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<document id="997D78289B4F346A67791C5F074D36D3" ID-DOI="10.5281/zenodo.11066801" ID-ISSN="0068-547X" ID-Zenodo-Dep="11066801" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1714049691827" checkinUser="felipe" docAuthor="Ceríaco, L. M. P., De Sá, S., Bandeira, S., Valério, H., Stanley, E. L., Kuhn, A. L., Marques, M. P., Vindum, J. V., Blackburn, D. C. &amp; Bauer, A. M." docDate="2016" docId="F67087FF53556D3AFF9753E6A849B3CA" docLanguage="en" docName="ProcCalifAcadSci.63.2.15-61.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (2)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.1:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="1" docTitle="Varanus albigularis subsp. angolensis Schmidt 1933" docType="treatment" docVersion="2" lastPageNumber="39" masterDocId="0A49FF87534D6D23FFEC5623AC39B238" masterDocTitle="Herpetological Survey of Iona National Park and Namibe Regional Natural Park, with a Synoptic List of the Amphibians and Reptiles of Namibe Province, Southwestern Angola" masterLastPageNumber="61" masterPageNumber="15" pageNumber="39" updateTime="1714656057459" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A9A08584AE33A47D5C36DE25445D8655">Herpetological Survey of Iona National Park and Namibe Regional Natural Park, with a Synoptic List of the Amphibians and Reptiles of Namibe Province, Southwestern Angola</mods:title>
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<mods:namePart id="252BCE2B1B06170A5F3471A89D2F7CCA">Ceríaco, L. M. P.</mods:namePart>
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<mods:namePart id="512982E3E9D61A701CFEF9B487562D55">De Sá, S.</mods:namePart>
<mods:affiliation id="637859FB1E4987098EDD2EDE9B606DF1">Instituto Nacional da Biodiversidade e Áreas de Conservação, Ministério do Ambiente de Angola, Centralidade do Kilamba, Rua 26 de Fevereiro, quarteirão Nimi ya Lukemi, edíficio Q 11, 3 ° andar, Angola.</mods:affiliation>
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<mods:namePart id="72C8EF9941A51147F425E9488AA6DADF">Bandeira, S.</mods:namePart>
<mods:affiliation id="0F6C8306FDCFDE758F6511D7AE9F101C">Instituto Nacional da Biodiversidade e Áreas de Conservação, Ministério do Ambiente de Angola, Centralidade do Kilamba, Rua 26 de Fevereiro, quarteirão Nimi ya Lukemi, edíficio Q 11, 3 ° andar, Angola.</mods:affiliation>
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<mods:affiliation id="A1D898FCF3CE30848FF31E7162ED0B97">Instituto Nacional da Biodiversidade e Áreas de Conservação, Ministério do Ambiente de Angola, Centralidade do Kilamba, Rua 26 de Fevereiro, quarteirão Nimi ya Lukemi, edíficio Q 11, 3 ° andar, Angola.</mods:affiliation>
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<mods:namePart id="86CB525E6A7427A890FB86F527697B4E">Stanley, E. L.</mods:namePart>
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<mods:namePart id="F47221D1CFD71C8621DAD16BBEEE608C">Kuhn, A. L.</mods:namePart>
<mods:affiliation id="B09B0B4AF8B952927273F1B8761C4649">American Museum of Natural History, Central Park West at 79 th Street, New York, New York 10024, USA. &amp; City University of New York, Graduate Center, 365 5 th Ave., New York, New York, 10016, USA.</mods:affiliation>
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<mods:namePart id="204B88DC3242C03658656E6A0E59EFF8">Marques, M. P.</mods:namePart>
<mods:affiliation id="7F19AEBE89E6ECB86E93F1C609CA69D7">Museu Nacional de História Natural e da Ciência, Universidade de Lisboa, Rua da Escola Politécnica, 58, 1269 - 102 Lisbon, Portugal.</mods:affiliation>
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<mods:namePart id="0A7D742C942792E09991245A4F0A39FF">Vindum, J. V.</mods:namePart>
<mods:affiliation id="582693DDA0F0D01C99E0B590D9EF32A0">Department of Herpetology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118, USA.</mods:affiliation>
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<mods:namePart id="3CF219A6BE3252766C92A89961B1FEC1">Blackburn, D. C.</mods:namePart>
<mods:affiliation id="9172EE842A8840EEFF7205F69F6571CE">Department of Natural History, Florida Museum of Natural History, Gainesville, FL, 32611, USA.</mods:affiliation>
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<mods:namePart id="6ACA6B3C50E697323CE9EAB7250526B1">Bauer, A. M.</mods:namePart>
<mods:affiliation id="3CED31AD04CEA9550FA1A295ED0F8D39">Department of Biology, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085 - 1699, USA.</mods:affiliation>
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<mods:date id="8D891FFF32E3A8347FC244F02FA14F82">2016</mods:date>
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<treatment id="F67087FF53556D3AFF9753E6A849B3CA" LSID="urn:lsid:plazi:treatment:F67087FF53556D3AFF9753E6A849B3CA" httpUri="http://treatment.plazi.org/id/F67087FF53556D3AFF9753E6A849B3CA" lastPageId="25" lastPageNumber="39" pageId="24" pageNumber="39">
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<taxonomicName id="B9D94D6A53556D3BFF9753C6AE43B7C5" ID-CoL="7J632" authority="Schmidt, 1933" authorityName="Schmidt" authorityYear="1933" box="[123,634,1508,1533]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="39" phylum="Chordata" rank="subSpecies" species="albigularis" subSpecies="angolensis">
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<emphasis id="4CADEAFB53556D3BFF9753C6ADE8B7C4" bold="true" box="[123,465,1508,1532]" italics="true" pageId="24" pageNumber="39">Varanus albigularis angolensis</emphasis>
<bibRefCitation id="1A484B1853556D3BFE3453C6AE43B7C5" author="SCHMIDT, K. P." box="[472,634,1509,1533]" pageId="24" pageNumber="39" pagination="1 - 15" refId="ref19666" refString="SCHMIDT, K. P. 1933. The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum 22: 1 - 15." type="journal article" year="1933">Schmidt, 1933</bibRefCitation>
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.—
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<quantity id="B9219B0C53556D3BFEA95003ADB6B40F" box="[325,399,1567,1591]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="24" pageNumber="39" unit="km" value="7.5">7.5 km</quantity>
NW (by road) of Pico Azevedo, adult, found in a rock crevice,
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,
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,
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,
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(
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.
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<emphasis id="4CADEAFB53556D3BFF47507CAD11B44E" bold="true" box="[171,296,1631,1655]" pageId="24" pageNumber="39">COMMENTS</emphasis>
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.— The subspecies was described from Gauca, Bihé (
<collectingRegion id="BC1DF80B53556D3BFC61507CAF8BB44F" box="[909,946,1631,1655]" country="Angola" name="Bie" pageId="24" pageNumber="39">Bié</collectingRegion>
) Province (
<bibRefCitation id="1A484B1853556D3BFBD0507CAC89B4A0" author="SCHMIDT, K. P." pageId="24" pageNumber="39" pagination="1 - 15" refId="ref19666" refString="SCHMIDT, K. P. 1933. The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum 22: 1 - 15." type="journal article" year="1933">Schmidt 1933</bibRefCitation>
). According to the original description, the subspecies differed from the nominotypic form by having larger scales everywhere on the body so that the scales around the body are about 125 instead of 150, and the transverse rows of scales from the collar to thighs are 75 instead of 100 (
<bibRefCitation id="1A484B1853546D3AFF6856D8AD23B32B" author="SCHMIDT, K. P." box="[132,282,251,275]" pageId="25" pageNumber="40" pagination="1 - 15" refId="ref19666" refString="SCHMIDT, K. P. 1933. The reptiles of the Pulitzer-Angola Expedition. Annals of the Carnegie Museum 22: 1 - 15." type="journal article" year="1933">Schmidt 1933</bibRefCitation>
). Laurent (1964b) noted that the morphological differences between the nominotypic form and
<taxonomicName id="B9D94D6A53546D3AFEF05738ADB7B30B" baseAuthorityName="Loveridge" baseAuthorityYear="1944" box="[284,398,283,307]" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="40" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="4CADEAFB53546D3AFEF05738ADB7B30B" box="[284,398,283,307]" italics="true" pageId="25" pageNumber="40">angolensis</emphasis>
</taxonomicName>
are quite subtle and that
<taxonomicName id="B9D94D6A53546D3AFD795738AF3EB30B" baseAuthorityName="Loveridge" baseAuthorityYear="1944" box="[661,775,283,307]" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="40" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="4CADEAFB53546D3AFD795738AF3EB30B" box="[661,775,283,307]" italics="true" pageId="25" pageNumber="40">angolensis</emphasis>
</taxonomicName>
may, in fact, be a synonym of
<taxonomicName id="B9D94D6A53546D3AFBA45738AC95B36B" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="40" phylum="Chordata" rank="species" species="albigularis">
<emphasis id="4CADEAFB53546D3AFBA45738AC95B36B" italics="true" pageId="25" pageNumber="40">albigularis</emphasis>
</taxonomicName>
. However,
<bibRefCitation id="1A484B1853546D3AFEC45718ADF3B36B" author="BAYLESS, M. K." box="[296,458,315,339]" pageId="25" pageNumber="40" pagination="1643 - 1701" refId="ref15448" refString="BAYLESS, M. K. 2002. Monitor lizards: a pan-African check-list of their zoogeography (Sauria: Varanidae: Polydaedalus). Journal of Biogeography 29: 1643 - 1701." type="journal article" year="2002">Bayless (2002)</bibRefCitation>
considered the material from nearby localities, such as Bibala and Caraculo as
<emphasis id="4CADEAFB53546D3AFF13577FAD9CB34B" box="[255,421,347,371]" italics="true" pageId="25" pageNumber="40">
V. a.
<taxonomicName id="B9D94D6A53546D3AFEDF5778AD9CB34B" baseAuthorityName="Loveridge" baseAuthorityYear="1944" box="[307,421,347,371]" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="40" phylum="Chordata" rank="species" species="angolensis">angolensis</taxonomicName>
</emphasis>
. Given the currently accepted distribution of
<taxonomicName id="B9D94D6A53546D3AFC675778AFC4B34B" baseAuthorityName="Loveridge" baseAuthorityYear="1944" box="[907,1021,347,371]" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="40" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="4CADEAFB53546D3AFC675778AFC4B34B" box="[907,1021,347,371]" italics="true" pageId="25" pageNumber="40">angolensis</emphasis>
</taxonomicName>
and the lower number of scales around the midbody and between the collar and the thighs, we tentatively identify our specimen as the Angolan subspecies. The subspecies appears to be the prevalent form in
<collectingCountry id="06CE767953546D3AFF975798ACF3B3EB" box="[123,202,443,467]" name="Angola" pageId="25" pageNumber="40">Angola</collectingCountry>
(and possibly extending to some neighboring regions of the
<collectingCountry id="06CE767953546D3AFCB65798AFA8B3EB" box="[858,913,443,467]" name="Democratic Republic of the Congo" pageId="25" pageNumber="40">DRC</collectingCountry>
, even if the nominotypic form occurs sympatrically, especially in the southern regions of the country (
<bibRefCitation id="1A484B1853546D3AFC3D57F8A859B3CB" author="BAYLESS, M. K." box="[977,1120,475,499]" pageId="25" pageNumber="40" pagination="1643 - 1701" refId="ref15448" refString="BAYLESS, M. K. 2002. Monitor lizards: a pan-African check-list of their zoogeography (Sauria: Varanidae: Polydaedalus). Journal of Biogeography 29: 1643 - 1701." type="journal article" year="2002">Bayless 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>