||
imagos same data as
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||
<typeStatus id="54E19DB3FFADFFCA7F8FF928F745DFC5" box="[484,590,1706,1732]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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;
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<specimenCount id="9D5CE898FFADFFCA7C31F928F7BCDFC5" box="[602,695,1706,1732]" pageId="20" pageNumber="21" type="male">10 male</specimenCount>
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||
and
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<specimenCount id="9D5CE898FFADFFCA7C9BF928F66FDFC5" box="[752,868,1706,1732]" pageId="20" pageNumber="21" type="female">26 female</specimenCount>
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||
imagos same data except Estero Cayapas,
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<quantity id="4CA28EF4FFADFFCA7B32F928F093DFC5" box="[1369,1432,1706,1732]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="21" unit="m" value="70.0">70 m</quantity>
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,
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<date id="FFE405D1FFADFFCA7EFCF952F428DFEB" box="[151,291,1744,1770]" pageId="20" pageNumber="21" value="2002-04-15">15-IV-2002</date>
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||
,
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||
<geoCoordinate id="EE6E45D6FFADFFCA7F44F952F4B7DFEB" box="[303,444,1744,1770]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-0.67194444">S 0º 40' 19"</geoCoordinate>
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||
<geoCoordinate id="EE6E45D6FFADFFCA7FAFF952F765DFEB" box="[452,622,1744,1770]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-78.99139">W 78º 59' 29"</geoCoordinate>
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, E. Domínguez col.;
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<specimenCount id="9D5CE898FFADFFCA7D00F952F6C1DFEB" box="[875,970,1744,1770]" pageId="20" pageNumber="21" type="male">11 male</specimenCount>
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||
and
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<specimenCount id="9D5CE898FFADFFCA7A6FF952F160DFEB" box="[1028,1131,1744,1770]" pageId="20" pageNumber="21" type="female">8 female</specimenCount>
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imagos same data except Estero Calle Mansa,
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||
<quantity id="4CA28EF4FFADFFCA7FE0F977F4C2DE0E" box="[395,457,1781,1807]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="50.0">50 m</quantity>
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,
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<date id="FFE405D1FFADFFCA7FBEF977F754DE0E" box="[469,607,1781,1807]" pageId="20" pageNumber="21" value="2002-04-17">17-IV-2002</date>
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||
,
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||
<geoCoordinate id="EE6E45D6FFADFFCA7C00F977F7FEDE0E" box="[619,757,1781,1807]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-0.70277774">S 0º 42' 10"</geoCoordinate>
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<geoCoordinate id="EE6E45D6FFADFFCA7C97F977F6A9DE0E" box="[764,930,1781,1807]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-78.96694">W 78º 58' 01"</geoCoordinate>
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, E. Domínguez col.; and
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<specimenCount id="9D5CE898FFADFFCA7AA0F977F04BDE0E" box="[1227,1344,1781,1807]" pageId="20" pageNumber="21" type="female">17 female</specimenCount>
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||
imagos same data except Estero Charco Vicente,
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||
<quantity id="4CA28EF4FFADFFCA7CE3F898F7C3DE35" box="[648,712,1818,1844]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="21" unit="m" value="40.0">40 m</quantity>
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||
,
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||
<date id="FFE405D1FFADFFCA7CBFF898F66ADE35" box="[724,865,1818,1844]" pageId="20" pageNumber="21" value="2002-04-14">14-IV-2002</date>
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||
,
|
||
<geoCoordinate id="EE6E45D6FFADFFCA7D06F898F6F0DE35" box="[877,1019,1818,1844]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-0.6933333">S 0º 41' 36"</geoCoordinate>
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||
<geoCoordinate id="EE6E45D6FFADFFCA7A6FF898F1A5DE35" box="[1028,1198,1818,1844]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="-78.909164">W 78º 54' 33"</geoCoordinate>
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||
, E. Domínguez col.
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<typeStatus id="54E19DB3FFADFFCA7EFCF8C2F40CDE5B" box="[151,263,1856,1882]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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||
,
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||
<specimenCount id="9D5CE898FFADFFCA7F79F8C2F46ADE5B" box="[274,353,1856,1882]" pageId="20" pageNumber="21" type="male">3 male</specimenCount>
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||
and
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<specimenCount id="9D5CE898FFADFFCA7FF2F8C2F4F5DE5B" box="[409,510,1856,1882]" pageId="20" pageNumber="21" type="female">3 female</specimenCount>
|
||
imagos
|
||
<typeStatus id="54E19DB3FFADFFCA7C0BF8C2F7DBDE5B" box="[608,720,1856,1882]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
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||
at
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<collectionCode id="ED4BBBD4FFADFFCA7C98F8C2F647DE5B" box="[755,844,1856,1882]" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" pageId="20" pageNumber="21">MECN</collectionCode>
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;
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<specimenCount id="9D5CE898FFADFFCA7D3CF8C2F6ADDE5B" box="[855,934,1856,1882]" pageId="20" pageNumber="21" type="male">2 male</specimenCount>
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||
and
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<specimenCount id="9D5CE898FFADFFCA7DB5F8C2F148DE5B" box="[990,1091,1856,1882]" pageId="20" pageNumber="21" type="female">2 female</specimenCount>
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imagos in
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<collectionCode id="ED4BBBD4FFADFFCA7AA8F8C2F002DE5B" box="[1219,1289,1856,1882]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="20" pageNumber="21">CUIC</collectionCode>
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and
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<collectionCode id="ED4BBBD4FFADFFCA7B2AF8C2F09CDE5B" box="[1345,1431,1856,1882]" pageId="20" pageNumber="21">FAMU</collectionCode>
|
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; remaining material deposited in
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<collectionCode id="ED4BBBD4FFADFFCA7C79F8E7F747DE7E" box="[530,588,1893,1919]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="20" pageNumber="21">IML</collectionCode>
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.
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</paragraph>
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<paragraph id="8BE52311FFADFFCB7EADF808F16DDD0D" blockId="20.[151,1436,1632,2031]" lastBlockId="21.[151,1437,152,1636]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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Male imago. Length (mm): body, 10.0–11.5; fore wing, 10.0–11.7; hind wing, 4.9–5.2; foreleg, 4.7–5.0; cerci, 26.0–30.0. General coloration yellowish white shaded with brown. Head shaded strongly with brownish gray on a band between ocelli, shading slighter posteriorly forming a profuse anastomosed pattern on occiput (
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<figureCitation id="13613F94FFACFFCB7ECBFF1AF5F2D9B3" box="[160,249,152,178]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
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); head not shaded ventrally and before to median ocellus. Antennae: scape and pedicel completely shaded with gray except at apical margin, flagellum hyaline. Thorax. Pronotum (
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<figureCitation id="13613F94FFACFFCB7A2FFF3FF193D9D6" box="[1092,1176,189,215]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
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) translucent brownish white shaded widely with brownish gray, anterior ring shaded stronger on submedian zones, paler posteriorly; posterior ring shaded strongly medially with a thin dark medial line and two paler sublateral spots; propleura and presternum translucent brownish white. Mesonotum yellowish white shaded brownish gray along medioparapsidal sutures forming a pair of bands that become darker and converge posteriorly; mesopleura and mesosternum shaded very slightly with gray on pleura, but strongly on basisternum; furcasternal protuberances diverging posteriorly from anterior margin (
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<figureCitation id="13613F94FFACFFCB7D04FE1FF6C6D8B6" box="[879,973,413,439]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 34</figureCitation>
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). Metanotum yellowish white shaded brownish gray widely in anterior zone, stronger shading restricted more medially towards posterior margin; metapleura and sternum brownish white shaded strongly with gray on a pair of submedian anterior marks. Legs. Fore legs shaded completely and strongly with brownish gray except on the very pale claws; middle and hind legs yellowish white slightly shaded gray. Wings. Membrane hyaline shaded brownish gray on C and Sc areas, longitudinal veins shaded brownish, cross veins pale. Abdomen whitish translucent shaded strongly with brownish gray, except on lateral areas. Tergum I shaded submedially, with a translucent medial band; terga II–VII with thin pale median and paramedian dashes, also with a pale oblique dash near lateral margins (
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||
<figureCitation id="13613F94FFACFFCB7EF4FD4AF5FCDBE3" box="[159,247,712,738]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 32</figureCitation>
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); terga VIII–X shaded more widely and strongly except on submedian and lateral pale marks; all terga except the first with a thin dark median line. Abdominal sterna (
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<figureCitation id="13613F94FFACFFCB7DF7FD6FF6FFDA06" box="[924,1012,749,775]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 33</figureCitation>
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) shaded strongly on median region except on pale medial line and paired submedial oval marks, shading darker around the medial line; indentations left by nymphal gill muscles (subcircular lateral areas on sterna II–VII) yellowish white shaded gray except on pale medial margin. Genitalia (
|
||
<figureCitation id="13613F94FFACFFCB7C8FFCDFF68BDA76" box="[740,896,861,887]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Figs. 36–40</figureCitation>
|
||
): sternum IX shaded strongly with gray on mediolongitudinal furrow but getting much lighter laterally; pedestals whitish turning yellowish towards apex of parastyli, parastyli relatively short and straight (
|
||
<figureCitation id="13613F94FFACFFCB7C93FC2AF68BDAC3" box="[760,896,936,962]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 36–38</figureCitation>
|
||
); small first forceps segment whitish, second segment strongly shaded gray; penes whitish, outer margin yellowish, shaded slightly gray on each penial arm. Caudal filaments whitish translucent shaded brownish gray except at joinings.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FFACFFCB7EADFB9AF7FFDC15" blockId="21.[151,1437,152,1636]" pageId="21" pageNumber="22">
|
||
Female imago. Length (mm): body, 9.6–13.0; fore wing, 11.2–14.5; hind wing, 4.5–6.2; cerci, 3.1–4.0. General coloration as male, eggs yellowish white. Metasternal black marks thinner and longer than in male. Wings darker than in male, membrane slightly tinged with yellow, longitudinal and cross veins shaded with brownish, more strongly on C, Sc and R1 veins; R3 and short intercalary vein before it present on fore wings. Brownish shading on body as in male except slightly paler on abdominal sterna; intersegmental membrane between sterna VII–VIII paler than the rest, sternum VIII with parastyli receptors located submedially (
|
||
<figureCitation id="13613F94FFACFFCB7B0AFB57F5BCDC15" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Figs. 35</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FFACFFCB7EA8FB78F405DC15" box="[195,270,1274,1300]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="21" pageNumber="22">70–71</figureCitation>
|
||
) and strongly shaded gray. Cerci whitish.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FFACFFCB7EADFAA2F7B9DC5E" blockId="21.[151,1437,152,1636]" pageId="21" pageNumber="22">
|
||
Eggs (
|
||
<figureCitation id="13613F94FFACFFCB7F78FAA2F465DC3B" box="[275,366,1312,1338]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="21" pageNumber="22">Fig. 72</figureCitation>
|
||
). Suboval, no attachment structures. Length, 400 µm; width, 330–345 µm. The relatively thick covering of the egg is relatively smooth.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FFACFFCB7EADFAE8F62FDC85" blockId="21.[151,1437,152,1636]" box="[198,804,1386,1412]" pageId="21" pageNumber="22">
|
||
Etymology: from the
|
||
<typeStatus id="54E19DB3FFACFFCB7FA8FAE8F4FDDC85" box="[451,502,1386,1412]" pageId="21" pageNumber="22">type</typeStatus>
|
||
locality, Estero
|
||
<taxonomicName id="4C5A5892FFACFFCB7CDFFAE8F62BDC85" box="[692,800,1386,1412]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="29" phylum="Arthropoda" rank="species" species="arenales">Arenales</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FFACFFCB7EADFA12F6B1DF65" blockId="21.[151,1437,152,1636]" pageId="21" pageNumber="22">
|
||
Discussion and diagnosis.
|
||
<taxonomicName id="4C5A5892FFACFFCB7F95FA12F7DADCA8" box="[510,721,1424,1449]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arenales" status="sp. nov.">
|
||
<emphasis id="B92EFF03FFACFFCB7F95FA12F7DADCA8" box="[510,721,1424,1449]" italics="true" pageId="21" pageNumber="22">Tortopus arenales</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D4278FFACFFCB7CB2FA12F63FDCAB" box="[729,820,1424,1450]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from all other species of the genus by the following combination of characters: 1) fore wing length 10.0–
|
||
<quantity id="4CA28EF4FFACFFCB7A64FA37F177DCCE" box="[1039,1148,1461,1487]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="21" pageNumber="22" unit="mm" value="11.7">11.7 mm</quantity>
|
||
(male),
|
||
<quantity id="4CA28EF4FFACFFCB7A83FA37F097DCCE" box="[1256,1436,1461,1487]" metricMagnitude="-2" metricUnit="m" metricValue="1.285" metricValueMax="1.45" metricValueMin="1.1199999999999999" pageId="21" pageNumber="22" unit="mm" value="12.85" valueMax="14.5" valueMin="11.2">11.2–14.5 mm</quantity>
|
||
(female); 2) parastyli with relatively wide base, rest very thin and acute, as long as pedestal (
|
||
<figureCitation id="13613F94FFACFFCB7A83FA58F066DCF5" box="[1256,1389,1498,1524]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 36–37</figureCitation>
|
||
); 3) ventral knob well developed (
|
||
<figureCitation id="13613F94FFACFFCB7F9FF982F747DF1B" box="[500,588,1536,1562]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 36</figureCitation>
|
||
); 4) penes relatively wide at base (fused portion), not strongly expanded distally (
|
||
<figureCitation id="13613F94FFACFFCB7E94F9A7F452DF3E" box="[255,345,1573,1599]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 36</figureCitation>
|
||
); 5) female sockets with oblique opening (
|
||
<figureCitation id="13613F94FFACFFCB7D3CF9A7F6A4DF3E" box="[855,943,1573,1599]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 35</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FFACFFCB7DD7F9A7F107DF3E" box="[956,1036,1573,1599]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="21" pageNumber="22">70–71</figureCitation>
|
||
); and 6) a relatively dark species, shading on occiput forming a profuse anastomosed pattern (
|
||
<figureCitation id="13613F94FFACFFCB7D3EF9C8F6A0DF65" box="[853,939,1610,1636]" captionStart="FIGURES 31 – 40" captionStartId="15.[151,269,1854,1878]" captionTargetBox="[191,1384,197,1829]" captionTargetId="figure@15.[180,1408,197,1829]" captionTargetPageId="15" captionText="FIGURES 31 – 40. Tortopus arenales. 31, male head, d. v.; 32, male abdominal terga VI – VII; 33, male abdominal sterna VII – VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v. v.; 37, same, l. v.; 38, penes after treatment with potash; 39 – 40, detail of apex of forceps." httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |