198 lines
33 KiB
XML
198 lines
33 KiB
XML
<document id="B1EEE4717C1F4F1E626035CFB8D8088E" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FF99FFFF7EFCFF15F73FD823" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis spatula Molineri, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="33" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="33" updateTime="1698238640945" updateUser="plazi">
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<mods:titleInfo id="82211B1803B272CA1A359E43BF8226CE">
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<mods:title id="C65788CBD514181FC52291C0E6EFBAE6">A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:name id="2B997FECA77DB29F07D5F504D1B6C050" type="personal">
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<mods:roleTerm id="DC3EC473FEB9F05F5E0F33FB8BB84BE6">Author</mods:roleTerm>
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<mods:namePart id="85AA459ABC09C8E182F96073C14336D1">Molineri, Carlos</mods:namePart>
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<mods:title id="C0C54693BE24DDEF3909D0486BE69F82">Zootaxa</mods:title>
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<mods:part id="3BEE67CD708322C9270AEBD6BFA38B3C">
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<mods:date id="296EE57236AF7DB3E4B67654029E465E">2010</mods:date>
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<mods:number id="BCF4772E7C8BFBF503CE1652FE52A7A3">2481</mods:number>
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<mods:classification id="720F9F48C76BA9A116F295E97B4E11B5">journal article</mods:classification>
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<mods:identifier id="8DFC1C02CA5D46CEB2C1DBD38E437278" type="DOI">10.5281/zenodo.195431</mods:identifier>
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<mods:identifier id="AC4C3C1C04EE0EF3AF92C6D2EEC7B4FA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F39207FF99FFFF7EFCFF15F73FD823" ID-DOI="http://doi.org/10.5281/zenodo.5690595" ID-GBIF-Taxon="119386675" ID-Zenodo-Dep="5690595" LSID="urn:lsid:plazi:treatment:03F39207FF99FFFF7EFCFF15F73FD823" httpUri="http://treatment.plazi.org/id/03F39207FF99FFFF7EFCFF15F73FD823" lastPageId="33" lastPageNumber="33" pageId="32" pageNumber="33">
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<subSubSection id="C340709AFF99FFFE7EFCFF15F4C3D9B0" box="[151,456,151,177]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BE52311FF99FFFE7EFCFF15F4C3D9B0" blockId="32.[151,456,151,177]" box="[151,456,151,177]" pageId="32" pageNumber="33">
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<heading id="D0AD947DFF99FFFE7EFCFF15F4C3D9B0" bold="true" box="[151,456,151,177]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
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<emphasis id="B92EFF03FF99FFFE7EFCFF15F4C3D9B0" bold="true" box="[151,456,151,177]" pageId="32" pageNumber="33">
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<taxonomicName id="4C5A5892FF99FFFE7EFCFF15F46FD9B0" ID-CoL="57H2K" box="[151,356,151,177]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
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<emphasis id="B92EFF03FF99FFFE7EFCFF15F46FD9B0" bold="true" box="[151,356,151,177]" italics="true" pageId="32" pageNumber="33">Tortopsis spatula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FF99FFFE7F00FF15F4C3D9B0" box="[363,456,151,177]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFF99FFFF7EFCFF62F73FD823" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BE52311FF99FFFE7EFCFF62F5DFDB25" blockId="32.[151,1437,224,2004]" pageId="32" pageNumber="33">
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Material:
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<typeStatus id="54E19DB3FF99FFFE7F62FF62F47BD9FB" box="[265,368,224,250]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
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male imago from
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<collectingCountry id="F34D6381FF99FFFE7C2DFF62F7B5D9FB" box="[582,702,224,250]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
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: Depto. Amazonas, Leticia, caño km 11 hacia Tarapacá,
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<geoCoordinate id="EE6E45D6FF99FFFE7B39FF62F5CBD81E" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="-4.125">S 4º 7' 30"</geoCoordinate>
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<geoCoordinate id="EE6E45D6FF99FFFE7EACFE87F467D81E" box="[199,364,261,287]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-69.95694">W 69º 57' 25"</geoCoordinate>
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,
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<quantity id="4CA28EF4FF99FFFE7F12FE87F4BCD81E" box="[377,439,261,287]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="32" pageNumber="33" unit="m" value="93.0">93 m</quantity>
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, light trap 18–20 h,
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<date id="FFE405D1FF99FFFE7CCAFE87F613D81E" box="[673,792,261,287]" pageId="32" pageNumber="33" value="1999-01-28">28-I-1999</date>
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, Z, D & M cols.
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<typeStatus id="54E19DB3FF99FFFE7DB5FE87F15ED81E" box="[990,1109,261,287]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="9D5CE898FF99FFFE7A34FE87F1A5D81E" box="[1119,1198,261,287]" pageId="32" pageNumber="33" type="male">3 male</specimenCount>
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and
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<specimenCount id="9D5CE898FF99FFFE7A8DFE87F047D81E" box="[1254,1356,261,287]" pageId="32" pageNumber="33" type="female">1 female</specimenCount>
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imago same data as
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<typeStatus id="54E19DB3FF99FFFE7F5AFEA8F490D845" box="[305,411,298,324]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
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;
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<specimenCount id="9D5CE898FF99FFFE7FCDFEA8F707D845" box="[422,524,298,324]" pageId="32" pageNumber="33" type="female">1 female</specimenCount>
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imago from Leticia, caño km 15 hacia Tarapacá,
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<quantity id="4CA28EF4FF99FFFE7A25FEA8F187D845" box="[1102,1164,298,324]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="32" pageNumber="33" unit="m" value="93.0">93 m</quantity>
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, light trap 18–20 h,
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<date id="FFE405D1FF99FFFE7B1EFEA8F5FED86B" pageId="32" pageNumber="33" value="1999-02-10">10- II-1999</date>
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,
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<geoCoordinate id="EE6E45D6FF99FFFE7F68FED2F48DD86B" box="[259,390,336,362]" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="-4.0947223">S 4º 5' 41"</geoCoordinate>
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<geoCoordinate id="EE6E45D6FF99FFFE7FE5FED2F726D86B" box="[398,557,336,362]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-69.983604">W 69º 59' 1"</geoCoordinate>
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, Z, D & M cols.;
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<specimenCount id="9D5CE898FF99FFFE7D60FED2F666D86B" box="[779,877,336,362]" pageId="32" pageNumber="33" type="male">26 male</specimenCount>
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and
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<specimenCount id="9D5CE898FF99FFFE7DC1FED2F11ED86B" box="[938,1045,336,362]" pageId="32" pageNumber="33" type="female">7 female</specimenCount>
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imagos from
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<collectingCountry id="F34D6381FF99FFFE7AAAFED2F035D86B" box="[1217,1342,336,362]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
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: Depto. Amazonas, Parque Nacional Amacayacu, río Amacayacu,
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<quantity id="4CA28EF4FF99FFFE7D2EFEF7F68FD88E" box="[837,900,373,399]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="32" pageNumber="33" unit="m" value="93.0">93 m</quantity>
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, light trap 18–20 h,
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<date id="FFE405D1FF99FFFE7A1BFEF7F1E9D88E" box="[1136,1250,373,399]" pageId="32" pageNumber="33" value="1999-02-03">3-II-1999</date>
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,
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<geoCoordinate id="EE6E45D6FF99FFFE7A84FEF7F073D88E" box="[1263,1400,373,399]" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="-3.8077776">S 3º 48' 28"</geoCoordinate>
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<geoCoordinate id="EE6E45D6FF99FFFE7BEBFEF7F410D8B5" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-70.25584">W 70º 15' 21"</geoCoordinate>
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, Z, D & M cols.;
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<specimenCount id="9D5CE898FF99FFFE7F85FE18F735D8B5" box="[494,574,410,436]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
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from
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<collectingCountry id="F34D6381FF99FFFE7CEDFE18F7F5D8B5" box="[646,766,410,436]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
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: Depto. Amazonas, Puerto Nariño, Loreto Yacu,
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<geoCoordinate id="EE6E45D6FF99FFFE7B2AFE18F5CAD8DB" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="-3.7405555">S 3º 44' 26"</geoCoordinate>
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<geoCoordinate id="EE6E45D6FF99FFFE7EA2FE42F47DD8DB" box="[201,374,448,474]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-70.45528">W 70º 27' 19"</geoCoordinate>
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, light trap 18–20 h,
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<date id="FFE405D1FF99FFFE7C07FE42F7E8D8DB" box="[620,739,448,474]" pageId="32" pageNumber="33" value="1999-02-05">5-II-1999</date>
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, Z, D & M cols.
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<typeStatus id="54E19DB3FF99FFFE7DD8FE42F12DD8DB" box="[947,1062,448,474]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
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, allotype and 5
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<typeStatus id="54E19DB3FF99FFFE7A8DFE42F052D8DB" box="[1254,1369,448,474]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
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male imagos at MUSENUV;
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<specimenCount id="9D5CE898FF99FFFE7FDCFE67F702D8FE" box="[439,521,485,511]" pageId="32" pageNumber="33" type="male">2 male</specimenCount>
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<typeStatus id="54E19DB3FF99FFFE7C7AFE67F78FD8FE" box="[529,644,485,511]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
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at
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<collectionCode id="ED4BBBD4FF99FFFE7CC0FE67F7F8D8FE" box="[683,755,485,511]" httpUri="http://grbio.org/cool/qxst-0310" name="Centro Interdipartimentale dell'Universita Museo di Storia Naturale e del Territorio" pageId="32" pageNumber="33">CIUC</collectionCode>
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and
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<specimenCount id="9D5CE898FF99FFFE7D44FE67F68AD8FE" box="[815,897,485,511]" pageId="32" pageNumber="33" type="male">2 male</specimenCount>
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||
<typeStatus id="54E19DB3FF99FFFE7DE2FE67F6F7D8FE" box="[905,1020,485,511]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
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at
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<collectionCode id="ED4BBBD4FF99FFFE7A4FFE67F170D8FE" box="[1060,1147,485,511]" pageId="32" pageNumber="33">FAMU</collectionCode>
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, remaining
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<typeStatus id="54E19DB3FF99FFFE7B61FE67F075D8FE" box="[1290,1406,485,511]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
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at
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<collectionCode id="ED4BBBD4FF99FFFE7EFCFD88F5DADB25" box="[151,209,522,548]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="32" pageNumber="33">IML</collectionCode>
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.
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</paragraph>
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<paragraph id="8BE52311FF99FFFE7EADFDB2F097DC35" blockId="32.[151,1437,224,2004]" pageId="32" pageNumber="33">
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Male imago. Length (mm): body, 9.0–12.0; fore wing, 9.5–11.2; hind wing, 4.1–5.3; foreleg, 4.2–5.8; cerci, 26.0–33.0. General coloration yellowish white shaded heavily with brownish gray (much lighter specimens were also collected, but the description below is based on those with the better marked color pattern). Head yellowish white shaded widely with gray on dorsum, except around median ocellus, shaded stronger between lateral ocelli (similar to
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<figureCitation id="13613F94FF99FFFE7CE1FD47F7E8DBDE" box="[650,739,709,735]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 60</figureCitation>
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). Antennae: scape and pedicel yellowish white completely shaded with gray, flagellum hyaline. Thorax. Pronotum with anterior ring relatively long, strongly shaded with gray dorsally; posterior ring shaded gray except on sublateral areas. Mesonotum whitish yellow shaded widely with gray along medioparapsidal sutures and medial line, and shaded with black on a V-shaped mark between posterior scutal protuberances. Metanotum whitish yellow shaded gray except laterally. Thoracic pleura and sterna whitish yellow, darker on carinae; metasternum shaded with gray in the middle. Legs yellowish white shaded with grayish on fore femur, tibia and first tarsal segment, rest of leg shaded much lighter; middle and hind legs yellowish shaded gray except tarsi whitish. Wings. Membrane hyaline, longitudinal and cross veins translucent shaded slightly with brownish. Abdomen. Tergum I shaded widely with brownish gray except medial band, terga II–VIII shaded more extensively but with a pair of submedian pale marks increasing in size towards rear segments, also with a pale oblique dash laterally; terga IX–X shaded more strongly, IX with pale marks submedially near fore margin and laterally, the lateral pale marks are surrounded with black. Abdominal sterna shaded widely with brownish gray except on pair of submedian spots and along medial and sublateral thin bands. Genitalia (
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<figureCitation id="13613F94FF99FFFE7D33FB28F6D0DDC5" box="[856,987,1194,1220]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 63–66</figureCitation>
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,
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<figureCitation id="13613F94FF99FFFE7D8DFB28F103DDC5" box="[998,1032,1194,1220]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">77</figureCitation>
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): sternum IX shaded very strongly in posterior margin; large parastyli orangeish, curved dorsally and with a longitudinal ventral furrow along its entire length (
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<figureCitation id="13613F94FF99FFFE7F50FB77F4B4DC0E" box="[315,447,1269,1295]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 63–65</figureCitation>
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,
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<figureCitation id="13613F94FF99FFFE7FA0FB77F4E7DC0E" box="[459,492,1269,1295]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">77</figureCitation>
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); forceps and penes translucent yellowish white, shaded slightly gray on second forceps segment; spine at apex of penes blunt, semicircular (
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<figureCitation id="13613F94FF99FFFE7D31FA98F6BADC35" box="[858,945,1306,1332]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 66</figureCitation>
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,
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<figureCitation id="13613F94FF99FFFE7DD6FA98F6D5DC35" box="[957,990,1306,1332]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">77</figureCitation>
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). Caudal filament whitish translucent.
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</paragraph>
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<paragraph id="8BE52311FF99FFFE7EADFAC2F641DF3B" blockId="32.[151,1437,224,2004]" pageId="32" pageNumber="33">
|
||
Female imago. Length (mm): body, 11.0–14.0; fore wing, 12.4–15.2; hind wing, 5.2–6.5; cerci, 4.5–5.5. General coloration as in male but more strongly marked, eggs yellowish white. Head black between ocelli, with gray anastomosed marks on occiput (
|
||
<figureCitation id="13613F94FF99FFFE7CC2FA08F602DCA5" box="[681,777,1418,1444]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 60</figureCitation>
|
||
). Pronotum with wide sublateral gray band, not well distinguished from the remaining gray marks (
|
||
<figureCitation id="13613F94FF99FFFE7CD2FA32F61ADCCB" box="[697,785,1456,1482]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 60</figureCitation>
|
||
). Wings membrane hyaline slightly tinged with yellow, veins more strongly shaded with brownish. Abdomen shaded uniformly with gray on terga, except on sublateral pale dashes. Sternum VIII with V-shaped parastyli receptors on sternum VIII (
|
||
<figureCitation id="13613F94FF99FFFE7AA7FA78F054DF15" box="[1228,1375,1530,1556]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Figs. 61–62</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FF99FFFE7B07FA78F086DF15" box="[1388,1421,1530,1556]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">78</figureCitation>
|
||
), margin of socket weakly sclerotized, depression elongated.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FF99FFFE7EADF9C7F1C2DF85" blockId="32.[151,1437,224,2004]" pageId="32" pageNumber="33">
|
||
Egg (
|
||
<figureCitation id="13613F94FF99FFFE7F6EF9C7F49EDF5E" box="[261,405,1605,1631]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">Figs. 79–80</figureCitation>
|
||
). Suboval, with a long thread coiled around the convex area of the egg. Length, 330–375 µm; width, 270–300 µm. The entire surface is covered by well marked subcircular grooves.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FF99FFFE7EADF912F639DFCE" blockId="32.[151,1437,224,2004]" pageId="32" pageNumber="33">Observations. The specimens from Amacayacu are very pale, the color pattern is the same but much less marked dorsally, and almost completely absent ventrally.</paragraph>
|
||
<paragraph id="8BE52311FF99FFFE7EADF958F421DE1B" blockId="32.[151,1437,224,2004]" pageId="32" pageNumber="33">
|
||
Etymology. From the latin "
|
||
<taxonomicName id="4C5A5892FF99FFFE7C70F958F77CDFF5" box="[539,631,1754,1780]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="spatula">spatula</taxonomicName>
|
||
", meaning spoon, an allusion to the flat and thin shape of penean apical spine.
|
||
</paragraph>
|
||
<paragraph id="8BE52311FF99FFFF7EADF8A7F73FD823" blockId="32.[151,1437,224,2004]" lastBlockId="33.[151,1436,152,290]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
Discussion and diagnosis.
|
||
<taxonomicName id="4C5A5892FF99FFFE7C73F8A7F7E1DE3F" box="[536,746,1829,1854]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
|
||
<emphasis id="B92EFF03FF99FFFE7C73F8A7F7E1DE3F" box="[536,746,1829,1854]" italics="true" pageId="32" pageNumber="33">Tortopsis spatula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D4278FF99FFFE7C91F8A7F66BDE3E" box="[762,864,1829,1855]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, known from imagos of both sexes, can be distinguished from all other species of the genus by: 1) fore wing length
|
||
<quantity id="4CA28EF4FF99FFFE7D99F8C8F186DE65" box="[1010,1165,1866,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.1199999999999999" metricValueMin="0.95" pageId="32" pageNumber="33" unit="mm" value="10.35" valueMax="11.2" valueMin="9.5">9.5–11.2 mm</quantity>
|
||
(male),
|
||
<quantity id="4CA28EF4FF99FFFE7A9AF8C8F097DE65" box="[1265,1436,1866,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" metricValueMax="1.52" metricValueMin="1.24" pageId="32" pageNumber="33" unit="mm" value="13.8" valueMax="15.2" valueMin="12.4">12.4–15.2 mm</quantity>
|
||
(female); 2) pale wings, veins translucent hyaline; 3) parastyli moderately curved, with a longitudinal ventral furrow (
|
||
<figureCitation id="13613F94FF99FFFE7E91F817F489DEAE" box="[250,386,1941,1967]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Fig. 63–65</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FF99FFFE7FE4F817F4BADEAE" box="[399,433,1941,1967]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">77</figureCitation>
|
||
); 4) apical spine of penes rounded and flattened (
|
||
<figureCitation id="13613F94FF99FFFE7A60F817F179DEAE" box="[1035,1138,1941,1967]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Figs. 66</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FF99FFFE7A14F817F1AADEAE" box="[1151,1185,1941,1967]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">77</figureCitation>
|
||
); 5) female parastyli receptors with V-shaped sockets (
|
||
<figureCitation id="13613F94FF99FFFE7C4AF838F7BBDED5" box="[545,688,1978,2004]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="32" pageNumber="33">Figs. 61–62</figureCitation>
|
||
,
|
||
<figureCitation id="13613F94FF99FFFE7CD7F838F7D6DED5" box="[700,733,1978,2004]" captionStart="FIGURES 73 – 80" captionStartId="27.[151,269,1875,1899]" captionTargetBox="[188,1398,197,1850]" captionTargetId="figure@27.[188,1398,197,1850]" captionTargetPageId="27" captionText="FIGURES 73 – 80. SEM photographs. Tortopsis limoncocha: 73, male genitalia, v. v.; 74, apex of penes, detail; 75, female abdominal sternum VIII, detail of left socket; 76, egg. Tortopsis spatula: 77, male genitalia, v. v. and detail of apex of penes; 78, female abdominal sternum VIII, detail of left socket; 79, egg; 80, egg, thread partially uncoiled." httpUri="https://zenodo.org/record/195443/files/figure.png" pageId="32" pageNumber="33">78</figureCitation>
|
||
); 6) head shaded black behind median ocellus, occiput with gray anastomosed marks (
|
||
<figureCitation id="13613F94FF98FFFF7FB7FF1AF731D9B3" box="[476,570,152,178]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="33" pageNumber="34">Fig. 60</figureCitation>
|
||
); 7) female pronotum as described in
|
||
<taxonomicName id="4C5A5892FF98FFFF7A7FFF1AF036D9B0" box="[1044,1341,152,177]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="obscuripennis">
|
||
<emphasis id="B92EFF03FF98FFFF7A7FFF1AF036D9B0" box="[1044,1341,152,177]" italics="true" pageId="33" pageNumber="34">Tortopsis obscuripennis</emphasis>
|
||
</taxonomicName>
|
||
but the sublateral gray mark is broad and not completely isolated from the main central shading (
|
||
<figureCitation id="13613F94FF98FFFF7ACFFF3FF1F0D9D6" box="[1188,1275,189,215]" captionStart="FIGURES 60 – 66" captionStartId="23.[151,270,1615,1639]" captionTargetBox="[267,1354,408,1573]" captionTargetId="figure@23.[229,1358,383,1589]" captionTargetPageId="23" captionText="FIGURES 60 – 66. Tortopsis spatula: 60, female head; 61, female abdominal sternum VIII, v. v.; 62, same, l. v.; 63, male genitalia after potash (left forceps and penes omitted); 64, male genitalia, v. v; 65, same, l. v.; 66, details of penes." httpUri="https://zenodo.org/record/195441/files/figure.png" pageId="33" pageNumber="34">Fig. 60</figureCitation>
|
||
). The form of the penean spine suggests a close relationship with
|
||
<taxonomicName id="4C5A5892FF98FFFF7C87FF60F694D9FA" box="[748,927,226,251]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="unguiculatus">
|
||
<emphasis id="B92EFF03FF98FFFF7C87FF60F694D9FA" box="[748,927,226,251]" italics="true" pageId="33" pageNumber="34">T. unguiculatus</emphasis>
|
||
</taxonomicName>
|
||
, as recovered in shortest phylogenetic trees with the reduced data set (
|
||
<figureCitation id="13613F94FF98FFFF7FA1FE8AF706D823" box="[458,525,264,290]" captionStart="FIGURE 1. A" captionStartId="4.[151,256,1731,1755]" captionTargetBox="[207,1379,197,1701]" captionTargetId="figure@4.[207,1380,197,1701]" captionTargetPageId="4" captionText="FIGURE 1. A, strict consensus of 234 shortest trees obtained with the complete data set (58 steps, Ci = 81, Ri = 93); B, strict consensus of 3 shortest trees obtained with the reduced data set (58 steps, Ci = 81, Ri = 92). Values near nodes indicate group support (frequency difference / absolute Bremer Support / relative BS). Black circles indicate unique apomorphies, white circles indicate parallelism or reversal, the small numbers above and below these marks are character and state numbers respectively." httpUri="https://zenodo.org/record/195432/files/figure.png" pageId="33" pageNumber="34">Fig. 1</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |