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<document id="2B4FA90FEEA5B0B09A9D1B535B1AB679" ID-CLB-Dataset="270518" ID-DOI="10.11646/phytotaxa.614.1.1" ID-GBIF-Dataset="143c8e81-3541-4298-b233-d3ce4e79d26c" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFBB1F2BFF50FF27FDF88DD3" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax orientalis" docType="treatment" docVersion="9" lastPageNumber="69" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="69" updateTime="1699690893055" updateUser="plazi">
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<mods:title id="2CE830881BB6F9E6BCC336004EDC4E78">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:namePart id="D5EDB28B8E1BD43533365CFC12F85B06">Henderson, Andrew</mods:namePart>
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<mods:date id="CA34B1BCBADDF94C46448D380833B7B7">2023</mods:date>
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<mods:classification id="2D3DE373C18B00DA59BEDF208B878B46">journal article</mods:classification>
<mods:identifier id="BC95B9D3EA5477212445F3A47CDA514B" type="CLB-Dataset">270518</mods:identifier>
<mods:identifier id="E1BDACDE46086BEC3FA92EDFF35FDB4F" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
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<treatment id="03B387DAFFBB1F2BFF50FF27FDF88DD3" ID-DOI="http://doi.org/10.5281/zenodo.8400399" ID-GBIF-Taxon="213635810" ID-Zenodo-Dep="8400399" LSID="urn:lsid:plazi:treatment:03B387DAFFBB1F2BFF50FF27FDF88DD3" httpUri="http://treatment.plazi.org/id/03B387DAFFBB1F2BFF50FF27FDF88DD3" lastPageNumber="69" pageId="68" pageNumber="69">
<subSubSection id="C3006547FFBB1F2BFF50FF27FBB588A3" box="[136,1100,157,185]" pageId="68" pageNumber="69" type="nomenclature">
<paragraph id="8BA536CCFFBB1F2BFF50FF27FBB588A3" blockId="68.[136,1452,157,293]" box="[136,1100,157,185]" pageId="68" pageNumber="69">
<emphasis id="B96EEADEFFBB1F2BFF50FF27FBB588A3" bold="true" box="[136,1100,157,185]" pageId="68" pageNumber="69">
1.27.
<taxonomicName id="4C1A4D4FFFBB1F2BFF14FF27FDC788A3" ID-CoL="WLZ7" authority="(Leon)" baseAuthorityName="Leon" box="[204,574,157,185]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B96EEADEFFBB1F2BFF14FF27FE1B88A2" bold="true" box="[204,482,158,185]" italics="true" pageId="68" pageNumber="69">Coccothrinax orientalis</emphasis>
(León)
</taxonomicName>
<bibRefCitation id="EF8B4B3DFFBB1F2BFD91FF27FC9688A3" author="Muniz, O. &amp; Borhidi, A." box="[585,879,158,184]" pageId="68" pageNumber="69" pagination="439 - 454" refId="ref63543" refString="Muniz, O. &amp; Borhidi, A. (1981 publ. 1982) Palmas nuevas del genero Coccothrinax Sarg. en Cuba. Acta Botanica Academiae Scientiarum Hungaricae 27: 439 - 454." type="journal article" year="1981">Muñiz &amp; Borhidi (1981</bibRefCitation>
publ. 1982: 451).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBB1F2BFB80FF27FED5891B" pageId="68" pageNumber="69" type="reference_group">
<paragraph id="8BA536CCFFBB1F2BFB80FF27FE3288C7" blockId="68.[136,1452,157,293]" pageId="68" pageNumber="69">
<taxonomicName id="4C1A4D4FFFBB1F2BFB80FF27FE3E88C7" ID-CoL="WLZ7" authority="Leon (1939: 121)" authorityName="Leon" authorityPageNumber="121" authorityYear="1939" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="variety" species="yuraguana" variety="orientalis">
<emphasis id="B96EEADEFFBB1F2BFB80FF27FA8F88A3" box="[1112,1398,158,184]" italics="true" pageId="68" pageNumber="69">Coccothrinax yuraguana</emphasis>
var.
<emphasis id="B96EEADEFFBB1F2BFF50FF7BFF0A88C7" box="[136,243,194,220]" italics="true" pageId="68" pageNumber="69">orientalis</emphasis>
<bibRefCitation id="EF8B4B3DFFBB1F2BFF24FF7BFE3E88C7" author="Leon" box="[252,455,194,220]" pageId="68" pageNumber="69" pagination="107 - 156" refId="ref62742" refString="Leon (1939) Contribucion al estudio de las palmas de Cuba. III. Memorias de la Sociedad Cubana de Historia Natural &quot; Felipe Poey &quot; 13: 107 - 156." type="journal article" year="1939">León (1939: 121)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA536CCFFBB1F2BFE0CFF7BFED5891B" blockId="68.[136,1452,157,293]" pageId="68" pageNumber="69">
<taxonomicName id="4C1A4D4FFFBB1F2BFE0CFF7BFBFE88C7" ID-CoL="WLZ7" authority="(Leon)" baseAuthorityName="Leon" box="[468,1031,194,220]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="subSpecies" species="yuraguana" subSpecies="orientalis">
<emphasis id="B96EEADEFFBB1F2BFE0CFF7BFD1788C7" box="[468,750,194,220]" italics="true" pageId="68" pageNumber="69">Coccothrinax yuraguana</emphasis>
subsp.
<emphasis id="B96EEADEFFBB1F2BFC9FFF7BFC4B88C7" box="[839,946,194,220]" italics="true" pageId="68" pageNumber="69">orientalis</emphasis>
(León)
</taxonomicName>
Borhidi in
<bibRefCitation id="EF8B4B3DFFBB1F2BFB55FF7BFA5588C7" author="Borhidi, A. &amp; Muniz, O." box="[1165,1452,194,220]" pageId="68" pageNumber="69" pagination="1 - 36" refId="ref61092" refString="Borhidi, A. &amp; Muniz, O. (1971 b publ. 1972) New plants in Cuba I. Acta Botanica Academiae Scientiarum Hungaricae 17: 1 - 36." type="journal article" year="1971">Borhidi &amp; Muñiz (1971b</bibRefCitation>
publ. 1972: 2).
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBB1F2BFEEBFF5EFC4D893F" pageId="68" pageNumber="69" type="materials_examined">
<paragraph id="8BA536CCFFBB1F2BFEEBFF5EFC4D893F" blockId="68.[136,1452,157,293]" pageId="68" pageNumber="69">
<typeStatus id="54A1886EFFBB1F2BFEEBFF5EFE5A891A" box="[307,419,231,257]" pageId="68" pageNumber="69" type="lectotype">Lectotype</typeStatus>
(designated by
<bibRefCitation id="EF8B4B3DFFBB1F2BFD8DFF5EFD21891B" author="Moya, C." box="[597,728,230,257]" pageId="68" pageNumber="69" pagination="1 - 63" refId="ref63249" refString="Moya, C. (2020) An annotated checklist of Cuban palms 2. Coccothrinax, Pt. 1: 1816 - 1939. Nomenclature, typification, and distribution. PalmArbor 4: 1 - 63." type="journal article" year="2020">Moya 2020</bibRefCitation>
):—
<collectingCountry id="F30D765CFFBB1F2BFCDAFF5FFCAD891B" box="[770,852,230,256]" name="Cuba" pageId="68" pageNumber="69">CUBA</collectingCountry>
. Oriente, Sierra de Nipe, cerca de Woodfred,
<quantity id="4CE29B29FFBB1F2BFA96FF5FFF59893E" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="68" pageNumber="69" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500-650 m</quantity>
,
<date id="FFA4100CFFBB1F2BFF74FEB3FE7D893F" box="[172,388,266,292]" pageId="68" pageNumber="69" value="1909-12-18">18 December 1909</date>
,
<emphasis id="B96EEADEFFBB1F2BFE57FEB2FDC8893F" box="[399,561,266,292]" italics="true" pageId="68" pageNumber="69">J. Shafer 3217</emphasis>
(
<typeStatus id="54A1886EFFBB1F2BFD98FEB3FD50893F" box="[576,681,266,292]" pageId="68" pageNumber="69" type="lectotype">lectotype</typeStatus>
NY!, isolectotype
<collectingCountry id="F30D765CFFBB1F2BFCA5FEB3FC5E893F" box="[893,935,266,292]" name="United States of America" pageId="68" pageNumber="69">US</collectingCountry>
!)
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBB1F2BFF50FEEFFBC889AF" pageId="68" pageNumber="69" type="reference_group">
<paragraph id="8BA536CCFFBB1F2BFF50FEEFFBC889AF" blockId="68.[136,1451,342,436]" pageId="68" pageNumber="69">
<taxonomicName id="4C1A4D4FFFBB1F2BFF50FEEFFD938977" ID-CoL="WLZ2" authority="(Borhidi &amp; Muniz) Muniz" authorityName="Muniz" baseAuthorityName="Borhidi &amp; Muniz" box="[136,618,342,364]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="moaensis">
<emphasis id="B96EEADEFFBB1F2BFF50FEEFFE908970" box="[136,361,342,363]" italics="true" pageId="68" pageNumber="69">Coccothrinax moaensis</emphasis>
(Borhidi &amp; Muñiz) Muñiz
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFBB1F2BFD5FFEEFFC938970" author="Muniz, O. &amp; Borhidi, A." box="[647,874,342,364]" pageId="68" pageNumber="69" pagination="439 - 454" refId="ref63543" refString="Muniz, O. &amp; Borhidi, A. (1981 publ. 1982) Palmas nuevas del genero Coccothrinax Sarg. en Cuba. Acta Botanica Academiae Scientiarum Hungaricae 27: 439 - 454." type="journal article" year="1981">Muñiz &amp; Borhidi (1981</bibRefCitation>
publ. 1982: 451).
<taxonomicName id="4C1A4D4FFFBB1F2BFBC2FEEFFE4D898B" ID-CoL="WLZ2" authority="Borhidi &amp; Muniz (1971 b" authorityName="Borhidi &amp; Muniz" authorityYear="1971" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="subSpecies" species="yuraguana" subSpecies="moaensis">
<emphasis id="B96EEADEFFBB1F2BFBC2FEEFFAF38977" box="[1050,1290,342,364]" italics="true" pageId="68" pageNumber="69">Coccothrinax yuraguana</emphasis>
subsp.
<emphasis id="B96EEADEFFBB1F2BFA8AFEEFFA528970" box="[1362,1451,342,363]" italics="true" pageId="68" pageNumber="69">moaensis</emphasis>
<bibRefCitation id="EF8B4B3DFFBB1F2BFF65FEC3FE4D898B" author="Borhidi, A. &amp; Muniz, O." box="[189,436,378,400]" pageId="68" pageNumber="69" pagination="1 - 36" refId="ref61092" refString="Borhidi, A. &amp; Muniz, O. (1971 b publ. 1972) New plants in Cuba I. Acta Botanica Academiae Scientiarum Hungaricae 17: 1 - 36." type="journal article" year="1971">Borhidi &amp; Muñiz (1971b</bibRefCitation>
</taxonomicName>
publ. 1972: 1). Type:—
<collectingCountry id="F30D765CFFBB1F2BFD7BFEC3FD118994" box="[675,744,378,399]" name="Cuba" pageId="68" pageNumber="69">CUBA</collectingCountry>
. Oriente,
<collectingRegion id="49DEF82EFFBB1F2BFC90FEC3FC3C8994" box="[840,965,378,399]" country="Cuba" name="Guantanamo" pageId="68" pageNumber="69">Guantánamo</collectingRegion>
, El Pino sur de la Reserva Cupeyal del Norte,
<date id="FFA4100CFFBB1F2BFA4CFEC3FEB589A8" pageId="68" pageNumber="69" value="1970-02-11">11 February 1970</date>
,
<emphasis id="B96EEADEFFBB1F2BFE8EFE27FD8D89A8" box="[342,628,414,435]" italics="true" pageId="68" pageNumber="69">A. Borhidi &amp; O. Muñiz 27934</emphasis>
(
<typeStatus id="54A1886EFFBB1F2BFD58FE27FD2F89AF" box="[640,726,414,436]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
HAC!, isotype BP
<emphasis id="B96EEADEFFBB1F2BFC48FE26FC4B89AF" box="[912,946,415,436]" italics="true" pageId="68" pageNumber="69">n.v.</emphasis>
, BP image!).
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBB1F2BFF50FE5BFDE58C27" pageId="68" pageNumber="69" type="description">
<paragraph id="8BA536CCFFBB1F2BFF50FE5BFDE58C27" blockId="68.[136,1452,482,1481]" pageId="68" pageNumber="69">
<emphasis id="B96EEADEFFBB1F2BFF50FE5BFF2989E7" bold="true" box="[136,208,482,508]" pageId="68" pageNumber="69">Stems</emphasis>
length not recorded, 9.0(2.9–15.0) cm diameter, solitary.
<emphasis id="B96EEADEFFBB1F2BFCB5FE5BFC4689E7" bold="true" box="[877,959,482,508]" pageId="68" pageNumber="69">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 1.2(0.6–1.8) mm diameter, stout, woody, loosely woven, ± joined or briefly free at the apices; petioles 7.9(5.5–11.8) mm diameter just below the apex; palmans 4.6(3.0–7.3) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 24(16–38) per leaf, the middle ones 33.8(24.4–42.0) cm long and 3.3(2.4–4.6) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively short and broad, abruptly narrowed (shoulder) toward the apex, otherwise parallel-sided, often strongly folded, stiff and leathery, the apices briefly splitting; middle leaf segment apices blunt and rounded; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially; leaf segments densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, light green to greenish-brown or reddish-brown center, sometimes without indumentum abaxially, without transverse veinlets.
<emphasis id="B96EEADEFFBB1F2BFCEBFCF3FC258B7F" bold="true" box="[819,988,842,868]" pageId="68" pageNumber="69">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few to numerous partial inflorescences; rachis bracts somewhat flattened, loosely sheathing, usually tomentose with a dense tuft of erect hairs at the apex; partial inflorescences number not recorded; proximalmost rachillae straight, 6.3(4.5–9.5) cm long and 1.2(0.8–1.6) mm diameter in fruit; rachillae glabrous at or near anthesis; stamens 8; fruit pedicels 1.9(1.1–2.6) mm long;
<emphasis id="B96EEADEFFBB1F2BFD68FC63FD088BEF" bold="true" box="[688,753,986,1012]" pageId="68" pageNumber="69">fruits</emphasis>
7.0(6.2–7.7) mm long and 7.3(6.6–8.3) mm diameter, purple or purplish; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBB1F2BFF65FBFFFEB78C9F" pageId="68" pageNumber="69" type="distribution">
<paragraph id="8BA536CCFFBB1F2BFF65FBFFFEB78C9F" blockId="68.[136,1452,482,1481]" pageId="68" pageNumber="69">
<emphasis id="B96EEADEFFBB1F2BFF65FBFFFDFA8C7B" bold="true" box="[189,515,1094,1120]" pageId="68" pageNumber="69">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFBB1F2BFDDBFBFFFDC78C7B" box="[515,574,1094,1120]" name="Cuba" pageId="68" pageNumber="69">Cuba</collectingCountry>
(
<collectingRegion id="49DEF82EFFBB1F2BFD94FBFFFD558C7B" box="[588,684,1094,1120]" country="Cuba" name="Holguin" pageId="68" pageNumber="69">Holguín</collectingRegion>
,
<collectingRegion id="49DEF82EFFBB1F2BFD6EFBFFFCBE8C7B" box="[694,839,1094,1120]" country="Cuba" name="Guantanamo" pageId="68" pageNumber="69">Guantánamo</collectingRegion>
) (
<figureCitation id="13212A49FFBB1F2BFC85FBFFFC548C7A" box="[861,941,1094,1121]" captionStart="FIGURE 15" captionStartId="67.[136,229,1957,1979]" captionTargetBox="[155,1429,649,1928]" captionTargetId="figure-194@67.[150,1434,644,1933]" captionTargetPageId="67" captionText="FIGURE 15. Distribution of Coccothrinax montgomeryana, C. munizii, C. orientalis, and C. pauciramosa." figureDoi="http://doi.org/10.5281/zenodo.8389401" httpUri="https://zenodo.org/record/8389401/files/figure.png" pageId="68" pageNumber="69">Fig. 15</figureCitation>
) in pine forest on serpentine soils at 701(500– 944) m elevation.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFBB1F2BFF65FB37FDF88DD3" pageId="68" pageNumber="69" type="discussion">
<paragraph id="8BA536CCFFBB1F2BFF65FB37FBE18D22" blockId="68.[136,1452,482,1481]" pageId="68" pageNumber="69">
<emphasis id="B96EEADEFFBB1F2BFF65FB37FE4A8CB3" bold="true" box="[189,435,1166,1192]" pageId="68" pageNumber="69">Taxonomic notes:—</emphasis>
Two preliminary species (
<emphasis id="B96EEADEFFBB1F2BFD35FB36FBDE8CB3" box="[749,1063,1166,1193]" italics="true" pageId="68" pageNumber="69">
<taxonomicName id="4C1A4D4FFFBB1F2BFD35FB36FC788CB3" box="[749,897,1167,1193]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="moaensis">C. moaensis</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFBB1F2BFC4CFB36FBDE8CB3" box="[916,1063,1166,1193]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="orientalis">C. orientalis</taxonomicName>
</emphasis>
) have a unique combination of qualitative character states except that
<taxonomicName id="4C1A4D4FFFBB1F2BFDE3FB0AFD3E8CD7" box="[571,711,1202,1229]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B96EEADEFFBB1F2BFDE3FB0AFD3E8CD7" box="[571,711,1202,1229]" italics="true" pageId="68" pageNumber="69">C. orientalis</emphasis>
</taxonomicName>
is polymorphic for abaxial segment indument. However, splitting the specimens into two species such that variables were consistent within each one led to improbable species that could not be justified morphologically or geographically. Therefore, specimens are recognized as a phylogenetic species, with one variable polymorphic. The earliest name for this species is
<taxonomicName id="4C1A4D4FFFBB1F2BFC5DFAA6FBE98D23" box="[901,1040,1310,1337]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B96EEADEFFBB1F2BFC5DFAA6FBE98D23" box="[901,1040,1310,1337]" italics="true" pageId="68" pageNumber="69">C. orientalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA536CCFFBB1F2BFF65FAFBFD3D8D9B" blockId="68.[136,1452,482,1481]" pageId="68" pageNumber="69">
<bibRefCitation id="EF8B4B3DFFBB1F2BFF65FAFBFEB08D47" author="Craft, P." box="[189,329,1346,1372]" pageId="68" pageNumber="69" refId="ref61560" refString="Craft, P. (2017) The Palms of Cuba. Palm Nut Pages, Florida, 232 pp." type="book" year="2017">Craft (2017)</bibRefCitation>
noted that
<taxonomicName id="4C1A4D4FFFBB1F2BFE1EFAFBFD298D47" box="[454,720,1346,1372]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B96EEADEFFBB1F2BFE1EFAFBFD298D47" box="[454,720,1346,1372]" italics="true" pageId="68" pageNumber="69">Coccothrinax orientalis</emphasis>
</taxonomicName>
could hybridize with
<taxonomicName id="4C1A4D4FFFBB1F2BFC1FFAFAFB808D47" authorityName="Burret" authorityYear="1929" box="[967,1145,1347,1373]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="pauciramosa">
<emphasis id="B96EEADEFFBB1F2BFC1FFAFAFB808D47" box="[967,1145,1347,1373]" italics="true" pageId="68" pageNumber="69">C. pauciramosa</emphasis>
</taxonomicName>
, and this may be the origin of the polymorphism in abaxial segment indument.
</paragraph>
<paragraph id="8BA536CCFFBB1F2BFF65FA33FDF88DD3" blockId="68.[136,1452,482,1481]" pageId="68" pageNumber="69">
There are two separate populations of
<taxonomicName id="4C1A4D4FFFBB1F2BFDB7FA33FC838DBF" box="[623,890,1418,1444]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B96EEADEFFBB1F2BFDB7FA33FC838DBF" box="[623,890,1418,1444]" italics="true" pageId="68" pageNumber="69">Coccothrinax orientalis</emphasis>
</taxonomicName>
. There are no differences between them, but only enough data to test five variables.
</paragraph>
</subSubSection>
</treatment>
</document>
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