273 lines
42 KiB
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273 lines
42 KiB
XML
<document id="EC0463116946F27B1782E08AC3F22041" ID-CLB-Dataset="4210" ID-DOI="10.5852/ejt.2021.767.1495" ID-GBIF-Dataset="67bc6a73-4137-4de3-9dc4-ce87972f742d" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514839" ID-ZooBank="9F019E02-95E7-4B6A-B5BC-01F8E26F3466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631927261920" checkinUser="felipe" docAuthor="Nahok, Benchawan, Tumpeesuwan, Sakboworn & Tumpeesuwan, Chanidaporn" docDate="2021" docId="03AC87E7C323FFC19F28FDB5972BBC2E" docLanguage="en" docName="ejt-767_142-166.pdf" docOrigin="European Journal of Taxonomy 767 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.767.1495" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria trochomorphoides Nahok & Tumpeesuwan & Tumpeesuwan 2021, sp. nov." docType="treatment" docUuid="B7DBFB2E-E7BF-46E5-AFB5-1B45CA4072DE" docUuidSource="ZooBank" docVersion="11" lastPageNumber="156" masterDocId="FF95FF9FC32EFFCF9D1FFF9B931BBE42" masterDocTitle="Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)" masterLastPageNumber="166" masterPageNumber="142" pageNumber="155" updateTime="1699449512379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8FCB2B8AC2DB505B6678DE27B3182728">Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)</mods:title>
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<mods:roleTerm id="EDB06380A41A53E21B1871C27ABBB62A">Author</mods:roleTerm>
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<mods:namePart id="0C7DFD4C46856353172FD3C7D5290E73">Nahok, Benchawan</mods:namePart>
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<mods:affiliation id="94A389455B3FAFEB9F69118D73F1A968">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Educational Research Development and Demonstration Institute, Srinakharinwirot University, Ongkharak District, Nakhon Nayok, 26120 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="507F509B1DFE280D9634EBF543AE54C1" type="email">nahok.b@gmail.com</mods:nameIdentifier>
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<mods:namePart id="159BF748F0B3DCA96AFE1C7944390995">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="3E783764D264837F9B8503A8CD42EA28">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Palaeontological Research and Education Centre, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="203749DAB108D8ACB36BB04CB64276AF" type="email">stumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="63CD67B6098B38BF58DC5C3663487C2E">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:affiliation id="87CBF60D0ACDBCF49A59B3D0557639E6">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="491D1A9DBF4A2E771E8E1C8471BB8BC0" type="email">ctumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:title id="F1366A8AC0E06414D53B574ABBEF86C3">European Journal of Taxonomy</mods:title>
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<mods:date id="BCD8A23B11DB9F2F997ED45A059B0213">2021</mods:date>
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<treatment id="03AC87E7C323FFC19F28FDB5972BBC2E" ID-DOI="http://doi.org/10.5281/zenodo.5514735" ID-GBIF-Taxon="188498239" ID-Zenodo-Dep="5514735" LSID="urn:lsid:zoobank.org:act:B7DBFB2E-E7BF-46E5-AFB5-1B45CA4072DE" httpUri="http://treatment.plazi.org/id/03AC87E7C323FFC19F28FDB5972BBC2E" lastPageId="14" lastPageNumber="156" pageId="13" pageNumber="155">
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<heading id="D0F2819DC323FFC29F28FDB590E6BC0B" box="[567,1021,558,585]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="155" reason="2">
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<taxonomicName id="4C054D72C323FFC29F28FDB59080BC0B" ID-CoL="8QH4H" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[567,923,558,585]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="155" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
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<emphasis id="B971EAE3C323FFC29F28FDB59080BC0B" bold="true" box="[567,923,558,585]" italics="true" pageId="13" pageNumber="155">Landouria trochomorphoides</emphasis>
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<taxonomicNameLabel id="A2425798C323FFC29EBDFDB490E6BC0B" box="[930,1021,559,585]" pageId="13" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<uuid id="FFA30C24C323FFC29F9EFDC997C9BC2F" box="[641,1234,594,621]" pageId="13" pageNumber="155">B7DBFB2E-E7BF-46E5-AFB5-1B45CA4072DE</uuid>
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<paragraph id="8BBA36F1C323FFC29F51FDEE90FEBCCD" blockId="13.[354,1234,558,656]" box="[590,997,629,656]" pageId="13" pageNumber="155">
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<figureCitation id="133E2A74C323FFC29F51FDEE91B7BCCD" box="[590,684,629,655]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="13" pageNumber="155">Figs 3C</figureCitation>
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,
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<figureCitation id="133E2A74C323FFC29FA6FDEE91C5BCCD" box="[697,734,629,655]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="13" pageNumber="155">4C</figureCitation>
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,
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<figureCitation id="133E2A74C323FFC29FF5FDEE9015BCD2" box="[746,782,629,656]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="13" pageNumber="155">5C</figureCitation>
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,
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<figureCitation id="133E2A74C323FFC29E04FDEE9041BCCD" box="[795,858,629,655]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="13" pageNumber="155">6G–I</figureCitation>
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;
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<tableCitation id="C687034AC323FFC29E79FDEE90FEBCCD" box="[870,997,629,655]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="2.[185,255,857,885]" captionStartId-1="4.[186,256,600,628]" captionStartId-2="6.[185,255,695,723]" captionStartId-3="10.[185,255,544,572]" captionTargetBox-0="[195,1921,987,1444]" captionTargetBox-1="[197,1907,827,1685]" captionTargetBox-2="[212,1909,868,1619]" captionTargetBox-3="[196,1908,758,1739]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="10" captionText-0="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." captionText-1="Table 2. Locality data and identification of specimens of Landouria Godwin-Austen, 1918 used for mtDNA sequencing, including collection catalogue numbers and GenBank accession numbers. Abbreviations: NHMSU = Natural History Museum, Mahasarakham University; HNHM = Hungarian Natural History Museum; AM = Australian Museum. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Hirano et al. (2014); 3 = Köhler et al. (2018)." captionText-2="Table 3. Pairwise p-distance values of Landouria Godwin-Austen, 1918 based on mitochondrial COI sequences." captionText-3="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri-0="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" httpUri-1="http://table.plazi.org/id/DF7A6679C32AFFCB9DA5FDC39249BC9F" httpUri-2="http://table.plazi.org/id/DF7A6679C328FFC99DA6FD2C96FFBC91" httpUri-3="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="13" pageNumber="155" tableUuid-0="DF7A6679C32CFFCD9DA6FCC295B0BD37" tableUuid-1="DF7A6679C32AFFCB9DA5FDC39249BC9F" tableUuid-2="DF7A6679C328FFC99DA6FD2C96FFBC91" tableUuid-3="DF7A6679C324FFC59DA6FDBB969CBCE7">Tables 1–4</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F657AC323FFC29DA2FD2D91E8BD04" pageId="13" pageNumber="155" type="diagnosis">
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<paragraph id="8BBA36F1C323FFC29DA2FD2D922DBC92" blockId="13.[189,310,694,720]" box="[189,310,694,720]" pageId="13" pageNumber="155">
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<heading id="D0F2819DC323FFC29DA2FD2D922DBC92" bold="true" box="[189,310,694,720]" fontSize="11" level="3" pageId="13" pageNumber="155" reason="3">
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<emphasis id="B971EAE3C323FFC29DA2FD2D922DBC92" bold="true" box="[189,310,694,720]" pageId="13" pageNumber="155">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA36F1C323FFC29DA2FD7E91E8BD04" blockId="13.[189,1399,741,838]" pageId="13" pageNumber="155">
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Shell with rather low spire, sharply keeled, with radial scaly processes (
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<figureCitation id="133E2A74C323FFC29929FD7E9783BCBD" box="[1078,1176,741,767]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="13" pageNumber="155">Fig. 4C</figureCitation>
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). Basal portion of flagellum thickened, tapering towards tip; penis narrow and cylindrical; base of gametolytic sac narrow. Radula, with lanceolate central and lateral teeth.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F657AC323FFC29DA2FCF69213BDBE" pageId="13" pageNumber="155" type="etymology">
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<paragraph id="8BBA36F1C323FFC29DA2FCF6925FBDC5" blockId="13.[189,324,877,903]" box="[189,324,877,903]" pageId="13" pageNumber="155">
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<heading id="D0F2819DC323FFC29DA2FCF6925FBDC5" bold="true" box="[189,324,877,903]" fontSize="11" level="3" pageId="13" pageNumber="155" reason="3">
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<emphasis id="B971EAE3C323FFC29DA2FCF6925FBDC5" bold="true" box="[189,324,877,903]" pageId="13" pageNumber="155">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA36F1C323FFC29DA2FC009213BDBE" blockId="13.[188,1399,923,1020]" pageId="13" pageNumber="155">
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The specific epithet ‘
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<taxonomicName id="4C054D72C323FFC29CA8FC009196BDF7" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[439,653,923,949]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="155" phylum="Mollusca" rank="species" species="trochomorphoides">
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<emphasis id="B971EAE3C323FFC29CA8FC009196BDF7" box="[439,653,923,949]" italics="true" pageId="13" pageNumber="155">trochomorphoides</emphasis>
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</taxonomicName>
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’ refers to the conchological similarity between the shell of the new species and that of the land snail genus
|
||
<taxonomicName id="4C054D72C323FFC29FA7FC249718BD98" authority="Albers, 1850" authorityName="Albers" authorityYear="1850" box="[696,1027,959,986]" class="Gastropoda" family="Trochomorphidae" genus="Trochomorpha" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="155" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C323FFC29FA7FC24907EBD9B" box="[696,869,959,985]" italics="true" pageId="13" pageNumber="155">Trochomorpha</emphasis>
|
||
Albers, 1850
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C054D72C323FFC2990FFC2493E0BDBE" authority="Mollendorf, 1890" authorityName="Mollendorf" authorityYear="1890" class="Gastropoda" family="Trochomorphidae" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="155" phylum="Mollusca" rank="family">Trochomorphidae Möllendorf, 1890</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC323FFC29DA2FBBF915CBB1C" pageId="13" pageNumber="155" type="materials_examined">
|
||
<paragraph id="8BBA36F1C323FFC29DA2FBBF9277BA7F" blockId="13.[189,364,1059,1086]" box="[189,364,1059,1086]" pageId="13" pageNumber="155">
|
||
<heading id="D0F2819DC323FFC29DA2FBBF9277BA7F" bold="true" box="[189,364,1059,1086]" fontSize="11" level="3" pageId="13" pageNumber="155" reason="3">
|
||
<emphasis id="B971EAE3C323FFC29DA2FBBF9277BA7F" bold="true" box="[189,364,1059,1086]" pageId="13" pageNumber="155">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C323FFC29DF5FBC99798BA94" blockId="13.[189,1399,1106,1238]" pageId="13" pageNumber="155">
|
||
<materialsCitation id="3B6D3CACC323FFC29DF5FBC99798BA94" ID-GBIF-Occurrence="3357769307" collectingDate="2017-10-06" collectionCode="UW" collectorName="B. Nahok & S. Tumpeesuwan & C. Tumpeesuwan" country="Thailand" elevation="359" latitude="17.119005" location="Pha Sawan Cave" longLatPrecision="1" longitude="101.93433" municipality="Pha Khao District" pageId="13" pageNumber="155" specimenCode="NHMSU-00027" specimenCount="1" stateProvince="Loei Province" typeStatus="holotype">
|
||
<emphasis id="B971EAE3C323FFC29DF5FBC99247BA2E" bold="true" box="[234,348,1106,1132]" pageId="13" pageNumber="155">
|
||
<typeStatus id="54BE8853C323FFC29DF5FBC99247BA2E" box="[234,348,1106,1132]" pageId="13" pageNumber="155" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3127661C323FFC29DA2FBEE924EBACD" box="[189,341,1141,1167]" name="Thailand" pageId="13" pageNumber="155">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C323FFC29C74FBEE92B9BACD" box="[363,418,1141,1167]" pageId="13" pageNumber="155">shell</specimenCount>
|
||
(SH = 10.0 mm, SW =
|
||
<quantity id="4CFD9B14C323FFC29FADFBEE9006BAD2" box="[690,797,1141,1168]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="13" pageNumber="155" unit="mm" value="21.3">21.3 mm</quantity>
|
||
, AH =
|
||
<quantity id="4CFD9B14C323FFC29E72FBEE90D2BAD2" box="[877,969,1141,1168]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="13" pageNumber="155" unit="mm" value="7.4">7.4 mm</quantity>
|
||
, AW =
|
||
<quantity id="4CFD9B14C323FFC29902FBEE9763BAD2" box="[1053,1144,1141,1168]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="13" pageNumber="155" unit="mm" value="9.1">9.1 mm</quantity>
|
||
, UW =
|
||
<quantity id="4CFD9B14C323FFC299CFFBEE9635BAD2" box="[1232,1326,1141,1168]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="13" pageNumber="155" unit="mm" value="6.2">6.2 mm</quantity>
|
||
);
|
||
<collectingRegion id="49C1F813C323FFC2985EFBEE9233BAF0" country="Thailand" name="Loei" pageId="13" pageNumber="155">Loei Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BDEAC8BC323FFC29C2BFB039110BAF0" box="[308,523,1176,1202]" pageId="13" pageNumber="155">Pha Khao District</collectingMunicipality>
|
||
,
|
||
<location id="8EDA602AC323FFC29F08FB0391FABAF0" LSID="urn:lsid:plazi:treatment:03AC87E7C323FFC19F28FDB5972BBC2E:8EDA602AC323FFC29F08FB0391FABAF0" box="[535,737,1176,1202]" country="Thailand" latitude="17.119005" longLatPrecision="1" longitude="101.93433" municipality="Pha Khao District" name="Pha Sawan Cave" pageId="13" pageNumber="155" stateProvince="Loei Province">Pha Sawan Cave</location>
|
||
;
|
||
<geoCoordinate id="EE315036C323FFC29FF2FB0C9090BAF1" box="[749,907,1175,1203]" degrees="17" direction="north" minutes="7" orientation="latitude" pageId="13" pageNumber="155" precision="1" seconds="8.42" value="17.119005">17°7′8.42″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE315036C323FFC29E88FB0C9754BAF1" box="[919,1103,1175,1203]" degrees="101" direction="east" minutes="56" orientation="longitude" pageId="13" pageNumber="155" precision="1" seconds="3.59" value="101.93433">101°56′3.59″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CFD9B14C323FFC29996FB0397CCBAF1" box="[1161,1239,1176,1203]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="13" pageNumber="155" unit="m" value="359.0">
|
||
<elevation id="0028D1C2C323FFC29996FB0397CCBAF1" box="[1161,1239,1176,1203]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="13" pageNumber="155" unit="m" value="359.0">359 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFBB1031C323FFC299FCFB039669BAF0" box="[1251,1394,1176,1202]" pageId="13" pageNumber="155" value="2017-10-06">
|
||
<collectingDate id="EFFFE9D9C323FFC299FCFB039669BAF0" box="[1251,1394,1176,1202]" pageId="13" pageNumber="155" value="2017-10-06">6 Oct. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F05327C323FFC29DA2FB27922BBA97" box="[189,304,1211,1238]" pageId="13" pageNumber="155">B. Nahok</collectorName>
|
||
,
|
||
<collectorName id="26F05327C323FFC29C23FB27911EBA94" box="[316,517,1212,1238]" pageId="13" pageNumber="155">S. Tumpeesuwan</collectorName>
|
||
and
|
||
<collectorName id="26F05327C323FFC29F21FB279012BA94" box="[574,777,1212,1238]" pageId="13" pageNumber="155">C. Tumpeesuwan</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBA39E8AC323FFC29E54FB279716BA94" box="[843,1037,1212,1238]" pageId="13" pageNumber="155">NHMSU-00027</specimenCode>
|
||
. (
|
||
<figureCitation id="133E2A74C323FFC2993FFB209762BA94" box="[1056,1145,1211,1238]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="13" pageNumber="155">Fig. 3C</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C323FFC29DF5FB66915CBB1C" blockId="13.[189,1399,1277,1374]" pageId="13" pageNumber="155">
|
||
<materialsCitation id="3B6D3CACC323FFC29DF5FB669158BB1C" ID-GBIF-Occurrence="3357769308" accessionNumber="MN449403 and MZ435748" collectingDate="2017-10-06" collectionCode="MN, MZ" collectorName="GenBank No." country="Thailand" elevation="359" latitude="17.119005" location="THAILAND" longLatPrecision="1" longitude="101.93433" municipality="Pha Khao District" pageId="13" pageNumber="155" specimenCode="NHMSU-00028" specimenCount="26" stateProvince="Loei" typeStatus="paratype">
|
||
<emphasis id="B971EAE3C323FFC29DF5FB66927CBB55" bold="true" box="[234,359,1277,1303]" pageId="13" pageNumber="155">
|
||
<typeStatus id="54BE8853C323FFC29DF5FB66927CBB55" box="[234,359,1277,1303]" pageId="13" pageNumber="155" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3127661C323FFC29DA2FABB924EBB78" box="[189,341,1312,1338]" name="Thailand" pageId="13" pageNumber="155">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C323FFC29C72FABB92CCBB78" box="[365,471,1312,1338]" pageId="13" pageNumber="155">17 shells</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD78C323FFC29CFDFABB919CBB79" box="[482,647,1312,1339]" pageId="13" pageNumber="155">9 living specs</specimenCount>
|
||
; same collection data as for holotype; GenBank No.
|
||
<accessionNumber id="9456AB12C323FFC299F0FABB966CBB78" box="[1263,1399,1312,1338]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN449403" pageId="13" pageNumber="155">MN449403</accessionNumber>
|
||
and
|
||
<accessionNumber id="9456AB12C323FFC29DF0FAD8926CBB1C" box="[239,375,1347,1374]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MZ435748" pageId="13" pageNumber="155" type="GenBank">MZ435748</accessionNumber>
|
||
;
|
||
<specimenCode id="DBA39E8AC323FFC29C9DFADF9158BB1C" box="[386,579,1348,1374]" pageId="13" pageNumber="155">NHMSU-00028</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC323FFC19DA2FA1E91C8BF05" lastPageId="14" lastPageNumber="156" pageId="13" pageNumber="155" type="description">
|
||
<paragraph id="8BBA36F1C323FFC29DA2FA1E9255BBDD" blockId="13.[189,334,1413,1439]" box="[189,334,1413,1439]" pageId="13" pageNumber="155">
|
||
<heading id="D0F2819DC323FFC29DA2FA1E9255BBDD" bold="true" box="[189,334,1413,1439]" fontSize="11" level="3" pageId="13" pageNumber="155" reason="3">
|
||
<emphasis id="B971EAE3C323FFC29DA2FA1E9255BBDD" bold="true" box="[189,334,1413,1439]" pageId="13" pageNumber="155">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C323FFC29DA2FA289712B875" blockId="13.[188,1399,1459,1592]" pageId="13" pageNumber="155">
|
||
SHELL (
|
||
<figureCitation id="133E2A74C323FFC29C0BFA28926BBB8F" box="[276,368,1459,1486]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="13" pageNumber="155">Fig. 3C</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC323FFC29C63FA2892C8BB8F" box="[380,467,1459,1485]" captionStart="Table 1" captionStartId="2.[185,255,857,885]" captionTargetBox="[195,1921,987,1444]" captionTargetPageId="2" captionText="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." httpUri="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" pageId="13" pageNumber="155" tableUuid="DF7A6679C32CFFCD9DA6FCC295B0BD37">Table 1</tableCitation>
|
||
). Dextral, relatively large-sized. Whorls six, suture shallow, spire only slightly elevated. Protoconch with radially elongated scars. Body whorl sharply keeled, brownish-corneous with a brown zone at periphery, scaly processes all over shell surface (
|
||
<figureCitation id="133E2A74C323FFC29EA1FA619707B856" box="[958,1052,1530,1556]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="13" pageNumber="155">Fig. 4C</figureCitation>
|
||
). Umbilicus moderately deep and very wide. Aperture lip rhombic, peristome reflexed, and expanded.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C323FFC29DA2F9C49178B90E" blockId="13.[187,1400,1630,1868]" pageId="13" pageNumber="155">
|
||
GENITAL SYSTEM (n = 3) (
|
||
<figureCitation id="133E2A74C323FFC29CCCF9C59137B83B" box="[467,556,1630,1657]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="13" pageNumber="155">Fig. 5C</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC323FFC29F2AF9C59196B83B" box="[565,653,1630,1657]" captionStart="Table 4" captionStartId="10.[185,255,544,572]" captionTargetBox="[196,1908,758,1739]" captionTargetPageId="10" captionText="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="13" pageNumber="155" tableUuid="DF7A6679C324FFC59DA6FDBB969CBCE7">Table 4</tableCitation>
|
||
). Penis slightly longer than vagina, slender cylindrical, internally with parallel, transverse robust folds; verge absent. Epiphallus slender cylindrical with thickened end. Flagellum as long as epiphallus, basal portion thick and progressively tapering towards tip, without nodes, inner surface smooth. Vas deferens is a thin cylindrical tube, entering epiphallus apically. Vagina thinner than penis, internally with parallel, transverse robust folds. Free oviduct very short. Gametolytic sac slightly narrower at base, with a long narrow cylindrical tube, and a small, oval sac at the distal end. Prostate gland and uterus very long.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C323FFC19DA2F8EF91C8BF05" blockId="13.[189,1399,1907,2004]" lastBlockId="14.[189,1398,265,327]" lastPageId="14" lastPageNumber="156" pageId="13" pageNumber="155">
|
||
RADULA (n = 3). Comprises 107–111 transverse rows with 65–73 teeth per row; radular formula: (24– 26)-(8–10)-1-(8–10)-(24–26). Central tooth unicuspid triangular (
|
||
<figureCitation id="133E2A74C323FFC29EC2F80D9759B9F3" box="[989,1090,1942,1969]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="13" pageNumber="155">Fig. 6G</figureCitation>
|
||
). Lateral teeth short and broad leaf-shaped, bicuspid, larger than central tooth, with small ectocone (
|
||
<figureCitation id="133E2A74C323FFC2992CF82297ACB996" box="[1075,1207,1977,2004]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="13" pageNumber="155">Fig. 6G–H</figureCitation>
|
||
). Marginal teeth asymmetric, tricuspid with small endocone; mesocone large and long, with curved margins; ectocone triangular with two to four cusps (
|
||
<figureCitation id="133E2A74C320FFC19F50FEB791DDBF05" box="[591,710,300,327]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="14" pageNumber="156">Fig. 6H–I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC320FFC19DA2FEF5914DBFBF" pageId="14" pageNumber="156" type="discussion">
|
||
<paragraph id="8BBA36F1C320FFC19DA2FEF59234BFCA" blockId="14.[189,303,366,392]" box="[189,303,366,392]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2FEF59234BFCA" bold="true" box="[189,303,366,392]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2FEF59234BFCA" bold="true" box="[189,303,366,392]" pageId="14" pageNumber="156">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2FE07914DBFBF" blockId="14.[189,1399,412,509]" pageId="14" pageNumber="156">
|
||
<taxonomicName id="4C054D72C320FFC19DA2FE07910EBFF4" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,533,412,438]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C320FFC19DA2FE07910EBFF4" box="[189,533,412,438]" italics="true" pageId="14" pageNumber="156">Landouria trochomorphoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C320FFC19F03FE06916EBFF5" box="[540,629,413,439]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from other taxa by its very large, rather flat, sharply keeled shell with scaly surface (
|
||
<figureCitation id="133E2A74C320FFC19CF5FE249157BF9B" box="[490,588,447,473]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="14" pageNumber="156">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A74C320FFC19F44FE249199BF9B" box="[603,642,447,473]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="14" pageNumber="156">4C</figureCitation>
|
||
). Lateral teeth of radula are stout bicuspid (
|
||
<figureCitation id="133E2A74C320FFC19989FE249614BF98" box="[1174,1295,447,474]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="14" pageNumber="156">Fig. 6G–I</figureCitation>
|
||
), genital system is small and slim (
|
||
<figureCitation id="133E2A74C320FFC19CF3FE799152BFBF" box="[492,585,482,509]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="14" pageNumber="156">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC320FFC19DA2FDBF972BBC2E" box="[189,1072,548,621]" pageId="14" pageNumber="156" type="distribution">
|
||
<paragraph id="8BBA36F1C320FFC19DA2FDBF924DBC7C" blockId="14.[189,342,548,574]" box="[189,342,548,574]" pageId="14" pageNumber="156">
|
||
<heading id="D0F2819DC320FFC19DA2FDBF924DBC7C" bold="true" box="[189,342,548,574]" fontSize="11" level="3" pageId="14" pageNumber="156" reason="3">
|
||
<emphasis id="B971EAE3C320FFC19DA2FDBF924DBC7C" bold="true" box="[189,342,548,574]" pageId="14" pageNumber="156">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C320FFC19DA2FDC9972BBC2E" blockId="14.[189,1072,594,621]" box="[189,1072,594,621]" pageId="14" pageNumber="156">
|
||
<taxonomicName id="4C054D72C320FFC19DA2FDC9910DBC2E" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[189,534,594,620]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="156" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C320FFC19DA2FDC9910DBC2E" box="[189,534,594,620]" italics="true" pageId="14" pageNumber="156">Landouria trochomorphoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C320FFC19F01FDC89163BC2F" box="[542,632,595,621]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus id="54BE8853C320FFC19E87FDC890D0BC2F" box="[920,971,595,621]" pageId="14" pageNumber="156">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |