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<document id="0B9A94991933A698E259FB6E10068E7D" ID-DOI="10.1093/zoolinnean/zlad078" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240120" ID-ZooBank="5D6C7EBF-E568-4100-9364-2DD357003878" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716361325736" checkinUser="plazi" docAuthor="Marceniuk, Alexandre Pires, Oliveira, Claudio &amp; Ferraris Jr, Carl J." docDate="2024" docId="03F32B77FFFBFFBC0EAAFA0DFBA54FFD" docLanguage="en" docName="zlad078.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad078" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Betancurichthys Marceniuk &amp; Oliveira &amp; Ferraris Jr 2024, gen. nov." docType="treatment" docVersion="2" lastPageNumber="455" masterDocId="FFCA530FFFE7FFA10D44FFD6FFAB4A1E" masterDocTitle="A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data" masterLastPageNumber="476" masterPageNumber="426" pageNumber="454" updateTime="1716362088827" updateUser="ExternalLinkService">
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<mods:titleInfo id="CD45AC7CD79A8E415A92FADB5B1C5158">
<mods:title id="7841AF625B6910387CBF8FD61A242E35">A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data</mods:title>
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<mods:roleTerm id="0AD646126E104951FD68710A2C2CA3FC">Author</mods:roleTerm>
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<mods:namePart id="50FD96E313B6843963DEFA88602FFD38">Marceniuk, Alexandre Pires</mods:namePart>
<mods:affiliation id="0CDCA91F5F25CBCCFC77358B9D83F846">Programa de Pós-Graduação em Ciências Biológicas, Universidade Federal da Paraíba, conjunto Presidente Castelo Branco III, João Pessoa, PB 58050 - 000, Brazil &amp; Programa de Pós-Graduação em Ecologia e Conservação, Universidade Estadual da Paraíba, Campina Grande, PB 58429 - 500, Brazil &amp; Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
<mods:nameIdentifier id="355E8D8E640143809F3BA11467B33C65" type="email">a_marceniuk@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="885A0F345D2D6752849DB64EDEC17740">Author</mods:roleTerm>
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<mods:namePart id="E9CCA0898E93BC1235FD203C82DA212D">Oliveira, Claudio</mods:namePart>
<mods:affiliation id="7E925E123CDDB9240FE6C29A2CEE6271">Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
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<mods:namePart id="97188312F89F4D25E335CF36BE787930">Ferraris Jr, Carl J.</mods:namePart>
<mods:affiliation id="403EC3F157C347798A12D7E813D3B69B">545 NE Floral Pl., Portland, OR 97232, USA</mods:affiliation>
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<mods:title id="C7104497B0A3F9AC7567BCA27AB7795F">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="97408E9C036323511F9ACDEA68BF3760">2024</mods:date>
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<mods:number id="B230B03F48EE97065AF675C29FC9BEE8">2023-08-21</mods:number>
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<mods:number id="2E6039DA545075DDCC4C9FE9AF9802AB">200</mods:number>
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<mods:number id="0486344A0DB624D9F6DC168474577512">2</mods:number>
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<mods:classification id="9AC3F0D365E1CBAD7FBD616B6C5D6B71">journal article</mods:classification>
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<mods:identifier id="33E86D1AF9C51144E054C38235875EE8" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F32B77FFFBFFBC0EAAFA0DFBA54FFD" LSID="urn:lsid:plazi:treatment:03F32B77FFFBFFBC0EAAFA0DFBA54FFD" httpUri="http://treatment.plazi.org/id/03F32B77FFFBFFBC0EAAFA0DFBA54FFD" lastPageId="29" lastPageNumber="455" pageId="28" pageNumber="454">
<subSubSection id="C340C9EAFFFBFFBD0EAAFA0DFB454FE8" box="[1006,1262,1499,1526]" pageId="28" pageNumber="454" type="nomenclature">
<paragraph id="8BE59A61FFFBFFBD0EAAFA0DFB454FE8" blockId="28.[1006,1262,1499,1526]" box="[1006,1262,1499,1526]" pageId="28" pageNumber="454">
<heading id="D0AD2D0DFFFBFFBD0EAAFA0DFB454FE8" bold="true" box="[1006,1262,1499,1526]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="454" reason="2">
<emphasis id="B92E4673FFFBFFBD0EAAFA0DFB454FE8" bold="true" box="[1006,1262,1499,1526]" pageId="28" pageNumber="454">
<taxonomicName id="4C5AE1E2FFFBFFBD0EAAFA0DFB254FE8" authority="Marceniuk &amp; Oliveira &amp; Ferraris Jr, 2024" authorityName="Marceniuk &amp; Oliveira &amp; Ferraris Jr" authorityYear="2024" box="[1006,1166,1499,1526]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="454" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B92E4673FFFBFFBD0EAAFA0DFB254FE8" bold="true" box="[1006,1166,1499,1526]" italics="true" pageId="28" pageNumber="454">Betancurichthys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21DFB08FFFBFFBD09D7FA0AFB454FE8" box="[1171,1262,1500,1526]" pageId="28" pageNumber="454" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C340C9EAFFFBFFBC0960F9DEFBA54FFD" lastPageId="29" lastPageNumber="455" pageId="28" pageNumber="454" type="description">
<paragraph id="8BE59A61FFFBFFBD0960F9DEFB124C3C" blockId="28.[1060,1209,1544,1570]" box="[1060,1209,1544,1570]" pageId="28" pageNumber="454">
<emphasis id="B92E4673FFFBFFBD0960F9DEFB124C3C" bold="true" box="[1060,1209,1544,1570]" pageId="28" pageNumber="454">
(
<figureCitation id="136186E4FFFBFFBD096BF9DEFB234C3C" box="[1071,1160,1544,1570]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[129,194,1805,1829]" captionStartId-1="3.[825,890,1803,1827]" captionStartId-2="4.[113,178,1813,1837]" captionTargetBox-1="[153,1434,148,1768]" captionTargetBox-2="[151,708,146,1775]" captionTargetId-2="figure-643@4.[149,717,144,1776]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-1="Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-2="Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." figureDoi-1="http://doi.org/10.5281/zenodo.11240122" figureDoi-2="http://doi.org/10.5281/zenodo.11240126" httpUri-1="https://zenodo.org/record/11240122/files/figure.png" httpUri-2="https://zenodo.org/record/11240126/files/figure.png" pageId="28" pageNumber="454">Figs 1–3</figureCitation>
,
<figureCitation id="136186E4FFFBFFBD09D7F9DEFB044C3C" box="[1171,1199,1544,1570]" captionStart="Figure 27" captionStartId="29.[129,194,1285,1309]" captionTargetBox="[129,1473,763,1257]" captionTargetId="figure-263@29.[129,1473,763,1257]" captionTargetPageId="29" captionText="Figure 27. Betancurichthys madagascariensis, MNHN 1894-0002, western Madagascar, type species of the genus." figureDoi="http://doi.org/10.5281/zenodo.11240175" httpUri="https://zenodo.org/record/11240175/files/figure.png" pageId="28" pageNumber="454">27</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BE59A61FFFBFFBD0E6EF9EAFB094C6D" blockId="28.[810,1459,1596,1651]" pageId="28" pageNumber="454">
<emphasis id="B92E4673FFFBFFBD0E6EF9EAFBBE4C4A" box="[810,1045,1596,1620]" italics="true" pageId="28" pageNumber="454">ZooBank registration:</emphasis>
<uri id="FFCB9663FFFBFFBD0976F9EAFB364C6D" pageId="28" pageNumber="454">
urn:lsid:zoobank.org:act:
<uuid id="FFFCA0B4FFFBFFBD0872F9EAFB364C6D" pageId="28" pageNumber="454">CF91C445- 9AA0-4852-8291-6F9FD57C9CD2</uuid>
</uri>
.
</paragraph>
<paragraph id="8BE59A61FFFBFFBD0E6EF95FFA8D4CBE" blockId="28.[810,1318,1672,1696]" box="[810,1318,1672,1696]" pageId="28" pageNumber="454">
<emphasis id="B92E4673FFFBFFBD0E6EF95FFC0D4CBE" box="[810,934,1673,1696]" italics="true" pageId="28" pageNumber="454">
<typeStatus id="54E124C3FFFBFFBD0E6EF95FFCF14CBE" box="[810,858,1673,1696]" pageId="28" pageNumber="454">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C5AE1E2FFFBFFBD0EEFF95EFA894CBE" authority="Vaillant, 1894" authorityName="Vaillant" authorityYear="1894" box="[939,1314,1672,1696]" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="28" pageNumber="454" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="B92E4673FFFBFFBD0EEFF95EFB264CBE" box="[939,1165,1672,1696]" italics="true" pageId="28" pageNumber="454">Arius madagascariensis</emphasis>
Vaillant, 1894
</taxonomicName>
.
</paragraph>
<paragraph id="8BE59A61FFFBFFBD0904F967FB364CD4" blockId="28.[1088,1181,1713,1738]" box="[1088,1181,1713,1738]" pageId="28" pageNumber="454">
<emphasis id="B92E4673FFFBFFBD0904F967FB364CD4" box="[1088,1181,1713,1738]" italics="true" pageId="28" pageNumber="454">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE59A61FFFBFFBD0E6EF901FAAB4DD4" blockId="28.[810,1459,1751,1994]" pageId="28" pageNumber="454">Bony bridge formed by lateral ethmoid and frontal cylindrical and thin (16, 0&gt; 1); frontal mesial laminar projection absent (23, 1&gt; 0); posterior cranial fontanel formed by frontals and parieto-supraoccipital (25, 1&gt; 0); otolith posterior margin rounded (74, 1&gt; 0); dorsal crest of premaxilla beginning between lateral one-third or one-half of anterior margin (124, 0&gt; 1); third basibranchial chalice shaped (168, 0&gt; 1); dorsal-fin spine prolonged into filament (221, 0&gt; 1).</paragraph>
<paragraph id="8BE59A61FFFAFFBC0E3DFF9CFA694A41" blockId="29.[889,1474,74,96]" box="[889,1474,74,96]" pageId="29" pageNumber="455">
Family
<taxonomicName id="4C5AE1E2FFFAFFBC0EFFFF9CFC564A7E" authorityName="Bleeker" authorityYear="1858" box="[955,1021,74,96]" family="Ariidae" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="family">Ariidae</taxonomicName>
classification: total-evidence hypothesis
<emphasis id="B92E4673FFFAFFBC0838FF9DFA694A41" bold="true" box="[1404,1474,74,96]" pageId="29" pageNumber="455">• 455</emphasis>
</paragraph>
<caption id="DF25CAE9FFFAFFBC0DC5FD4EFF1548D2" ID-DOI="http://doi.org/10.5281/zenodo.11240173" ID-Zenodo-Dep="11240173" httpUri="https://zenodo.org/record/11240173/files/figure.png" pageId="29" pageNumber="455" startId="29.[129,194,664,688]" targetBox="[129,1473,135,636]" targetPageId="29" targetType="figure">
<paragraph id="8BE59A61FFFAFFBC0DC5FD4EFF1548D2" blockId="29.[129,1413,664,716]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0DC5FD4EFF4D48AE" bold="true" box="[129,230,664,688]" pageId="29" pageNumber="455">Figure 26.</emphasis>
<taxonomicName id="4C5AE1E2FFFAFFBC0DAEFD4EFDA448B1" authority=", FMNH" authorityName="FMNH" box="[234,527,664,688]" family="Ariidae" genus="Batrachocephalus" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="mino">
<emphasis id="B92E4673FFFAFFBC0DAEFD4EFE1748AE" box="[234,444,664,688]" italics="true" pageId="29" pageNumber="455">Batrachocephalus mino</emphasis>
, FMNH
</taxonomicName>
68052 (senior synonym of
<taxonomicName id="4C5AE1E2FFFAFFBC0E56FD4EFBB848AE" authorityName="Bleeker" authorityYear="1846" box="[786,1043,664,688]" family="Ariidae" genus="Batrachocephalus" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="ageneiosus">
<emphasis id="B92E4673FFFAFFBC0E56FD4EFBB848AE" box="[786,1043,664,688]" italics="true" pageId="29" pageNumber="455">Batrachocephalus ageneiosus</emphasis>
</taxonomicName>
), Indo-West Pacific, type species of the genus.
</paragraph>
</caption>
<caption id="DF25CAE9FFFAFFBC0DC5FAD3FB344F00" ID-DOI="http://doi.org/10.5281/zenodo.11240175" ID-Zenodo-Dep="11240175" box="[129,1183,1285,1310]" httpUri="https://zenodo.org/record/11240175/files/figure.png" pageId="29" pageNumber="455" startId="29.[129,194,1285,1309]" targetBox="[129,1473,763,1257]" targetPageId="29" targetType="figure">
<paragraph id="8BE59A61FFFAFFBC0DC5FAD3FB344F00" blockId="29.[129,1183,1285,1310]" box="[129,1183,1285,1310]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0DC5FAD3FF4E4F00" bold="true" box="[129,229,1285,1310]" pageId="29" pageNumber="455">Figure 27.</emphasis>
<taxonomicName id="4C5AE1E2FFFAFFBC0DADFAD0FD0D4F00" authority=", MNHN 1894" authorityName=", MNHN" authorityYear="1894" box="[233,678,1286,1310]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="B92E4673FFFAFFBC0DADFAD0FDB24F00" box="[233,537,1286,1310]" italics="true" pageId="29" pageNumber="455">Betancurichthys madagascariensis</emphasis>
, MNHN 1894
</taxonomicName>
-0002, western Madagascar, type species of the genus.
</paragraph>
</caption>
<paragraph id="8BE59A61FFFAFFBC0DC5FA98FE3A4F9B" blockId="29.[129,777,1358,1413]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0DC5FA98FE2F4F7B" box="[129,388,1358,1382]" italics="true" pageId="29" pageNumber="455">Ambiguous optimization:</emphasis>
Sesamoid bone I very long and subtriangular (145, 0&gt; 1).
</paragraph>
<paragraph id="8BE59A61FFFAFFBC0CD6FA72FE534FA0" blockId="29.[402,504,1444,1470]" box="[402,504,1444,1470]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0CD6FA72FE534FA0" box="[402,504,1444,1470]" italics="true" pageId="29" pageNumber="455">Etymology</emphasis>
</paragraph>
<paragraph id="8BE59A61FFFAFFBC0DC5FA1DFD094C3C" blockId="29.[129,777,1483,1570]" pageId="29" pageNumber="455">Named after Colombian ichthyologist Ricardo Betancur-R. for his dedication to the knowledge of the taxonomy, evolution, and biogeography of marine catfishes. Gender: masculine.</paragraph>
<paragraph id="8BE59A61FFFAFFBC0C39F997FDA64C45" blockId="29.[381,525,1601,1627]" box="[381,525,1601,1627]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0C39F997FDA64C45" box="[381,525,1601,1627]" italics="true" pageId="29" pageNumber="455">Included species</emphasis>
</paragraph>
<paragraph id="8BE59A61FFFAFFBC0DC5F9BEFCAF4CA0" blockId="29.[129,772,1640,1727]" pageId="29" pageNumber="455">
<taxonomicName id="4C5AE1E2FFFAFFBC0DC5F9BEFD874C9E" authority="Ng &amp; Sparks, 2003" authorityName="Ng &amp; Sparks" authorityYear="2003" box="[129,556,1640,1664]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="festinus">
<emphasis id="B92E4673FFFAFFBC0DC5F9BEFECC4C9E" box="[129,359,1640,1664]" italics="true" pageId="29" pageNumber="455">Betancurichthys festinus</emphasis>
Ng &amp; Sparks, 2003
</taxonomicName>
*
<emphasis id="B92E4673FFFAFFBC0F04F9BEFD674C9E" box="[576,716,1640,1664]" italics="true" pageId="29" pageNumber="455">sedis mutabilis</emphasis>
<taxonomicName id="4C5AE1E2FFFAFFBC0DD8F951FDDD4C81" authority="Vaillant, 1894" authorityName="Vaillant" authorityYear="1894" box="[156,630,1671,1695]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="B92E4673FFFAFFBC0DD8F951FE484C81" box="[156,483,1671,1695]" italics="true" pageId="29" pageNumber="455">Betancurichthys madagascariensis</emphasis>
Vaillant, 1894
</taxonomicName>
<taxonomicName id="4C5AE1E2FFFAFFBC0DD8F970FDF54CA0" authority="Ng &amp; Sparks, 2003" authorityName="Ng &amp; Sparks" authorityYear="2003" box="[156,606,1702,1727]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="uncinatus">
<emphasis id="B92E4673FFFAFFBC0DD8F970FE324CA0" box="[156,409,1702,1726]" italics="true" pageId="29" pageNumber="455">Betancurichthys uncinatus</emphasis>
Ng &amp; Sparks, 2003
</taxonomicName>
*
<emphasis id="B92E4673FFFAFFBC0F36F970FD554CA0" box="[626,766,1702,1726]" italics="true" pageId="29" pageNumber="455">sedis mutabilis</emphasis>
.
</paragraph>
<paragraph id="8BE59A61FFFAFFBC0DC5F936FEF44D09" blockId="29.[129,778,1760,1815]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0DC5F936FE294CE6" box="[129,386,1760,1784]" italics="true" pageId="29" pageNumber="455">Habitat and distribution:</emphasis>
Fresh and brackish waters, western
<collectingCountry id="F34DDAF1FFFAFFBC0DC5F929FF514D09" box="[129,250,1791,1815]" name="Madagascar" pageId="29" pageNumber="455">Madagascar</collectingCountry>
(
<figureCitation id="136186E4FFFAFFBC0C4DF929FEE54D09" box="[265,334,1791,1815]" captionStart="Figure 24" captionStartId="27.[129,194,1851,1875]" captionTargetBox="[259,1343,151,1822]" captionTargetId="figure-8@27.[257,1345,145,1823]" captionTargetPageId="27" captionText="Figure 24. Representative skulls, in dorsal view, and distribution of the subtribe Ariina.A, Arius maculatus, AUM 44875. B, Cephalocassis melanochir, CAS 49426. C, Cryptarius truncatus, AUM 50243. D, Hexanematichthys sagor, AUM 50242. E, Kyataphisa nenga, AUM 46280. F, Osteogeneiosus militaris, AUM 44880. G, Pseudosciades sona, AUM 44861. * marine, ** brackish water, *** freshwater." figureDoi="http://doi.org/10.5281/zenodo.11240169" httpUri="https://zenodo.org/record/11240169/files/figure.png" pageId="29" pageNumber="455">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8BE59A61FFFAFFBC0CDFF8E0FE444D4E" blockId="29.[411,495,1846,1872]" box="[411,495,1846,1872]" pageId="29" pageNumber="455">
<emphasis id="B92E4673FFFAFFBC0CDFF8E0FE444D4E" box="[411,495,1846,1872]" italics="true" pageId="29" pageNumber="455">Remarks</emphasis>
</paragraph>
<paragraph id="8BE59A61FFFAFFBC0DC5F88BFBA54FFD" blockId="29.[129,778,1885,1972]" lastBlockId="29.[825,1474,1358,1507]" pageId="29" pageNumber="455">
The genus
<taxonomicName id="4C5AE1E2FFFAFFBC0DB9F88BFE3C4D6B" authorityName="Marceniuk &amp; Oliveira &amp; Ferraris Jr" authorityYear="2024" box="[253,407,1885,1909]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFFAFFBC0DB9F88BFE3C4D6B" box="[253,407,1885,1909]" italics="true" pageId="29" pageNumber="455">Betancurichthys</emphasis>
</taxonomicName>
was established to accommodate species endemic to
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, in the western Indian Ocean, corroborating molecular results presented by Betancur-R. (2009). Its inclusion in the
<taxonomicName id="4C5AE1E2FFFAFFBC091BFA98FB344F78" authorityName="Bleeker" authorityYear="1858" box="[1119,1183,1358,1382]" family="Ariidae" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="458" phylum="Chordata" rank="subTribe" subTribe="Ariina">Ariina</taxonomicName>
is supported by the BI and ML analyses. The inclusion of
<taxonomicName id="4C5AE1E2FFFAFFBC09D9FABBFA264F9B" authorityName="Ng &amp; Sparks" authorityYear="2003" box="[1181,1421,1389,1413]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="festinus">
<emphasis id="B92E4673FFFAFFBC09D9FABBFA264F9B" box="[1181,1421,1389,1413]" italics="true" pageId="29" pageNumber="455">Betancurichthys festinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE1E2FFFAFFBC0E7DFA5AFB914FBA" authorityName="Ng &amp; Sparks" authorityYear="2003" box="[825,1082,1420,1444]" family="Ariidae" genus="Betancurichthys" kingdom="Animalia" order="Siluriformes" pageId="29" pageNumber="455" phylum="Chordata" rank="species" species="uncinatus">
<emphasis id="B92E4673FFFAFFBC0E7DFA5AFB914FBA" box="[825,1082,1420,1444]" italics="true" pageId="29" pageNumber="455">Betancurichthys uncinatus</emphasis>
</taxonomicName>
in the genus is provisional, based on the distribution and life habits of the species restricted to freshwater in
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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