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<document id="E7D4A6DB7295FCE6DA14CBDAF54F9A30" ID-CLB-Dataset="31388" ID-DOI="10.11646/zootaxa.4083.4.7" ID-GBIF-Dataset="f2c3d1e3-2e79-4957-9cd2-e6c322bf1654" ID-ISSN="1175-5326" ID-Zenodo-Dep="1053923" ID-ZooBank="1B3E9DB6-7E5F-4AF7-82DF-21E71E2D8349" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510926575616" checkinUser="plazi" docAuthor="Pati, S. K., Thackeray, T. &amp; Khaire, A." docDate="2016" docId="03EC87B9FFD2FFF1FF2F677CFBC4D484" docLanguage="en" docName="zootaxa.4083.4.7.pdf" docOrigin="Zootaxa 4083 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gubernatoriana waghi Pati &amp; Thackeray &amp; Khaire 2016, n. sp." docType="treatment" docUuid="4576553B-2232-4DA0-89CD-EEA2B6485308" docUuidSource="ZooBank" docVersion="8" lastPageNumber="585" masterDocId="FFD5FFC1FFDCFFE1FFB86558FFD3D144" masterDocTitle="Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India" masterLastPageNumber="586" masterPageNumber="569" pageNumber="583" updateTime="1698500881418" updateUser="ExternalLinkService">
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<mods:title id="759AED3DFEDC71F42D790D28882A7811">Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India</mods:title>
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<mods:namePart id="BCB7FE4B93EE4C6A99283363C4CD34F9">Pati, S. K.</mods:namePart>
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<mods:namePart id="08F93FECCEB5D8EC2FC3BF7E622E43AE">Thackeray, T.</mods:namePart>
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<mods:namePart id="219BF3958F11BC666EB7F013DC83A25D">Khaire, A.</mods:namePart>
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<mods:date id="17C17E3BF2A99AB76602EAD24962A823">2016</mods:date>
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<treatment id="03EC87B9FFD2FFF1FF2F677CFBC4D484" ID-DOI="http://doi.org/10.5281/zenodo.6069956" ID-GBIF-Taxon="137453399" ID-Zenodo-Dep="6069956" LSID="urn:lsid:zoobank.org:act:4576553B-2232-4DA0-89CD-EEA2B6485308" httpUri="http://treatment.plazi.org/id/03EC87B9FFD2FFF1FF2F677CFBC4D484" lastPageId="16" lastPageNumber="585" pageId="14" pageNumber="583">
<subSubSection id="C35F6524FFD2FFEFFF2F677CFEFFD31B" pageId="14" pageNumber="583" type="nomenclature">
<paragraph id="8BFA36AFFFD2FFEFFF2F677CFDFDD37A" blockId="14.[151,558,548,607]" box="[151,558,548,574]" pageId="14" pageNumber="583">
<heading id="D0B281C3FFD2FFEFFF2F677CFDFDD37A" bold="true" box="[151,558,548,574]" fontSize="11" level="1" pageId="14" pageNumber="583" reason="1">
<emphasis id="B931EABDFFD2FFEFFF2F677CFDFDD37A" bold="true" box="[151,558,548,574]" pageId="14" pageNumber="583">
<taxonomicName id="4C454D2CFFD2FFEFFF2F677CFE36D37A" ID-CoL="84FVP" authority="Pati" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[151,485,548,574]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFD2FFEFFF2F677CFE78D37A" bold="true" box="[151,427,548,574]" italics="true" pageId="14" pageNumber="583">Gubernatoriana waghi</emphasis>
Pati
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD2FFEFFE54677CFDFDD37A" box="[492,558,548,574]" pageId="14" pageNumber="583" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA36AFFFD2FFEFFF2F671EFEFFD31B" blockId="14.[151,558,548,607]" box="[151,300,582,607]" pageId="14" pageNumber="583">
(
<figureCitation id="137E2A2AFFD2FFEFFF27671EFF28D31B" box="[159,251,582,607]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="14" pageNumber="583">Figs. 11</figureCitation>
,
<figureCitation id="137E2A2AFFD2FFEFFEBF671FFEF7D31B" box="[263,292,583,607]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="14" pageNumber="583">12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD2FFEFFF2F67D6FE90D273" pageId="14" pageNumber="583" type="materials_examined">
<paragraph id="8BFA36AFFFD2FFEFFF2F67D6FE90D273" blockId="14.[151,1436,654,823]" pageId="14" pageNumber="583">
<emphasis id="B931EABDFFD2FFEFFF2F67D6FE9BD3E3" bold="true" box="[151,328,654,679]" pageId="14" pageNumber="583">
<typeStatus id="54FE880DFFD2FFEFFF2F67D6FF01D3E3" box="[151,210,654,679]" pageId="14" pageNumber="583">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F352763FFFD2FFEFFEEA67D7FE75D3E3" box="[338,422,655,679]" name="India" pageId="14" pageNumber="583">INDIA</collectingCountry>
: adult male,
<typeStatus id="54FE880DFFD2FFEFFD8567D6FD4CD3E3" box="[573,671,654,679]" pageId="14" pageNumber="583" type="holotype">holotype</typeStatus>
(cw
<quantity id="4CBD9B4AFFD2FFEFFD6667D6FC87D3E2" box="[734,852,654,679]" metricMagnitude="-2" metricUnit="m" metricValue="1.664" pageId="14" pageNumber="583" unit="mm" value="16.64">16.64 mm</quantity>
, cl
<quantity id="4CBD9B4AFFD2FFEFFC3A67D7FC2AD3E2" box="[898,1017,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="14" pageNumber="583" unit="mm" value="12.9">12.90 mm</quantity>
, ch 8.00 mm, fw
<quantity id="4CBD9B4AFFD2FFEFFB7467D6FAE4D3E3" box="[1228,1335,654,679]" metricMagnitude="-3" metricUnit="m" metricValue="6.58" pageId="14" pageNumber="583" unit="mm" value="6.58">6.58 mm</quantity>
), On the trekking route for Harishchandragad from Paachnai, Akole taluk, Ahmednagar district,
<collectingRegion id="4981F84DFFD2FFEFFB3F67EAFAC0D38F" box="[1159,1299,690,715]" country="India" name="Maharashtra" pageId="14" pageNumber="583">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE715068FFD2FFEFFA9D67EBFA4FD38E" box="[1317,1436,691,715]" direction="north" orientation="latitude" pageId="14" pageNumber="583" precision="55" value="19.404">19.404° N</geoCoordinate>
and
<geoCoordinate id="EE715068FFD2FFEFFF7E678FFEE9D3AB" box="[198,314,727,751]" direction="east" orientation="longitude" pageId="14" pageNumber="583" precision="55" value="73.78">73.780° E</geoCoordinate>
), altitude
<quantity id="4CBD9B4AFFD2FFEFFE12678EFDD2D3AA" box="[426,513,726,751]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="14" pageNumber="583" unit="m" value="1065.0">1065 m</quantity>
,
<date id="FFFB106FFFD2FFEFFDB5678FFD6DD3AB" box="[525,702,726,751]" pageId="14" pageNumber="583" value="2014-10-01">1 October 2014</date>
, coll. P.L. Wagh (ZSI, WRC-C.1517);
<typeStatus id="54FE880DFFD2FFEFFBC9678FFB0CD3AB" box="[1137,1247,727,751]" pageId="14" pageNumber="583" type="paratype">paratypes</typeStatus>
, three males (cw
<quantity id="4CBD9B4AFFD2FFEFFF2F67A2FE8BD256" box="[151,344,762,787]" metricMagnitude="-2" metricUnit="m" metricValue="1.424" metricValueMax="1.536" metricValueMin="1.3119999999999998" pageId="14" pageNumber="583" unit="mm" value="14.239999999999998" valueMax="15.36" valueMin="13.12">13.1215.36 mm</quantity>
, cl
<quantity id="4CBD9B4AFFD2FFEFFEC767A3FDE2D256" box="[383,561,763,787]" metricMagnitude="-2" metricUnit="m" metricValue="1.078" metricValueMax="1.174" metricValueMin="0.982" pageId="14" pageNumber="583" unit="mm" value="10.78" valueMax="11.74" valueMin="9.82">9.8211.74 mm</quantity>
, ch
<quantity id="4CBD9B4AFFD2FFEFFDE767A2FCD7D256" box="[607,772,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="6.98" metricValueMax="7.88" metricValueMin="6.08" pageId="14" pageNumber="583" unit="mm" value="6.98" valueMax="7.88" valueMin="6.08">6.087.88 mm</quantity>
, fw
<quantity id="4CBD9B4AFFD2FFEFFC8C67A2FC08D257" box="[820,987,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="5.56" metricValueMax="5.96" metricValueMin="5.16" pageId="14" pageNumber="583" unit="mm" value="5.56" valueMax="5.96" valueMin="5.16">5.165.96 mm</quantity>
), same collection data as
<typeStatus id="54FE880DFFD2FFEFFB4167A2FA88D257" box="[1273,1371,762,787]" pageId="14" pageNumber="583" type="holotype">holotype</typeStatus>
(ZSI, WRC-C.1518).
</paragraph>
</subSubSection>
<caption id="DF3A6627FFD2FFEFFF2F62D3FCE4D69B" httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="14" pageNumber="583" targetBox="[340,1247,842,1907]" targetPageId="14">
<paragraph id="8BFA36AFFFD2FFEFFF2F62D3FCE4D69B" blockId="14.[151,1436,1931,2015]" pageId="14" pageNumber="583">
<emphasis id="B931EABDFFD2FFEFFF2F62D3FECCD6E4" bold="true" box="[151,287,1931,1953]" pageId="14" pageNumber="583">FIGURE 11.</emphasis>
<taxonomicName id="4C454D2CFFD2FFEFFE9E62D3FDC2D6E5" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[294,529,1931,1953]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="583" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFD2FFEFFE9E62D3FDC2D6E5" box="[294,529,1931,1953]" italics="true" pageId="14" pageNumber="583">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD2FFEFFDA062D4FD81D6E5" box="[536,594,1932,1953]" pageId="14" pageNumber="583" rank="species">
<emphasis id="B931EABDFFD2FFEFFDA062D4FD81D6E5" bold="true" box="[536,594,1932,1953]" pageId="14" pageNumber="583">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (ZSI, WRC—C.1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, DG), 1 mm (H).
</paragraph>
</caption>
<caption id="DF3A6627FFD3FFEEFF2F6658FBCAD270" httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="15" pageNumber="584" targetBox="[355,1227,268,683]" targetPageId="15">
<paragraph id="8BFA36AFFFD3FFEEFF2F6658FBCAD270" blockId="15.[151,1435,768,820]" pageId="15" pageNumber="584">
<emphasis id="B931EABDFFD3FFEEFF2F6658FEF3D251" bold="true" box="[151,288,768,790]" pageId="15" pageNumber="584">FIGURE 12.</emphasis>
<taxonomicName id="4C454D2CFFD3FFEEFE9F6658FDC3D252" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[295,528,768,790]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="584" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFD3FFEEFE9F6658FDC3D252" box="[295,528,768,790]" italics="true" pageId="15" pageNumber="584">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD3FFEEFDAF6658FD83D251" box="[535,592,768,789]" pageId="15" pageNumber="584" rank="species">
<emphasis id="B931EABDFFD3FFEEFDAF6658FD83D251" bold="true" box="[535,592,768,789]" pageId="15" pageNumber="584">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (ZSI, WRC—C.1517). A, dorsal view of right G1; B, ventral view of right G1; C, dorsal view of right G1 terminal article; D, right G2. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C35F6524FFD3FFEEFF7F6639FBFCD5A6" pageId="15" pageNumber="584" type="diagnosis">
<paragraph id="8BFA36AFFFD3FFEEFF7F6639FBFCD5A6" blockId="15.[151,1437,865,2006]" pageId="15" pageNumber="584">
<emphasis id="B931EABDFFD3FFEEFF7F6639FE92D23E" bold="true" box="[199,321,865,890]" pageId="15" pageNumber="584">Diagnosis.</emphasis>
Carapace quadrate (cw/cl = 1.3); dorsal surface smooth, burnt orange coloured; anterolateral margin short, smooth, cristiform; posterolateral margin slightly convergent, with fine, oblique striations; front strongly deflexed, narrow (fw/cw = 0.4); epigastric cristae faintly visible; postorbital cristae indistinct; exorbital angle triangular, distinct, with long outer margin, ca. 2 times length of inner margin; epibranchial tooth visible as weak notch; frontal median triangle incomplete; epistomal median lobe broadly triangular with acute apex, lacking median tooth (
<figureCitation id="137E2A2AFFD3FFEEFE81614DFE18D56A" box="[313,459,1045,1070]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Figs. 11A, B</figureCitation>
). Suture between male thoracic sternites s2/s3 indistinct, suture between sternites s3/s4 missing except for two short lateral grooves (
<figureCitation id="137E2A2AFFD3FFEEFD1C6161FCE4D516" box="[676,823,1081,1106]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Figs. 11C, F</figureCitation>
). Male abdomen narrowly T-shaped; sixth abdominal somite slightly broader than long, nearly equal to telson length; telson short; male sternoabdominal cavity deep, long, extending beyond level of bases of third maxillipeds (
<figureCitation id="137E2A2AFFD3FFEEFCF661D9FC34D5DE" box="[846,999,1153,1178]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Figs. 11C, G</figureCitation>
). G1 with slender, cone-shaped, short terminal article, ca. 0.3 times length of subterminal segment, with distal portion slightly curved inwards, tip straight (
<figureCitation id="137E2A2AFFD3FFEEFF426191FE5DD5A6" box="[250,398,1225,1250]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="15" pageNumber="584">Figs. 12AC</figureCitation>
). G2 with very short or vestigial terminal article (
<figureCitation id="137E2A2AFFD3FFEEFC016191FBF1D5A6" box="[953,1058,1225,1250]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="15" pageNumber="584">Fig. 12D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD3FFF1FF7F61B5FB25D0B1" lastPageId="16" lastPageNumber="585" pageId="15" pageNumber="584" type="description">
<paragraph id="8BFA36AFFFD3FFEEFF7F61B5FF25D762" blockId="15.[151,1437,865,2006]" pageId="15" pageNumber="584">
<emphasis id="B931EABDFFD3FFEEFF7F61B5FE23D442" bold="true" box="[199,496,1261,1286]" pageId="15" pageNumber="584">
Description of
<typeStatus id="54FE880DFFD3FFEEFE3A61B5FE38D442" box="[386,491,1261,1286]" pageId="15" pageNumber="584" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace quadrate (cw/cl = 1.3); dorsal surface smooth; anterolateral carapace gently inflated in frontal view; anterolateral margin short, smooth, cristiform; posterolateral margin slightly convergent, with fine, oblique striations; front strongly deflexed, squarecut, nearly flat, frontal margin straight, narrow (fw/cw = 0.4); epigastric cristae faintly visible, smooth; postorbital cristae indistinct; exorbital angle triangular, distinct, with long outer margin, ca. 2 times length of inner margin; epibranchial tooth visible as weak notch; postorbital region shallow; branchial regions gently inflated; subhepatic region slightly rugose; cervical grooves indistinct; mesogastric groove long, with slight bifurcation posteriorly; H-groove visible, shallow; frontal median triangle incomplete; epistomal median lobe broadly triangular with acute apex, lacking median tooth (
<figureCitation id="137E2A2AFFD3FFEEFADC60B1FF3AD762" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Figs. 11A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD3FFEEFF7F6369FD35D79E" blockId="15.[151,1437,865,2006]" pageId="15" pageNumber="584">
Chelipeds unequal; right chela larger, smooth, fingers with two or three larger teeth, distinct gape when tips in contact (
<figureCitation id="137E2A2AFFD3FFEEFF4F630DFE8DD72A" box="[247,350,1621,1646]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Fig. 11D</figureCitation>
); length of dactylus greater than length of upper margin of palm; carpal spine small, blunt; outer surface of merus rugose. Ambulatory legs (p2p5) long with fine, long tuft of brownish bristles with yellowish tips on anterior, posterior margins of propodus, dactylus; dactylus (p2p5) slightly longer than propodus; longest propodus (p3) two times as long as broad (
<figureCitation id="137E2A2AFFD3FFEEFDCD6399FD0BD79E" box="[629,728,1729,1754]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Fig. 11E</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD3FFEEFF7F63BEFDF5D666" blockId="15.[151,1437,865,2006]" pageId="15" pageNumber="584">
Suture between male thoracic sternites s2/s3 indistinct, suture between sternites s3/s4 missing except for two short lateral grooves (
<figureCitation id="137E2A2AFFD3FFEEFE336251FDCBD666" box="[395,536,1801,1826]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Figs. 11C, F</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD3FFEEFF7F6275FBC7D6F6" blockId="15.[151,1437,865,2006]" pageId="15" pageNumber="584">
Male abdomen narrowly T-shaped; fifth abdominal somite distinctly broader than long, with concave lateral margins; sixth abdominal somite slightly broader than long, nearly equal in length to telson, with slightly concave lateral margins; telson short, broadly triangular with almost straight outer margins; male sternoabdominal cavity deep, long, extending beyond level of bases of third maxillipeds (
<figureCitation id="137E2A2AFFD3FFEEFCCC62C1FBD4D6F6" box="[884,1031,1945,1970]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="15" pageNumber="584">Figs. 11C, G</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD3FFF1FF7F62E5FBB1D190" blockId="15.[151,1437,865,2006]" lastBlockId="16.[151,1437,151,1472]" lastPageId="16" lastPageNumber="585" pageId="15" pageNumber="584">
Mandibular palp with two joints, terminal joint bilobed; exopods of first, second maxillipeds with long flagellum; exopod of third maxilliped lacking flagellum, longer than ischium; ventral sulcus on ischium faint; merus with rounded anterior external angle, depression towards inner sides (
<figureCitation id="137E2A2AFFCCFFF1FC5665E4FB86D190" box="[1006,1109,188,213]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="16" pageNumber="585">Fig. 11H</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFCCFFF1FF7F65B8FC84D0CC" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
G1 with slender, cone-shaped, short terminal article, ca. 0.3 times length of subterminal segment, with distal portion slightly curved inwards, tip broad, straight, joint between terminal article, subterminal segment clear; subterminal segment distinctly broader than terminal article, basal two-thirds distinctly broader than distal onethird (
<figureCitation id="137E2A2AFFCCFFF1FF646414FEA2D020" box="[220,369,332,357]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="16" pageNumber="585">Figs. 12AC</figureCitation>
). G2 conspicuously short with very short or vestigial terminal article; proximal half portion of basal segment much stouter than distal half portion (
<figureCitation id="137E2A2AFFCCFFF1FD5A6437FC98D0CC" box="[738,843,367,392]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="16" pageNumber="585">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFCCFFF1FF7F64CCFA99D0E8" blockId="16.[151,1437,151,1472]" box="[199,1354,404,429]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F64CCFE80D0E9" bold="true" box="[199,339,404,429]" pageId="16" pageNumber="585">Live colour.</emphasis>
Carapace is burnt orange, and chelipeds and ambulatory legs are ivory coloured (
<figureCitation id="137E2A2AFFCCFFF1FB5E64CCFAEDD0E8" box="[1254,1342,404,429]" captionStart="FIGURE 2" captionStartId="2.[151,250,1654,1676]" captionTargetBox="[340,1247,758,1632]" captionTargetId="figure@2.[340,1247,758,1632]" captionTargetPageId="2" captionText="FIGURE 2. Live crabs of new species of Ghatiana and Gubernatoriana. A, Ghatiana aurantiaca n. sp. from Amboli; B, Ghatiana splendida n. sp. from Chaukul; C, Gubernatoriana alcocki n. sp. from Vankusawade; D, Gubernatoriana thackerayi n. sp. from Raghuvir Ghat; C, Gubernatoriana waghi n. sp. from Malshej Ghat. Photo credits: A, B, Arjun Kamdar; C, B. V. Jadhav; D, Shailesh Bhosale; E, Rachit Shah." httpUri="https://zenodo.org/record/1053927/files/figure.png" pageId="16" pageNumber="585">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFCCFFF1FF7F64EFFB25D0B1" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F64EFFE95D094" bold="true" box="[199,326,439,464]" pageId="16" pageNumber="585">
<typeStatus id="54FE880DFFCCFFF1FF7F64EFFE92D094" box="[199,321,439,464]" pageId="16" pageNumber="585" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<typeStatus id="54FE880DFFCCFFF1FEF464E0FE6AD094" box="[332,441,440,464]" pageId="16" pageNumber="585" type="paratype">Paratypes</typeStatus>
are consistent with the
<typeStatus id="54FE880DFFCCFFF1FD0664EFFCF3D094" box="[702,800,439,464]" pageId="16" pageNumber="585" type="holotype">holotype</typeStatus>
in carapace and gonopod features except for the smallest male (ZSI, WRC- C.1518), which has three or four larger teeth on the fingers of the major cheliped.
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFCCFFF1FF7F64A7FBE3D35C" box="[199,1072,511,536]" pageId="16" pageNumber="585" type="etymology">
<paragraph id="8BFA36AFFFCCFFF1FF7F64A7FBE3D35C" blockId="16.[151,1437,151,1472]" box="[199,1072,511,536]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F64A7FE9DD35C" bold="true" box="[199,334,511,536]" pageId="16" pageNumber="585">Etymology.</emphasis>
Species is named after its collector Dr. Prashant Wagh, zoologist.
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFCCFFF1FF7F677CFD47D324" pageId="16" pageNumber="585" type="materials_examined">
<paragraph id="8BFA36AFFFCCFFF1FF7F677CFD47D324" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F677CFEBFD379" bold="true" box="[199,364,548,573]" pageId="16" pageNumber="585">
<typeStatus id="54FE880DFFCCFFF1FF7F677CFED1D379" box="[199,258,548,573]" pageId="16" pageNumber="585">Type</typeStatus>
locality.
</emphasis>
Between Harishchandragad and Paachnai, Akole taluk, Ahmednagar district,
<collectingRegion id="4981F84DFFCCFFF1FAA8677CFA4FD379" box="[1296,1436,548,573]" country="India" name="Maharashtra" pageId="16" pageNumber="585">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE715068FFCCFFF1FF276710FEC0D31B" box="[159,275,584,608]" direction="north" orientation="latitude" pageId="16" pageNumber="585" precision="55" value="19.404">19.404° N</geoCoordinate>
and
<geoCoordinate id="EE715068FFCCFFF1FEF26710FE68D324" box="[330,443,584,608]" direction="east" orientation="longitude" pageId="16" pageNumber="585" precision="55" value="73.78">73.780° E</geoCoordinate>
) (altitude
<quantity id="4CBD9B4AFFCCFFF1FD97671FFD5BD324" box="[559,648,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="16" pageNumber="585" unit="m" value="1065.0">1065 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFCCFFF1FF7F6734FC72D250" pageId="16" pageNumber="585" type="discussion">
<paragraph id="8BFA36AFFFCCFFF1FF7F6734FC72D250" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F6734FEE8D3C1" bold="true" box="[199,315,620,645]" pageId="16" pageNumber="585">Remarks.</emphasis>
<taxonomicName id="4C454D2CFFCCFFF1FEFA6734FD90D3C0" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[322,579,620,644]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FEFA6734FD90D3C0" box="[322,579,620,644]" italics="true" pageId="16" pageNumber="585">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FDF36734FD59D3C1" box="[587,650,620,645]" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FDF36734FD59D3C1" bold="true" box="[587,650,620,645]" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
shares many morphological and gonopod features with its congeners. Nevertheless,
<taxonomicName id="4C454D2CFFCCFFF1FE8367C8FE71D3EC" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[315,418,656,680]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FE8367C8FE71D3EC" box="[315,418,656,680]" italics="true" pageId="16" pageNumber="585">G. waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FE1767D7FE20D3EC" box="[431,499,655,680]" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FE1767D7FE20D3EC" bold="true" box="[431,499,655,680]" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
can be distinguished from them by the following suite of characters: carapace quadrate-shaped; frontal margin narrow; tips of the cheliped fingers pointed; G1 terminal article short; G2 terminal article very short or vestigial (
<figureCitation id="137E2A2AFFCCFFF1FE5F678FFD48D3AB" box="[487,667,727,752]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="16" pageNumber="585">Figs. 11A, B, D</figureCitation>
,
<figureCitation id="137E2A2AFFCCFFF1FD1E6780FD2FD3B4" box="[678,764,728,752]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="16" pageNumber="585">12A, D</figureCitation>
). In live crabs, the colour pattern is diagnostic, burnt orange carapace and ivory coloured chelipeds and ambulatory legs (
<figureCitation id="137E2A2AFFCCFFF1FC8567A4FC46D250" box="[829,917,764,789]" captionStart="FIGURE 2" captionStartId="2.[151,250,1654,1676]" captionTargetBox="[340,1247,758,1632]" captionTargetId="figure@2.[340,1247,758,1632]" captionTargetPageId="2" captionText="FIGURE 2. Live crabs of new species of Ghatiana and Gubernatoriana. A, Ghatiana aurantiaca n. sp. from Amboli; B, Ghatiana splendida n. sp. from Chaukul; C, Gubernatoriana alcocki n. sp. from Vankusawade; D, Gubernatoriana thackerayi n. sp. from Raghuvir Ghat; C, Gubernatoriana waghi n. sp. from Malshej Ghat. Photo credits: A, B, Arjun Kamdar; C, B. V. Jadhav; D, Shailesh Bhosale; E, Rachit Shah." httpUri="https://zenodo.org/record/1053927/files/figure.png" pageId="16" pageNumber="585">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFCCFFF1FF7F6678FBCFD5AC" pageId="16" pageNumber="585" type="reference_group">
<paragraph id="8BFA36AFFFCCFFF1FF7F6678FEC9D554" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
<taxonomicName id="4C454D2CFFCCFFF1FF7F6678FE1AD27C" ID-CoL="84FVP" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[199,457,800,824]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FF7F6678FE1AD27C" box="[199,457,800,824]" italics="true" pageId="16" pageNumber="585">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FE696647FDC2D27C" box="[465,529,799,824]" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FE696647FDC2D27C" bold="true" box="[465,529,799,824]" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
is more similar to
<taxonomicName id="4C454D2CFFCCFFF1FD516678FCA8D27C" ID-CoL="7WDYV" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[745,891,800,824]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFCCFFF1FD516678FCA8D27C" box="[745,891,800,824]" italics="true" pageId="16" pageNumber="585">G. triangulus</emphasis>
</taxonomicName>
in the structure of its G1 due to proportionately stouter appearance of the subterminal segment as compared to the terminal article (
<figureCitation id="137E2A2AFFCCFFF1FBE8661CFB69D218" box="[1104,1210,836,861]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="16" pageNumber="585">Fig. 12A</figureCitation>
; see
<bibRefCitation id="EFD44B5EFFCCFFF1FB4C661CFF33D2C4" author="Pati, S. K. &amp; Sharma, R. M." journalOrPublisher="India. Journal of Natural History" pageId="16" pageNumber="585" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014" yearSuffix="a">Pati &amp; Sharma 2014a</bibRefCitation>
: fig. 7E for
<taxonomicName id="4C454D2CFFCCFFF1FEDB6630FE27D2C4" ID-CoL="7WDYV" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[355,500,872,896]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFCCFFF1FEDB6630FE27D2C4" box="[355,500,872,896]" italics="true" pageId="16" pageNumber="585">G. triangulus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C454D2CFFCCFFF1FDB36630FCDFD2C4" ID-CoL="84FVP" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[523,780,872,896]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FDB36630FCDFD2C4" box="[523,780,872,896]" italics="true" pageId="16" pageNumber="585">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FCAA663FFC82D2C4" box="[786,849,871,896]" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FCAA663FFC82D2C4" bold="true" box="[786,849,871,896]" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
can be easily differentiated from
<taxonomicName id="4C454D2CFFCCFFF1FB7F6630FA8BD2C4" ID-CoL="7WDYV" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[1223,1368,872,896]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFCCFFF1FB7F6630FA8BD2C4" box="[1223,1368,872,896]" italics="true" pageId="16" pageNumber="585">G. triangulus</emphasis>
</taxonomicName>
by its short G1 terminal article, ca. 0.3 times the length of the subterminal segment (vs. long G1 terminal article, ca. 0.5 times the length of the subterminal segment) and inwardly curved distal portion of the terminal article (vs. outwardly curved distal portion of the terminal article) (
<figureCitation id="137E2A2AFFCCFFF1FCB5668CFC71D2A8" box="[781,930,980,1005]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="16" pageNumber="585">Figs. 12A, C</figureCitation>
; see
<bibRefCitation id="EFD44B5EFFCCFFF1FC61668CFB03D2A8" author="Pati, S. K. &amp; Sharma, R. M." box="[985,1232,980,1005]" journalOrPublisher="India. Journal of Natural History" pageId="16" pageNumber="585" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014" yearSuffix="a">Pati &amp; Sharma 2014a</bibRefCitation>
: figs. 7E, G for
<taxonomicName id="4C454D2CFFCCFFF1FA3C668CFED9D554" ID-CoL="7WDYV" authorityName="Pati &amp; Sharma" authorityYear="2014" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFCCFFF1FA3C668CFED9D554" italics="true" pageId="16" pageNumber="585">G. triangulus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BFA36AFFFCCFFF1FF7F6144FBCFD5AC" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
The G1 structure of
<taxonomicName id="4C454D2CFFCCFFF1FE146144FDDCD570" ID-CoL="84FVP" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[428,527,1052,1076]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FE146144FDDCD570" box="[428,527,1052,1076]" italics="true" pageId="16" pageNumber="585">G. waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FDA06144FD84D571" box="[536,599,1052,1077]" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FDA06144FD84D571" bold="true" box="[536,599,1052,1077]" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
is almost identical to that of
<taxonomicName id="4C454D2CFFCCFFF1FC1A6144FBF6D570" ID-CoL="7WDYT" authorityName="Alcock" authorityYear="1909" box="[930,1061,1052,1076]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B931EABDFFCCFFF1FC1A6144FBF6D570" box="[930,1061,1052,1076]" italics="true" pageId="16" pageNumber="585">G. pilosipes</emphasis>
</taxonomicName>
as both have a short G1 terminal article (
<figureCitation id="137E2A2AFFCCFFF1FF546167FE86D51C" box="[236,341,1087,1112]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="16" pageNumber="585">Fig. 12A</figureCitation>
; see
<bibRefCitation id="EFD44B5EFFCCFFF1FE336167FD51D51C" author="Pati, S. K. &amp; Sharma, R. M." box="[395,642,1087,1112]" journalOrPublisher="Occasional Paper, No. 363" pageId="16" pageNumber="585" pagination="1 - 44" refString="Pati, S. K. &amp; Sharma, R. M. (2014 b) Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India, Occasional Paper, No. 363, 1 - 44. [published by the Director, Zoological Survey of India, Kolkata]" title="Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India" type="book chapter" year="2014" yearSuffix="b">Pati &amp; Sharma 2014b</bibRefCitation>
: pl. VIII, fig. 8D for
<taxonomicName id="4C454D2CFFCCFFF1FCD66118FC22D51C" ID-CoL="7WDYT" authorityName="Alcock" authorityYear="1909" box="[878,1009,1088,1112]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B931EABDFFCCFFF1FCD66118FC22D51C" box="[878,1009,1088,1112]" italics="true" pageId="16" pageNumber="585">G. pilosipes</emphasis>
</taxonomicName>
). However,
<taxonomicName id="4C454D2CFFCCFFF1FBC46118FAACD51C" ID-CoL="84FVP" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[1148,1407,1088,1112]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FBC46118FAACD51C" box="[1148,1407,1088,1112]" italics="true" pageId="16" pageNumber="585">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FA3D6167FF6BD539" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FA3D6167FF6BD539" bold="true" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
is immediately distinguished from
<taxonomicName id="4C454D2CFFCCFFF1FDE8613CFD06D538" ID-CoL="7WDYT" authorityName="Alcock" authorityYear="1909" box="[592,725,1124,1148]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B931EABDFFCCFFF1FDE8613CFD06D538" box="[592,725,1124,1148]" italics="true" pageId="16" pageNumber="585">G. pilosipes</emphasis>
</taxonomicName>
by its pointed cheliped fingers (vs. broadly rounded or spoon tipped cheliped fingers) and long male sternoabdominal cavity, extending beyond the level of third maxilliped bases (vs. short male sternoabdominal cavity, extending beyond the level of cheliped bases) (
<figureCitation id="137E2A2AFFCCFFF1FB1C61F4FAE5D580" box="[1188,1334,1196,1221]" captionStart="FIGURE 11" captionStartId="14.[151,250,1931,1953]" captionTargetBox="[340,1247,842,1907]" captionTargetId="figure@14.[340,1247,842,1907]" captionTargetPageId="14" captionText="FIGURE 11. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D G), 1 mm (H)." httpUri="https://zenodo.org/record/1053945/files/figure.png" pageId="16" pageNumber="585">Figs. 11C, D</figureCitation>
; see
<bibRefCitation id="EFD44B5EFFCCFFF1FAD461F5FF00D5AC" author="Bott, R." journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="16" pageNumber="585" pagination="1 - 338" part="526" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." title="Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)" type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 34, fig. 34 and
<bibRefCitation id="EFD44B5EFFCCFFF1FE136197FD72D5AC" author="Pati, S. K. &amp; Sharma, R. M." box="[427,673,1231,1256]" journalOrPublisher="Occasional Paper, No. 363" pageId="16" pageNumber="585" pagination="1 - 44" refString="Pati, S. K. &amp; Sharma, R. M. (2014 b) Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India, Occasional Paper, No. 363, 1 - 44. [published by the Director, Zoological Survey of India, Kolkata]" title="Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India" type="book chapter" year="2014" yearSuffix="b">Pati &amp; Sharma 2014b</bibRefCitation>
: pl. VIII, fig. 8C for
<taxonomicName id="4C454D2CFFCCFFF1FC326188FBDFD5AC" ID-CoL="7WDYT" authorityName="Alcock" authorityYear="1909" box="[906,1036,1232,1256]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B931EABDFFCCFFF1FC326188FBDFD5AC" box="[906,1036,1232,1256]" italics="true" pageId="16" pageNumber="585">G. pilosipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFCCFFF1FF7F61ACFBC4D484" pageId="16" pageNumber="585" type="description">
<paragraph id="8BFA36AFFFCCFFF1FF7F61ACFBD8D474" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F61ACFE5DD449" bold="true" box="[199,398,1268,1293]" pageId="16" pageNumber="585">Ecological notes.</emphasis>
Individuals were collected under stones and small boulders of a narrow passage on the edge of mountain cliffs. Crabs may also occur inside the crevices of mountain cliffs.
</paragraph>
<paragraph id="8BFA36AFFFCCFFF1FF7F6064FBC4D484" blockId="16.[151,1437,151,1472]" pageId="16" pageNumber="585">
<emphasis id="B931EABDFFCCFFF1FF7F6064FDC1D411" bold="true" box="[199,530,1340,1365]" pageId="16" pageNumber="585">Geographical distribution.</emphasis>
<taxonomicName id="4C454D2CFFCCFFF1FD9F6064FCE5D410" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[551,822,1340,1364]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="585" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFCCFFF1FD9F6064FCE5D410" box="[551,822,1340,1364]" italics="true" pageId="16" pageNumber="585">Gubernatoriana waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFCCFFF1FCF36064FC44D411" box="[843,919,1340,1365]" pageId="16" pageNumber="585" rank="species">
<emphasis id="B931EABDFFCCFFF1FCF36064FC44D411" bold="true" box="[843,919,1340,1365]" pageId="16" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
is known only from the
<typeStatus id="54FE880DFFCCFFF1FB466065FAFDD411" box="[1278,1326,1341,1365]" pageId="16" pageNumber="585">type</typeStatus>
locality, Harishchandragad in Ahmednagar district of
<collectingRegion id="4981F84DFFCCFFF1FD196007FCE2D43C" box="[673,817,1375,1400]" country="India" name="Maharashtra" pageId="16" pageNumber="585">Maharashtra</collectingRegion>
. This species was seen in Pune district from Malshej Ghat, which is about
<quantity id="4CBD9B4AFFCCFFF1FE3060DDFE11D4D9" box="[392,450,1412,1437]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="585" unit="km" value="7.0">7 km</quantity>
south of Harishchandragad (R. Shah, personal communication). Its occurrence in other parts of the
<collectingRegion id="4981F84DFFCCFFF1FEA460F1FEA4D484" box="[284,375,1449,1472]" country="Sri Lanka" name="Western" pageId="16" pageNumber="585">Western</collectingRegion>
Ghats with similar habitats, i.e. mountain cliffs, is possible.
</paragraph>
</subSubSection>
</treatment>
</document>