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<mods:title id="05737D7EB90808093BA227CE5E83406D">A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<treatment id="03F39207FFA5FFC17EFCF93AF02FD823" ID-DOI="http://doi.org/10.5281/zenodo.5690591" ID-GBIF-Taxon="119386669" ID-Zenodo-Dep="5690591" LSID="urn:lsid:plazi:treatment:03F39207FFA5FFC17EFCF93AF02FD823" httpUri="http://treatment.plazi.org/id/03F39207FFA5FFC17EFCF93AF02FD823" lastPageId="31" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C340709AFFA5FFC27EFCF93AF7DCDFD3" box="[151,727,1720,1746]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BE52311FFA5FFC27EFCF93AF7DCDFD3" blockId="28.[151,727,1720,1746]" box="[151,727,1720,1746]" pageId="28" pageNumber="29">
<heading id="D0AD947DFFA5FFC27EFCF93AF7DCDFD3" bold="true" box="[151,727,1720,1746]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B92EFF03FFA5FFC27EFCF93AF7DCDFD3" bold="true" box="[151,727,1720,1746]" pageId="28" pageNumber="29">
<taxonomicName id="4C5A5892FFA5FFC27EFCF93AF4FEDFD3" ID-CoL="57H2J" authority="Dominguez" authorityName="Dominguez" box="[151,501,1720,1746]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sarae" status="comb. nov.">
<emphasis id="B92EFF03FFA5FFC27EFCF93AF446DFD3" bold="true" box="[151,333,1720,1746]" italics="true" pageId="28" pageNumber="29">Tortopsis sarae</emphasis>
(Domínguez)
</taxonomicName>
<taxonomicNameLabel id="A21D4278FFA5FFC27F97F93AF7DCDFD3" box="[508,727,1720,1746]" pageId="28" pageNumber="29">New combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFA5FFC27EFCF882F7FEDE16" box="[151,757,1792,1815]" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8BE52311FFA5FFC27EFCF882F7FEDE16" blockId="28.[151,757,1792,1815]" box="[151,757,1792,1815]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB4A043FFFA5FFC27EFCF882F7FEDE16" box="[151,757,1792,1815]" pageId="28" pageNumber="29">
<taxonomicName id="4C5A5892FFA5FFC27EFCF882F72ADE16" authority="Dominguez 1985: 71" authorityName="Dominguez" authorityPageNumber="71" authorityYear="1985" box="[151,545,1792,1815]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sarae">
<emphasis id="B92EFF03FFA5FFC27EFCF882F43DDE16" box="[151,310,1792,1815]" italics="true" pageId="28" pageNumber="29">Tortopus sarae</emphasis>
<treatmentCitation id="0AFB0500FFA5FFC27F56F882F72ADE16" author="Dominguez" box="[317,545,1792,1815]" page="71" pageId="28" pageNumber="29" year="1985">
<bibRefCitation id="EFCB5EE0FFA5FFC27F56F882F4F2DE16" author="Dominguez" box="[317,505,1792,1815]" pageId="28" pageNumber="34" refString="Dominguez, E. (1985) El genero Tortopus Needham y Murphy (Ephemeroptera: Polymitarcyidae) en la Argentina. Physis, B 43, 69 - 72." type="journal article" year="1985">Domínguez 1985</bibRefCitation>
: 71
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AFB0500FFA5FFC27C47F882F7FADE16" author="Molineri" box="[556,753,1792,1815]" page="13" pageId="28" pageNumber="29" year="2008">
<bibRefCitation id="EFCB5EE0FFA5FFC27C47F882F7C2DE16" author="Molineri" box="[556,713,1792,1815]" pageId="28" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
: 13
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFA5FFC17EFCF8C7F02FD823" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BE52311FFA5FFC27EFCF8C7F056DEF4" blockId="28.[151,1436,1861,2037]" pageId="28" pageNumber="29">
Material (
<collectionCode id="ED4BBBD4FFA5FFC27F62F8C7F44DDE5E" box="[265,326,1861,1887]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="28" pageNumber="29">IML</collectionCode>
):
<typeStatus id="54E19DB3FFA5FFC27F33F8C7F4B4DE5E" box="[344,447,1861,1887]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
and 16
<typeStatus id="54E19DB3FFA5FFC27C70F8C7F780DE5E" box="[539,651,1861,1887]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
female imagos from
<collectingCountry id="F34D6381FFA5FFC27DE8F8C7F6F7DE5E" box="[899,1020,1861,1887]" name="Argentina" pageId="28" pageNumber="29">Argentina</collectingCountry>
: Jujuy,
<quantity id="4CA28EF4FFA5FFC27A39F8C7F197DE5E" box="[1106,1180,1861,1887]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="km" value="10.0">10 km</quantity>
N Ledesma, río Zora,
<date id="FFE405D1FFA5FFC27EFCF8E8F425DE85" box="[151,302,1898,1924]" pageId="28" pageNumber="29" value="1983-12-14">14-XII-1983</date>
, E. Domínguez col. Other material reported in
<bibRefCitation id="EFCB5EE0FFA5FFC27D35F8E8F12BDE85" author="Molineri" box="[862,1056,1898,1924]" pageId="28" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri (2008)</bibRefCitation>
. New localities:
<specimenCount id="9D5CE898FFA5FFC27A8EF8E8F04FDE85" box="[1253,1348,1898,1924]" pageId="28" pageNumber="29" type="male">10 male</specimenCount>
and
<specimenCount id="9D5CE898FFA5FFC27B14F8E8F5ECDEAB" pageId="28" pageNumber="29" type="female">10 female</specimenCount>
imagos from
<collectingCountry id="F34D6381FFA5FFC27FE1F812F708DEAB" box="[394,515,1936,1962]" name="Argentina" pageId="28" pageNumber="29">Argentina</collectingCountry>
: Tucumán, Acheral, río Aranillas,
<quantity id="4CA28EF4FFA5FFC27DF2F812F6EDDEAB" box="[921,998,1936,1962]" metricMagnitude="2" metricUnit="m" metricValue="3.66" pageId="28" pageNumber="29" unit="m" value="366.0">366 m</quantity>
,
<geoCoordinate id="EE6E45D6FFA5FFC27D99F812F198DEAB" box="[1010,1171,1936,1962]" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="-27.11664">S 27º 6' 59.9&quot;</geoCoordinate>
-
<geoCoordinate id="EE6E45D6FFA5FFC27AC0F812F063DEAB" box="[1195,1384,1936,1962]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="-65.46219">W 65º 27' 43.9&quot;</geoCoordinate>
,
<date id="FFE405D1FFA5FFC27B1FF812F40DDECE" pageId="28" pageNumber="29" value="2007-12-29">29- XII-2007</date>
, C. Molineri &amp; J. Giordano cols.; 1 reared male adult and 15 nymphs from
<collectingCountry id="F34D6381FFA5FFC27A11F837F1F8DECE" box="[1146,1267,1973,1999]" name="Argentina" pageId="28" pageNumber="29">Argentina</collectingCountry>
: Tucumán, ca. Manuela Pedraza, río Balderrama,
<quantity id="4CA28EF4FFA5FFC27C44F859F777DEF4" box="[559,636,2011,2037]" metricMagnitude="2" metricUnit="m" metricValue="3.35" pageId="28" pageNumber="29" unit="m" value="335.0">335 m</quantity>
,
<geoCoordinate id="EE6E45D6FFA5FFC27CE3F859F623DEF4" box="[648,808,2011,2037]" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="-27.200167">S 27º 12' 0.6&quot;</geoCoordinate>
-
<geoCoordinate id="EE6E45D6FFA5FFC27D2BF859F6F7DEF4" box="[832,1020,2011,2037]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="-65.35841">W 65º 21' 30.3&quot;</geoCoordinate>
,
<date id="FFE405D1FFA5FFC27A63F859F19EDEF4" box="[1032,1173,2011,2037]" pageId="28" pageNumber="29" value="2009-11-14">14-XI-2009</date>
, C. Molineri col.
</paragraph>
<caption id="DF257399FFA4FFC37EFCF9BBF7FBDF73" httpUri="https://zenodo.org/record/195445/files/figure.png" pageId="29" pageNumber="30" targetBox="[348,1239,197,1568]" targetPageId="29">
<paragraph id="8BE52311FFA4FFC37EFCF9BBF7FBDF73" blockId="29.[151,1435,1593,1650]" pageId="29" pageNumber="30">
<emphasis id="B92EFF03FFA4FFC37EFCF9BBF465DF50" bold="true" box="[151,366,1593,1617]" pageId="29" pageNumber="30">FIGURES 8792.</emphasis>
<taxonomicName id="4C5A5892FFA4FFC37F1DF9B8F4E8DF50" box="[374,483,1594,1617]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFA4FFC37F1DF9B8F4E8DF50" box="[374,483,1594,1617]" italics="true" pageId="29" pageNumber="30">Tortopsis</emphasis>
</taxonomicName>
, female adults, head and pronotum. 87,
<taxonomicName id="4C5A5892FFA4FFC37DA8F9B8F16FDF50" box="[963,1124,1594,1617]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bruchianus">
<emphasis id="B92EFF03FFA4FFC37DA8F9B8F16FDF50" box="[963,1124,1594,1617]" italics="true" pageId="29" pageNumber="30">T. bruchianus</emphasis>
</taxonomicName>
; 88,
<taxonomicName id="4C5A5892FFA4FFC37AC8F9B8F042DF50" box="[1187,1353,1594,1617]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="limoncocha">
<emphasis id="B92EFF03FFA4FFC37AC8F9B8F042DF50" box="[1187,1353,1594,1617]" italics="true" pageId="29" pageNumber="30">T. limoncocha</emphasis>
</taxonomicName>
; 89,
<taxonomicName id="4C5A5892FFA4FFC37BECF9B8F427DF73" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B92EFF03FFA4FFC37BECF9B8F427DF73" italics="true" pageId="29" pageNumber="30">T. obscuripennis</emphasis>
</taxonomicName>
; 90,
<taxonomicName id="4C5A5892FFA4FFC37F09F9D9F4B4DF73" box="[354,447,1627,1650]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="B92EFF03FFA4FFC37F09F9D9F4B4DF73" box="[354,447,1627,1650]" italics="true" pageId="29" pageNumber="30">T. puella</emphasis>
</taxonomicName>
; 91,
<taxonomicName id="4C5A5892FFA4FFC37F9EF9D9F741DF73" box="[501,586,1627,1650]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sarae">
<emphasis id="B92EFF03FFA4FFC37F9EF9D9F741DF73" box="[501,586,1627,1650]" italics="true" pageId="29" pageNumber="30">T. sarae</emphasis>
</taxonomicName>
; 92,
<taxonomicName id="4C5A5892FFA4FFC37CEAF9D9F7E2DF73" box="[641,745,1627,1650]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="spatula">
<emphasis id="B92EFF03FFA4FFC37CEAF9D9F7E2DF73" box="[641,745,1627,1650]" italics="true" pageId="29" pageNumber="30">T. spatula</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE52311FFA4FFC37EADF918F161DE9A" blockId="29.[151,1436,1690,2021]" pageId="29" pageNumber="30">
Female imago. Head shaded black between ocelli, occiput without marks. Pronotum with sublateral gray lines not completely separated from the rest of the pigmented area. Furcasternal plates basally contiguous (
<figureCitation id="13613F94FFA4FFC37ECBF967F5F5DFFE" box="[160,254,1765,1791]" captionStart="FIGURES 44 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="29" pageNumber="30">Fig. 45</figureCitation>
). Wings: veins C, Sc y R1 (and membrane in between) shaded with brownish gray, rest of wing whitish. Vein R3 and short intercalary before it absent in fore wings but some females present a weak vein in zig-zag formed by coalescence of cross veins, with a length of 3 cells or less. Abdominal terga widely pigmented except on sublateral dashes on terga IIVII, black thin median line present on terga IVIII, tergum IX with a thin pale median line. Sternum VIII with C-shaped sockets (
<figureCitation id="13613F94FFA4FFC37DA6F803F156DE9A" box="[973,1117,1921,1947]" captionStart="FIGURES 44 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="29" pageNumber="30">Figs. 4647</figureCitation>
).
</paragraph>
<paragraph id="8BE52311FFA4FFC37EADF824F5FEDEE4" blockId="29.[151,1436,1690,2021]" pageId="29" pageNumber="30">Egg. Suboval, no attachment structures. Length, 355375 µm; width, 290305 µm. The chorionic cover is smooth.</paragraph>
<caption id="DF257399FFA7FFC07EFCF8D2F710DE89" httpUri="https://zenodo.org/record/195446/files/figure.png" pageId="30" pageNumber="31" targetBox="[188,1398,197,1845]" targetPageId="30">
<paragraph id="8BE52311FFA7FFC07EFCF8D2F710DE89" blockId="30.[151,1437,1872,1928]" pageId="30" pageNumber="31">
<emphasis id="B92EFF03FFA7FFC07EFCF8D2F468DE69" bold="true" box="[151,355,1872,1896]" pageId="30" pageNumber="31">FIGURES 9396.</emphasis>
<taxonomicName id="4C5A5892FFA7FFC07F01F8D2F72ADE66" box="[362,545,1872,1895]" class="Insecta" family="Polymitarcyidae" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="family">Polymitarcyidae</taxonomicName>
, nymphal head: 93,
<taxonomicName id="4C5A5892FFA7FFC07D6AF8D2F674DE66" box="[769,895,1872,1895]" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFA7FFC07D6AF8D2F674DE66" box="[769,895,1872,1895]" italics="true" pageId="30" pageNumber="31">Asthenopus</emphasis>
</taxonomicName>
sp.; 94,
<taxonomicName id="4C5A5892FFA7FFC07D8BF8D2F156DE66" box="[992,1117,1872,1895]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFA7FFC07D8BF8D2F156DE66" box="[992,1117,1872,1895]" italics="true" pageId="30" pageNumber="31">Campsurus</emphasis>
</taxonomicName>
near
<taxonomicName id="4C5A5892FFA7FFC07AF0F8D2F008DE66" box="[1179,1283,1872,1895]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="violaceus">
<emphasis id="B92EFF03FFA7FFC07AF0F8D2F008DE66" box="[1179,1283,1872,1895]" italics="true" pageId="30" pageNumber="31">violaceus</emphasis>
</taxonomicName>
; 95,
<taxonomicName id="4C5A5892FFA7FFC07B57F8D2F5EBDE89" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFA7FFC07B57F8D2F5EBDE89" italics="true" pageId="30" pageNumber="31">Tortopus harrisi</emphasis>
</taxonomicName>
; 96,
<taxonomicName id="4C5A5892FFA7FFC07F7EF8F3F718DE89" box="[277,531,1905,1928]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B92EFF03FFA7FFC07F7EF8F3F718DE89" box="[277,531,1905,1928]" italics="true" pageId="30" pageNumber="31">Tortopsis obscuripennis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE52311FFA7FFC17EADF834F02FD823" blockId="30.[151,1437,1974,2038]" lastBlockId="31.[151,1436,152,290]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
Discussion and diagnosis.
<taxonomicName id="4C5A5892FFA7FFC07C66F835F7CCDED1" box="[525,711,1975,2000]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sarae">
<emphasis id="B92EFF03FFA7FFC07C66F835F7CCDED1" box="[525,711,1975,2000]" italics="true" pageId="30" pageNumber="31">Tortopsis sarae</emphasis>
</taxonomicName>
is currently known from all the stages (
<bibRefCitation id="EFCB5EE0FFA7FFC07AA8F834F09CDED1" author="Dominguez" box="[1219,1431,1974,2000]" pageId="30" pageNumber="34" refString="Dominguez, E. (1985) El genero Tortopus Needham y Murphy (Ephemeroptera: Polymitarcyidae) en la Argentina. Physis, B 43, 69 - 72." type="journal article" year="1985">Domínguez 1985</bibRefCitation>
,
<bibRefCitation id="EFCB5EE0FFA7FFC07EFCF85EF44FDEF7" author="Molineri" box="[151,324,2012,2038]" pageId="30" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
). A short description of the female imago is given to complete its characterization. This species can be recognized by: 1) fore wing length 13.013.5 (male), 17.518.0 (female); 2) wings whitish translucent except C and Sc areas brownish; 3) parastyli moderately curved; 4) penes cylindrical; 5) female parastylus receptor C-shaped (
<figureCitation id="13613F94FFA6FFC17F17FF60F706D9FD" box="[380,525,226,252]" captionStart="FIGURES 44 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="31" pageNumber="32">Figs. 4647</figureCitation>
); 6) head shaded black behind median ocellus, occiput very pale, at most with a small grayish netting near hind margin; 7) female pronotum shaded as in
<taxonomicName id="4C5A5892FFA6FFC17A6DFE8AF016D820" box="[1030,1309,264,289]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B92EFF03FFA6FFC17A6DFE8AF016D820" box="[1030,1309,264,289]" italics="true" pageId="31" pageNumber="32">Tortopsis obscuripennis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>