treatments-xml/data/03/AC/87/03AC87D7FFBD33249E85FD06FD1BFD3F.xml

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<mods:title id="56CD31781F88E8039FDBA6D9E5ABADBD">Cambarus (Jugicambarus) pauleyi, a new species of crayfish (Decapoda: Cambaridae) endemic to southcentral West Virginia, USA, with a re-description of Cambarus (J.) dubius</mods:title>
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<mods:namePart id="2DCD78A9ECB073D9F9D0BEB4B1FD3D25">Loughman, Zachary J.</mods:namePart>
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<mods:namePart id="AD649D0319DA73F216BEFD4E05F86319">Thoma, Roger F.</mods:namePart>
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<mods:namePart id="88C1EC3B02B4408783416C72A922B441">Fetzner Jr, James W.</mods:namePart>
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<mods:namePart id="3DA3CBCA49B600A7416FB5A5CB26F698">Stocker, G. Whitney</mods:namePart>
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<treatment id="03AC87D7FFBD33249E85FD06FD1BFD3F" ID-DOI="http://doi.org/10.5281/zenodo.5681281" ID-GBIF-Taxon="119634476" ID-Zenodo-Dep="5681281" LSID="urn:lsid:zoobank.org:act:A004D13C-1588-4713-9BE6-BDFB5E402CA2" httpUri="http://treatment.plazi.org/id/03AC87D7FFBD33249E85FD06FD1BFD3F" lastPageId="16" lastPageNumber="542" pageId="8" pageNumber="534">
<subSubSection id="C31F654AFFBD333C9E85FD06FE13FD55" pageId="8" pageNumber="534" type="nomenclature">
<paragraph id="8BBA36C1FFBD333C9E85FD06FB5AFD74" blockId="8.[151,1249,692,751]" box="[151,1249,692,718]" pageId="8" pageNumber="534">
<heading id="D0F281ADFFBD333C9E85FD06FB5AFD74" bold="true" box="[151,1249,692,718]" fontSize="11" level="1" pageId="8" pageNumber="534" reason="1">
<emphasis id="B971EAD3FFBD333C9E85FD06FB5AFD74" bold="true" box="[151,1249,692,718]" pageId="8" pageNumber="534">
<taxonomicName id="4C054D42FFBD333C9E85FD06FD87FD74" ID-CoL="7RV3Y" box="[151,572,692,718]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="534" phylum="Arthropoda" rank="species" species="pauleyi" status="sp. nov." subGenus="Jugicambarus">
<emphasis id="B971EAD3FFBD333C9E85FD06FEAFFD74" bold="true" box="[151,276,692,718]" italics="true" pageId="8" pageNumber="534">Cambarus</emphasis>
(
<emphasis id="B971EAD3FFBD333C9F37FD06FE6AFD74" bold="true" box="[293,465,692,718]" italics="true" pageId="8" pageNumber="534">Jugicambarus</emphasis>
) 
<emphasis id="B971EAD3FFBD333C9FF1FD06FD87FD74" bold="true" box="[483,572,692,718]" italics="true" pageId="8" pageNumber="534">pauleyi</emphasis>
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, 
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Loughman, Thoma, Fetzner, and Stocker
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<figureCitation id="133E2A44FFBD333C9E85FD64FF59FD55" box="[151,226,726,751]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="8" pageNumber="534">Figs. 5</figureCitation>
A–I, 6, 7; 
<tableCitation id="C687037AFFBD333C9F47FD64FE13FD55" box="[341,424,726,751]" captionStart="TABLE 1" captionStartId="12.[151,242,152,175]" captionTargetBox="[156,1289,202,1367]" captionTargetPageId="12" captionText="TABLE 1. Measurements (mm) of Cambarus pauleyi, n. sp." httpUri="http://table.plazi.org/id/DF7A6649FFB933389E85FF2AFC9AFF15" pageId="8" pageNumber="534" tableUuid="DF7A6649FFB933389E85FF2AFC9AFF15">Table 1</tableCitation>
</paragraph>
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<subSubSection id="C31F654AFFBD333C9E85FCACFC9BFC15" pageId="8" pageNumber="534" type="reference_group">
<paragraph id="8BBA36C1FFBD333C9E85FCACFDB3FCE9" blockId="8.[151,1436,798,943]" pageId="8" pageNumber="534">
<treatmentCitationGroup id="AB1511EFFFBD333C9E85FCACFDB3FCE9" infered="true" pageId="8" pageNumber="534">
<taxonomicName id="4C054D42FFBD333C9E85FCACFE47FC8E" ID-CoL="7RTY5" authority="Faxon, 1884:114" authorityName="Faxon" authorityPageNumber="114" authorityYear="1884" box="[151,508,798,820]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="534" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBD333C9E85FCACFEF0FC8E" box="[151,331,798,820]" italics="true" pageId="8" pageNumber="534">Cambarus dubius</emphasis>
<treatmentCitation id="0AA410D0FFBD333C9F43FCADFE47FC8E" author="Faxon" box="[337,508,798,820]" page="114" pageId="8" pageNumber="534" year="1884">
<bibRefCitation id="EF944B30FFBD333C9F43FCADFE69FC8E" author="Faxon" box="[337,466,798,820]" pageId="8" pageNumber="534" refString="Faxon, W. (1884) Descriptions of new species of Cambarus, to which is added a synonymical list of the known species of Cambarus and Astacus. Proceedings of the American Academy of Arts and Sciences, 20, 107 - 158. http: // dx. doi. org / 10.2307 / 25138768" type="journal article" year="1884">Faxon, 1884</bibRefCitation>
:114
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</taxonomicName>
[in part]. 
<treatmentCitation id="0AA410D0FFBD333C9C70FCADFD46FC8E" author="Faxon" box="[610,765,798,820]" page="70" pageId="8" pageNumber="534" year="1885">
<bibRefCitation id="EF944B30FFBD333C9C70FCADFD65FC8E" author="Faxon" box="[610,734,798,820]" pageId="8" pageNumber="534" refString="Faxon, W. (1885) A revision of the Astacidae. Part I. The genera Cambarus and Astacus. Memoirs of the Museum of Comparative Zoology at Harvard College, 10, 1 - 186." type="journal article" year="1885">Faxon 1885</bibRefCitation>
:70
</treatmentCitation>
, pl. 4: fig. 3; pl. 8: figs. 7, 7' [in part]; 
<treatmentCitation id="0AA410D0FFBD333C9A97FCADFA91FC8E" author="Dewees" box="[1157,1322,798,820]" page="1" pageId="8" pageNumber="534" year="1972">
<bibRefCitation id="EF944B30FFBD333C9A97FCADFAA6FC8E" author="Dewees" box="[1157,1309,798,820]" pageId="8" pageNumber="534" refString="Dewees, J. P. (1972) Geographic variation in the primary burrowing crayfish, Cambarus dubius and Cambarus carolinus in Tennessee with notes on ecology and life history. M. S. Thesis University of Tennessee, Knoxville, Tennessee, 76 pp." type="book" year="1972">Dewees, 1972</bibRefCitation>
:1
</treatmentCitation>
, figs. 1b–l, 2c–h, 3, 4, 5b–h, 6–16 [in part].
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</paragraph>
<paragraph id="8BBA36C1FFBD333C9E85FCEEFD41FCC8" blockId="8.[151,1436,798,943]" box="[151,762,860,882]" pageId="8" pageNumber="534">
<treatmentCitationGroup id="AB1511EFFFBD333C9E85FCEEFD41FCC8" box="[151,762,860,882]" pageId="8" pageNumber="534">
<taxonomicName id="4C054D42FFBD333C9E85FCEEFDD1FCC8" ID-CoL="7RTY5" authority="Faxon, 1914:396" authorityName="Faxon" authorityPageNumber="396" authorityYear="1914" box="[151,618,860,882]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="534" phylum="Arthropoda" rank="subSpecies" species="carolinus" subSpecies="dubius">
<emphasis id="B971EAD3FFBD333C9E85FCEEFE14FCC8" box="[151,431,860,882]" italics="true" pageId="8" pageNumber="534">Cambarus carolinus dubius</emphasis>
: 
<treatmentCitation id="0AA410D0FFBD333C9FAEFCEFFDD1FCC8" author="Faxon" box="[444,618,860,882]" page="396" pageId="8" pageNumber="534" year="1914">
<bibRefCitation id="EF944B30FFBD333C9FAEFCEFFD85FCC8" author="Faxon" box="[444,574,860,882]" pageId="8" pageNumber="534" refString="Faxon, W. (1914) Notes on the crayfishes in the United States National Museum and the Museum of Comparative Zoology with descriptions of new species and subspecies to which is appended a catalogue of the known species and subspecies. Memoirs of the Museum of Comparative Zoology at Harvard College, 40, 351 - 427." type="journal article" year="1914">Faxon, 1914</bibRefCitation>
:396
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</taxonomicName>
, 425 [in part].
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</paragraph>
<paragraph id="8BBA36C1FFBD333C9E85FCC8FC9BFC15" blockId="8.[151,1436,798,943]" pageId="8" pageNumber="534">
<treatmentCitationGroup id="AB1511EFFFBD333C9E85FCC8FC9BFC15" pageId="8" pageNumber="534">
<taxonomicName id="4C054D42FFBD333C9E85FCC8FC8DFC2A" ID-CoL="7RTY5" authority="Hobbs and Bouchard, 1973:62" authorityName="Hobbs and Bouchard" authorityPageNumber="62" authorityYear="1973" box="[151,822,890,912]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="534" phylum="Arthropoda" rank="species" species="dubius" subGenus="Jugicambarus">
<emphasis id="B971EAD3FFBD333C9E85FCC8FE41FC2A" box="[151,506,890,912]" italics="true" pageId="8" pageNumber="534">Cambarus (Jugicambarus) dubius:</emphasis>
<treatmentCitation id="0AA410D0FFBD333C9C12FCC8FC8DFC2A" author="Hobbs" box="[512,822,890,912]" page="62" pageId="8" pageNumber="534" year="1973">
<bibRefCitation id="EF944B30FFBD333C9C12FCC8FCA3FC2A" author="Hobbs" box="[512,792,890,912]" pageId="8" pageNumber="534" refString="Hobbs, H. H. Jr. &amp; Bouchard, R. W. (1973) A new crayfish from the Cumberland Plateau System with notes on Cambarus carolinus. Proceedings of the Biological Society of Washington, 86, 41 - 68." type="journal article" year="1973">Hobbs and Bouchard, 1973</bibRefCitation>
:62
</treatmentCitation>
</taxonomicName>
[in part]; 
<treatmentCitation id="0AA410D0FFBD333C9D8EFCC8FBFBFC2A" author="Hobbs" box="[924,1088,890,912]" page="18" pageId="8" pageNumber="534" year="1974">
<bibRefCitation id="EF944B30FFBD333C9D8EFCC8FB98FC2A" author="Hobbs" box="[924,1059,890,912]" pageId="8" pageNumber="534" refString="Hobbs, H. H. Jr. (1974) A checklist of North and Middle America crayfishes (Decapoda: Astacidae and Cambaridae). Smithsonian Contribution to Zoology, 166, 1 - 161." type="journal article" year="1974">Hobbs, 1974</bibRefCitation>
:18
</treatmentCitation>
, fig. 60 [in part]; 
<treatmentCitation id="0AA410D0FFBD333C9AE6FCC8FA23FC2A" author="Hobbs" box="[1268,1432,890,912]" page="22" pageId="8" pageNumber="534" year="1989">
<bibRefCitation id="EF944B30FFBD333C9AE6FCC8FAC2FC2A" author="Hobbs" box="[1268,1401,890,912]" pageId="8" pageNumber="534" refString="Hobbs, H. H. Jr. (1989) An illustrated checklist of the American crayfishes (Decapoda: Astacidae, Cambaridae and Parastacidae). Smithsonian Contribution to Zoology, 480, 1 - 236." type="journal article" year="1989">Hobbs, 1989</bibRefCitation>
:22
</treatmentCitation>
, fig. 78 [in part]; Jezerinac 
<emphasis id="B971EAD3FFBD333C9FC1FC28FDBBFC15" box="[467,512,921,943]" italics="true" pageId="8" pageNumber="534">et al</emphasis>
., 1995:121, fig. 61 [in part].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31F654AFFBD333C9E85FC6EFC56F885" pageId="8" pageNumber="534" type="diagnosis">
<paragraph id="8BBA36C1FFBD333C9E85FC6EFC56F885" blockId="8.[151,1437,988,2035]" pageId="8" pageNumber="534">
<emphasis id="B971EAD3FFBD333C9E85FC6EFEAAFC49" bold="true" box="[151,273,988,1013]" pageId="8" pageNumber="534">Diagnosis.</emphasis>
Body and eyes pigmented. Posterior region of rostrum concave and deflected anteriorly. Rostral margins thickened, parallel to slightly converging at acumen. Rostrum excavated. Acumen distinctly triangular with prominent dorsally deflected spiniform tubercle at terminus. Areola obliterated. Cervical spines always absent. Mandibular, branchiostegal, and orbital regions of carapace with numerous well-developed tubercles. Postorbital ridges short; spiniform tubercle present in juveniles and subadults; adult postorbital ridge lacks tubercle. Suborbital angle acute to subacute. Carapace slightly vaulted in lateral profile. Antennal scale widest in middle, 1.3–3.6 (
<emphasis id="B971EAD3FFBD333C9F49FB08FED2FB6C" bold="true" box="[347,361,1210,1238]" pageId="8" pageNumber="534">x</emphasis>
<emphasis id="B971EAD3FFBD333C9F7FFB07FE3BFB76" box="[365,384,1205,1228]" italics="true" pageId="8" pageNumber="534">=</emphasis>
1.8, 
<emphasis id="B971EAD3FFBD333C9FAAFB07FE7DFB76" box="[440,454,1205,1228]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 35, 
<emphasis id="B971EAD3FFBD333C9C1FFB07FD94FB76" box="[525,559,1205,1228]" italics="true" pageId="8" pageNumber="534">SD</emphasis>
= 0.5) times as long as wide. Total carapace length (TCL) 2.0–2.2 (
<emphasis id="B971EAD3FFBD333C9B38FB08FA83FB6C" bold="true" box="[1322,1336,1210,1238]" pageId="8" pageNumber="534">x</emphasis>
<emphasis id="B971EAD3FFBD333C9B51FB07FAEDFB76" box="[1347,1366,1205,1228]" italics="true" pageId="8" pageNumber="534">=</emphasis>
2.1, 
<emphasis id="B971EAD3FFBD333C9B9CFB07FA27FB76" box="[1422,1436,1205,1228]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 35, 
<emphasis id="B971EAD3FFBD333C9EC5FB6EFEA8FB49" box="[215,275,1244,1267]" italics="true" pageId="8" pageNumber="534">SD =</emphasis>
0.1) times longer than width. Form I and II males possessing hook on ischium of third pereiopods only; hook gently curved at apex, overarching basioischial joint in form I males, not reaching basioischial joint in form II males; hooks not opposed by tubercle on basis. Mesial surface of chelae with two rows of tubercles; mesial most row with 5–9 (
<emphasis id="B971EAD3FFBD333C9F5EFAFFFEE1FAD3" bold="true" box="[332,346,1357,1385]" pageId="8" pageNumber="534">x</emphasis>
<emphasis id="B971EAD3FFBD333C9F4DFAFAFEC9FAE5" box="[351,370,1352,1375]" italics="true" pageId="8" pageNumber="534">=</emphasis>
6.7, 
<emphasis id="B971EAD3FFBD333C9FA3FAFAFE04FAE5" box="[433,447,1352,1375]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 35, 
<emphasis id="B971EAD3FFBD333C9C00FAF5FD8FFAE4" box="[530,564,1351,1374]" italics="true" pageId="8" pageNumber="534">SD</emphasis>
= 0.9) tubercles, second dorsal row with 2–7 (
<emphasis id="B971EAD3FFBD333C9A76FAFFFBC9FAD3" bold="true" box="[1124,1138,1357,1385]" pageId="8" pageNumber="534">x</emphasis>
= 5.1, 
<emphasis id="B971EAD3FFBD333C9AD5FAFAFB6EFAE5" box="[1223,1237,1352,1375]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 35, 
<emphasis id="B971EAD3FFBD333C9B3AFAF5FAF1FAE4" box="[1320,1354,1351,1374]" italics="true" pageId="8" pageNumber="534">SD</emphasis>
= 1.1) tubercles. Tubercles do not extend from mesial surface of chela to body of palm. One or two subpalmar tubercles present at junction of dactyl with propodus. Longitudinal ridge of dactyl consisting of several well developed scattered tubercles. Dorsomedian ridge of fixed finger of propodus pronounced. Ventral margin of propodus costate. Chelae lack defined lateral impression at the junction of the fixed finger with the propodus base. Dactyl and fixed finger with sharp corneous subterminal tip forming spine. Form I male palm length 73.9–77.5% (
<emphasis id="B971EAD3FFBD333C9B62F9B6FAC5F99A" bold="true" box="[1392,1406,1540,1568]" pageId="8" pageNumber="534">x</emphasis>
= 75.7%, 
<emphasis id="B971EAD3FFBD333C9EFFF994FF40F987" box="[237,251,1574,1597]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 3, 
<emphasis id="B971EAD3FFBD333C9F25F997FEE2F986" box="[311,345,1573,1596]" italics="true" pageId="8" pageNumber="534">SD</emphasis>
= 3.5%) of palm width, form I male palm length 35.5–37.7% (
<emphasis id="B971EAD3FFBD333C9A34F999FB8FF9FD" bold="true" box="[1062,1076,1579,1607]" pageId="8" pageNumber="534">x</emphasis>
= 36.6%, 
<emphasis id="B971EAD3FFBD333C9ABDF994FB06F987" box="[1199,1213,1574,1597]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 3, 
<emphasis id="B971EAD3FFBD333C9AEAF997FAA1F986" box="[1272,1306,1573,1596]" italics="true" pageId="8" pageNumber="534">SD</emphasis>
= 1.6%) of total propodus length; female dactyl length 56.0–62.0% (
<emphasis id="B971EAD3FFBD333C9D3AF9E0FC8DF9D4" bold="true" box="[808,822,1618,1646]" pageId="8" pageNumber="534">x</emphasis>
<emphasis id="B971EAD3FFBD333C9D29F9FFFCF5F9DE" box="[827,846,1613,1636]" italics="true" pageId="8" pageNumber="534">=</emphasis>
58.6, 
<emphasis id="B971EAD3FFBD333C9D85F9FFFC1EF9DE" box="[919,933,1613,1636]" italics="true" pageId="8" pageNumber="534">n</emphasis>
= 19, 
<emphasis id="B971EAD3FFBD333C9DE1F9FEFBAEF9D9" box="[1011,1045,1612,1635]" italics="true" pageId="8" pageNumber="534">SD</emphasis>
= 2.0%) of total propodus length. One or two large carpal spines present. First pleopod of form I male with short terminal elements. Central projection not tapering distally; recurved&gt; 90º to main shaft of gonopod, with distinct subapical notch. Mesial process directed 90° to shaft, bent cephalolaterally; inflated cephalically, tapering to distinct caudal point beyond terminance of central projection; mesial process terminating in a split tip. Annulus ventralis immovable; distinctly asymmetrical posteriorly; cephalic portion with median trough leading to strongly sculptured central fossa; exaggerated “S” bend in sinus terminating at caudal edge. Fossa sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C31F654AFFBD333F9ED5F8F8FD9AFA40" lastPageId="11" lastPageNumber="537" pageId="8" pageNumber="534" type="description">
<paragraph id="8BBA36C1FFBD333D9ED5F8F8FDC9FEFA" blockId="8.[151,1437,988,2035]" lastBlockId="9.[151,1437,151,1943]" lastPageId="9" lastPageNumber="535" pageId="8" pageNumber="534">
<emphasis id="B971EAD3FFBD333C9ED5F8F8FD1CF8D9" bold="true" box="[199,679,1866,1891]" pageId="8" pageNumber="534">Description of holotypic male, form I.</emphasis>
(
<figureCitation id="133E2A44FFBD333C9CA9F8F8FCB6F8D9" box="[699,781,1866,1891]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="8" pageNumber="534">Figs. 5</figureCitation>
A–G). Body somewhat compressed laterally (
<figureCitation id="133E2A44FFBD333C9B3CF8F8FACFF8D9" box="[1326,1396,1866,1891]" captionStart="FIGURE 1" captionStartId="3.[151,250,1879,1901]" captionTargetBox="[214,1369,215,1831]" captionTargetId="figure@3.[196,1389,195,1848]" captionTargetPageId="3" captionText="FIGURE 1. Illustration of various morphological features of Cambarus dubius sensu stricto, as herein defined. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from WLU 2050 b (female); I from WLU 2050 a (form II male)." httpUri="https://zenodo.org/record/239949/files/figure.png" pageId="8" pageNumber="534">Fig. 1</figureCitation>
A); carapace posterior to cervical groove slightly wider than abdomen. Carapace depth greater than carapace width at caudodorsal margin of cervical groove. Total carapace length 
<quantity id="4CFD9B24FFBD333C9D7BF820FC6FF810" box="[873,980,1938,1963]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="8" pageNumber="534" unit="mm" value="35.4">35.4 mm</quantity>
; post orbital carapace length 
<quantity id="4CFD9B24FFBD333C9B3CF821FA22F810" box="[1326,1433,1939,1963]" metricMagnitude="-2" metricUnit="m" metricValue="3.07" pageId="8" pageNumber="534" unit="mm" value="30.7">30.7 mm</quantity>
. Areola length 
<quantity id="4CFD9B24FFBD333C9F28F804FE1FF875" box="[314,420,1974,1999]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="8" pageNumber="534" unit="mm" value="14.6">14.6 mm</quantity>
: areola 41.2% of TCL (47.6% of postorbital carapace length (PCL). Rostrum moderately excavated, more so proximally than distally; margins thickened, parallel and continuous to base of acumen where margins becomes subparallel; floor of rostrum with numerous punctations. Rostrum 1.4 times longer than wide. Acumen broad and triangular, ending in dorsally upturned corneous tip (
<figureCitation id="133E2A44FFBC333D9DD1FF0EFBBDFF6F" box="[963,1030,188,213]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
A). Postorbital ridges short and well developed, lacking cephalic tubercles or spines. Suborbital angle subacute, lacking tubercle (
<figureCitation id="133E2A44FFBC333D9AD8FF6DFAAAFF42" box="[1226,1297,223,248]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
B). Cervical spine absent. Mandibular, branchiostegal, and orbital regions of carapace punctated with well-developed tubercles; greatest tubercle density in hepatic region.
</paragraph>
<paragraph id="8BBA36C1FFBC333D9ED5FEFEFA8CFD12" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="535">
Abdomen supraequal in length to carapace, pleura rounded cephaloventrally, angled distoventrally. Lateral margin of terga angulate; lateral margin of second pleuron deeply furrowed. Cephalic section of telson with 2 large spines in each caudolateral corner. Proximal podomere of uropod with distal spine on mesial lobe; mesial ramus of uropod with median ridge ending distally in distomedian spine not overreaching margin of ramus; laterodistal spine pronounced. Distal margin of proximal segment of lateral ramus of right uropod having 13 immovable, small spines and 1 lateral, large, movable spine. Cephalomedian lobe of epistome subtriangular, zygoma moderately arched (
<figureCitation id="133E2A44FFBC333D9EE2FD96FE88FD87" box="[240,307,548,573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
C); cephalolateral margins thickened, forming sharp angle at junction with endostyle (
<figureCitation id="133E2A44FFBC333D9AE6FD96FA8CFD87" box="[1268,1335,548,573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
C). Body of epistome possessing prominent cephalomedian fovea. Antennal scale broadest in middle; lateral margin thickened, terminating in large corneous spine; mesial margin setiferous (
<figureCitation id="133E2A44FFBC333D9DF3FDDEFB9DFD3F" box="[993,1062,620,645]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
D). Right antennal scale 
<quantity id="4CFD9B24FFBC333D9B56FDDEFA26FD3E" box="[1348,1437,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="535" unit="mm" value="4.9">4.9 mm</quantity>
long, 
<quantity id="4CFD9B24FFBC333D9EC5FD3DFE96FD1D" box="[215,301,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="535" unit="mm" value="5.5">5.5 mm</quantity>
wide (
<figureCitation id="133E2A44FFBC333D9F6BFD3DFE00FD12" box="[377,443,655,680]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
D). Tip of right antenna reaching posterior portion of carapace when adpressed.
</paragraph>
<paragraph id="8BBA36C1FFBC333D9ED5FD06FCD2FC3A" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="535">
Mesial surface of right chela with 2 rows of tubercles; mesial most row with 8 tubercles, second dorsal row with 5 tubercles (
<figureCitation id="133E2A44FFBC333D9F70FD65FE1CFD4A" box="[354,423,727,752]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
E). Palm length 39.0 % of palm width; depth of palm 
<quantity id="4CFD9B24FFBC333D9A31FD65FB3BFD55" box="[1059,1152,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="535" unit="mm" value="6.5">6.5 mm</quantity>
. Ventral surface of palm containing 2 subpalmar tubercles. Dorsal longitudinal ridge of dactyl developed and possessing moderate sized tubercles (
<figureCitation id="133E2A44FFBC333D9F1DFCADFEEFFC82" box="[271,340,799,824]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
E); dactyl terminating in large corneous spine. Dorsomedian ridge of fixed finger of propodus pronounced. Propodus terminating in sharp, corneous tip. Lateral impression at junction of fixed figure and propodus absent. All measurements and counts from right chela.
</paragraph>
<paragraph id="8BBA36C1FFBC333D9ED5FC3EFC80FBC7" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="535">
Carpus with prominent dorsal furrow and 3 weak dorsomesial tubercles; remainder of surface with some setiferous punctations; mesial margin with 2 large, procurved spines at about midlength. Distodorsal surface of merus with 9 spiniform tubercles; ventrolateral ridge with 5 small spines and large, corneous distal spine; ventromesial ridge with 3 well-developed spines; ventrolateral margin of ischium with 2 small, spiniform tubercles. Hook on ischium of third pereiopods only (
<figureCitation id="133E2A44FFBC333D9CFBFBAEFC90FB8F" box="[745,811,1052,1077]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[217,1374,227,1828]" captionTargetId="figure@10.[205,1388,193,1843]" captionTargetPageId="10" captionText="FIGURE 5. Illustration of various morphological features of Cambarus pauleyi, new species. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from allotype; I from morphotype." httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="9" pageNumber="535">Fig. 5</figureCitation>
F); hook gently curved at apex, overarching basioischial joint, not opposed by tubercle on basis. Form I gonopod as described in diagnosis (
<figureCitation id="133E2A44FFBC333D9A54FB8DFB30FBE2" box="[1094,1163,1087,1112]" captionStart="FIGURE 4" captionStartId="7.[151,250,1597,1619]" captionTargetBox="[240,1346,193,1575]" captionTargetId="figure@7.[240,1346,193,1575]" captionTargetPageId="7" captionText="FIGURE 4. Known range of Cambarus dubius sensu stricto in Maryland, Pennsylvania and West Virginia." httpUri="https://zenodo.org/record/239952/files/figure.png" pageId="9" pageNumber="535">Fig. 4</figureCitation>
G); tip reaching anterior margin of fourth caudomesial boss when abdomen is flexed.
</paragraph>
<paragraph id="8BBA36C1FFBC333D9ED5FB35FBBCFA27" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="535">
<emphasis id="B971EAD3FFBC333D9ED5FB35FDF6FB1A" bold="true" box="[199,589,1159,1184]" pageId="9" pageNumber="535">Description of allotypic female.</emphasis>
(
<figureCitation id="133E2A44FFBC333D9C73FB35FD10FB1A" box="[609,683,1159,1184]" captionStart="FIGURE 4" captionStartId="7.[151,250,1597,1619]" captionTargetBox="[240,1346,193,1575]" captionTargetId="figure@7.[240,1346,193,1575]" captionTargetPageId="7" captionText="FIGURE 4. Known range of Cambarus dubius sensu stricto in Maryland, Pennsylvania and West Virginia." httpUri="https://zenodo.org/record/239952/files/figure.png" pageId="9" pageNumber="535">Fig. 4</figureCitation>
H, 
<tableCitation id="C687037AFFBC333D9CC0FB35FC92FB1A" box="[722,809,1159,1184]" captionStart="TABLE 1" captionStartId="12.[151,242,152,175]" captionTargetBox="[156,1289,202,1367]" captionTargetPageId="12" captionText="TABLE 1. Measurements (mm) of Cambarus pauleyi, n. sp." httpUri="http://table.plazi.org/id/DF7A6649FFB933389E85FF2AFC9AFF15" pageId="9" pageNumber="535" tableUuid="DF7A6649FFB933389E85FF2AFC9AFF15">Table 1</tableCitation>
). Differing from 
<typeStatus id="54BE8863FFBC333D9DEDFB35FBDAFB1A" box="[1023,1121,1159,1184]" pageId="9" pageNumber="535" type="holotype">holotype</typeStatus>
in the following respects; carapace height less than carapace width (12.6 and 
<quantity id="4CFD9B24FFBC333D9CCEFB1EFCF8FB7E" box="[732,835,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="9" pageNumber="535" unit="mm" value="13.1">13.1 mm</quantity>
, respectively, 96.2%); TCL 
<quantity id="4CFD9B24FFBC333D9A90FB1EFB52FB7E" box="[1154,1257,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="9" pageNumber="535" unit="mm" value="28.2">28.2 mm</quantity>
, PCL 
<quantity id="4CFD9B24FFBC333D9B23FB1EFA22FB7E" box="[1329,1433,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="9" pageNumber="535" unit="mm" value="24.9">24.9 mm</quantity>
. Areola 41.1% of TCL (46.6% of PCL). Posterior portion of rostrum more excavated than anterior portion; rostrum as long as wide. Abdomen length 25.0 mm. Mesial surface of chelae with 2 rows of tubercles; mesial most row with 7, and second dorsal row with 4. Palm length (
<quantity id="4CFD9B24FFBC333D9CC0FAA5FC90FA8A" box="[722,811,1303,1328]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="9" pageNumber="535" unit="mm" value="6.7">6.7 mm</quantity>
) 75.2% of palm width (
<quantity id="4CFD9B24FFBC333D9A20FAAAFB37FA8A" box="[1074,1164,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="9" pageNumber="535" unit="mm" value="8.9">8.9 mm</quantity>
); depth of palm 
<quantity id="4CFD9B24FFBC333D9B52FAA5FA22FA8A" box="[1344,1433,1303,1328]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="535" unit="mm" value="5.6">5.6 mm</quantity>
. Antennal scale 4.0 mm long, 
<quantity id="4CFD9B24FFBC333D9FF8FA8EFDF9FAEE" box="[490,578,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="535" unit="mm" value="4.4">4.4 mm</quantity>
wide. All measurements and counts from right chela. Annulus ventralis as described in diagnosis (
<figureCitation id="133E2A44FFBC333D9FB0FAEDFE59FAC2" box="[418,482,1375,1400]" captionStart="FIGURE 1" captionStartId="3.[151,250,1879,1901]" captionTargetBox="[214,1369,215,1831]" captionTargetId="figure@3.[196,1389,195,1848]" captionTargetPageId="3" captionText="FIGURE 1. Illustration of various morphological features of Cambarus dubius sensu stricto, as herein defined. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from WLU 2050 b (female); I from WLU 2050 a (form II male)." httpUri="https://zenodo.org/record/239949/files/figure.png" pageId="9" pageNumber="535">Fig. 1</figureCitation>
H); width of postannular sclerite half total width of annulus ventralis; first pleopods uniramous, reaching central region of annulus ventralis when abdomen flexed.
</paragraph>
<paragraph id="8BBA36C1FFBC333D9ED5FA15FCCAF959" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="535">
<emphasis id="B971EAD3FFBC333D9ED5FA15FD6AFA7A" bold="true" box="[199,721,1447,1472]" pageId="9" pageNumber="535">Description of morphotypic male, form II.</emphasis>
(
<figureCitation id="133E2A44FFBC333D9CF7FA15FC90FA7A" box="[741,811,1447,1472]" captionStart="FIGURE 4" captionStartId="7.[151,250,1597,1619]" captionTargetBox="[240,1346,193,1575]" captionTargetId="figure@7.[240,1346,193,1575]" captionTargetPageId="7" captionText="FIGURE 4. Known range of Cambarus dubius sensu stricto in Maryland, Pennsylvania and West Virginia." httpUri="https://zenodo.org/record/239952/files/figure.png" pageId="9" pageNumber="535">Fig. 4</figureCitation>
G, 
<tableCitation id="C687037AFFBC333D9D5CFA15FC19FA7A" box="[846,930,1447,1472]" captionStart="TABLE 1" captionStartId="12.[151,242,152,175]" captionTargetBox="[156,1289,202,1367]" captionTargetPageId="12" captionText="TABLE 1. Measurements (mm) of Cambarus pauleyi, n. sp." httpUri="http://table.plazi.org/id/DF7A6649FFB933389E85FF2AFC9AFF15" pageId="9" pageNumber="535" tableUuid="DF7A6649FFB933389E85FF2AFC9AFF15">Table 1</tableCitation>
). Differing from 
<typeStatus id="54BE8863FFBC333D9A66FA15FB6DFA7A" box="[1140,1238,1447,1472]" pageId="9" pageNumber="535" type="holotype">holotype</typeStatus>
in the following respects: Carapace wider than abdomen (16.4 and 
<quantity id="4CFD9B24FFBC333D9CDAFA7EFC95FA5E" box="[712,814,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="9" pageNumber="535" unit="mm" value="13.3">13.3 mm</quantity>
, respectively); carapace height less than carapace width (15.9 and 
<quantity id="4CFD9B24FFBC333D9F1BFA5DFED5F9B2" box="[265,366,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="9" pageNumber="535" unit="mm" value="16.4">16.4 mm</quantity>
respectively, 97%); TCL 32.0 mm and PCL 
<quantity id="4CFD9B24FFBC333D9D78FA5DFC69F9B2" box="[874,978,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="9" pageNumber="535" unit="mm" value="28.6">28.6 mm</quantity>
. Areola length 40.0% of TCL (46.4% of PCL). Rostrum margins subparallel to base of acumen; rostrum 1.3 times as long as wide. Abdomen 
<quantity id="4CFD9B24FFBC333D9AE8F9A6FAE6F996" box="[1274,1373,1556,1581]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" pageId="9" pageNumber="535" unit="mm" value="30.5">30.5 mm</quantity>
long. Mesial row of tubercles on palm of chela with 7 tubercles; second dorsal row with 6. Palm length (
<quantity id="4CFD9B24FFBC333D9AFEF98AFAFDF9EA" box="[1260,1350,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="535" unit="mm" value="7.8">7.8 mm</quantity>
) 98.7% of palm width (
<quantity id="4CFD9B24FFBC333D9F59F9EDFE11F9CD" box="[331,426,1631,1655]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="9" pageNumber="535" unit="mm" value="7.9">7.9 mm</quantity>
). Antennal scale 
<quantity id="4CFD9B24FFBC333D9C60F9ECFD70F9CD" box="[626,715,1630,1655]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="535" unit="mm" value="5.4">5.4 mm</quantity>
long, 
<quantity id="4CFD9B24FFBC333D9D05F9EDFCD4F9CD" box="[791,879,1631,1655]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="535" unit="mm" value="4.9">4.9 mm</quantity>
wide. Gonopods reaching anterior margin of 4th pereiopod caudomesial boss. Central projection curved 90° to shaft, with complete apex; rounded (
<figureCitation id="133E2A44FFBC333D9AF7F930FA98F921" box="[1253,1315,1666,1691]" captionStart="FIGURE 1" captionStartId="3.[151,250,1879,1901]" captionTargetBox="[214,1369,215,1831]" captionTargetId="figure@3.[196,1389,195,1848]" captionTargetPageId="3" captionText="FIGURE 1. Illustration of various morphological features of Cambarus dubius sensu stricto, as herein defined. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A – G from holotype; H from WLU 2050 b (female); I from WLU 2050 a (form II male)." httpUri="https://zenodo.org/record/239949/files/figure.png" pageId="9" pageNumber="535">Fig. 1</figureCitation>
C). Mesial process tapered, bulbous, directed caudolaterally. Hook on ischium of third pereiopod small, not reaching basioischial joint. All measurements and counts from right chela.
</paragraph>
<paragraph id="8BBA36C1FFBC333D9ED5F95DFB55F82D" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="535">
<emphasis id="B971EAD3FFBC333D9ED5F95DFF46F8BD" bold="true" box="[199,253,1775,1799]" italics="true" pageId="9" pageNumber="535">Size.</emphasis>
Form I male (n = 3) TCL ranges in size from 
<quantity id="4CFD9B24FFBC333D9D1DF95CFC08F8BD" box="[783,947,1774,1799]" metricMagnitude="-2" metricUnit="m" metricValue="3.395" metricValueMax="3.54" metricValueMin="3.25" pageId="9" pageNumber="535" unit="mm" value="33.95" valueMax="35.4" valueMin="32.5">32.5–35.4 mm</quantity>
(PCL 29.0– 
<quantity id="4CFD9B24FFBC333D9A2DF95DFB1FF8BD" box="[1087,1188,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="3.07" pageId="9" pageNumber="535" unit="mm" value="30.7">30.7 mm</quantity>
) with a mean TCL of 
<quantity id="4CFD9B24FFBC333D9E85F8A0FEBBF890" box="[151,256,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.36" pageId="9" pageNumber="535" unit="mm" value="33.6">33.6 mm</quantity>
. Form II male (n = 17) mean TCL is 
<quantity id="4CFD9B24FFBC333D9CA6F8A1FCA1F890" box="[692,794,1811,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.19" pageId="9" pageNumber="535" unit="mm" value="31.9">31.9 mm</quantity>
and ranges in size from 28.0– 
<quantity id="4CFD9B24FFBC333D9A6BF8A0FB65F890" box="[1145,1246,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="9" pageNumber="535" unit="mm" value="35.5">35.5 mm</quantity>
(PCL 
<quantity id="4CFD9B24FFBC333D9B3EF8A1FF7CF8F5" metricMagnitude="-2" metricUnit="m" metricValue="2.785" metricValueMax="3.1399999999999997" metricValueMin="2.43" pageId="9" pageNumber="535" unit="mm" value="27.85" valueMax="31.4" valueMin="24.3">24.3–31.4 mm</quantity>
). Non-ovigerous female (n =10) TCL mean is 
<quantity id="4CFD9B24FFBC333D9CC0F885FC8DF8F5" box="[722,822,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.03" pageId="9" pageNumber="535" unit="mm" value="30.3">30.3 mm</quantity>
and ranges from 
<quantity id="4CFD9B24FFBC333D9DE9F885FB25F8F5" box="[1019,1182,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" metricValueMax="4.02" metricValueMin="2.82" pageId="9" pageNumber="535" unit="mm" value="34.2" valueMax="40.2" valueMin="28.2">28.2–40.2 mm</quantity>
(PCL 
<quantity id="4CFD9B24FFBC333D9AFBF884FA2BF8F5" box="[1257,1424,1846,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.0349999999999997" metricValueMax="3.5799999999999996" metricValueMin="2.4899999999999998" pageId="9" pageNumber="535" unit="mm" value="30.349999999999998" valueMax="35.8" valueMin="24.9">24.9–35.8 mm</quantity>
). Ovigerous female (n = 4) TCL ranges in size from 32.2–40.0 mm (PCL 
<quantity id="4CFD9B24FFBC333D9DD8F8E8FBCBF8C9" box="[970,1136,1882,1907]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.5799999999999996" metricValueMin="2.82" pageId="9" pageNumber="535" unit="mm" value="32.0" valueMax="35.8" valueMin="28.2">28.2–35.8 mm</quantity>
) with a mean of 
<quantity id="4CFD9B24FFBC333D9B23F8E8FA22F8C8" box="[1329,1433,1882,1907]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="9" pageNumber="535" unit="mm" value="35.6">35.6 mm</quantity>
. The largest specimen examined was a nonovigerous female with TCL of 
<quantity id="4CFD9B24FFBC333D9DD9F8CDFB94F82D" box="[971,1071,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="9" pageNumber="535" unit="mm" value="40.2">40.2 mm</quantity>
(PCL 
<quantity id="4CFD9B24FFBC333D9A6BF8CCFB59F82D" box="[1145,1250,1918,1943]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="9" pageNumber="535" unit="mm" value="35.8">35.8 mm</quantity>
).
</paragraph>
<caption id="DF7A6649FFBF333E9E85F8E7FEEBF85F" httpUri="https://zenodo.org/record/239953/files/figure.png" pageId="10" pageNumber="536" targetBox="[217,1374,227,1828]" targetPageId="10">
<paragraph id="8BBA36C1FFBF333E9E85F8E7FEEBF85F" blockId="10.[151,1436,1877,2021]" pageId="10" pageNumber="536">
<emphasis id="B971EAD3FFBF333E9E85F8E7FEAFF8D1" bold="true" box="[151,276,1877,1899]" pageId="10" pageNumber="536">FIGURE 5.</emphasis>
Illustration of various morphological features of 
<emphasis id="B971EAD3FFBF333E9D04F8E7FC62F8D1" box="[790,985,1877,1899]" italics="true" pageId="10" pageNumber="536">
<taxonomicName id="4C054D42FFBF333E9D04F8E7FC6EF8D1" box="[790,981,1877,1899]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="536" phylum="Arthropoda" rank="species" species="pauleyi" status="sp. nov.">Cambarus pauleyi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A24257A8FFBF333E9DF0F8E4FBDBF8D1" box="[994,1120,1877,1899]" pageId="10" pageNumber="536" rank="species">new species</taxonomicNameLabel>
. (A.) lateral view of carapace; (B.) dorsal view of carapace; (C.) epistome; (D.) antennal scale; (E.) dorsal view of distal podomere of right cheliped of form I male; (F.) ischial hook of form I male; (G.) lateral (left) and mesial (right) view of left first gonopod of form I male; (H.) annulus ventralis; (I.) lateral (left) and mesial (right) view of form II male gonopod. A–G from holotype; H from allotype; I from morphotype.
</paragraph>
</caption>
<caption id="DF7A6649FFBE333F9E85FCD1FBD4FCC3" box="[151,1135,867,889]" httpUri="https://zenodo.org/record/239954/files/figure.png" pageId="11" pageNumber="537" targetBox="[156,1436,196,848]" targetPageId="11">
<paragraph id="8BBA36C1FFBE333F9E85FCD1FBD4FCC3" blockId="11.[151,1135,867,889]" box="[151,1135,867,889]" pageId="11" pageNumber="537">
<emphasis id="B971EAD3FFBE333F9E85FCD1FEA9FCC3" bold="true" box="[151,274,867,889]" pageId="11" pageNumber="537">FIGURE 6.</emphasis>
A specimen of 
<taxonomicName id="4C054D42FFBE333F9FA1FCD1FDD5FCC3" box="[435,622,867,889]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="537" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBE333F9FA1FCD1FDD5FCC3" box="[435,622,867,889]" italics="true" pageId="11" pageNumber="537">Cambarus pauleyi</emphasis>
</taxonomicName>
from the type location showing typical life colors.
</paragraph>
</caption>
<paragraph id="8BBA36C1FFBE333F9ED5FC13FD9AFA40" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="537">
<emphasis id="B971EAD3FFBE333F9ED5FC13FEABFC00" bold="true" box="[199,272,929,954]" pageId="11" pageNumber="537">Color.</emphasis>
Carapace ground color deep blue to navy blue; posterior margin of carapace navy blue (
<figureCitation id="133E2A44FFBE333F9AFEFC13FA8BFC00" box="[1260,1328,929,954]" captionStart="FIGURE 6. A" captionStartId="11.[151,250,867,889]" captionTargetBox="[156,1436,196,848]" captionTargetId="figure@11.[156,1440,196,849]" captionTargetPageId="11" captionText="FIGURE 6. A specimen of Cambarus pauleyi from the type location showing typical life colors." httpUri="https://zenodo.org/record/239954/files/figure.png" pageId="11" pageNumber="537">Fig. 6</figureCitation>
). Hepatic and antennal region of carapace punctuated with cream, light blue, or white tubercles. Postorbital ridge color blue, to navy blue. Rostrum margins and acumen maroon or red, rarely blue. Cephalic section of carapace immediately anterior to and including cervical groove blue to indigo; mandibular abductor scars ranging from indigo to blue. Lateral margin of antennal scale light blue to blue; body of antennal scale light blue to blue. Antennal flagellum and antennules light blue; dorsal surface of lamellae blue to light blue; ventral surface cream to light blue. Dorsal surface of chelae blue to navy blue with white highlights; mesial, second dorsal row, and dorsal surface of dactyl tubercles blue or navy blue. Denticles on opposable surfaces of fingers cream, white, or tan. Ventral surface of chelae white or cream. Dorsal surface of carpus blue, navy blue or light blue; occasionally light purple; region adjacent to and including furrow navy blue to blue; carpus spine cream. Merus indigo to blue, or light blue. Podomeres of pereiopods light blue, blue-green, or blue; joints of pereiopodomeres cream or white. Dorsal and dorsolateral surface of abdomen same color as carapace; anterior region of abdomen with slight indigo tint; tergal margins red to reddish brown. Uropods navy blue, blue, light blue, with purple tint; margins white to cream. Ventral surface of abdomen and carapace cream or white. Dorsal ridge of form I gonopod central projection amber; body of central projection, gonopod, and mesial process tan. Form II gonopod and all associated processes cream. Cephalic portion of annulus ventralis pink to pink-cream; ridge of fossa pink; caudal region of annulus ventralis ranges from pink to cream colored.
</paragraph>
</subSubSection>
<subSubSection id="C31F654AFFBE333A9ED5F9B7FB26F9EA" lastPageId="14" lastPageNumber="540" pageId="11" pageNumber="537" type="materials_examined">
<paragraph id="8BBA36C1FFBE333F9ED5F9B7FAB2F8D8" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="537">
<emphasis id="B971EAD3FFBE333F9ED5F9B7FEDEF9A4" bold="true" box="[199,357,1541,1566]" pageId="11" pageNumber="537">
<typeStatus id="54BE8863FFBE333F9ED5F9B7FEB9F9A4" box="[199,258,1541,1566]" pageId="11" pageNumber="537">Type</typeStatus>
locality.
</emphasis>
Seepage stream adjacent to Moncove Lake State Park Playground, 2.0 km (
<quantity id="4CFD9B24FFBE333F9AA5F9B7FABAF9A4" box="[1207,1281,1541,1566]" metricMagnitude="3" metricUnit="m" metricValue="2.0921472" pageId="11" pageNumber="537" unit="mi" value="1.3">1.3 mi</quantity>
) northwest of Roxalia, Monroe County, West Virginia (
<geoCoordinate id="EE315006FFBE333F9C64F99BFCAAF9FB" box="[630,785,1577,1602]" direction="north" orientation="latitude" pageId="11" pageNumber="537" precision="1" value="37.620747">37.620745 N</geoCoordinate>
, - 
<geoCoordinate id="EE315006FFBE333F9D3AF99BFC71F9FB" box="[808,970,1577,1602]" direction="west" orientation="longitude" pageId="11" pageNumber="537" precision="1" value="-83.35299">83.352988 W</geoCoordinate>
). All animals were extracted from deep within their burrows, and did not rise to the surface during extraction efforts. Crayfish were collected from burrows in streambanks and seepage wetlands associated with a small headwater stream coursing through a mixed mesophyitc forest composed of American Beech (
<taxonomicName id="4C054D42FFBE333F9CC7F924FC57F914" authority="Ehrh." authorityName="Ehrh." box="[725,1004,1685,1710]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="537" phylum="Tracheophyta" rank="species" species="grandifolia">
<emphasis id="B971EAD3FFBE333F9CC7F924FC19F914" box="[725,930,1685,1710]" italics="true" pageId="11" pageNumber="537">Fagus grandifolia</emphasis>
Ehrh.
</taxonomicName>
), Red Oak (
<taxonomicName id="4C054D42FFBE333F9A6BF924FA9BF914" box="[1145,1312,1685,1710]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="537" phylum="Tracheophyta" rank="species" species="rubra">
<emphasis id="B971EAD3FFBE333F9A6BF924FA9BF914" box="[1145,1312,1685,1710]" italics="true" pageId="11" pageNumber="537">Quercus rubra</emphasis>
</taxonomicName>
L.), Black Oak (
<taxonomicName id="4C054D42FFBE333F9EC5F908FE66F96B" authority="Lam." authorityName="Lam." box="[215,477,1721,1746]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="537" phylum="Tracheophyta" rank="species" species="velutina">
<emphasis id="B971EAD3FFBE333F9EC5F908FE2CF968" box="[215,407,1721,1746]" italics="true" pageId="11" pageNumber="537">Quercus velutina</emphasis>
Lam.
</taxonomicName>
), and Tulip Poplar (
<taxonomicName id="4C054D42FFBE333F9CD0F90BFC7DF968" box="[706,966,1721,1746]" class="Magnoliopsida" family="Magnoliaceae" genus="Liriodendron" kingdom="Plantae" order="Magnoliales" pageId="11" pageNumber="537" phylum="Tracheophyta" rank="species" species="tulipifera">
<emphasis id="B971EAD3FFBE333F9CD0F90BFC7DF968" box="[706,966,1721,1746]" italics="true" pageId="11" pageNumber="537">Liriodendron tulipifera</emphasis>
</taxonomicName>
L.). The 
<typeStatus id="54BE8863FFBE333F9A21F90BFB23F968" box="[1075,1176,1721,1746]" pageId="11" pageNumber="537" type="holotype">holotype</typeStatus>
, allotype, morphotype, and 
<typeStatus id="54BE8863FFBE333F9EDBF96CFE8FF94C" box="[201,308,1758,1782]" pageId="11" pageNumber="537" type="paratype">paratypes</typeStatus>
were all excavated from burrows ranging from 0.6 to 1.3 meters deep in iron-rich clay-loam soil. Burrow architecture was simple, consisting of one or two surface tunnels unifying into a central tunnel, terminating in an enlarged resting chamber. The 
<typeStatus id="54BE8863FFBE333F9C23F894FDDAF884" box="[561,609,1830,1854]" pageId="11" pageNumber="537">type</typeStatus>
series and 
<typeStatus id="54BE8863FFBE333F9CF2F894FCF0F884" box="[736,843,1830,1854]" pageId="11" pageNumber="537" type="paratype">paratypes</typeStatus>
were all collected on 
<date id="FFBB1001FFBE333F9A56F897FB61F887" box="[1092,1242,1829,1853]" pageId="11" pageNumber="537" value="2014-06-28">28 June 2014</date>
by Spencer Bell, Zachary Dillard, ZJL, Luke Sadecky, Nicole Sadecky, R. Katie Scott, Eric Tennant and Eric Tidmore.
</paragraph>
<paragraph id="8BBA36C1FFBE333F9ED5F8DFFDB9F84B" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="537">
<emphasis id="B971EAD3FFBE333F9ED5F8DFFE79F83C" bold="true" box="[199,450,1901,1926]" pageId="11" pageNumber="537">
Disposition of 
<typeStatus id="54BE8863FFBE333F9F6FF8DFFE06F83C" box="[381,445,1901,1926]" pageId="11" pageNumber="537">types</typeStatus>
.
</emphasis>
The 
<typeStatus id="54BE8863FFBE333F9C14F8DFFDD0F83C" box="[518,619,1901,1926]" pageId="11" pageNumber="537" type="holotype">holotype</typeStatus>
, allotype, and morphotype are deposited in the National Museum of Natural History (
<collectionCode id="ED14AE04FFBE333F9F71F823FE01F813" LSID="urn:lsid:biocol.org:col:34871" box="[355,442,1937,1961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="537">USNM</collectionCode>
), Smithsonian Institution, Washington, D. C. (catalogue numbers 
<collectionCode id="ED14AE04FFBE333F9AC6F823FA9EF813" LSID="urn:lsid:biocol.org:col:34871" box="[1236,1317,1937,1961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="537">USNM</collectionCode>
1283506, 1283507, 1283508, respectively). 
<typeStatus id="54BE8863FFBE333F9C0EF804FD32F874" box="[540,649,1974,1998]" pageId="11" pageNumber="537" type="paratype">Paratypes</typeStatus>
are deposited in the West Liberty University Astacology Collection, West Liberty, WV (WLU 2001).
</paragraph>
<caption id="DF7A6649FFB933389E85FF2AFC9AFF15" ID-Table-UUID="DF7A6649FFB933389E85FF2AFC9AFF15" box="[151,801,151,175]" httpUri="http://table.plazi.org/id/DF7A6649FFB933389E85FF2AFC9AFF15" pageId="12" pageNumber="538" targetBox="[156,1289,202,1367]" targetIsTable="true" targetPageId="12">
<paragraph id="8BBA36C1FFB933389E85FF2AFC9AFF15" blockId="12.[151,801,151,175]" box="[151,801,151,175]" pageId="12" pageNumber="538">
<emphasis id="B971EAD3FFB933389E85FF2AFEB7FF15" bold="true" box="[151,268,152,175]" pageId="12" pageNumber="538">TABLE 1.</emphasis>
Measurements (mm) of 
<taxonomicName id="4C054D42FFB933389C01FF25FD6CFF15" box="[531,727,151,175]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="538" phylum="Arthropoda" rank="species" species="pauleyi" status="sp. nov.">
<emphasis id="B971EAD3FFB933389C01FF25FD6CFF15" box="[531,727,151,175]" italics="true" pageId="12" pageNumber="538">Cambarus pauleyi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24257A8FFB933389CF7FF2AFC9AFF15" box="[741,801,152,175]" pageId="12" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBA36C1FFB933389E8EFF43FB17FAED" pageId="12" pageNumber="538">
<table id="F905C461FFB9CCCB9E8EFF78FAB2FAED" box="[156,1289,202,1367]" gridcols="4" gridrows="19" pageId="12" pageNumber="538">
<tr id="35353483FFB9CCCB9E8EFF78FAB2FEBD" box="[156,1289,202,263]" gridrow="0" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFF78FE7BFEBD" box="[156,448,202,263]" gridcol="0" gridrow="0" pageId="12" pageNumber="538">Carapace</th>
<th id="76E45DFFFFB9CCCB9CC7FF78FC88FEBD" box="[725,819,202,263]" gridcol="1" gridrow="0" pageId="12" pageNumber="538">Holotype</th>
<th id="76E45DFFFFB9CCCB9DA7FF78FBB5FEBD" box="[949,1038,202,263]" gridcol="2" gridrow="0" pageId="12" pageNumber="538">Allotype</th>
<th id="76E45DFFFFB9CCCB9A9EFF78FAB2FEBD" box="[1164,1289,202,263]" gridcol="3" gridrow="0" pageId="12" pageNumber="538">Morphotype</th>
</tr>
<tr id="35353483FFB9CCCB9E8EFEABFAB2FE95" box="[156,1289,281,303]" gridrow="1" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFEABFE7BFE95" box="[156,448,281,303]" gridcol="0" gridrow="1" pageId="12" pageNumber="538">Total carapace length</th>
<td id="76E45DFFFFB9CCCB9CC7FEABFC88FE95" box="[725,819,281,303]" gridcol="1" gridrow="1" pageId="12" pageNumber="538">35.4</td>
<td id="76E45DFFFFB9CCCB9DA7FEABFBB5FE95" box="[949,1038,281,303]" gridcol="2" gridrow="1" pageId="12" pageNumber="538">28.2</td>
<td id="76E45DFFFFB9CCCB9A9EFEABFAB2FE95" box="[1164,1289,281,303]" gridcol="3" gridrow="1" pageId="12" pageNumber="538">32.0</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFEF3FAB2FEC4" box="[156,1289,321,382]" gridrow="2" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFEF3FE7BFEC4" box="[156,448,321,382]" gridcol="0" gridrow="2" pageId="12" pageNumber="538">Postorbital length Length cephalic section</th>
<td id="76E45DFFFFB9CCCB9CC7FEF3FC88FEC4" box="[725,819,321,382]" gridcol="1" gridrow="2" pageId="12" pageNumber="538">30.7 20.7</td>
<td id="76E45DFFFFB9CCCB9DA7FEF3FBB5FEC4" box="[949,1038,321,382]" gridcol="2" gridrow="2" pageId="12" pageNumber="538">24.9 16.4</td>
<td id="76E45DFFFFB9CCCB9A9EFEF3FAB2FEC4" box="[1164,1289,321,382]" gridcol="3" gridrow="2" pageId="12" pageNumber="538">28.6 18.7</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFE22FAB2FE1C" box="[156,1289,400,422]" gridrow="3" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFE22FE7BFE1C" box="[156,448,400,422]" gridcol="0" gridrow="3" pageId="12" pageNumber="538">Width</th>
<td id="76E45DFFFFB9CCCB9CC7FE22FC88FE1C" box="[725,819,400,422]" gridcol="1" gridrow="3" pageId="12" pageNumber="538">14.5</td>
<td id="76E45DFFFFB9CCCB9DA7FE22FBB5FE1C" box="[949,1038,400,422]" gridcol="2" gridrow="3" pageId="12" pageNumber="538">13.1</td>
<td id="76E45DFFFFB9CCCB9A9EFE22FAB2FE1C" box="[1164,1289,400,422]" gridcol="3" gridrow="3" pageId="12" pageNumber="538">16.4</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFE0AFAB2FE74" box="[156,1289,440,462]" gridrow="4" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFE0AFE7BFE74" box="[156,448,440,462]" gridcol="0" gridrow="4" pageId="12" pageNumber="538">Depth</th>
<td id="76E45DFFFFB9CCCB9CC7FE0AFC88FE74" box="[725,819,440,462]" gridcol="1" gridrow="4" pageId="12" pageNumber="538">16.5</td>
<td id="76E45DFFFFB9CCCB9DA7FE0AFBB5FE74" box="[949,1038,440,462]" gridcol="2" gridrow="4" pageId="12" pageNumber="538">12.6</td>
<td id="76E45DFFFFB9CCCB9A9EFE0AFAB2FE74" box="[1164,1289,440,462]" gridcol="3" gridrow="4" pageId="12" pageNumber="538">15.9</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFE52FAB2FDA7" box="[156,1289,480,541]" gridrow="5" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFE52FE7BFDA7" box="[156,448,480,541]" gridcol="0" gridrow="5" pageId="12" pageNumber="538">Length rostrum Length acumen</th>
<td id="76E45DFFFFB9CCCB9CC7FE52FC88FDA7" box="[725,819,480,541]" gridcol="1" gridrow="5" pageId="12" pageNumber="538">6.4 2.3</td>
<td id="76E45DFFFFB9CCCB9DA7FE52FBB5FDA7" box="[949,1038,480,541]" gridcol="2" gridrow="5" pageId="12" pageNumber="538">5.5 1.9</td>
<td id="76E45DFFFFB9CCCB9A9EFE52FAB2FDA7" box="[1164,1289,480,541]" gridcol="3" gridrow="5" pageId="12" pageNumber="538">6.0 1.8</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFD9DFAB2FDFF" box="[156,1289,559,581]" gridrow="6" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFD9DFE7BFDFF" box="[156,448,559,581]" gridcol="0" gridrow="6" pageId="12" pageNumber="538">Length areola</th>
<td id="76E45DFFFFB9CCCB9CC7FD9DFC88FDFF" box="[725,819,559,581]" gridcol="1" gridrow="6" pageId="12" pageNumber="538">14.6</td>
<td id="76E45DFFFFB9CCCB9DA7FD9DFBB5FDFF" box="[949,1038,559,581]" gridcol="2" gridrow="6" pageId="12" pageNumber="538">11.6</td>
<td id="76E45DFFFFB9CCCB9A9EFD9DFAB2FDFF" box="[1164,1289,559,581]" gridcol="3" gridrow="6" pageId="12" pageNumber="538">12.8</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFDE5FAB2FDD7" box="[156,1289,599,621]" gridrow="7" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFDE5FE7BFDD7" box="[156,448,599,621]" gridcol="0" gridrow="7" pageId="12" pageNumber="538">Width areola</th>
<td id="76E45DFFFFB9CCCB9CC7FDE5FC88FDD7" box="[725,819,599,621]" gridcol="1" gridrow="7" pageId="12" pageNumber="538">0</td>
<td id="76E45DFFFFB9CCCB9DA7FDE5FBB5FDD7" box="[949,1038,599,621]" gridcol="2" gridrow="7" pageId="12" pageNumber="538">0</td>
<td id="76E45DFFFFB9CCCB9A9EFDE5FAB2FDD7" box="[1164,1289,599,621]" gridcol="3" gridrow="7" pageId="12" pageNumber="538">0</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFD14FAB2FD06" box="[156,1289,678,700]" gridrow="8" pageId="12" pageNumber="538" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76E45DFFFFB9CCCB9E8EFD14FE7BFD06" box="[156,448,678,700]" gridcol="0" gridrow="8" pageId="12" pageNumber="538">Antennal scale</th>
</tr>
<tr id="35353483FFB9CCCB9E8EFD7CFAB2FD5E" box="[156,1289,718,740]" gridrow="9" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFD7CFE7BFD5E" box="[156,448,718,740]" gridcol="0" gridrow="9" pageId="12" pageNumber="538">Width</th>
<td id="76E45DFFFFB9CCCB9CC7FD7CFC88FD5E" box="[725,819,718,740]" gridcol="1" gridrow="9" pageId="12" pageNumber="538">5.5</td>
<td id="76E45DFFFFB9CCCB9DA7FD7CFBB5FD5E" box="[949,1038,718,740]" gridcol="2" gridrow="9" pageId="12" pageNumber="538">4.4</td>
<td id="76E45DFFFFB9CCCB9A9EFD7CFAB2FD5E" box="[1164,1289,718,740]" gridcol="3" gridrow="9" pageId="12" pageNumber="538">4.9</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFCAFFAB2FC89" box="[156,1289,797,819]" gridrow="10" pageId="12" pageNumber="538" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76E45DFFFFB9CCCB9E8EFCAFFE7BFC89" box="[156,448,797,819]" gridcol="0" gridrow="10" pageId="12" pageNumber="538">Abdomen</th>
</tr>
<tr id="35353483FFB9CCCB9E8EFCF7FAB2FCE1" box="[156,1289,837,859]" gridrow="11" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFCF7FE7BFCE1" box="[156,448,837,859]" gridcol="0" gridrow="11" pageId="12" pageNumber="538">Width</th>
<td id="76E45DFFFFB9CCCB9CC7FCF7FC88FCE1" box="[725,819,837,859]" gridcol="1" gridrow="11" pageId="12" pageNumber="538">12.1</td>
<td id="76E45DFFFFB9CCCB9DA7FCF7FBB5FCE1" box="[949,1038,837,859]" gridcol="2" gridrow="11" pageId="12" pageNumber="538">10.8</td>
<td id="76E45DFFFFB9CCCB9A9EFCF7FAB2FCE1" box="[1164,1289,837,859]" gridcol="3" gridrow="11" pageId="12" pageNumber="538">13.3</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFC27FAB2FC4B" box="[156,1289,917,1009]" gridrow="12" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFC27FE7BFC4B" box="[156,448,917,1009]" gridcol="0" gridrow="12" pageId="12" pageNumber="538">Cheliped Length mesial margin palm</th>
<td id="76E45DFFFFB9CCCB9CC7FC27FC88FC4B" box="[725,819,917,1009]" gridcol="1" gridrow="12" pageId="12" pageNumber="538">8.4</td>
<td id="76E45DFFFFB9CCCB9DA7FC27FBB5FC4B" box="[949,1038,917,1009]" gridcol="2" gridrow="12" pageId="12" pageNumber="538">6.7</td>
<td id="76E45DFFFFB9CCCB9A9EFC27FAB2FC4B" box="[1164,1289,917,1009]" gridcol="3" gridrow="12" pageId="12" pageNumber="538">7.8</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFBB1FAB2FBA3" box="[156,1289,1027,1049]" gridrow="13" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFBB1FE7BFBA3" box="[156,448,1027,1049]" gridcol="0" gridrow="13" pageId="12" pageNumber="538">Width palm</th>
<td id="76E45DFFFFB9CCCB9CC7FBB1FC88FBA3" box="[725,819,1027,1049]" gridcol="1" gridrow="13" pageId="12" pageNumber="538">3.2</td>
<td id="76E45DFFFFB9CCCB9DA7FBB1FBB5FBA3" box="[949,1038,1027,1049]" gridcol="2" gridrow="13" pageId="12" pageNumber="538">8.9</td>
<td id="76E45DFFFFB9CCCB9A9EFBB1FAB2FBA3" box="[1164,1289,1027,1049]" gridcol="3" gridrow="13" pageId="12" pageNumber="538">7.9</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFB99FAB2FBD2" box="[156,1289,1067,1128]" gridrow="14" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFB99FE7BFBD2" box="[156,448,1067,1128]" gridcol="0" gridrow="14" pageId="12" pageNumber="538">Depth palm Length dactyl</th>
<td id="76E45DFFFFB9CCCB9CC7FB99FC88FBD2" box="[725,819,1067,1128]" gridcol="1" gridrow="14" pageId="12" pageNumber="538">6.5 14.3</td>
<td id="76E45DFFFFB9CCCB9DA7FB99FBB5FBD2" box="[949,1038,1067,1128]" gridcol="2" gridrow="14" pageId="12" pageNumber="538">5.6 9.4</td>
<td id="76E45DFFFFB9CCCB9A9EFB99FAB2FBD2" box="[1164,1289,1067,1128]" gridcol="3" gridrow="14" pageId="12" pageNumber="538">6.4 11.5</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFBC8FAB2FB2A" box="[156,1289,1146,1168]" gridrow="15" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFBC8FE7BFB2A" box="[156,448,1146,1168]" gridcol="0" gridrow="15" pageId="12" pageNumber="538">Length carpus</th>
<td id="76E45DFFFFB9CCCB9CC7FBC8FC88FB2A" box="[725,819,1146,1168]" gridcol="1" gridrow="15" pageId="12" pageNumber="538">11.1</td>
<td id="76E45DFFFFB9CCCB9DA7FBC8FBB5FB2A" box="[949,1038,1146,1168]" gridcol="2" gridrow="15" pageId="12" pageNumber="538">7.3</td>
<td id="76E45DFFFFB9CCCB9A9EFBC8FAB2FB2A" box="[1164,1289,1146,1168]" gridcol="3" gridrow="15" pageId="12" pageNumber="538">9.7</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFB10FAB2FB02" box="[156,1289,1186,1208]" gridrow="16" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFB10FE7BFB02" box="[156,448,1186,1208]" gridcol="0" gridrow="16" pageId="12" pageNumber="538">Width carpus</th>
<td id="76E45DFFFFB9CCCB9CC7FB10FC88FB02" box="[725,819,1186,1208]" gridcol="1" gridrow="16" pageId="12" pageNumber="538">6.6</td>
<td id="76E45DFFFFB9CCCB9DA7FB10FBB5FB02" box="[949,1038,1186,1208]" gridcol="2" gridrow="16" pageId="12" pageNumber="538">5.7</td>
<td id="76E45DFFFFB9CCCB9A9EFB10FAB2FB02" box="[1164,1289,1186,1208]" gridcol="3" gridrow="16" pageId="12" pageNumber="538">6.7</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFB78FAB2FABD" box="[156,1289,1226,1287]" gridrow="17" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFB78FE7BFABD" box="[156,448,1226,1287]" gridcol="0" gridrow="17" pageId="12" pageNumber="538">Length dorsal margin merus Depth merus</th>
<td id="76E45DFFFFB9CCCB9CC7FB78FC88FABD" box="[725,819,1226,1287]" gridcol="1" gridrow="17" pageId="12" pageNumber="538">14.3 6.8</td>
<td id="76E45DFFFFB9CCCB9DA7FB78FBB5FABD" box="[949,1038,1226,1287]" gridcol="2" gridrow="17" pageId="12" pageNumber="538">11.2 5.8</td>
<td id="76E45DFFFFB9CCCB9A9EFB78FAB2FABD" box="[1164,1289,1226,1287]" gridcol="3" gridrow="17" pageId="12" pageNumber="538">14.3 6.7</td>
</tr>
<tr id="35353483FFB9CCCB9E8EFAF3FAB2FAED" box="[156,1289,1345,1367]" gridrow="18" pageId="12" pageNumber="538">
<th id="76E45DFFFFB9CCCB9E8EFAF3FE7BFAED" box="[156,448,1345,1367]" gridcol="0" gridrow="18" pageId="12" pageNumber="538">Gonopod length</th>
<td id="76E45DFFFFB9CCCB9CC7FAF3FC88FAED" box="[725,819,1345,1367]" gridcol="1" gridrow="18" pageId="12" pageNumber="538">8.2</td>
<td id="76E45DFFFFB9CCCB9DA7FAF3FBB5FAED" box="[949,1038,1345,1367]" gridcol="2" gridrow="18" pageId="12" pageNumber="538">N.A.</td>
<td id="76E45DFFFFB9CCCB9A9EFAF3FAB2FAED" box="[1164,1289,1345,1367]" gridcol="3" gridrow="18" pageId="12" pageNumber="538">5.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBA36C1FFB933389ED5FA3EFC72F8BA" blockId="12.[151,1436,1417,2034]" pageId="12" pageNumber="538">
<emphasis id="B971EAD3FFB933389ED5FA3EFDEAFA1F" bold="true" box="[199,593,1420,1445]" pageId="12" pageNumber="538">Range and specimens examined.</emphasis>
<taxonomicName id="4C054D42FFB933389C48FA3EFC90FA1F" box="[602,811,1420,1445]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="538" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB933389C48FA3EFC90FA1F" box="[602,811,1420,1445]" italics="true" pageId="12" pageNumber="538">Cambarus pauleyi</emphasis>
</taxonomicName>
is endemic to an 
<quantity id="4CFD9B24FFB933389DE9FA3EFBEAFA1F" box="[1019,1105,1420,1445]" metricMagnitude="5" metricUnit="m" metricValue="8.459999999999999" pageId="12" pageNumber="538" unit="km" value="846.0">846 km</quantity>
2/ 
<quantity id="4CFD9B24FFB933389A70FA3FFB08FA1F" box="[1122,1203,1420,1445]" metricMagnitude="5" metricUnit="m" metricValue="5.21427456" pageId="12" pageNumber="538" unit="mi" value="324.0">324 mi</quantity>
2 section of the high elevation (&gt; 
<quantity id="4CFD9B24FFB933389F39FA1DFEC3FA7D" box="[299,376,1455,1480]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="538" unit="m" value="762.0">762 m</quantity>
; 
<quantity id="4CFD9B24FFB933389F95FA1DFE58FA72" box="[391,483,1455,1480]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="538" unit="ft" value="2500.0">2,500 ft</quantity>
) Appalachian Plateau physiographic province in Greenbrier and Monroe counties, West Virginia and confined to the Meadow and Greenbrier River basins. The entirety of its range is close to the junction of the Appalachian Plateau, Allegheny Mountain, and Ridge and Valley physiographic provinces, with the eastern extent limited to the western front of the Ridge and Valley, and the northern extent limited to the southern foothills of the Allegheny Mountain provinces (
<figureCitation id="133E2A44FFB933389CA9F98BFCB9F9E8" box="[699,770,1593,1618]" captionStart="FIGURE 7" captionStartId="13.[151,250,1178,1200]" captionTargetBox="[167,1430,193,1156]" captionTargetId="figure@13.[151,1436,193,1157]" captionTargetPageId="13" captionText="FIGURE 7. Known range of Cambarus pauleyi, new species in West Virginia." httpUri="https://zenodo.org/record/239955/files/figure.png" pageId="12" pageNumber="538">Fig. 7</figureCitation>
). The distribution of 
<taxonomicName id="4C054D42FFB933389DE7F98BFB7EF9E8" box="[1013,1221,1593,1618]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="538" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB933389DE7F98BFB7EF9E8" box="[1013,1221,1593,1618]" italics="true" pageId="12" pageNumber="538">Cambarus pauleyi</emphasis>
</taxonomicName>
also appears to be limited to elevations between 
<quantity id="4CFD9B24FFB933389FE0F9EFFD34F9CE" box="[498,655,1629,1653]" metricMagnitude="2" metricUnit="m" metricValue="7.6" metricValueMax="7.9" metricValueMin="7.3" pageId="12" pageNumber="538" unit="m" value="760.0" valueMax="790.0" valueMin="730.0">730 to 790 m</quantity>
(
<quantity id="4CFD9B24FFB933389C8DF9EEFCF7F9CF" box="[671,844,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="7.5971400000000004" metricValueMax="7.8943200000000004" metricValueMin="7.2999600000000004" pageId="12" pageNumber="538" unit="ft" value="2492.5" valueMax="2590.0" valueMin="2395.0">2,395–2,590 ft</quantity>
), with the majority of known extant occurrences in wetlands adjacent to Meadow River in Greenbrier County, West Virginia. Collecting efforts between the 
<typeStatus id="54BE8863FFB933389B7EF932FA27F922" box="[1388,1436,1664,1688]" pageId="12" pageNumber="538">type</typeStatus>
locality and the apparent core of the 
<taxonomicName id="4C054D42FFB933389C26F910FD1CF900" box="[564,679,1697,1722]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="538" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB933389C26F910FD1CF900" box="[564,679,1697,1722]" italics="true" pageId="12" pageNumber="538">C. pauleyi</emphasis>
</taxonomicName>
range only produced 
<taxonomicName id="4C054D42FFB933389D8EF910FB8EF900" box="[924,1077,1697,1722]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="538" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFB933389D8EF910FC14F903" box="[924,943,1698,1721]" italics="true" pageId="12" pageNumber="538">C</emphasis>
. aff. 
<emphasis id="B971EAD3FFB933389DF8F913FB8EF900" box="[1002,1077,1697,1722]" italics="true" pageId="12" pageNumber="538">dubius</emphasis>
</taxonomicName>
. Over 95% of historic wetlands in this region had been ditch-drained, and converted to agricultural fields. It is possible that populations persist in unsampled areas in this region, warranting future surveys for the species.
</paragraph>
<paragraph id="8BBA36C1FFB933389ED5F8B8FEB9F848" blockId="12.[151,1436,1417,2034]" pageId="12" pageNumber="538">
Due to its limited distribution, and potential conservation concerns associated with over collecting, specific locations will not be given following collection accession numbers, but are available from the primary author upon request. All West Liberty University collections (WLU) were made by ZJL and the 2014 West Liberty University Astacology Laboratory summer field crew, and are housed in the West Liberty University Astacology Collection, denoted with the prefix WLU. Additional specimens were examined from the Ohio State University Museum of Biological Diversity Crustacean Collection, denoted with the prefix 
<collectionCode id="ED14AE04FFB933389D80F805FC75F875" box="[914,974,1975,1999]" httpUri="http://grbio.org/cool/9iaf-dk63" name="Ohio State University" pageId="12" pageNumber="538">OSU</collectionCode>
. MI = form I male, MII = form II male, F = female.
</paragraph>
<caption id="DF7A6649FFB833399E85FB28FC0DFB0A" box="[151,950,1178,1200]" httpUri="https://zenodo.org/record/239955/files/figure.png" pageId="13" pageNumber="539" targetBox="[167,1430,193,1156]" targetPageId="13">
<paragraph id="8BBA36C1FFB833399E85FB28FC0DFB0A" blockId="13.[151,950,1178,1200]" box="[151,950,1178,1200]" pageId="13" pageNumber="539">
<emphasis id="B971EAD3FFB833399E85FB28FEA9FB0A" bold="true" box="[151,274,1178,1200]" pageId="13" pageNumber="539">FIGURE 7.</emphasis>
Known range of 
<taxonomicName id="4C054D42FFB833399FD6FB28FDC5FB0A" box="[452,638,1178,1200]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="539" phylum="Arthropoda" rank="species" species="pauleyi" status="sp. nov.">
<emphasis id="B971EAD3FFB833399FD6FB28FDC5FB0A" box="[452,638,1178,1200]" italics="true" pageId="13" pageNumber="539">Cambarus pauleyi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24257A8FFB833399C98FB29FCBFFB0A" box="[650,772,1178,1200]" pageId="13" pageNumber="539" rank="species">new species</taxonomicNameLabel>
in West Virginia.
</paragraph>
</caption>
<paragraph id="8BBA36C1FFB833399ED5FB6CFC51FA15" blockId="13.[151,1437,1245,2032]" pageId="13" pageNumber="539">
A total of 
<specimenCount id="9D03FD48FFB833399F2FFB6FFE6CFB4C" box="[317,471,1245,1270]" pageId="13" pageNumber="539" type="generic">62 specimens</specimenCount>
were examined from 8 localities that compose the following 12 lots: WLU 1997, 
<date id="FFBB1001FFB833399B92FB6FFEB7FAA3" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 6 MII, 3 F; WLU 1998, 
<date id="FFBB1001FFB833399C31FAB3FD00FAA3" box="[547,699,1281,1306]" pageId="13" pageNumber="539" value="2014-06-25">25 June 2014</date>
, 3 MII, 1 F; WLU 1999, 
<date id="FFBB1001FFB833399DC0FAB3FBD1FAA3" box="[978,1130,1281,1305]" pageId="13" pageNumber="539" value="2014-06-27">27 June 2014</date>
, 1 MI; WLU 2000, 
<date id="FFBB1001FFB833399B54FAB3FF69FA84" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 1 F ovig; WLU 2001, 
<date id="FFBB1001FFB833399FABFA94FDEAFA84" box="[441,593,1318,1342]" pageId="13" pageNumber="539" value="2014-06-28">28 June 2014</date>
, 6 MII, 2 F; WLU 2002, 
<date id="FFBB1001FFB833399D7EFA97FBBFFA84" box="[876,1028,1317,1342]" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 1 F ovig, WLU 2003, 
<date id="FFBB1001FFB833399AF8FA97FA39FA84" box="[1258,1410,1317,1342]" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 1 F ovig; WLU 2004, 
<date id="FFBB1001FFB833399F4DFAFEFE42FADE" box="[351,505,1356,1380]" pageId="13" pageNumber="539" value="2014-06-27">27 June 2014</date>
, 2 MII, 2 F; WLU 2005, 
<date id="FFBB1001FFB833399D04FAFEFC0BFADE" box="[790,944,1356,1381]" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 1 MII; WLU 2006, 
<date id="FFBB1001FFB833399A88FAFEFA88FADE" box="[1178,1331,1356,1381]" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 4 MII, 4 F; WLU 2007, 
<date id="FFBB1001FFB833399F53FAC0FE62FA31" box="[321,473,1394,1419]" pageId="13" pageNumber="539" value="2014-06-26">26 June 2014</date>
, 2 MII, 4 F; WLU 2008, 
<date id="FFBB1001FFB833399CE2FAC1FC33FA31" box="[752,904,1395,1419]" pageId="13" pageNumber="539" value="2008-05-24">24 May 2008</date>
, 2 F, ZJL and Christopher G. Vopal; 
<collectionCode id="ED14AE04FFB833399B0CFAC1FAEDFA31" box="[1310,1366,1395,1419]" httpUri="http://grbio.org/cool/9iaf-dk63" name="Ohio State University" pageId="13" pageNumber="539">OSU</collectionCode>
9183, 
<date id="FFBB1001FFB833399E85FA25FEF6FA15" box="[151,333,1431,1455]" pageId="13" pageNumber="539" value="2013-08-30">30 August 2013</date>
, 1 MI, 1 MII, 7 F, 7 Juv, R. F. Thoma and Mitchel E. Farley
</paragraph>
<paragraph id="8BBA36C1FFB833399ED5FA08FC0FF9D9" blockId="13.[151,1437,1245,2032]" pageId="13" pageNumber="539">
<emphasis id="B971EAD3FFB833399ED5FA08FE7AFA69" bold="true" box="[199,449,1466,1491]" pageId="13" pageNumber="539">Conservation status.</emphasis>
<taxonomicName id="4C054D42FFB833399FDCFA08FD18FA69" box="[462,675,1466,1491]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="539" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB833399FDCFA08FD18FA69" box="[462,675,1466,1491]" italics="true" pageId="13" pageNumber="539">Cambarus pauleyi</emphasis>
</taxonomicName>
should be listed as Endangered (E) according to the American Fisheries Society criteria (
<bibRefCitation id="EF944B30FFB833399FD4FA6CFD25FA4D" author="Taylor" box="[454,670,1502,1527]" pageId="13" pageNumber="539" refString="Taylor, C. A., Schuster, G. A., Cooper, J. E., DiStefano, R. J., Eversole, A. G., Hamr, P., Hobbs, H. H. III, Robison, H. W., Skelton, C. E. &amp; Thoma, R. F. (2007) Reassessment of the conservation status of crayfishes of the United States and Canada after 10 + years of increased awareness. Fisheries, 32, 372 - 389. http: // dx. doi. org / 10.1577 / 1548 - 8446 (2007) 32 [372: AROTCS] 2.0. CO; 2" type="journal article" year="2007">
Taylor 
<emphasis id="B971EAD3FFB833399C09FA52FDF4FA4D" box="[539,591,1502,1527]" italics="true" pageId="13" pageNumber="539">et al</emphasis>
. 2007
</bibRefCitation>
), and assigned a G2 ranking according to the global conservation criteria (
<bibRefCitation id="EF944B30FFB833399EE7F9B1FE2BF9A1" author="Masters" box="[245,400,1539,1563]" pageId="13" pageNumber="539" refString="Masters, L. (1991) Assessing threats and setting priorities in conservation. Conservation Biology, 5, 559 - 563." type="journal article" year="1991">Masters 1991</bibRefCitation>
) for conservation listing due to its limited range, and reduction of range. 
<taxonomicName id="4C054D42FFB833399ADFF9B0FA27F9A1" box="[1229,1436,1538,1563]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="539" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB833399ADFF9B0FA27F9A1" box="[1229,1436,1538,1563]" italics="true" pageId="13" pageNumber="539">Cambarus pauleyi</emphasis>
</taxonomicName>
should be listed as near threatened (VU) using the International Union for the Conservation of Nature (
<collectionCode id="ED14AE04FFB833399B1CF995FAEFF985" box="[1294,1364,1575,1599]" pageId="13" pageNumber="539">IUCN</collectionCode>
2001) criteria due to its narrow distribution and reduction of its natural range.
</paragraph>
<paragraph id="8BBA36C1FFB833399ED5F9DCFE7DF8E5" blockId="13.[151,1437,1245,2032]" pageId="13" pageNumber="539">
Preferred habitats for 
<taxonomicName id="4C054D42FFB833399FC2F9DCFD1FF93D" box="[464,676,1646,1671]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="539" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB833399FC2F9DCFD1FF93D" box="[464,676,1646,1671]" italics="true" pageId="13" pageNumber="539">Cambarus pauleyi</emphasis>
</taxonomicName>
appear to be upland wetlands, seepage stream banks, and wet meadows at the base of hillsides. These habitats are also prime agricultural sites in a state dominated by mountainous topography. Within the upper Meadow River Valley, and specifically within Meadow River Wildlife Management Area (
<collectionCode id="ED14AE04FFB833399F66F969FE57F949" box="[372,492,1755,1779]" pageId="13" pageNumber="539">MRWMA</collectionCode>
), 
<bibRefCitation id="EF944B30FFB833399FEDF969FD1DF949" author="Putnam" box="[511,678,1754,1779]" pageId="13" pageNumber="539" refString="Putnam, N. J. (1995) Plant communities of the Meadow River Wetlands. Final project report submitted to the West Virginia Division of Natural Resources Heritage Program, 17 pp." type="book" year="1995">Putnam (1995)</bibRefCitation>
cited conversion of wetlands to agricultural sites as the number one threat to unique floristic communities in the area. Currently the 
<collectionCode id="ED14AE04FFB833399D7EF94DFC65F8AD" box="[876,990,1791,1815]" pageId="13" pageNumber="539">MRWMA</collectionCode>
is the stronghold for the species, given the majority of wetlands and meadows that abut the 
<collectionCode id="ED14AE04FFB833399CF3F891FCE8F881" box="[737,851,1827,1851]" pageId="13" pageNumber="539">MRWMA</collectionCode>
parcel have been converted to pasture and row crop agriculture (
<bibRefCitation id="EF944B30FFB833399F0CF8F5FE02F8E5" author="Putnam" box="[286,441,1862,1887]" pageId="13" pageNumber="539" refString="Putnam, N. J. (1995) Plant communities of the Meadow River Wetlands. Final project report submitted to the West Virginia Division of Natural Resources Heritage Program, 17 pp." type="book" year="1995">Putnam 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BBA36C1FFB8333A9ED5F8D8FCBFFF6F" blockId="13.[151,1437,1245,2032]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="540" pageId="13" pageNumber="539">
Given the rate of conversion of wetlands to farmland in the upper Meadow River Valley, and the lack of any specimens procured from this habitat by us, it is highly likely that 
<taxonomicName id="4C054D42FFB833399D88F83DFBB5F81D" box="[922,1038,1934,1959]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="539" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFB833399D88F83DFBB5F81D" box="[922,1038,1934,1959]" italics="true" pageId="13" pageNumber="539">C. pauleyi</emphasis>
</taxonomicName>
has experienced greater than 50% range reduction across its limited natural range in Greenbrier and Monroe County, West Virginia. West Virginia currently is experiencing a boom in natural gas extraction. Associated with this industry is a burgeoning network of natural gas pipelines, some of which have been proposed to run through the core range of 
<taxonomicName id="4C054D42FFBB333A9A9EFF2AFB44FF0A" box="[1164,1279,151,176]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9A9EFF2AFB44FF0A" box="[1164,1279,151,176]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
. The ultimate impact of this practice on 
<taxonomicName id="4C054D42FFBB333A9FA8FF0FFD95FF6F" box="[442,558,188,213]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9FA8FF0FFD95FF6F" box="[442,558,188,213]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
remains unknown.
</paragraph>
<paragraph id="8BBA36C1FFBB333A9ED5FF6DFAD6FD12" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="540">
<emphasis id="B971EAD3FFBB333A9ED5FF6DFE48FF42" bold="true" box="[199,499,223,248]" pageId="14" pageNumber="540">Ecology and life history.</emphasis>
<taxonomicName id="4C054D42FFBB333A9FECFF6DFD74FF42" box="[510,719,223,248]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9FECFF6DFD74FF42" box="[510,719,223,248]" italics="true" pageId="14" pageNumber="540">Cambarus pauleyi</emphasis>
</taxonomicName>
should be classified as a primary burrower following 
<bibRefCitation id="EF944B30FFBB333A9B41FF6DFF5AFEA6" author="Hobbs" pageId="14" pageNumber="540" refString="Hobbs, H. H. Jr. (1981) The Crayfishes of Georgia. Smithsonian Contributions to Zoology, 318, 1 - 549. http: // dx. doi. org / 10.5479 / si. 00810282.318" type="journal article" year="1981">Hobbs (1981)</bibRefCitation>
due to its utilization of burrows for all facets of its ecology and life history. No animals were observed in surface waters, though several colonies were in direct contact with such habitats (ZJL, personal observation). Seepage wetlands at slightly higher (
<quantity id="4CFD9B24FFBB333A9C2CFEFEFD1EFEDE" box="[574,677,332,357]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="14" pageNumber="540" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
) elevations immediately adjacent to large Pin Oak/Maple swamps appear to be the preferred habitat for 
<taxonomicName id="4C054D42FFBB333A9C2CFEC2FD08FE32" box="[574,691,367,392]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9C2CFEC2FD08FE32" box="[574,691,367,392]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
. Forested seeps on hillsides and along ridgelines within the range of the species did not harbor any colonies of 
<taxonomicName id="4C054D42FFBB333A9C9CFE27FCB9FE17" box="[654,770,404,429]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9C9CFE27FCB9FE17" box="[654,770,404,429]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
, but we did find populations of an unnamed member of the 
<taxonomicName id="4C054D42FFBB333A9E85FE0AFE8DFE6A" box="[151,310,439,464]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBB333A9E85FE0AFF0BFE75" box="[151,176,440,463]" italics="true" pageId="14" pageNumber="540">C.</emphasis>
aff. 
<emphasis id="B971EAD3FFBB333A9EF9FE05FE8DFE6A" box="[235,310,439,464]" italics="true" pageId="14" pageNumber="540">dubius</emphasis>
</taxonomicName>
complex, which we refer to as the black-body orange-claw (
<collectionCode id="ED14AE04FFBB333A9DEFFE0AFBF7FE6A" box="[1021,1100,440,464]" pageId="14" pageNumber="540">BBOC</collectionCode>
) morph, in these habitats. In addition to seepage wetlands, 
<taxonomicName id="4C054D42FFBB333A9FF5FE6FFDE0FE4F" box="[487,603,476,501]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9FF5FE6FFDE0FE4F" box="[487,603,476,501]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
also occurs in low-gradient headwater seepage streams. In these environs, 
<taxonomicName id="4C054D42FFBB333A9E85FDB2FEABFDA2" box="[151,272,511,536]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9E85FDB2FEABFDA2" box="[151,272,511,536]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
burrows both in floodplains and occasionally in stream banks. 
<taxonomicName id="4C054D42FFBB333A9A05FE4DFB50FDA2" box="[1047,1259,511,536]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9A05FE4DFB50FDA2" box="[1047,1259,511,536]" italics="true" pageId="14" pageNumber="540">Cambarus pauleyi</emphasis>
</taxonomicName>
does occur in anthropogenic habitats, although at much lower densities when compared to population levels in natural undisturbed situations (ZJL, personal observation). Anthropogenic habitats utilized by 
<taxonomicName id="4C054D42FFBB333A9A6DFDFAFB4EFDDA" box="[1151,1269,583,608]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9A6DFDFAFB4EFDDA" box="[1151,1269,583,608]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
included both residential yards and roadside ditches, with higher densities being found in the latter. In disturbed situations, 
<taxonomicName id="4C054D42FFBB333A9B90FDDFFF52FD12" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9B90FDDFFF52FD12" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
was subordinate to 
<taxonomicName id="4C054D42FFBB333A9FDBFD22FD8CFD12" box="[457,567,655,680]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="thomai">
<emphasis id="B971EAD3FFBB333A9FDBFD22FD8CFD12" box="[457,567,655,680]" italics="true" pageId="14" pageNumber="540">C. thomai</emphasis>
</taxonomicName>
, the latter being more prevalent in wetlands associated with such habitat.
</paragraph>
<paragraph id="8BBA36C1FFBB333A9ED5FD06FE91FBAA" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="540">
Compared to other members of the 
<taxonomicName id="4C054D42FFBB333A9C61FD07FCAEFD77" box="[627,789,692,717]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBB333A9C61FD07FD37FD76" box="[627,652,693,716]" italics="true" pageId="14" pageNumber="540">C.</emphasis>
aff. 
<emphasis id="B971EAD3FFBB333A9CD8FD06FCAEFD77" box="[714,789,692,717]" italics="true" pageId="14" pageNumber="540">dubius</emphasis>
</taxonomicName>
complex, 
<taxonomicName id="4C054D42FFBB333A9D86FD07FBB0FD77" box="[916,1035,692,717]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9D86FD07FBB0FD77" box="[916,1035,692,717]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
burrow morphology is simplistic. Burrows maintain one or two entrances leading to vertical tunnels, which are frequently capped, plugged or possess well-constructed chimneys. Burrows with two openings maintained bifurcate tunnels that unify into single central tunnels after 
<quantity id="4CFD9B24FFBB333A9F3BFCADFE27FC8D" box="[297,412,799,824]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="14" pageNumber="540" unit="cm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 cm</quantity>
. Central tunnels continue an additional 
<quantity id="4CFD9B24FFBB333A9D4BFCADFC72FC8D" box="[857,969,799,824]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="14" pageNumber="540" unit="cm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 cm</quantity>
vertically, ultimately ending in simplistic resting chambers. Ancillary tunnels running laterally and along resting chamber floors are rarely encountered in this species (ZJL, personal observation). Burrows of syntopic 
<taxonomicName id="4C054D42FFBB333A9D4DFCDAFC74FC3A" box="[863,975,871,896]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="thomai">
<emphasis id="B971EAD3FFBB333A9D4DFCDAFC74FC3A" box="[863,975,871,896]" italics="true" pageId="14" pageNumber="540">C. thomai</emphasis>
</taxonomicName>
often maintain several ancillary tunnels extending from both the central tunnel and resting chambers of the burrow (ZJL, personal observation). Burrow morphology for the 
<collectionCode id="ED14AE04FFBB333A9F65FC02FE78FC72" box="[375,451,944,968]" pageId="14" pageNumber="540">BBOC</collectionCode>
morph differs from 
<taxonomicName id="4C054D42FFBB333A9CB4FC02FCA1FC72" box="[678,794,943,968]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9CB4FC02FCA1FC72" box="[678,794,943,968]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
in the orientation of the initial burrow tunnel, which runs laterally into hillsides for 
<quantity id="4CFD9B24FFBB333A9FCBFC66FDEBFC56" box="[473,592,980,1004]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="14" pageNumber="540" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
before adopting a more vertical orientation (ZJL and RFT, personal observation).
</paragraph>
<paragraph id="8BBA36C1FFBB333A9ED5FBAEFB26F9EA" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="540">
Surface activity in this species was not relegated solely to periods of darkness. On three occasions, 
<taxonomicName id="4C054D42FFBB333A9B3AFBAFFA20FB8F" box="[1320,1435,1052,1077]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9B3AFBAFFA20FB8F" box="[1320,1435,1052,1077]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
was observed positioned within 
<quantity id="4CFD9B24FFBB333A9C1DFB8DFDF1FBED" box="[527,586,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="540" unit="cm" value="5.0">5 cm</quantity>
of burrow entrances at midafternoon; two of these observation occurred during periods of bright sunshine. Though observed during periods of daylight, 
<taxonomicName id="4C054D42FFBB333A9A0BFBD7FB36FBC7" box="[1049,1165,1124,1149]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9A0BFBD7FB36FBC7" box="[1049,1165,1124,1149]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
surface activity reaches its highest levels during the first three hours following twilight (ZJL, personal observation). Fifty plus nocturnal observations were made of this species resting at the entrance to its burrow (ZJL, personal observation). Observed surface behaviors included plugging burrows, maintaining chimneys, grooming, and foraging. Foraging behavior included positioning of antennae at both 45° and 135°, with chelae slightly elevated and agape. Animals encountered in this position frequently lunged at any object that made contact with their primary antennae (ZJL, personal observation). After a brief period following grasping of an object, objects deemed edible were drug down into the confines of the burrow. Defensive behaviors were also observed for 
<taxonomicName id="4C054D42FFBB333A9A13FAD2FBCDFAC2" box="[1025,1142,1375,1400]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9A13FAD2FBCDFAC2" box="[1025,1142,1375,1400]" italics="true" pageId="14" pageNumber="540">C. pauleyi</emphasis>
</taxonomicName>
resting at burrow portals. Animals engaged at the burrow entrance that were not interested in food would most frequently flee down their burrows, but occasionally would position their antennae over their carapace, at the same time gaping their chelae. Once this posture was adopted, animals deliberately moved towards stimuli, slashing their agape chelae at the perceived threat. After two or three bouts of this behavior, most animals would quickly fall down their burrows by pulling their pereiopods tightly against their bodies. Individuals that displayed this behavior were not observed at burrow openings for the remaining period of a collection effort (ZJL, personal observation).
</paragraph>
</subSubSection>
<subSubSection id="C31F654AFFBB333B9ED5F9EEFACBFE4E" lastPageId="15" lastPageNumber="541" pageId="14" pageNumber="540" type="reference_group">
<paragraph id="8BBA36C1FFBB333B9ED5F9EEFE0EFE17" blockId="14.[151,1437,151,2013]" lastBlockId="15.[151,1437,151,2027]" lastPageId="15" lastPageNumber="541" pageId="14" pageNumber="540">
<taxonomicName id="4C054D42FFBB333A9ED5F9EEFE22F9CF" ID-CoL="7RV3Y" box="[199,409,1628,1653]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="540" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBB333A9ED5F9EEFE22F9CF" box="[199,409,1628,1653]" italics="true" pageId="14" pageNumber="540">Cambarus pauleyi</emphasis>
</taxonomicName>
life history is poorly understood and in need of further research. Based on specimens captured with this effort, as well as Jezerinac 
<emphasis id="B971EAD3FFBB333A9CB8F933FD66F922" box="[682,733,1663,1688]" italics="true" pageId="14" pageNumber="540">et al</emphasis>
. (1995), males appear to molt from form II to form I during August through September, and maintain a form I condition from fall through mid-May to early June, when they molt from form I back to form II. This is evidenced by only one form I male (5.5 %) collected in 
<date id="FFBB1001FFBB333A9B0DF97BFA23F95A" box="[1311,1432,1736,1760]" pageId="14" pageNumber="540" value="2014-06">June 2014</date>
, compared to 17 form II males (94.5%) from across 6 sites. Carapaces were immaculate for several males captured during this time indicating recent molting activity (ZJL, personal observation). Jezerinac 
<emphasis id="B971EAD3FFBB333A9A8FF8A3FB69F892" box="[1181,1234,1807,1832]" italics="true" pageId="14" pageNumber="540">et al</emphasis>
. (1995) collected form I males on 
<date id="FFBB1001FFBB333A9F5DF886FDBFF8F6" box="[335,516,1844,1868]" pageId="14" pageNumber="540" value="1989-08-19">19 August 1989</date>
, further supporting this contention. The 
<typeStatus id="54BE8863FFBB333A9DD4F886FB93F8F7" box="[966,1064,1844,1869]" pageId="14" pageNumber="540" type="holotype">holotype</typeStatus>
molted from form II to form I on 
<date id="FFBB1001FFBB333A9E85F8EAFEC6F8CA" box="[151,381,1879,1904]" pageId="14" pageNumber="540" value="2014-09-17">17 September 2014</date>
, adding support. Glared females were observed in August, September and October (ZJL, personal observation). Glare glands of two females maintained in captivity steadily developed beginning in August, and reached full development by early October (ZJL, personal observation). It is theorized that egg deposition in this species occurs April through June. A single ovigerous female with 
<specimenCount id="9D03FD48FFBB333A9A5EF876FB19F867" box="[1100,1186,1988,2013]" pageId="14" pageNumber="540" type="egg">31 eggs</specimenCount>
was excavated from a burrow on 
<date id="FFBB1001FFBA333B9F08FF25FE07FF0A" box="[282,444,151,176]" pageId="15" pageNumber="541" value="2014-06-26">26 June 2014</date>
. Two females maintaining instars excavated from burrows on 
<date id="FFBB1001FFBA333B9A89FF25FA2CFF0A" box="[1179,1431,151,176]" pageId="15" pageNumber="541" value="2014-06-26" valueMax="2014-06-28" valueMin="2014-06-26">26 and 28 June 2014</date>
; average instar compliment for the females was 26.0 instars (SE = 5.67 instars). Both females were collected when chimneys were removed from burrows. In both instances females were resting inside the chimney, and did not attempt to flee once observed (ZJL, personal observation). The third female responded aggressively to plunging of her burrow, rising from the confines of the resting chamber with chelae agape. In several instances, juveniles were encountered inside burrows alongside females, but never alongside males. This behavior has been observed for several primary burrowing crayfish (Richardson 2007; 
<bibRefCitation id="EF944B30FFBA333B9D1BFEDDFC69FE32" author="Loughman" box="[777,978,367,392]" pageId="15" pageNumber="541" refString="Loughman, Z. J. (2010 b) Ecology of Cambarus dubius in north-central West Virginia. Southeastern Naturalist, 9 (Special Issue 3), 217 - 230." type="journal article" year="2010" yearSuffix="b">Loughman 2010b</bibRefCitation>
) including nominate 
<taxonomicName id="4C054D42FFBA333B9AD7FEC2FA89FE32" ID-CoL="7RTY5" box="[1221,1330,367,392]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9AD7FEC2FA89FE32" box="[1221,1330,367,392]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF944B30FFBA333B9B50FEC2FF68FE16" author="Dewees" pageId="15" pageNumber="541" refString="Dewees, J. P. (1972) Geographic variation in the primary burrowing crayfish, Cambarus dubius and Cambarus carolinus in Tennessee with notes on ecology and life history. M. S. Thesis University of Tennessee, Knoxville, Tennessee, 76 pp." type="book" year="1972">Dewees 1972</bibRefCitation>
; 
<bibRefCitation id="EF944B30FFBA333B9ECFFE26FE13FE17" author="Loughman" box="[221,424,404,429]" pageId="15" pageNumber="541" refString="Loughman, Z. J. (2010 b) Ecology of Cambarus dubius in north-central West Virginia. Southeastern Naturalist, 9 (Special Issue 3), 217 - 230." type="journal article" year="2010" yearSuffix="b">Loughman 2010b</bibRefCitation>
).
</paragraph>
<paragraph id="8BBA36C1FFBA333B9ED5FE05FACBFE4E" blockId="15.[151,1437,151,2027]" pageId="15" pageNumber="541">
<emphasis id="B971EAD3FFBA333B9ED5FE05FE02FE6A" bold="true" box="[199,441,439,464]" pageId="15" pageNumber="541">Crayfish associates.</emphasis>
<taxonomicName id="4C054D42FFBA333B9FD5FE05FD69FE6A" ID-CoL="7RV3Y" box="[455,722,439,464]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi" subGenus="Jugicambarus">
<emphasis id="B971EAD3FFBA333B9FD5FE05FD69FE6A" box="[455,722,439,464]" italics="true" pageId="15" pageNumber="541">Cambarus (J.) pauleyi</emphasis>
</taxonomicName>
has been collected in areas also harboring 
<taxonomicName id="4C054D42FFBA333B9AFEFE05FE24FE4E" ID-CoL="7RTWH" authority="Hay, 1914" authorityName="Hay" authorityYear="1914" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="carinirostris" subGenus="C.">
<emphasis id="B971EAD3FFBA333B9AFEFE05FADAFE6A" box="[1260,1377,439,464]" italics="true" pageId="15" pageNumber="541">Cambarus</emphasis>
(
<emphasis id="B971EAD3FFBA333B9B6BFE0AFA37FE75" box="[1401,1420,440,463]" italics="true" pageId="15" pageNumber="541">C</emphasis>
.) 
<emphasis id="B971EAD3FFBA333B9E85FE6FFE99FE4E" box="[151,290,477,500]" italics="true" pageId="15" pageNumber="541">carinirostris</emphasis>
<bibRefCitation id="EF944B30FFBA333B9F3BFE6FFE24FE4E" author="Hay" box="[297,415,476,501]" pageId="15" pageNumber="541" refString="Hay, W. P. (1914) Cambarus bartonii carinirostris Hay. In: Faxon W (1914) Notes on the crayfishes in the United States National Museum and the Museum of Comparative Zoology, with descriptions of new species and subspecies to which is appended a catalogue of the known species and subspecies. Memoirs of the Museum of Comparative Zoology at Harvard College, 40, 351 - 427." type="journal article" year="1914">Hay, 1914</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C054D42FFBA333B9FB8FE6EFCE1FE4E" ID-CoL="7RV5R" authority="Jezerinac, 1993" authorityName="Jezerinac" authorityYear="1993" box="[426,858,476,501]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="thomai" subGenus="T.">
<emphasis id="B971EAD3FFBA333B9FB8FE6EFDA4FE4F" box="[426,543,476,501]" italics="true" pageId="15" pageNumber="541">Cambarus</emphasis>
(
<emphasis id="B971EAD3FFBA333B9C3DFE6FFD87FE4E" box="[559,572,477,500]" italics="true" pageId="15" pageNumber="541">T</emphasis>
.) 
<emphasis id="B971EAD3FFBA333B9C46FE6EFD19FE4F" box="[596,674,476,501]" italics="true" pageId="15" pageNumber="541">thomai</emphasis>
<bibRefCitation id="EF944B30FFBA333B9CBBFE6EFCE1FE4E" author="Jezerinac" box="[681,858,476,501]" pageId="15" pageNumber="541" refString="Jezerinac, R. F. (1993) A new subgenus and new species of crayfish (Decapoda: Cambaridae) of the genus Cambarus with an amended description of the subgenus Lacunicambarus. Proceedings of the Biological Society of Washington, 106, 532 - 544." type="journal article" year="1993">Jezerinac, 1993</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C054D42FFBA333B9D87FE6EFB97FE4F" ID-CoL="7PV9X" box="[917,1068,476,501]" class="Malacostraca" family="Cambaridae" genus="Procambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="B971EAD3FFBA333B9D87FE6EFB97FE4F" box="[917,1068,476,501]" italics="true" pageId="15" pageNumber="541">Procambarus</emphasis>
</taxonomicName>
(
<collectingCountry id="F3127651FFBA333B9A2FFE6FFBEAFE4E" box="[1085,1105,477,500]" name="Iceland" pageId="15" pageNumber="541">
<emphasis id="B971EAD3FFBA333B9A2FFE6FFBEAFE4E" box="[1085,1105,477,500]" italics="true" pageId="15" pageNumber="541">O</emphasis>
</collectingCountry>
.) 
<emphasis id="B971EAD3FFBA333B9A7AFE6FFBCDFE4E" box="[1128,1142,477,500]" italics="true" pageId="15" pageNumber="541">a</emphasis>
. 
<emphasis id="B971EAD3FFBA333B9A96FE6FFB76FE4E" box="[1156,1229,477,500]" italics="true" pageId="15" pageNumber="541">acutus</emphasis>
<bibRefCitation id="EF944B30FFBA333B9AC6FE6EFAD6FE4E" author="Fowler" box="[1236,1389,476,501]" pageId="15" pageNumber="541" refString="Fowler, H. W. (1912) The Crustacea of New Jersey. In: Annual Report of the New Jersey State Museum for 1911. Trenton, New Jersey, pp. 29 - 651." type="book chapter" year="1912">Fowler, 1912</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F654AFFBA33249ED5FE4DFE67FEDE" lastPageId="16" lastPageNumber="542" pageId="15" pageNumber="541" type="description">
<paragraph id="8BBA36C1FFBA333B9ED5FE4DFDE3FD3F" blockId="15.[151,1437,151,2027]" pageId="15" pageNumber="541">
<emphasis id="B971EAD3FFBA333B9ED5FE4DFE80FDA2" bold="true" box="[199,315,511,536]" pageId="15" pageNumber="541">Variation.</emphasis>
Very little morphological variation was observed between sites and life stages, with the only noticeable variation being juvenile carapaces that are less laterally inflated than adults. Tubercle shape and degree of supination did not display ontogenic differences in this species; both juvenile and adult 
<taxonomicName id="4C054D42FFBA333B9AF4FDFAFAE4FDDA" box="[1254,1375,583,608]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9AF4FDFAFAE4FDDA" box="[1254,1375,583,608]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
lack postorbital, cervical, and hepatic spines.
</paragraph>
<paragraph id="8BBA36C1FFBA333B9ED5FD3DFACAFC48" blockId="15.[151,1437,151,2027]" pageId="15" pageNumber="541">
<emphasis id="B971EAD3FFBA333B9ED5FD3DFDF2FD12" bold="true" box="[199,585,655,680]" pageId="15" pageNumber="541">Relationships and comparisons.</emphasis>
<taxonomicName id="4C054D42FFBA333B9C41FD3DFC9FFD12" box="[595,804,655,680]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9C41FD3DFC9FFD12" box="[595,804,655,680]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
is placed in the subgenus 
<emphasis id="B971EAD3FFBA333B9A72FD3DFB44FD12" box="[1120,1279,655,680]" italics="true" pageId="15" pageNumber="541">Jugicambarus</emphasis>
based on the presence of cristiform tubercles on the mesial margin of the chelae (
<bibRefCitation id="EF944B30FFBA333B9DB3FD06FB8BFD77" author="Hobbs" box="[929,1072,692,717]" pageId="15" pageNumber="541" refString="Hobbs, H. H. Jr. (1969) On the distribution and phylogeny of the crayfish genus Cambarus. In: Holt, P. C. &amp; Hoffman, R. L. (Eds.), The Distributional History of the Biota of the Southern Appalachians. Part I: Invertebrates. Research Division Monograph 1. Virginia Polytechnic Institution, Blacksburg, Virginia, pp. 93 - 178." type="journal article" year="1969">Hobbs 1969</bibRefCitation>
). Among described members of the subgenus, 
<taxonomicName id="4C054D42FFBA333B9F2AFD6AFE17FD4A" box="[312,428,727,752]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9F2AFD6AFE17FD4A" box="[312,428,727,752]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
is most similar to nominate 
<taxonomicName id="4C054D42FFBA333B9CFCFD6AFCE0FD4A" box="[750,859,727,752]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9CFCFD6AFCE0FD4A" box="[750,859,727,752]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
and several undescribed members of the 
<taxonomicName id="4C054D42FFBA333B9B22FD6AFA27FD4A" box="[1328,1436,727,752]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9B22FD6AFA27FD4A" box="[1328,1436,727,752]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
complex. Color in life, differences in palm depth/palm length and rostral width/rostral length ratios, as well as morphological differences in the chelae and carapace anatomy can all be used to distinguish 
<taxonomicName id="4C054D42FFBA333B9AD6FC92FA82FC82" box="[1220,1337,799,824]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9AD6FC92FA82FC82" box="[1220,1337,799,824]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C054D42FFBA333B9B90FC92FF59FCE7" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="dubius">
<emphasis id="B971EAD3FFBA333B9B90FC92FF59FCE7" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257A8FFBA333B9EFEFCF7FEAAFCE6" box="[236,273,837,860]" pageId="15" pageNumber="541" sensu="stricto">s.s.</taxonomicNameLabel>
, as well as undescribed members of the complex. The dorsal carapace, chelae, and abdomen of 
<taxonomicName id="4C054D42FFBA333B9B90FCF7FF52FC3A" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9B90FCF7FF52FC3A" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
all display various shades of blue in living specimens, whereas 
<taxonomicName id="4C054D42FFBA333B9DA2FCDAFBA0FC3A" box="[944,1051,871,896]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="dubius">
<emphasis id="B971EAD3FFBA333B9DA2FCDAFBA0FC3A" box="[944,1051,871,896]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257A8FFBA333B9A33FCDAFBFEFCC5" box="[1057,1093,872,895]" pageId="15" pageNumber="541" sensu="stricto">s.s.</taxonomicNameLabel>
is predominately composed of oranges and orange-reds. 
<taxonomicName id="4C054D42FFBA333B9FA7FC3EFD38FC1F" box="[437,643,908,933]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9FA7FC3EFD38FC1F" box="[437,643,908,933]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
palm depth/palm length (
<emphasis id="B971EAD3FFBA333B9DBAFC20FC0DFC14" bold="true" box="[936,950,914,942]" pageId="15" pageNumber="541">x</emphasis>
= 79.8%; range 74.7–84.9%) is larger than 
<taxonomicName id="4C054D42FFBA333B9E85FC06FEBCFC76" box="[151,263,947,972]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="dubius">
<emphasis id="B971EAD3FFBA333B9E85FC06FEBCFC76" box="[151,263,947,972]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257A8FFBA333B9F03FC06FE8EFC71" box="[273,309,948,971]" pageId="15" pageNumber="541" sensu="stricto">s.s.</taxonomicNameLabel>
(
<emphasis id="B971EAD3FFBA333B9F5FFC0BFEE0FC6F" bold="true" box="[333,347,953,981]" pageId="15" pageNumber="541">x</emphasis>
= 71.0%, range 64.0–77%). 
<taxonomicName id="4C054D42FFBA333B9CBEFC01FCC5FC76" box="[684,894,947,972]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9CBEFC01FCC5FC76" box="[684,894,947,972]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
rostral width/rostral length ratios (
<emphasis id="B971EAD3FFBA333B9B0CFC0BFA97FC6F" bold="true" box="[1310,1324,953,981]" pageId="15" pageNumber="541">x</emphasis>
= 66.7%; range 60.9–73.8%) are smaller than 
<taxonomicName id="4C054D42FFBA333B9C3FFC68FD22FC48" box="[557,665,985,1010]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="dubius">
<emphasis id="B971EAD3FFBA333B9C3FFC68FD22FC48" box="[557,665,985,1010]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257A8FFBA333B9CB2FC68FD7FFC4B" box="[672,708,986,1009]" pageId="15" pageNumber="541" sensu="stricto">s.s.</taxonomicNameLabel>
(rostral length/rostral width (
<emphasis id="B971EAD3FFBA333B9A01FC52FB9AFC46" bold="true" box="[1043,1057,992,1020]" pageId="15" pageNumber="541">x</emphasis>
= 73.5%; range 67.3–79.6%).
</paragraph>
<paragraph id="8BBA36C1FFBA333B9ED5FBB2FE51FA82" blockId="15.[151,1437,151,2027]" pageId="15" pageNumber="541">
The number of subpalmar tubercles is an important character that separates 
<taxonomicName id="4C054D42FFBA333B9A3AFBB3FB26FBA3" box="[1064,1181,1024,1049]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9A3AFBB3FB26FBA3" box="[1064,1181,1024,1049]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
from both typical and undescribed members of the complex. 
<taxonomicName id="4C054D42FFBA333B9C59FB97FCA2FB84" box="[587,793,1061,1086]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9C59FB97FCA2FB84" box="[587,793,1061,1086]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
always has at least 1, and sometimes as many as three (
<emphasis id="B971EAD3FFBA333B9B9DFB94FA20FB87" box="[1423,1435,1062,1085]" italics="true" pageId="15" pageNumber="541">x</emphasis>
= 1.2) subpalmar tubercles; all other populations of 
<taxonomicName id="4C054D42FFBA333B9CECFBFBFC24FBDB" box="[766,927,1096,1121]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9CECFBFBFCACFBDA" box="[766,791,1097,1120]" italics="true" pageId="15" pageNumber="541">C.</emphasis>
aff. 
<emphasis id="B971EAD3FFBA333B9D46FBFAFC24FBDB" box="[852,927,1096,1121]" italics="true" pageId="15" pageNumber="541">dubius</emphasis>
</taxonomicName>
lack subpalmar tuberlces. This character is particularly important when dealing with preserved specimens once they have been bleached following preservation (ZJL, personal observation). In addition to the subpalmar tubercles, the primary carpal spines frequently (82% of specimens examined) are bifurcated into two large spines in 
<taxonomicName id="4C054D42FFBA333B9A23FB04FB1DFB74" box="[1073,1190,1205,1230]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9A23FB04FB1DFB74" box="[1073,1190,1205,1230]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
. To date, no other 
<taxonomicName id="4C054D42FFBA333B9B90FB04FF59FB4B" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9B90FB04FF59FB4B" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
population analyzed in West Virginia, or elsewhere by the authors, possess either of these characters states. Finally, the 
<taxonomicName id="4C054D42FFBA333B9F0CFB4CFE29FAAC" box="[286,402,1277,1302]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9F0CFB4CFE29FAAC" box="[286,402,1277,1302]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
carapace is more dorsally flattened and less vaulted compared to the laterally compressed, vaulted carapace of 
<taxonomicName id="4C054D42FFBA333B9F65FA93FE58FA83" box="[375,483,1312,1337]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9F65FA93FE58FA83" box="[375,483,1312,1337]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBA36C1FFBA333B9ED5FAF7FD2FF8A8" blockId="15.[151,1437,151,2027]" pageId="15" pageNumber="541">
The BBOC morph is the closest geographic member of the 
<taxonomicName id="4C054D42FFBA333B9D8EFAF4FBFBFAE4" box="[924,1088,1349,1374]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B971EAD3FFBA333B9D8EFAF4FC0DFAE7" box="[924,950,1350,1373]" italics="true" pageId="15" pageNumber="541">C.</emphasis>
aff. 
<emphasis id="B971EAD3FFBA333B9DE7FAF7FBFBFAE4" box="[1013,1088,1349,1374]" italics="true" pageId="15" pageNumber="541">dubius</emphasis>
</taxonomicName>
complex to 
<taxonomicName id="4C054D42FFBA333B9ACFFAF4FAEDFAE4" box="[1245,1366,1349,1374]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9ACFFAF4FAEDFAE4" box="[1245,1366,1349,1374]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
, with populations occurring within 1.0 km of a site harboring 
<taxonomicName id="4C054D42FFBA333B9D19FADBFCC5FA3B" box="[779,894,1384,1409]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9D19FADBFCC5FA3B" box="[779,894,1384,1409]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
(ZJL, unpublished data). 
<taxonomicName id="4C054D42FFBA333B9AB2FADAFAD5FA3B" box="[1184,1390,1384,1409]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9AB2FADAFAD5FA3B" box="[1184,1390,1384,1409]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
can be differentiated from the BBOC morph by overall coloration, palm depth/palm length ratio, and rostral and carapace morphology. The dorsal coloration of the carapace and abdomen of the BBOC morph ranges from black to brown, with lateral carapace colors ranging from orange to red, which differs from the blue carapace of 
<taxonomicName id="4C054D42FFBA333B9B90FA64FF52F9AB" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9B90FA64FF52F9AB" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
. Dorsal surfaces of the palm in the BBOC morph are composed primarily of black/browns, with both the dactyl and propodus primarily consisting of oranges and reds. 
<taxonomicName id="4C054D42FFBA333B9D40F9AFFB9BF98C" box="[850,1056,1565,1590]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9D40F9AFFB9BF98C" box="[850,1056,1565,1590]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
chelae coloration always consists of blues. Unlike comparison with 
<taxonomicName id="4C054D42FFBA333B9C31F9F3FD2AF9E3" box="[547,657,1600,1625]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="dubius">
<emphasis id="B971EAD3FFBA333B9C31F9F3FD2AF9E3" box="[547,657,1600,1625]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257A8FFBA333B9C89F9F3FD7BF9E2" box="[667,704,1601,1624]" pageId="15" pageNumber="541" sensu="stricto">s.s.</taxonomicNameLabel>
, the palm length/palm depth ratio in 
<taxonomicName id="4C054D42FFBA333B9A61F9F3FB53F9E3" box="[1139,1256,1600,1625]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9A61F9F3FB53F9E3" box="[1139,1256,1600,1625]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
is smaller (
<emphasis id="B971EAD3FFBA333B9B6BF9F5FA3CF9D9" bold="true" box="[1401,1415,1607,1635]" pageId="15" pageNumber="541">x</emphasis>
= 79.8%; range 74.7–84.9%) than the BBOC morph (
<emphasis id="B971EAD3FFBA333B9CCDF9DCFD56F930" bold="true" box="[735,749,1646,1674]" pageId="15" pageNumber="541">x</emphasis>
<emphasis id="B971EAD3FFBA333B9CE0F9DBFCBEF93A" box="[754,773,1641,1664]" italics="true" pageId="15" pageNumber="541">=</emphasis>
83.1%; range 80.7–88.0%, n = 22) ratio. Morphologically, both the BBOC morph and 
<taxonomicName id="4C054D42FFBA333B9FC3F93DFD85F91D" box="[465,574,1678,1703]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" sensu="stricto" species="dubius">
<emphasis id="B971EAD3FFBA333B9FC3F93DFD85F91D" box="[465,574,1678,1703]" italics="true" pageId="15" pageNumber="541">C. dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24257A8FFBA333B9C54F922FDD1F91D" box="[582,618,1680,1703]" pageId="15" pageNumber="541" sensu="stricto">s.s.</taxonomicNameLabel>
lack subpalmar tubercles and possess a more dorsolaterally compressed carapace compared to 
<taxonomicName id="4C054D42FFBA333B9FB9F901FD9DF971" box="[427,550,1714,1739]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9FB9F901FD9DF971" box="[427,550,1714,1739]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
, the latter of which always maintains subpalmar tubercles as well as a dorsolventrally compressed carapace. The rostrum of 
<taxonomicName id="4C054D42FFBA333B9CE2F964FC05F955" box="[752,958,1750,1775]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9CE2F964FC05F955" box="[752,958,1750,1775]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
is moderately excavated, while the BBOC morph possesses a deeply excavated rostrum.
</paragraph>
<paragraph id="8BBA36C1FFBA333B9ED5F8ACFA26F851" blockId="15.[151,1437,151,2027]" pageId="15" pageNumber="541">
The dorsal and lateral carapace and abdominal coloration in 
<taxonomicName id="4C054D42FFBA333B9D83F8ACFAC5F88D" authority="Ortmann, 1905" authorityName="Ortmann" authorityYear="1905" box="[913,1406,1822,1847]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="monongalensis">
<emphasis id="B971EAD3FFBA333B9D83F8ACFB02F88D" box="[913,1209,1822,1847]" italics="true" pageId="15" pageNumber="541">Cambarus monongalensis</emphasis>
<bibRefCitation id="EF944B30FFBA333B9AD6F8ADFAC5F88D" author="Ortmann" box="[1220,1406,1822,1847]" pageId="15" pageNumber="541" refString="Ortmann, A. E. (1905 a) The crawfishes of Western Pennsylvania. Annals of the Carnegie Museum, 3, 387 - 406." type="journal article" year="1905" yearSuffix="a">Ortmann, 1905a</bibRefCitation>
</taxonomicName>
is similar to 
<taxonomicName id="4C054D42FFBA333B9F15F8F1FEC1F8E1" box="[263,378,1858,1883]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9F15F8F1FEC1F8E1" box="[263,378,1858,1883]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
, consisting of several shades of blue. The latter can be distinguished from 
<taxonomicName id="4C054D42FFBA333B9AABF8F1FA97F8E1" box="[1209,1324,1858,1883]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9AABF8F1FA97F8E1" box="[1209,1324,1858,1883]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
by chelae and rostral morphology. The margins of the propodus in 
<emphasis id="B971EAD3FFBA333B9D3AF8D4FC42F8C5" box="[808,1017,1894,1919]" italics="true" pageId="15" pageNumber="541">
Cambaurs 
<taxonomicName id="4C054D42FFBA333B9DB5F8D4FC42F8C5" box="[935,1017,1894,1919]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">pauleyi</taxonomicName>
</emphasis>
are costate compared to the smooth margins of 
<taxonomicName id="4C054D42FFBA333B9F07F839FE66F819" box="[277,477,1930,1955]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="monongalensis">
<emphasis id="B971EAD3FFBA333B9F07F839FE66F819" box="[277,477,1930,1955]" italics="true" pageId="15" pageNumber="541">C. monongalensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C054D42FFBA333B9FF9F838FD03F819" box="[491,696,1930,1955]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9FF9F838FD03F819" box="[491,696,1930,1955]" italics="true" pageId="15" pageNumber="541">Cambarus pauleyi</emphasis>
</taxonomicName>
also has two rows of tubercles on the mesial margin of the chelae, compared to a single row in 
<taxonomicName id="4C054D42FFBA333B9FC0F81DFD21F87D" box="[466,666,1966,1991]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="monongalensis">
<emphasis id="B971EAD3FFBA333B9FC0F81DFD21F87D" box="[466,666,1966,1991]" italics="true" pageId="15" pageNumber="541">C. monongalensis</emphasis>
</taxonomicName>
. Rostral anatomy can also be used to differentiate these species; with 
<taxonomicName id="4C054D42FFBA333B9E85F861FEABF851" box="[151,272,2002,2027]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFBA333B9E85F861FEABF851" box="[151,272,2002,2027]" italics="true" pageId="15" pageNumber="541">C. pauleyi</emphasis>
</taxonomicName>
having a more elongate rostrum while in 
<taxonomicName id="4C054D42FFBA333B9D0BF861FC53F851" box="[793,1000,2002,2027]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="541" phylum="Arthropoda" rank="species" species="monongalensis">
<emphasis id="B971EAD3FFBA333B9D0BF861FC53F851" box="[793,1000,2002,2027]" italics="true" pageId="15" pageNumber="541">C. monongalensis</emphasis>
</taxonomicName>
the rostrum is truncated, and short.
</paragraph>
<paragraph id="8BBA36C1FFA533249E85FF25FE67FEDE" blockId="16.[151,1437,151,645]" pageId="16" pageNumber="542">
Furthermore, the two species can be separated in the field by the location of collection, given that the ranges of 
<taxonomicName id="4C054D42FFA533249B90FF2AFF52FF6F" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFA533249B90FF2AFF52FF6F" italics="true" pageId="16" pageNumber="542">C. pauleyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C054D42FFA533249F37FF0FFE4BFF6F" box="[293,496,188,213]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="monongalensis">
<emphasis id="B971EAD3FFA533249F37FF0FFE4BFF6F" box="[293,496,188,213]" italics="true" pageId="16" pageNumber="542">C. monongalensis</emphasis>
</taxonomicName>
do not overlap. 
<taxonomicName id="4C054D42FFA533249CA3FF0EFC3AFF6F" box="[689,897,188,213]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFA533249CA3FF0EFC3AFF6F" box="[689,897,188,213]" italics="true" pageId="16" pageNumber="542">Cambarus pauleyi</emphasis>
</taxonomicName>
is relegated to the high elevation Appalachian Plateau in Greenbrier and Monroe County, West Virginia. 
<taxonomicName id="4C054D42FFA533249D26FF6DFBE1FF42" box="[820,1114,223,248]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="monongalensis">
<emphasis id="B971EAD3FFA533249D26FF6DFBE1FF42" box="[820,1114,223,248]" italics="true" pageId="16" pageNumber="542">Cambarus monongalensis</emphasis>
</taxonomicName>
has a range that consists of two disjunct populations, one in the Allegheny Mountains of West Virginia and Virginia, and another present in the low elevation Appalachian Plateau of West Virginia and Pennsylvania (
<bibRefCitation id="EF944B30FFA533249DCBFE9AFB28FEFA" author="Ortmann" box="[985,1171,295,320]" pageId="16" pageNumber="542" refString="Ortmann, A. E. (1905 b) The mutual affinities of the species of the genus Cambarus and their dispersal over the United States. Proceedings of the American Philosophical Society, 44, 91 - 136." type="journal article" year="1905" yearSuffix="b">Ortmann 1905b</bibRefCitation>
; Jezerinac 
<emphasis id="B971EAD3FFA533249B07FE9BFAF2FEFA" box="[1301,1353,295,320]" italics="true" pageId="16" pageNumber="542">et al</emphasis>
. 1995; Loughman and 
<collectingCountry id="F3127651FFA533249F54FEFEFE30FEDF" box="[326,395,332,357]" name="United Kingdom" pageId="16" pageNumber="542">Welsh</collectingCountry>
2013).
</paragraph>
</subSubSection>
<subSubSection id="C31F654AFFA533249ED5FEDDFD1BFD3F" pageId="16" pageNumber="542" type="etymology">
<paragraph id="8BBA36C1FFA533249ED5FEDDFCECFDDA" blockId="16.[151,1437,151,645]" pageId="16" pageNumber="542">
<emphasis id="B971EAD3FFA533249ED5FEDDFEF5FE32" bold="true" box="[199,334,367,392]" pageId="16" pageNumber="542">Etymology.</emphasis>
It is with great pleasure, specifically from the primary author, that this crayfish be named in honor of Dr. Thomas K. Pauley, Professor Emeritus, 
<collectingCountry id="F3127651FFA533249CA6FE26FCA3FE17" box="[692,792,404,429]" name="Marshall Islands" pageId="16" pageNumber="542">Marshall</collectingCountry>
University. 
<collectingCountry id="F3127651FFA533249DB5FE26FC7DFE16" box="[935,966,404,428]" name="American Samoa" pageId="16" pageNumber="542">As</collectingCountry>
ZJL’s graduate advisor, you let his little “crayfish distraction” away from the herpetological focus of his thesis occur without objection, thus changing his professional focus, and academic life forever. Furthermore, it is also fitting that 
<taxonomicName id="4C054D42FFA533249A08FE6FFB35FE4F" box="[1050,1166,476,501]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFA533249A08FE6FFB35FE4F" box="[1050,1166,476,501]" italics="true" pageId="16" pageNumber="542">C. pauleyi</emphasis>
</taxonomicName>
bare Professor Pauley’s name, given 
<taxonomicName id="4C054D42FFA533249F3CFDB2FE1EFDA2" box="[302,421,511,536]" class="Malacostraca" family="Cambaridae" genus="Cambarus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="542" phylum="Arthropoda" rank="species" species="pauleyi">
<emphasis id="B971EAD3FFA533249F3CFDB2FE1EFDA2" box="[302,421,511,536]" italics="true" pageId="16" pageNumber="542">C. pauleyi</emphasis>
</taxonomicName>
burrows provide refuge and safety for countless reptiles and amphibians that share its range, much like Pauley’s research efforts to conserve West Virginia’s reptiles and amphibians have undoubtedly aided in the conservation of reptiles and amphibians statewide.
</paragraph>
<paragraph id="8BBA36C1FFA533249ED5FDDEFD1BFD3F" blockId="16.[151,1437,151,645]" box="[199,672,620,645]" pageId="16" pageNumber="542">
<emphasis id="B971EAD3FFA533249ED5FDDEFE3EFD3F" bold="true" box="[199,389,620,645]" pageId="16" pageNumber="542">Common name.</emphasis>
Meadow River Mudbug.
</paragraph>
</subSubSection>
</treatment>
</document>