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<document id="E01B64002A23D715268E84A7DF348B4E" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFA6FFC17EFCFEF5F137DEB4" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis limoncocha Molineri, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="32" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="32" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="E36FB43E3303F8F2E1AE8CBAC885FCF7">A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:date id="09451D601B12A0787F964560432D41BE">2010</mods:date>
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<subSubSection id="C340709AFFA6FFC17EFCFEF5F4F5D890" box="[151,510,375,401]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BE52311FFA6FFC17EFCFEF5F4F5D890" blockId="31.[151,510,375,401]" box="[151,510,375,401]" pageId="31" pageNumber="32">
<heading id="D0AD947DFFA6FFC17EFCFEF5F4F5D890" bold="true" box="[151,510,375,401]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B92EFF03FFA6FFC17EFCFEF5F4F5D890" bold="true" box="[151,510,375,401]" pageId="31" pageNumber="32">
<taxonomicName id="4C5A5892FFA6FFC17EFCFEF5F491D890" ID-CoL="57H2D" box="[151,410,375,401]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="limoncocha" status="sp. nov.">
<emphasis id="B92EFF03FFA6FFC17EFCFEF5F491D890" bold="true" box="[151,410,375,401]" italics="true" pageId="31" pageNumber="32">Tortopsis limoncocha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D4278FFA6FFC17FCAFEF5F4F5D890" box="[417,510,375,401]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFA6FFC17EFCFE42F137DEB4" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BE52311FFA6FFC17EFCFE42F16DDB4B" blockId="31.[151,1437,448,1973]" pageId="31" pageNumber="32">
Material:
<typeStatus id="54E19DB3FFA6FFC17F66FE42F47DD8DB" box="[269,374,448,474]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
male imago from
<collectingCountry id="F34D6381FFA6FFC17C3CFE42F7CBD8DB" box="[599,704,448,474]" name="Ecuador" pageId="31" pageNumber="32">Ecuador</collectingCountry>
: Limon Cocha, río Jivino,
<quantity id="4CA28EF4FFA6FFC17A6FFE42F158D8DB" box="[1028,1107,448,474]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="32" unit="m" value="250.0">250 m</quantity>
,
<date id="FFE405D1FFA6FFC17A0BFE42F1E9D8DB" box="[1120,1250,448,474]" pageId="31" pageNumber="32" value="1984-06-06">6-VI-1984</date>
, E. Domínguez col.; allotype female imago, and
<specimenCount id="9D5CE898FFA6FFC17C5AFE67F79FD8FE" box="[561,660,485,511]" pageId="31" pageNumber="32" type="male">36 male</specimenCount>
and
<specimenCount id="9D5CE898FFA6FFC17CBAFE67F637D8FE" box="[721,828,485,511]" pageId="31" pageNumber="32" type="female">2 female</specimenCount>
imagos
<typeStatus id="54E19DB3FFA6FFC17DCCFE67F116D8FE" box="[935,1053,485,511]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
same data as
<typeStatus id="54E19DB3FFA6FFC17AA7FE67F037D8FE" box="[1228,1340,485,511]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D5CE898FFA6FFC17B22FE67F096D8FE" box="[1353,1437,485,511]" pageId="31" pageNumber="32" type="male">1 male</specimenCount>
<typeStatus id="54E19DB3FFA6FFC17EFCFD88F5F6DB25" box="[151,253,522,548]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
from
<collectingCountry id="F34D6381FFA6FFC17F2DFD88F4A6DB25" box="[326,429,522,548]" name="Ecuador" pageId="31" pageNumber="32">Ecuador</collectingCountry>
: Coca,
<date id="FFE405D1FFA6FFC17C6EFD88F789DB25" box="[517,642,522,548]" pageId="31" pageNumber="32" value="1984-04-04">4-IV-1984</date>
, E. Domínguez col. All the material in
<collectionCode id="ED4BBBD4FFA6FFC17A33FD88F185DB25" box="[1112,1166,522,548]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="31" pageNumber="32">IML</collectionCode>
except
<typeStatus id="54E19DB3FFA6FFC17A80FD88F059DB25" box="[1259,1362,522,548]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D5CE898FFA6FFC17BE6FD88F5DBDB4B" pageId="31" pageNumber="32" type="male">3 male</specimenCount>
<typeStatus id="54E19DB3FFA6FFC17EBCFDB2F44CDB4B" box="[215,327,560,586]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED4BBBD4FFA6FFC17F00FDB2F4C8DB4B" box="[363,451,560,586]" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" pageId="31" pageNumber="32">MECN</collectionCode>
,
<specimenCount id="9D5CE898FFA6FFC17FA5FDB2F715DB4B" box="[462,542,560,586]" pageId="31" pageNumber="32" type="male">1 male</specimenCount>
<typeStatus id="54E19DB3FFA6FFC17C4EFDB2F781DB4B" box="[549,650,560,586]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED4BBBD4FFA6FFC17CC6FDB2F7F8DB4B" box="[685,755,560,586]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="31" pageNumber="32">CUIC</collectionCode>
and
<specimenCount id="9D5CE898FFA6FFC17D47FDB2F670DB4B" box="[812,891,560,586]" pageId="31" pageNumber="32" type="male">1 male</specimenCount>
<typeStatus id="54E19DB3FFA6FFC17DE9FDB2F6ECDB4B" box="[898,999,560,586]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED4BBBD4FFA6FFC17A60FDB2F169DB4B" box="[1035,1122,560,586]" pageId="31" pageNumber="32">FAMU</collectionCode>
.
</paragraph>
<paragraph id="8BE52311FFA6FFC17EADFDD7F1DBDC5D" blockId="31.[151,1437,448,1973]" pageId="31" pageNumber="32">
Male imago. Length (mm): body, 11.011.8; fore wings, 10.512.5; hind wing, 4.86.0, fore leg, 4.85.5; cerci, 28.030.0. General coloration yellowish white. Head whitish shaded gray strongly between ocelli, shaded lighter on occiput except pale median area (
<figureCitation id="13613F94FFA6FFC17C86FD22F64FDBBB" box="[749,836,672,698]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 53</figureCitation>
). Antennae: scape and pedicel whitish shaded gray, flagellum hyaline. Thorax. Anterior ring of pronotum hyaline shaded widely with gray posteriorly, a whitish tranversal line separate it from the posterior ring; posterior ring hyaline becoming whitish laterally, widely shaded gray; hind margin shaded black on a transversal thin line (
<figureCitation id="13613F94FFA6FFC17DCDFC92F6F4DA2B" box="[934,1023,784,810]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 53</figureCitation>
); propleura and sternum yellowish white. Mesonotum yellowish white shaded gray except on posteromedian membranous area, shaded stronger on carinae and V-shaped mark between posterior scutal protuberances; mesopleura and sternum yellowish white shaded slightly gray on basisternum (in some males, not shaded) and margin of sclerites; furcasternal protuberances with medial margins parallel anteriorly (
<figureCitation id="13613F94FFA6FFC17D53FC27F69DDABE" box="[824,918,933,959]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 54</figureCitation>
). Metanotum yellowish white shaded gray medially, pleura and sternum paler, shaded gray medially on sternum. Legs yellowish white shaded gray on fore leg, stronger on tibiae and tarsomeres 14, claws pale. Wings membrane hyaline, longitudinal veins whitish translucent shaded gray, stronger on veins Sc and R1, cross veins pale. Abdomen whitish translucent with a thin medial black line in all terga except I and X; tergum I shaded gray laterally, terga IIVII shaded gray almost completely except on submedial elongated pale marks; terga VIIIX shaded more strongly except submedial and sublateral pale marks on terga VIIIIX. Abdominal sterna shaded gray uniformly (some males not shaded at all) except medial line and paramedian posterior spots, pale; sterna shaded strongly on indentations of nymphal gill muscles; sternum VIII with sublateral blackish longitudinal lines. Genitalia (
<figureCitation id="13613F94FFA6FFC17ECBFB7AF43BDC13" box="[160,304,1272,1298]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Figs. 5759</figureCitation>
,
<figureCitation id="13613F94FFA6FFC17F55FB7AF485DC13" box="[318,398,1272,1298]" captionStart="FIGURES 73 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="31" pageNumber="32">7374</figureCitation>
): sternum IX completely shaded gray; parastyli yellowish, long and curved at the apex (
<figureCitation id="13613F94FFA6FFC17ECBFA9FF5FCDC36" box="[160,247,1309,1335]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 58</figureCitation>
,
<figureCitation id="13613F94FFA6FFC17F6EFA9FF42DDC36" box="[261,294,1309,1335]" captionStart="FIGURES 73 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="31" pageNumber="32">73</figureCitation>
); forceps and penes whitish translucent shaded light gray on penes, penes divided from the base with strongly sclerotized and distal hook (
<figureCitation id="13613F94FFA6FFC17CE8FAC0F7D1DC5D" box="[643,730,1346,1372]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 59</figureCitation>
,
<figureCitation id="13613F94FFA6FFC17C8DFAC0F60CDC5D" box="[742,775,1346,1372]" captionStart="FIGURES 73 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="31" pageNumber="32">74</figureCitation>
). Caudal filaments whitish translucent.
</paragraph>
<paragraph id="8BE52311FFA6FFC17EADFAEAF087DF41" blockId="31.[151,1437,448,1973]" pageId="31" pageNumber="32">
Female imago. Length (mm): body, 13.015.0; fore wings, 14.016.0; hind wing, 6.06.5; cerci, 5.0. Similar to male except wing membrane slightly yellowish and longitudinal and cross veins shaded stronger with gray; vein R3 and short intercalary before it absent. Pronotum without well marked sublateral gray lines, mesosternum with relatively narrow basisternum. Abdominal terga with a pattern similar to males except on terga VIIIIX shaded more widely; the wide median band as dark as the rest. Sternum VIII with lateral Cshaped parastyli receptors (
<figureCitation id="13613F94FFA6FFC17FBCF9A4F76EDF41" box="[471,613,1574,1600]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Figs. 5556</figureCitation>
,
<figureCitation id="13613F94FFA6FFC17C1AF9A4F79ADF41" box="[625,657,1574,1600]" captionStart="FIGURES 73 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="31" pageNumber="32">75</figureCitation>
), shaded gray on inner margins, depression somewhat elongated.
</paragraph>
<paragraph id="8BE52311FFA6FFC17EADF9C9F125DFB1" blockId="31.[151,1437,448,1973]" pageId="31" pageNumber="32">
Eggs. Suboval, one of the sides slightly more pronounced than others, no attachment structures. Length, 385400 µm; width, 330 µm. Chorion slightly sculptured with hexagonal plates almost entirely, and with shallow grooves on the concave area near the more pronounced pole (
<figureCitation id="13613F94FFA6FFC17DA2F914F12ADFB1" box="[969,1057,1686,1712]" captionStart="FIGURES 73 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="31" pageNumber="32">Fig. 76</figureCitation>
).
</paragraph>
<paragraph id="8BE52311FFA6FFC17EADF939F601DFD4" blockId="31.[151,1437,448,1973]" box="[198,778,1723,1749]" pageId="31" pageNumber="32">
Etymology: from the
<typeStatus id="54E19DB3FFA6FFC17FA8F939F4FEDFD4" box="[451,501,1723,1749]" pageId="31" pageNumber="32">type</typeStatus>
locality, Limon Cocha.
</paragraph>
<paragraph id="8BE52311FFA6FFC17EADF962F137DEB4" blockId="31.[151,1437,448,1973]" pageId="31" pageNumber="32">
Discussion and diagnosis.
<taxonomicName id="4C5A5892FFA6FFC17F94F963F7EBDFFB" box="[511,736,1761,1786]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="limoncha" status="sp. nov.">
<emphasis id="B92EFF03FFA6FFC17F94F963F7EBDFFB" box="[511,736,1761,1786]" italics="true" pageId="31" pageNumber="32">Tortopsis limoncha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D4278FFA6FFC17C8CF962F649DFFB" box="[743,834,1760,1786]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished by: 1) fore wing length
<quantity id="4CA28EF4FFA6FFC17B32F962F40BDE21" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.25" metricValueMin="1.05" pageId="31" pageNumber="32" unit="mm" value="11.5" valueMax="12.5" valueMin="10.5">10.5 12.5 mm</quantity>
(male), 14.016.0 mm (female); 2) wings whitish translucent, darker on costal margin and veins; 3) parastyli relatively straight, curved apically (
<figureCitation id="13613F94FFA6FFC17CCAF8A9F7F3DE44" box="[673,760,1835,1861]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 58</figureCitation>
); 4) penes cylindrical (
<figureCitation id="13613F94FFA6FFC17A68F8A9F151DE44" box="[1027,1114,1835,1861]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Fig. 57</figureCitation>
); 5) parastylus receptors Cshaped (
<figureCitation id="13613F94FFA6FFC17E90F8D2F486DE6B" box="[251,397,1872,1898]" captionStart="FIGURES 53 59" captionStartId="18.[151,272,1342,1366]" captionTargetBox="[202,1387,197,1318]" captionTargetId="figure@18.[189,1398,194,1321]" captionTargetPageId="18" captionText="FIGURES 53 59. Tortopsis limoncocha. 53, male head, v. v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l. v.; 56, same, v. v.; 57, male genitalia, v. v.; 58, same, l. v.; 59, detail of apex of penes." httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="31" pageNumber="32">Figs. 5556</figureCitation>
,
<figureCitation id="13613F94FFA6FFC17FF2F8D2F4B0DE6B" box="[409,443,1872,1898]" captionStart="FIGURES 73 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="31" pageNumber="32">75</figureCitation>
); 6) head shaded black behind median ocellus, shaded gray on occiput except pale median area; 7) pronotum shaded rather similar to
<taxonomicName id="4C5A5892FFA6FFC17D67F8F4F131DE8E" box="[780,1082,1910,1935]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B92EFF03FFA6FFC17D67F8F4F131DE8E" box="[780,1082,1910,1935]" italics="true" pageId="31" pageNumber="32">Tortopsis obscuripennis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A5892FFA6FFC17A13F8F4F035DE8E" box="[1144,1342,1910,1935]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sarae">
<emphasis id="B92EFF03FFA6FFC17A13F8F4F035DE8E" box="[1144,1342,1910,1935]" italics="true" pageId="31" pageNumber="32">Tortopsis sarae</emphasis>
</taxonomicName>
but the sublateral isolated gray marks on posterior ring of female pronotum are thinner.
</paragraph>
</subSubSection>
</treatment>
</document>