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<mods:title id="0421BB45BE05B28C153668D1B05E3D1F">Taxonomy and palaeobiogeography of charophytes from the Upper Eocene- Lower Oligocene of the Eastern Ebro Basin (Catalonia, NE Spain)</mods:title>
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<treatment id="D25987DDFFF6FFA9FF7AEA3E2CD88C69" ID-DOI="http://doi.org/10.5281/zenodo.4836378" ID-GBIF-Taxon="182403506" ID-Zenodo-Dep="4836378" LSID="urn:lsid:plazi:treatment:D25987DDFFF6FFA9FF7AEA3E2CD88C69" httpUri="http://treatment.plazi.org/id/D25987DDFFF6FFA9FF7AEA3E2CD88C69" lastPageNumber="404" pageId="19" pageNumber="404">
<subSubSection id="12EA6540FFF6FFA9FF7AEA3E2E998ACE" pageId="19" pageNumber="404" type="nomenclature">
<paragraph id="5A4F36CBFFF6FFA9FF7AEA3E2E998ACE" blockId="19.[171,590,627,717]" pageId="19" pageNumber="404">
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<taxonomicName id="9DF04D48FFF6FFA9FF7AEA3E2E998ACE" authority="(Groves, 1926) Soulie-Marsche, 1989" authorityName="Soulie-Marsche" authorityYear="1989" baseAuthorityName="Groves" baseAuthorityYear="1926" class="Charophyceae" family="Characeae" genus="Lychnothamnus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="19" pageNumber="404" phylum="Charophyta" rank="species" species="vectensis">
<emphasis id="6884EAD9FFF6FFA9FF7AEA3E2ED88AED" bold="true" box="[171,428,627,654]" italics="true" pageId="19" pageNumber="404">Lychnothamnus vectensis</emphasis>
(
<bibRefCitation id="3E614B3AFFF6FFA9FE69EA392D328AEC" author="GROVES J." box="[440,582,628,655]" pageId="19" pageNumber="404" pagination="165 - 173" refId="ref17584" refString="GROVES J. 1926. - Charophyta, in REID E. M. &amp; CHAN- DLER M. E. J. (eds), hve Bembridge Flora. British Museum Catalogue, Cainozoicum Plants, London 1: 165 - 173." type="journal article" year="1926">Groves, 1926</bibRefCitation>
) Soulié-Märsche, 1989
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</heading>
</paragraph>
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<subSubSection id="12EA6540FFF6FFA9FEECEAFE2ECF8AAE" box="[317,443,691,717]" pageId="19" pageNumber="404" type="description">
<paragraph id="5A4F36CBFFF6FFA9FEECEAFE2ECF8AAE" blockId="19.[171,590,627,717]" box="[317,443,691,717]" pageId="19" pageNumber="404">
(
<figureCitation id="C2CB2A4EFFF6FFA9FE95EAFE2E0A8AAE" box="[324,382,691,717]" captionStart="FIG" captionStartId="20.[113,124,1498,1516]" captionTargetBox="[125,1193,221,1389]" captionTargetId="figure-226@20.[116,662,761,1310]" captionTargetPageId="20" captionText="FIG. 8.— Charophytes from the Artés and lateral south-western Formations,Ebro Basin:A-C, Lychnothamnus grambastii (Feist-Castel, 1971), Torre Casanova section; A, apical view, no. 80100 MGSCB, sample TC-9; B, lateral view, no. 80101 MGSCB, sample TC-9; C, basal view,no.80102 MGSCB,sample TC-9;D-F, Lychnothamnus vectensis (Groves,1926),Santpedor and Rubió sections;D, apical view, no. 80103 MGSCB, sample RB-3; E, oblicuous view, no. 80104 MGSCB, sample SP-18; F, lateral view, no. 80104 MGSCB, sample SP-18; G, basal view, no. 80105 MGSCB, sample SP-4; H-K, Lychnothamnus major (Grambast &amp; Paul, 1965), Sarral section and El Talladell outcrop; H, apex, no. 80106 MGSCB, sample BO-1; I, lateral view, no. 80107 MGSCB, sample BO-1; J, lateral view, no. 80108 MGSCB, sample SA-19; K, basal view, no. 80109 MGSCB 09, sample SA-19; L-N, Nitellopsis (Tectochara) merianii (Al. Braun ex Unger, 1852), Vinaixa section; L, apical view, no. 80110 MGSCB, sample VI-5; M, lateral view, no. 80111 MGSCB, sample VI-5; N, basal view, no. 80112 MGSCB, sample VI-5. Scale bar: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4538545" httpUri="https://zenodo.org/record/4538545/files/figure.png" pageId="19" pageNumber="404">Fig. 8</figureCitation>
D-G)
</paragraph>
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<subSubSection id="12EA6540FFF6FFA9FFA0EAA12D178B00" pageId="19" pageNumber="404" type="reference_group">
<paragraph id="5A4F36CBFFF6FFA9FFA0EAA12D4E8B67" blockId="19.[113,570,748,772]" box="[113,570,748,772]" pageId="19" pageNumber="404">
<treatmentCitationGroup id="7AE011E5FFF6FFA9FFA0EAA12D4E8B67" box="[113,570,748,772]" pageId="19" pageNumber="404">
<treatmentCitation id="DB5110DAFFF6FFA9FFA0EAA12EDB8B67" author="GROVES J." box="[113,431,748,772]" page="172" pageId="19" pageNumber="404" year="1926">
<taxonomicName id="9DF04D48FFF6FFA9FFA0EAA12EDB8B67" authority="Groves, 1926: 172" authorityName="Groves" authorityPageNumber="172" authorityYear="1926" box="[113,431,748,772]" class="Charophyceae" family="Characeae" genus="Chara" kingdom="Plantae" order="Charales" pageId="19" pageNumber="404" phylum="Charophyta" rank="species" species="vectensis">
<emphasis id="6884EAD9FFF6FFA9FFA0EAA12F8D8B67" box="[113,249,748,772]" italics="true" pageId="19" pageNumber="404">Chara vectensis</emphasis>
<bibRefCitation id="3E614B3AFFF6FFA9FF2FEAA12EDB8B67" author="GROVES J." box="[254,431,748,772]" pageId="19" pageNumber="404" pagination="165 - 173" refId="ref17584" refString="GROVES J. 1926. - Charophyta, in REID E. M. &amp; CHAN- DLER M. E. J. (eds), hve Bembridge Flora. British Museum Catalogue, Cainozoicum Plants, London 1: 165 - 173." type="journal article" year="1926">Groves, 1926: 172</bibRefCitation>
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, pl. 2, figs 2-8.
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</paragraph>
<paragraph id="5A4F36CBFFF6FFA9FFA0EB512D508B50" blockId="19.[113,548,795,820]" box="[113,548,795,820]" pageId="19" pageNumber="404">
<treatmentCitationGroup id="7AE011E5FFF6FFA9FFA0EB512D508B50" box="[113,548,795,820]" pageId="19" pageNumber="404">
<taxonomicName id="9DF04D48FFF6FFA9FFA0EB512E368B50" box="[113,322,796,820]" class="Charophyceae" family="Characeae" genus="Stephanochara" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="19" pageNumber="404" phylum="Charophyta" rank="species" species="vectensis">
<emphasis id="6884EAD9FFF6FFA9FFA0EB512E368B50" box="[113,322,796,820]" italics="true" pageId="19" pageNumber="404">Stephanochara vectensis</emphasis>
</taxonomicName>
<treatmentCitation id="DB5110DAFFF6FFA9FE8BEB512D558B50" author="GRAMBAST L." box="[346,545,795,820]" page="158" pageId="19" pageNumber="404" year="1958">
<bibRefCitation id="3E614B3AFFF6FFA9FE8BEB512D558B50" author="GRAMBAST L." box="[346,545,795,820]" pageId="19" pageNumber="404" refId="ref17420" refString="GRAMBAST L. 1958. - Etude sur les charophytes tertiaires d'Europe Occidentale et leurs rapports avec les formes actuelles. PhD dissertation Universite de Paris, 258 p." type="book" year="1958">Grambast 1958: 158</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="5A4F36CBFFF6FFA9FFA0EB062D178B00" blockId="19.[113,611,843,867]" box="[113,611,843,867]" pageId="19" pageNumber="404">
<taxonomicName id="9DF04D48FFF6FFA9FFA0EB062E398B00" authorityName="Soulie-Marsche" authorityYear="1989" baseAuthorityName="Groves" baseAuthorityYear="1926" box="[113,333,843,867]" class="Charophyceae" family="Characeae" genus="Lychnothamnus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="19" pageNumber="404" phylum="Charophyta" rank="species" species="vectensis">
<emphasis id="6884EAD9FFF6FFA9FFA0EB062E398B00" box="[113,333,843,867]" italics="true" pageId="19" pageNumber="404">Lychnothamnus vectensis</emphasis>
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Soulié-Märsche 1989: 160.
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</subSubSection>
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<paragraph id="5A4F36CBFFF6FFA9FFA0EB362E8E8B9E" blockId="19.[111,647,891,1021]" pageId="19" pageNumber="404">
DISTRIBUTION. —
<taxonomicName id="9DF04D48FFF6FFA9FEFBEB362D7F8BF0" authorityName="Soulie-Marsche" authorityYear="1989" baseAuthorityName="Groves" baseAuthorityYear="1926" box="[298,523,891,915]" class="Charophyceae" family="Characeae" genus="Lychnothamnus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Charales" pageId="19" pageNumber="404" phylum="Charophyta" rank="species" species="vectensis">
<emphasis id="6884EAD9FFF6FFA9FEFBEB362D7F8BF0" box="[298,523,891,915]" italics="true" pageId="19" pageNumber="404">Lychnothamnus vectensis</emphasis>
</taxonomicName>
has been recorded in the
<collectingRegion id="9834F829FFF6FFA9FF21EBD82E2E8BCE" box="[240,346,917,941]" country="United Kingdom" name="Hampshire" pageId="19" pageNumber="404">Hampshire</collectingRegion>
(
<bibRefCitation id="3E614B3AFFF6FFA9FEB5EBD82D538BCE" author="REID C. &amp; GROVES J." box="[356,551,917,941]" pageId="19" pageNumber="404" pagination="175 - 192" refId="ref18702" refString="REID C. &amp; GROVES J. 1921. - Ne Charophyta of the Lower Headon Beds of Hordle (Hordwell) Cliffs (South Hampshire). Quarterly Journal of the Geological Society of London 77: 175 - 192." type="journal article" year="1921">Reid &amp; Groves 1921</bibRefCitation>
) and Paris (
<bibRefCitation id="3E614B3AFFF6FFA9FFA7EBFD2F828BAB" author="RIVELINE J." box="[118,246,944,968]" pageId="19" pageNumber="404" refId="ref18744" refString="RIVELINE J. 1986. - Les Charophytes du Paleogene et du Miocene inferieur d'Europe Occidentale. Cahiers de Paleontologie, Centre National de la Recherche Scientifique, 227 p." type="book" year="1986">Riveline 1986</bibRefCitation>
) basins.In the Ebro Basin,this species occurs within Uppermost Priabonian-Lower Rupelian deposits in Santpedor, Rubió and Sarral (Tables 3, 4).
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF6FFA9FFA0EC7D2EB48E46" pageId="19" pageNumber="404" type="description">
<paragraph id="5A4F36CBFFF6FFA9FFA0EC7D2F8A8C2B" blockId="19.[111,648,1072,1573]" box="[113,254,1072,1098]" pageId="19" pageNumber="404">DESCRIPTION</paragraph>
<paragraph id="5A4F36CBFFF6FFA9FFA0EC022EB48E46" blockId="19.[111,648,1072,1573]" pageId="19" pageNumber="404">
Medium sized gyrogonite, variable in size,
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in high (mean
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), and
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in width (mean
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), normally ellipsoidal but sometimes ovoidal in shape with an isopolarity index of 103-145 (mean 121). Spiral cells about
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wide, normally flat to convex. Seven to nine (frequently eight), convolutions are visible laterally. Apex flat or slightly convex. Spiral cells do not show constrictions in the apical periphery but display marked thinning in the apical zone, resulting in an apical to periapical depression. In some gyrogonites, the apex is ornamented by tubercles that vary notably in size depending on the specimen. Ŋe base is rounded with a large and superficial basal pore about
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across.
</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF6FFA9FFA0EE072FA18E01" box="[113,213,1610,1636]" pageId="19" pageNumber="404" type="discussion">
<paragraph id="5A4F36CBFFF6FFA9FFA0EE072FA18E01" blockId="19.[113,648,1610,1700]" box="[113,213,1610,1636]" pageId="19" pageNumber="404">REMARKS</paragraph>
</subSubSection>
<subSubSection id="12EA6540FFF6FFA9FFA0EE242CD88C69" pageId="19" pageNumber="404" type="reference_group">
<paragraph id="5A4F36CBFFF6FFA9FFA0EE242CD88C69" blockId="19.[113,648,1610,1700]" lastBlockId="19.[674,1212,216,1034]" pageId="19" pageNumber="404">
<taxonomicName id="9DF04D48FFF6FFA9FFA0EE242E6D8EE0" authorityName="Groves" authorityYear="1926" box="[113,281,1641,1667]" class="Charophyceae" family="Characeae" genus="Chara" kingdom="Plantae" order="Charales" pageId="19" pageNumber="404" phylum="Charophyta" rank="species" species="vectensis">
<emphasis id="6884EAD9FFF6FFA9FFA0EE242E6D8EE0" box="[113,281,1641,1667]" italics="true" pageId="19" pageNumber="404">Chara vectensis</emphasis>
</taxonomicName>
was first determined by
<bibRefCitation id="3E614B3AFFF6FFA9FDE8EE272FC98EC7" author="GROVES J." pageId="19" pageNumber="404" pagination="165 - 173" refId="ref17584" refString="GROVES J. 1926. - Charophyta, in REID E. M. &amp; CHAN- DLER M. E. J. (eds), hve Bembridge Flora. British Museum Catalogue, Cainozoicum Plants, London 1: 165 - 173." type="journal article" year="1926">Groves (1926)</bibRefCitation>
based on ellipsoidal gyrogonites extremely variable in size,
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in high and
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in width, with 9-10 concave spiral cells usually swollen at the apex and often forming prominent rosettes. Ŋe
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designated and illustrated by
<bibRefCitation id="3E614B3AFFF6FFA9FC93E91A2C9C8912" author="GROVES J." box="[834,1000,342,369]" pageId="19" pageNumber="404" pagination="165 - 173" refId="ref17584" refString="GROVES J. 1926. - Charophyta, in REID E. M. &amp; CHAN- DLER M. E. J. (eds), hve Bembridge Flora. British Museum Catalogue, Cainozoicum Plants, London 1: 165 - 173." type="journal article" year="1926">Groves (1926)</bibRefCitation>
, from the ACourt Smith collection at the Natural History Museum, London, belonged precisely to the few specimens which displayed a prominent apical rosette and this character was further considered as diagnostic of the species, rather than as merely an extreme version of its total morphological variation. Taking into account this range of variation, the main difference between the material from the Ebro Basin and the type material is the size of the gyrogonites, which are about
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smaller in the Ebro Basin. Gyrogonites of this species from the Paris Basin gathered in the collection created by J. Riveline (Laboratoire de Biominéralization et énvironnement sedimentaires, Université Pierre et Marie Curie, Paris) show biometric parameters (mean values of
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high,
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wide, isopolarity index of 115 and 8 convolutions) similar to the type material. Furthermore, most of the gyrogonites from the Paris Basin show a swollen apex and concave spiral cells whilst the presence of prominent apical nodules is not common.
</paragraph>
</subSubSection>
</treatment>
</document>