203 lines
26 KiB
XML
203 lines
26 KiB
XML
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<mods:title id="DEE934656C7DD03636D0904CE7E5DD30">On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands</mods:title>
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<mods:title id="1089387F85274A19FA25736E8749CF31">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="8B92DF10924D106E91700D84E21B7A1D">2014</mods:date>
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<treatment id="03F3175FEF6C5E46FC7A3C94FC3AF9F2" ID-DOI="http://doi.org/10.5281/zenodo.4775629" ID-GBIF-Taxon="182237992" ID-Zenodo-Dep="4775629" LSID="urn:lsid:plazi:treatment:03F3175FEF6C5E46FC7A3C94FC3AF9F2" httpUri="http://treatment.plazi.org/id/03F3175FEF6C5E46FC7A3C94FC3AF9F2" lastPageNumber="118" pageId="3" pageNumber="118">
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<subSubSection id="C340F5C2EF6C5E46FC7A3C94FAE6FF29" box="[932,1332,191,215]" pageId="3" pageNumber="118" type="nomenclature">
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<paragraph id="8BE5A649EF6C5E46FC7A3C94FAE6FF29" blockId="3.[932,1332,191,215]" box="[932,1332,191,215]" pageId="3" pageNumber="118">
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<heading id="D0AD1125EF6C5E46FC7A3C94FAE6FF29" bold="true" box="[932,1332,191,215]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="118" reason="2">
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<taxonomicName id="4C5ADDCAEF6C5E46FC7A3C94FAE6FF29" ID-CoL="4RYCR" authority="(Martens, 1863)" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[932,1332,191,215]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
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<emphasis id="B92E7A5BEF6C5E46FC7A3C94FBACFF29" bold="true" box="[932,1150,191,215]" italics="true" pageId="3" pageNumber="118">Rhagada solorensis</emphasis>
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(Martens, 1863)
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C340F5C2EF6C5E46FCF33CD5FC70FE78" pageId="3" pageNumber="118" type="reference_group">
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<paragraph id="8BE5A649EF6C5E46FCF33CD5FACBFEB3" blockId="3.[813,1453,254,390]" pageId="3" pageNumber="118">
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<taxonomicName id="4C5ADDCAEF6C5E46FCF33CD5FBADFEED" ID-CoL="6LM2Q" authority="Martens, 1863: 179" authorityName="Martens" authorityPageNumber="179" authorityYear="1863" box="[813,1151,254,275]" class="Gastropoda" family="Camaenidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
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<emphasis id="B92E7A5BEF6C5E46FCF33CD5FC11FEED" box="[813,963,254,275]" italics="true" pageId="3" pageNumber="118">Helix solorensis</emphasis>
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<materialsCitation id="3B32AC14EF6C5E46FC163CD5FBADFEED" ID-GBIF-Occurrence="3114366306" box="[968,1151,254,275]" collectionCode="ZMB" latitude="-8.433333" longLatPrecision="21" longitude="123.166664" pageId="3" pageNumber="118" specimenCount="1" typeStatus="syntype">Martens, 1863: 179</materialsCitation>
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</taxonomicName>
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(from ‘Lamakera, Solor’,
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<geoCoordinate id="EE6EC08EEF6C5E46FAAB3CD5FCB6FECE" degrees="8" direction="south" minutes="25" orientation="latitude" pageId="3" pageNumber="118" precision="15" seconds="60" value="-8.433333">8° 25' 60" S</geoCoordinate>
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,
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<geoCoordinate id="EE6EC08EEF6C5E46FCAE3D30FBD0FECE" box="[880,1026,283,304]" degrees="123" direction="east" minutes="10" orientation="longitude" pageId="3" pageNumber="118" precision="15" seconds="00" value="123.166664">123° 10' 00" E</geoCoordinate>
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; nine
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<typeStatus id="54E118EBEF6C5E46FBE03D30FB40FECE" box="[1086,1170,283,304]" pageId="3" pageNumber="118" type="syntype">syntypes</typeStatus>
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ZMB Moll. 5587, not seen).
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<taxonomicName id="4C5ADDCAEF6C5E46FCF33D13FBE5FEB3" ID-CoL="6LM2Q" authorityName="Martens" authorityYear="1863" box="[813,1079,312,333]" class="Gastropoda" family="Camaenidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis" subGenus="Rhagada">
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<emphasis id="B92E7A5BEF6C5E46FCF33D13FCB3FEB3" box="[813,865,312,333]" italics="true" pageId="3" pageNumber="118">Helix</emphasis>
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(
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<emphasis id="B92E7A5BEF6C5E46FCAE3D13FC15FEB3" box="[880,967,312,333]" italics="true" pageId="3" pageNumber="118">Rhagada</emphasis>
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)
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<emphasis id="B92E7A5BEF6C5E46FC093D13FBE5FEB3" box="[983,1079,312,333]" italics="true" pageId="3" pageNumber="118">solorensis</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="EFCBDBB8EF6C5E46FB8D3D13FAC7FEB3" author="Martens E von" box="[1107,1301,312,333]" pageId="3" pageNumber="118" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891: 235</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BE5A649EF6C5E46FCF33D7FFC70FE78" blockId="3.[813,1453,254,390]" pageId="3" pageNumber="118">
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<treatmentCitationGroup id="AB4A8167EF6C5E46FCF33D7FFC70FE78" pageId="3" pageNumber="118">
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<taxonomicName id="4C5ADDCAEF6C5E46FCF33D7FFB9CFE97" ID-CoL="4RYCR" authority="solorensis" box="[813,1102,340,361]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="subSpecies" species="solorensis" subSpecies="solorensis">
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<emphasis id="B92E7A5BEF6C5E46FCF33D7FFB9CFE97" box="[813,1102,340,361]" italics="true" pageId="3" pageNumber="118">Rhagada solorensis solorensis</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AFB8058EF6C5E46FBBB3D7FFADEFE97" author="Rensch B" box="[1125,1292,340,361]" page="95" pageId="3" pageNumber="118" year="1931">
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<bibRefCitation id="EFCBDBB8EF6C5E46FBBB3D7FFADEFE97" author="Rensch B" box="[1125,1292,340,361]" pageId="3" pageNumber="118" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch, 1931: 95</bibRefCitation>
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||
</treatmentCitation>
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,
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<treatmentCitation id="0AFB8058EF6C5E46FAC83D7FFC97FE78" author="Maassen WJM" page="66" pageId="3" pageNumber="118" year="2009">
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||
<bibRefCitation id="EFCBDBB8EF6C5E46FAC83D7FFC97FE78" author="Maassen WJM" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009: 66</bibRefCitation>
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</treatmentCitation>
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(partim).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C340F5C2EF6C5E46FCF33D80FC72FE23" pageId="3" pageNumber="118" type="materials_examined">
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||
<paragraph id="8BE5A649EF6C5E46FCF33D80FC72FE23" blockId="3.[813,1451,427,477]" pageId="3" pageNumber="118">
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<emphasis id="B92E7A5BEF6C5E46FCF33D80FC2EFE3E" bold="true" box="[813,1020,427,448]" pageId="3" pageNumber="118">Material examined.</emphasis>
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||
<collectionCode id="ED4B3E8CEF6C5E46FBDA3D80FBF9FE3E" box="[1028,1067,427,448]" country="Australia" name="Australian Museum" pageId="3" pageNumber="118" type="Museum">AM</collectionCode>
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||
C.471154 (
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<collectingCountry id="F34DE6D9EF6C5E46FB7A3D80FADAFE3E" box="[1188,1288,427,448]" name="Indonesia" pageId="3" pageNumber="118">Indonesia</collectingCountry>
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, Solor Island, 3 wet, 2 dry).
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</paragraph>
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</subSubSection>
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<subSubSection id="C340F5C2EF6C5E46FCF33E2EFB76FB6C" pageId="3" pageNumber="118" type="discussion">
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<paragraph id="8BE5A649EF6C5E46FCF33E2EFB76FB6C" blockId="3.[813,1452,517,1170]" pageId="3" pageNumber="118">
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<emphasis id="B92E7A5BEF6C5E46FCF33E2EFBC6FDE2" bold="true" box="[813,1044,517,540]" pageId="3" pageNumber="118">Taxonomic history.</emphasis>
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Representing the first named non-Australian species,
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<taxonomicName id="4C5ADDCAEF6C5E46FBD03E0EFB73FDC2" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1038,1185,549,572]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
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<emphasis id="B92E7A5BEF6C5E46FBD03E0EFB73FDC2" box="[1038,1185,549,572]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
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</taxonomicName>
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has continuously been maintained as a distinct species in previous taxonomic treatments.
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<bibRefCitation id="EFCBDBB8EF6C5E46FC6B3E4FFAD8FD85" author="Vermeulen JJ & Whitten AJ" box="[949,1290,612,635]" pageId="3" pageNumber="118" pagination="1 - 164" refId="ref6974" refString="Vermeulen JJ & Whitten AJ (1998) Fauna Malesiana. Guide to the Land Snails of Bali. Backhuys Publishers, Leiden. Pp. 1 - 164." type="book chapter" year="1998">Vermeulen & Whitten (1998)</bibRefCitation>
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and
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<bibRefCitation id="EFCBDBB8EF6C5E46FA943E4FFCA1FD64" author="Maassen WJM" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
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reported
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<taxonomicName id="4C5ADDCAEF6C5E46FC3E3EA8FBBEFD64" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[992,1132,643,666]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
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<emphasis id="B92E7A5BEF6C5E46FC3E3EA8FBBEFD64" box="[992,1132,643,666]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
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</taxonomicName>
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from
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<collectingRegion id="499E68ABEF6C5E46FB6A3EA8FB32FD64" box="[1204,1248,643,666]" country="Indonesia" name="Bali" pageId="3" pageNumber="118">Bali</collectingRegion>
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and Nusa Penida, apparently unaware of the description of
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<taxonomicName id="4C5ADDCAEF6C5E46FAC83E88FA79FD44" authorityName="Falconieri" authorityYear="1995" box="[1302,1451,675,698]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="marghitae">
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<emphasis id="B92E7A5BEF6C5E46FAC83E88FA79FD44" box="[1302,1451,675,698]" italics="true" pageId="3" pageNumber="118">R. marghitae</emphasis>
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</taxonomicName>
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from these islands. In fact,
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<bibRefCitation id="EFCBDBB8EF6C5E46FBBE3EE9FAF2FD27" author="Maassen WJM" box="[1120,1312,706,729]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen’s (2009)</bibRefCitation>
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report of
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<taxonomicName id="4C5ADDCAEF6C5E46FA4A3EE9FC49FD07" baseAuthorityName="Martens" baseAuthorityYear="1863" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
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<emphasis id="B92E7A5BEF6C5E46FA4A3EE9FC49FD07" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
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</taxonomicName>
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is entirely based on material of
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<taxonomicName id="4C5ADDCAEF6C5E46FACD3EC9FA76FD07" authorityName="Falconieri" authorityYear="1995" box="[1299,1444,738,761]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="marghitae">
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||
<emphasis id="B92E7A5BEF6C5E46FACD3EC9FA76FD07" box="[1299,1444,738,761]" italics="true" pageId="3" pageNumber="118">R. marghitae</emphasis>
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||
</taxonomicName>
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.
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<bibRefCitation id="EFCBDBB8EF6C5E46FCF33F2AFC1AFCE6" author="Rensch B" box="[813,968,769,792]" pageId="3" pageNumber="118" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch (1931)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FC273F2AFB70FCE6" author="Maassen WJM" box="[1017,1186,769,792]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
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||
assigned
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<taxonomicName id="4C5ADDCAEF6C5E46FAD43F2AFA5DFCE6" baseAuthorityName="Martens" baseAuthorityYear="1891" box="[1290,1423,769,792]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="floresiana">
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||
<emphasis id="B92E7A5BEF6C5E46FAD43F2AFA5DFCE6" box="[1290,1423,769,792]" italics="true" pageId="3" pageNumber="118">R. floresiana</emphasis>
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||
</taxonomicName>
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||
as a second subspecies for its similar shell. However,
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<bibRefCitation id="EFCBDBB8EF6C5E46FA833F0AFCA6FCA9" author="Rensch B" pageId="3" pageNumber="118" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch (1931)</bibRefCitation>
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||
indiscriminately lumped together most Lesser Sunda species while
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<bibRefCitation id="EFCBDBB8EF6C5E46FC1F3F4BFBBEFC89" author="Maassen WJM" box="[961,1132,864,887]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
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||
confused
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FB073F4BFAB3FC89" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1241,1377,864,887]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
|
||
<emphasis id="B92E7A5BEF6C5E46FB073F4BFAB3FC89" box="[1241,1377,864,887]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FA4A3F4BFC48FC68" authorityName="Falconieri" authorityYear="1995" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="marghitae">
|
||
<emphasis id="B92E7A5BEF6C5E46FA4A3F4BFC48FC68" italics="true" pageId="3" pageNumber="118">R. marghitae</emphasis>
|
||
</taxonomicName>
|
||
. The proposals of both authors are therefore not well-substantiated. In light of generally widespread narrowrange endemism in
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FBDD3F95FBB1FC2B" authorityName="Albers" authorityYear="1860" box="[1027,1123,958,981]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92E7A5BEF6C5E46FBDD3F95FBB1FC2B" box="[1027,1123,958,981]" italics="true" pageId="3" pageNumber="118">Rhagada</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FBAC3F95FAD6FC2B" author="Kohler F" box="[1138,1284,958,981]" pageId="3" pageNumber="118" pagination="203 - 406" refId="ref6101" refString="Kohler F (2011) The camaenid species of the Kimberley Islands, Western Australia (Stylommatophora: Helicoidea). Malacologia, 54: 203 - 406." type="journal article" year="2011">Köhler, 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FACE3F95FCB4FC0A" author="Johnson MS & Hamilton ZR & Teale ROY & Kendrick PG" pageId="3" pageNumber="118" pagination="316 - 327" refId="ref6053" refString="Johnson MS, Hamilton ZR, Teale ROY & Kendrick PG (2012) Endemic evolutionary radiation of Rhagada land snails (Pulmonata: Camaenidae) in a continental archipelago in northern Western Australia. Biological Journal of the Linnean Society, 106: 316 - 327." type="journal article" year="2012">Johnson et al., 2012</bibRefCitation>
|
||
), I prefer maintaining the original treatment of Martens (1863, 1891) and consider
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FB943FD6FB00FBEA" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1098,1234,1021,1044]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
|
||
<emphasis id="B92E7A5BEF6C5E46FB943FD6FB00FBEA" box="[1098,1234,1021,1044]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
|
||
</taxonomicName>
|
||
as endemic to Solor Island. Populations from other islands should not be lumped with
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FCBD3817FC3EFBAD" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[867,1004,1084,1107]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
|
||
<emphasis id="B92E7A5BEF6C5E46FCBD3817FC3EFBAD" box="[867,1004,1084,1107]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
|
||
</taxonomicName>
|
||
in absence of evidence from comparative studies. Consequently,
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FBFB3870FB7FFB8C" baseAuthorityName="Martens" baseAuthorityYear="1891" box="[1061,1197,1115,1138]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="floresiana">
|
||
<emphasis id="B92E7A5BEF6C5E46FBFB3870FB7FFB8C" box="[1061,1197,1115,1138]" italics="true" pageId="3" pageNumber="118">R. floresiana</emphasis>
|
||
</taxonomicName>
|
||
is also maintained as a distinct species endemic to Flores.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340F5C2EF6C5E46FCF33891FBD4FA90" pageId="3" pageNumber="118" type="diagnosis">
|
||
<paragraph id="8BE5A649EF6C5E46FCF33891FBD4FA90" blockId="3.[813,1452,1210,1390]" pageId="3" pageNumber="118">
|
||
<emphasis id="B92E7A5BEF6C5E46FCF33891FC73FB2F" bold="true" box="[813,929,1210,1233]" pageId="3" pageNumber="118">Diagnosis.</emphasis>
|
||
Shell comparatively large (D =
|
||
<quantity id="4CA20BACEF6C5E46FB2C3891FAB5FB2F" box="[1266,1383,1210,1233]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="3" pageNumber="118" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
|
||
;
|
||
<tableCitation id="C6D893F2EF6C5E46FAAF3891FCEBFB0E" captionStart="Table 1" captionStartId="4.[136,189,191,212]" captionTargetPageId="4" captionText="Table 1. Shell measurements for N shells of different Rhagada species and subspecies. H = height; D = diameter; W = no. of whorls." httpUri="http://table.plazi.org/id/DF25F6C1EF6B5E41FF563C94FA79FF2A" pageId="3" pageNumber="118" tableUuid="DF25F6C1EF6B5E41FF563C94FA79FF2A">Table 1</tableCitation>
|
||
), smooth or with fine axial ribs only, umbilicus forming a chink or narrowly open, peripheral band particularly welldeveloped (
|
||
<figureCitation id="1361BACCEF6C5E46FC6C3933FBDEFAD1" box="[946,1036,1304,1327]" captionStart="Fig" captionStartId="1.[113,148,1918,1939]" captionTargetBox="[113,752,690,1894]" captionTargetId="figure-175@1.[113,752,690,1894]" captionTargetPageId="1" captionText="Fig. 1. Shells (1.5× of natural size). A, Rhagada solorensis, Solor Island, AM C.471154; B, Rhagada setzeri setzeri, Alor Island, AM C.471156; C, D, Rhagada setzeri atauroensis n. subsp., Atauro Island: C, Paratype AM C.468711; D, Paratype AM C.468709; E–H, Rhagada abbasi n. sp., Lembata Island: E, Holotype AM C.478280; F–H: Paratypes AM C.471153, showing variation in sculpture and colour pattern. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4502136" httpUri="https://zenodo.org/record/4502136/files/figure.png" pageId="3" pageNumber="118">Fig. 1A</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FBF63933FB00FAD1" author="Martens E von" box="[1064,1234,1304,1327]" pageId="3" pageNumber="118" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891</bibRefCitation>
|
||
). Penis with large, smooth penial verge with rounded tip, epiphallus with short flagellum (
|
||
<figureCitation id="1361BACCEF6C5E46FC7C397CFC28FA90" box="[930,1018,1367,1390]" captionStart="Fig" captionStartId="2.[136,171,2006,2027]" captionTargetBox="[143,1467,293,1913]" captionTargetId="figure-6@2.[129,1480,291,1924]" captionTargetPageId="2" captionText="Fig. 2. Penial anatomy, with penial chamber opened showing internal anatomy. A, Rhagada solorensis, Solor Island, AM C.471154; B, Rhagada marghitae, Nusa Penida AM C.327851; C, Rhagada setzeri atauroensis n. subsp., Atauro Island, holotype AM C.478282; D, Rhagada abbasi n. sp., Lembata Island, holotype AM C.478280. Abbreviations: ep = epiphallus; fl = epiphallic flagellum; gp = genital pore; lwp = longitudinal wall pilaster; pv = penial verge; rm = penial retractor muscle; va = vagina; vd = vas deferens. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4502138" httpUri="https://zenodo.org/record/4502138/files/figure.png" pageId="3" pageNumber="118">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340F5C2EF6C5E46FCF339BDFC3AF9F2" pageId="3" pageNumber="118" type="discussion">
|
||
<paragraph id="8BE5A649EF6C5E46FCF339BDFC3AF9F2" blockId="3.[813,1452,1430,1548]" pageId="3" pageNumber="118">
|
||
<emphasis id="B92E7A5BEF6C5E46FCF339BDFBE0FA53" bold="true" box="[813,1074,1430,1453]" pageId="3" pageNumber="118">Comparative remarks.</emphasis>
|
||
Shells figured by
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FB2039BDFA79FA53" author="Maassen WJM" box="[1278,1451,1430,1453]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
|
||
from Nusa Penida are misidentified specimens of
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FA4D399EFC4BFA12" authorityName="Falconieri" authorityYear="1995" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="marghitae">
|
||
<emphasis id="B92E7A5BEF6C5E46FA4D399EFC4BFA12" italics="true" pageId="3" pageNumber="118">R. marghitae</emphasis>
|
||
</taxonomicName>
|
||
. Most typical characteristic is smooth penial verge with rounded tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |